165 lines
18 KiB
XML
165 lines
18 KiB
XML
<document id="DC27E52FA798A70192712C74379CFE2B" ID-DOI="10.12782/sd.21.1.071" ID-ISSN="2189-7301" ID-Zenodo-Dep="5737668" ID-ZooBank="8C100709-5B8B-42CB-BF9E-60B6F55F6045" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638226989121" checkinUser="felipe" docAuthor="Prokofiev, Artem M." docDate="2016" docId="554887B6E2497920FEB1473FFD1EB0AA" docLanguage="en" docName="SpeciesDiversity.21.1.71-77.pdf" docOrigin="Species Diversity 21" docStyle="DocumentStyle:27FECD2C3CD153DA405029F2D33C9514.7:SpeciesDiversity.2012-.journal_article" docStyleId="27FECD2C3CD153DA405029F2D33C9514" docStyleName="SpeciesDiversity.2012-.journal_article" docStyleVersion="7" docTitle="Engertia amboinae" docType="treatment" docVersion="3" lastPageNumber="73" masterDocId="A971FFCEE24B7922FFA8436AFFFBB730" masterDocTitle="On the Phytophagous Scarabs of the Subfamilies Dynastinae, Rutelinae, and Melolonthinae from the Schouten Islands (Kepulauan Biak), Indonesian Papua (Coleoptera: Scarabaeidae)" masterLastPageNumber="77" masterPageNumber="71" pageNumber="73" updateTime="1699433237484" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="1494A4445E3C88448061357F5E85B850">
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<mods:title id="66E0405CFDB6790418B0C0929C75B632">On the Phytophagous Scarabs of the Subfamilies Dynastinae, Rutelinae, and Melolonthinae from the Schouten Islands (Kepulauan Biak), Indonesian Papua (Coleoptera: Scarabaeidae)</mods:title>
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<mods:name id="AE985F456EA12731B0C20991D5884D0E" type="personal">
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<mods:roleTerm id="CD90A7388EA6251E8FE2668C38FD4C8E">Author</mods:roleTerm>
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<mods:namePart id="170CDED3CC2F3BF2881045262DD63931">Prokofiev, Artem M.</mods:namePart>
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<mods:typeOfResource id="FC3FDC29229DB530B17A10393998CCEC">text</mods:typeOfResource>
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<mods:titleInfo id="842E0924FDF5A257803E3F2473294ED5">
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<mods:title id="1B9F0197FD55DD8E873672F4250A7E41">Species Diversity</mods:title>
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</mods:titleInfo>
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<mods:part id="A639F483E3AA13C23CDAB505BB6553E6">
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<mods:date id="C8FFA1836A6748B03D07BA2F8E4A9DE5">2016</mods:date>
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<mods:detail id="453921913F7894C8ED4D03C2F1B17BB1" type="pubDate">
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<mods:number id="82A90FD1A75383FC33C5E333ECCC1D50">2016-05-25</mods:number>
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</mods:detail>
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<mods:detail id="D3D3C081D0CF25421DAAD40398F23C84" type="volume">
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<mods:number id="0038538EC16522F0BB8CF929A5BCC098">21</mods:number>
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<mods:start id="065F846E718F945F29F883014871B112">71</mods:start>
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<mods:end id="8A45C177DA8FBFCE531C59F220621281">77</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="72F5B576712BF9CD22C37DD464AE2B2D">journal article</mods:classification>
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<mods:identifier id="82B1E0B1E3C83DAB9CFE0BFD2060AC7E" type="DOI">10.12782/sd.21.1.071</mods:identifier>
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<mods:identifier id="9F9F1B79C000D0C47D43FC5FA259A98D" type="ISSN">2189-7301</mods:identifier>
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<treatment id="554887B6E2497920FEB1473FFD1EB0AA" ID-DOI="http://doi.org/10.5281/zenodo.5737636" ID-Zenodo-Dep="5737636" LSID="urn:lsid:plazi:treatment:554887B6E2497920FEB1473FFD1EB0AA" httpUri="http://treatment.plazi.org/id/554887B6E2497920FEB1473FFD1EB0AA" lastPageNumber="73" pageId="2" pageNumber="73">
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<subSubSection id="95FB652BE2497920FEB1473FFD79B35D" box="[281,642,1109,1133]" pageId="2" pageNumber="73" type="nomenclature">
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<paragraph id="DD5E36A0E2497920FEB1473FFD79B35D" blockId="2.[281,642,1109,1133]" box="[281,642,1109,1133]" pageId="2" pageNumber="73">
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<heading id="861681CCE2497920FEB1473FFD79B35D" box="[281,642,1109,1133]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="73" reason="2">
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<taxonomicName id="1AE14D23E2497920FEB1473FFD79B35D" authority="(Brenske, 1897)" baseAuthorityName="Brenske" baseAuthorityYear="1897" box="[281,642,1109,1133]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="amboinae">
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<emphasis id="EF95EAB2E2497920FEB1473FFE24B35D" bold="true" box="[281,479,1109,1133]" italics="true" pageId="2" pageNumber="73">Engertia amboinae</emphasis>
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(
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||
<bibRefCitation id="B9704B51E2497920FE43473FFD80B35D" author="Brenske, E." box="[491,635,1109,1133]" pageId="2" pageNumber="73" pagination="96 - 120" refId="ref6020" refString="Brenske, E. 1897. Neue Gattungen und Arten der Melolonthiden aus Afrika und Asien. Stettiner Entomologische Zeitung 58: 96 - 120." type="journal article" year="1897">Brenske, 1897</bibRefCitation>
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||
)
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</taxonomicName>
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</heading>
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||
</paragraph>
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</subSubSection>
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||
<subSubSection id="95FB652BE2497920FF3447FCFDFBB39E" box="[156,512,1174,1198]" pageId="2" pageNumber="73" type="reference_group">
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<paragraph id="DD5E36A0E2497920FF3447FCFDFBB39E" blockId="2.[156,512,1174,1198]" box="[156,512,1174,1198]" pageId="2" pageNumber="73">
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<treatmentCitationGroup id="FDF1118EE2497920FF3447FCFDFBB39E" box="[156,512,1174,1198]" pageId="2" pageNumber="73">
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<treatmentCitation id="5C4010B1E2497920FF3447FCFE07B39E" author="Brenske, E." box="[156,508,1174,1198]" page="111" pageId="2" pageNumber="73" year="1897">
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<taxonomicName id="1AE14D23E2497920FF3447FCFE07B39E" authority="Brenske, 1897: 111" authorityName="Brenske" authorityPageNumber="111" authorityYear="1897" box="[156,508,1174,1198]" class="Insecta" family="Melolonthidae" genus="Phila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="amboinae">
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<emphasis id="EF95EAB2E2497920FF3447FCFEC3B39E" box="[156,312,1174,1198]" italics="true" pageId="2" pageNumber="73">Phila amboinae</emphasis>
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<bibRefCitation id="B9704B51E2497920FE9547FCFE07B39E" author="Brenske, E." box="[317,508,1174,1198]" pageId="2" pageNumber="73" pagination="96 - 120" refId="ref6020" refString="Brenske, E. 1897. Neue Gattungen und Arten der Melolonthiden aus Afrika und Asien. Stettiner Entomologische Zeitung 58: 96 - 120." type="journal article" year="1897">Brenske, 1897: 111</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="95FB652BE2497920FF1D47BDFD32B261" pageId="2" pageNumber="73" type="materials_examined">
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<paragraph id="DD5E36A0E2497920FF1D47BDFD32B261" blockId="2.[156,767,1239,1946]" pageId="2" pageNumber="73">
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<emphasis id="EF95EAB2E2497920FF1D47BDFEEEB3DF" bold="true" box="[181,277,1239,1263]" pageId="2" pageNumber="73">Material.</emphasis>
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<materialsCitation id="6D893CFDE2497920FE8747B2FD51B3DF" box="[303,682,1239,1264]" country="Indonesia" location="Indonesia" pageId="2" pageNumber="73" specimenCount="1" specimenCount-female="1">
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<specimenCount id="CBE7FD29E2497920FE8747B2FE72B3DF" box="[303,393,1239,1264]" pageId="2" pageNumber="73" type="female">1 female</specimenCount>
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,
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<collectingCountry id="A5F67630E2497920FE3F47BDFE06B3DF" box="[407,509,1239,1263]" name="Indonesia" pageId="2" pageNumber="73">Indonesia</collectingCountry>
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,
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<taxonomicName id="1AE14D23E2497920FDA247B2FD51B3DF" authority=", Yapen I." authorityName="Yapen I." box="[522,682,1240,1263]" class="Insecta" family="Melolonthidae" genus="Apogonia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="papua">Papua, Yapen I</taxonomicName>
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</materialsCitation>
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.,
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<materialsCitation id="6D893CFDE2497920FD1447BDFDCEB220" country="Indonesia" location="Rosbori village" pageId="2" pageNumber="73" specimenCount="1">
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northern coast,
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<location id="D83E607BE2497920FEAD4792FE64B220" LSID="urn:lsid:plazi:treatment:554887B6E2497920FEB1473FFD1EB0AA:D83E607BE2497920FEAD4792FE64B220" box="[261,415,1272,1296]" country="Indonesia" name="Rosbori village" pageId="2" pageNumber="73">Rosbori village</location>
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, at light,
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<date id="A95F1060E2497920FE564792FDCEB220" box="[510,565,1272,1296]" pageId="2" pageNumber="73" value="2012-11-14">14.11</date>
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</materialsCitation>
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.2012, A
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<materialsCitation id="6D893CFDE2497920FD234792FD55B23F" box="[651,686,1272,1296]" country="Indonesia" isTailEnumeration="true" location="Rosbori village" pageId="2" pageNumber="73" specimenCount="1">. M</materialsCitation>
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. Prokofiev leg
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<materialsCitation id="6D893CFDE2497920FF424672FF0EB260" country="Indonesia" location="Biak" municipality="Yapen Is" pageId="2" pageNumber="73" specimenCount="1">
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.; 1 isolated elyton, Miosindi Islet between
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<location id="D83E607BE2497920FD0B4672FD2BB200" LSID="urn:lsid:plazi:treatment:554887B6E2497920FEB1473FFD1EB0AA:D83E607BE2497920FD0B4672FD2BB200" box="[675,720,1304,1328]" country="Indonesia" municipality="Yapen Is" name="Biak" pageId="2" pageNumber="73">Biak</location>
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and
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<collectingMunicipality id="3D3AACDAE2497920FF344653FF0EB260" box="[156,245,1337,1360]" pageId="2" pageNumber="73">Yapen Is</collectingMunicipality>
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</materialsCitation>
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., on sea coast,
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<date id="A95F1060E2497920FE2F4653FE0DB261" box="[391,502,1337,1361]" pageId="2" pageNumber="73" value="2012-11-15">15.11.2012</date>
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, A
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<materialsCitation id="6D893CFDE2497920FDBC4653FDCEB260" box="[532,565,1337,1361]" country="Indonesia" isTailEnumeration="true" location="Biak" municipality="Yapen Is" pageId="2" pageNumber="73" specimenCount="1">. M</materialsCitation>
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. Prokofiev leg.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="95FB652BE2497920FF1D4630FD1EB0AA" pageId="2" pageNumber="73" type="discussion">
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<paragraph id="DD5E36A0E2497920FF1D4630FD1EB0AA" blockId="2.[156,767,1239,1946]" pageId="2" pageNumber="73">
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<emphasis id="EF95EAB2E2497920FF1D4630FEE2B242" bold="true" box="[181,281,1370,1394]" pageId="2" pageNumber="73">Remarks.</emphasis>
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Five known species of
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<taxonomicName id="1AE14D23E2497920FD8D4630FF36B2A2" authority="Dalla Torre, 1912" authorityName="Dalla Torre" authorityYear="1912" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="genus">
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<emphasis id="EF95EAB2E2497920FD8D4630FD82B241" box="[549,633,1370,1393]" italics="true" pageId="2" pageNumber="73">Engertia</emphasis>
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Dalla Torre, 1912
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</taxonomicName>
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are distributed in the
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<collectingCountry id="A5F67630E2497920FE1D4610FDD2B2A2" box="[437,553,1402,1426]" name="Philippines" pageId="2" pageNumber="73">Philippines</collectingCountry>
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,
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<collectingCountry id="A5F67630E2497920FD9C4610FD62B2A2" box="[564,665,1402,1426]" name="Sri Lanka" pageId="2" pageNumber="73">Sri Lanka</collectingCountry>
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, southern
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<collectingCountry id="A5F67630E2497920FF3446F1FF05B283" box="[156,254,1435,1459]" name="Indonesia" pageId="2" pageNumber="73">Moluccas</collectingCountry>
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, and New
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<collectingCountry id="A5F67630E2497920FEDA46F1FE47B283" box="[370,444,1435,1459]" name="Guinea-Bissau" pageId="2" pageNumber="73">Guinea</collectingCountry>
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(
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<bibRefCitation id="B9704B51E2497920FE6346F1FDB8B282" author="Keith, D." box="[459,579,1434,1459]" pageId="2" pageNumber="73" pagination="90 - 94" refId="ref6134" refString="Keith, D. 2006. Reflexions sur le genre Engertia et description d'une nouvelle espece des Philippines. Lambillionea 106: 90 - 94." type="journal article" year="2006">Keith 2006</bibRefCitation>
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). Surprisingly, the Schouten specimens represent not
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<taxonomicName id="1AE14D23E2497920FE5346D6FD04B2E3" authority="(Moser, 1913)" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[507,767,1467,1491]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="papuana">
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<emphasis id="EF95EAB2E2497920FE5346D6FD96B2E3" box="[507,621,1468,1491]" italics="true" pageId="2" pageNumber="73">E. papuana</emphasis>
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(Moser, 1913)
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</taxonomicName>
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known from the main island of New
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<collectingCountry id="A5F67630E2497920FD9A46B6FD85B2C4" box="[562,638,1500,1524]" name="Guinea-Bissau" pageId="2" pageNumber="73">Guinea</collectingCountry>
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, but
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<taxonomicName id="1AE14D23E2497920FD1046B6FF25B124" authorityName="Brenske" authorityYear="1897" class="Insecta" family="Melolonthidae" genus="Phila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="amboinae">
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<emphasis id="EF95EAB2E2497920FD1046B6FF25B124" italics="true" pageId="2" pageNumber="73">E. amboinae</emphasis>
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</taxonomicName>
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which has been known hitherto only from
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<collectingCountry id="A5F67630E2497920FD194696FD05B124" box="[689,766,1532,1556]" name="Indonesia" pageId="2" pageNumber="73">Ambon</collectingCountry>
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I. (
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<bibRefCitation id="B9704B51E2497920FF124577FEC9B105" author="Keith, D." box="[186,306,1565,1589]" pageId="2" pageNumber="73" pagination="90 - 94" refId="ref6134" refString="Keith, D. 2006. Reflexions sur le genre Engertia et description d'une nouvelle espece des Philippines. Lambillionea 106: 90 - 94." type="journal article" year="2006">Keith 2006</bibRefCitation>
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). The females of
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<taxonomicName id="1AE14D23E2497920FE4F4577FD9DB105" authorityName="Brenske" authorityYear="1897" box="[487,614,1565,1589]" class="Insecta" family="Melolonthidae" genus="Phila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="amboinae">
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<emphasis id="EF95EAB2E2497920FE4F4577FD9DB105" box="[487,614,1565,1589]" italics="true" pageId="2" pageNumber="73">E. amboinae</emphasis>
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</taxonomicName>
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can be distinguished easily from those of
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<taxonomicName id="1AE14D23E2497920FE664554FDB8B165" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[462,579,1598,1621]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="papuana">
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<emphasis id="EF95EAB2E2497920FE664554FDB8B165" box="[462,579,1598,1621]" italics="true" pageId="2" pageNumber="73">E. papuana</emphasis>
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</taxonomicName>
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by the shape and density of the scales on the elytra which are larger, ovate, and more numerous and more closely packed in contrast to the sparse lanceolate scales of
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<taxonomicName id="1AE14D23E2497920FE7145CAFDB5B187" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[473,590,1696,1719]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="papuana">
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<emphasis id="EF95EAB2E2497920FE7145CAFDB5B187" box="[473,590,1696,1719]" italics="true" pageId="2" pageNumber="73">E. papuana</emphasis>
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||
</taxonomicName>
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||
(
|
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<figureCitation id="45DA2A25E2497920FDF545F5FD42B187" box="[605,697,1695,1719]" captionStart="Figs 1–3" captionStartId="2.[157,194,1006,1028]" captionTargetBox="[244,1343,205,968]" captionTargetId="figure-447@2.[244,1343,205,968]" captionTargetPageId="2" captionText="Figs 1–3. Scalation of the elytral interspaces in Engertia amboinae, Yapen I. (1); E. papuana, Papua New Guinea: Sentani (2), and E. setifera, Ceram I. (3). Scale bar (common)=1mm." figureDoi="http://doi.org/10.5281/zenodo.5737670" httpUri="https://zenodo.org/record/5737670/files/figure.png" pageId="2" pageNumber="73">Figs 1–2</figureCitation>
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||
). Both Schouten specimens agree well in this character with the
|
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<typeStatus id="025A8802E2497920FF34458AFF33B1C8" box="[156,200,1760,1784]" pageId="2" pageNumber="73">type</typeStatus>
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||
of
|
||
<taxonomicName id="1AE14D23E2497920FF41458BFE9EB1C8" authorityName="Brenske" authorityYear="1897" box="[233,357,1760,1784]" class="Insecta" family="Melolonthidae" genus="Phila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="amboinae">
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||
<emphasis id="EF95EAB2E2497920FF41458BFE9EB1C8" box="[233,357,1760,1784]" italics="true" pageId="2" pageNumber="73">E. amboinae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="EF95EAB2E2497920FEDC458AFE77B1C8" box="[372,396,1760,1784]" italics="true" pageId="2" pageNumber="73">cf.</emphasis>
|
||
<bibRefCitation id="B9704B51E2497920FE39458AFDF8B1C8" author="Keith, D." box="[401,515,1760,1784]" pageId="2" pageNumber="73" pagination="90 - 94" refId="ref6134" refString="Keith, D. 2006. Reflexions sur le genre Engertia et description d'une nouvelle espece des Philippines. Lambillionea 106: 90 - 94." type="journal article" year="2006">Keith 2006</bibRefCitation>
|
||
). The third Indo-Australian species of this genus,
|
||
<taxonomicName id="1AE14D23E2497920FE18446BFD55B029" authority="(Moser, 1913)" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[432,686,1792,1817]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="setifera">
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<emphasis id="EF95EAB2E2497920FE18446BFDEFB028" box="[432,532,1792,1816]" italics="true" pageId="2" pageNumber="73">E. setifera</emphasis>
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(Moser, 1913)
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</taxonomicName>
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, can be easily distinguished by the presence of setae instead of scales on the elytra (
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<figureCitation id="45DA2A25E2497920FE9C442BFE88B06A" box="[308,371,1857,1882]" captionStart="Figs 1–3" captionStartId="2.[157,194,1006,1028]" captionTargetBox="[244,1343,205,968]" captionTargetId="figure-447@2.[244,1343,205,968]" captionTargetPageId="2" captionText="Figs 1–3. Scalation of the elytral interspaces in Engertia amboinae, Yapen I. (1); E. papuana, Papua New Guinea: Sentani (2), and E. setifera, Ceram I. (3). Scale bar (common)=1mm." figureDoi="http://doi.org/10.5281/zenodo.5737670" httpUri="https://zenodo.org/record/5737670/files/figure.png" pageId="2" pageNumber="73">Fig. 3</figureCitation>
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). The Yapen I. and Miosindi Islet records greatly extend the known distribution of this species, which is probably widely distributed in eastern
|
||
<collectingCountry id="A5F67630E2497920FDD244E8FD1AB0AA" box="[634,737,1922,1946]" name="Indonesia" pageId="2" pageNumber="73">Indonesia</collectingCountry>
|
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.
|
||
</paragraph>
|
||
</subSubSection>
|
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</treatment>
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</document> |