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<document id="9EBECE38635EAB6B6AA657F086027034" ID-CLB-Dataset="25410" ID-DOI="10.11646/zootaxa.4674.2.6" ID-GBIF-Dataset="98247436-820b-480f-9488-86b44184cfd6" ID-ISSN="1175-5326" ID-Zenodo-Dep="3458973" ID-ZooBank="C3601614-6287-4F18-9684-ED84C6BE705E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1569310420439" checkinUser="plazi" docAuthor="Shibatta, Oscar A." docDate="2019" docId="5541526FFFC0FF9A9E84F8C5FAFEFA0A" docLanguage="en" docName="zootaxa.4674.2.6.pdf" docOrigin="Zootaxa 4674 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Batrochoglanis castaneus Shibatta 2019, new species" docType="treatment" docVersion="9" lastPageNumber="256" masterDocId="A9782A17FFC1FF979E13FFF1FFE4FFAE" masterDocTitle="New species of bumblebee catfish of the genus Batrochoglanis Gill, 1858 (Siluriformes: Pseudopimelodidae) from the Aripuanã River basin, Mato Grosso, Brazil" masterLastPageNumber="263" masterPageNumber="243" pageNumber="244" updateTime="1698756334229" updateUser="ExternalLinkService">
<mods:mods id="3DE6896C818D7C661BA9458A390B0501" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6CB38283CDC95340B90FCEBBC1969D67">
<mods:title id="BA510B23B4EE84EEB47E2D3298A37AEC">New species of bumblebee catfish of the genus Batrochoglanis Gill, 1858 (Siluriformes: Pseudopimelodidae) from the Aripuanã River basin, Mato Grosso, Brazil</mods:title>
</mods:titleInfo>
<mods:name id="F27955C6F7B0B4F1CCE8A00EC84EE84A" type="personal">
<mods:role id="A93824B4DEA7F1BF96FB75AF7EA10B62">
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<mods:namePart id="AB43B5635770ACD0A92EFB6406F8D2CB">Shibatta, Oscar A.</mods:namePart>
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<mods:title id="BA914D5B7B2BC124EB30D675119F5160">Zootaxa</mods:title>
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<mods:part id="DC372FEADB7CD61B5D20C1CF3121D59D">
<mods:date id="E70560420F93C8FC52854E9CF1B6F7E0">2019</mods:date>
<mods:detail id="A87DC5D3D5287421EAA5BA2D3E107FBE" type="pubDate">
<mods:number id="4A09460D40CF1D631CE4B5D17D549121">2019-09-24</mods:number>
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<mods:number id="8B52EABE1A22A85D029CA9398933E97C">4674</mods:number>
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<mods:number id="55A6346D431E07DE188BB0844CBFDD80">2</mods:number>
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<mods:classification id="0969793191568CDBC9C4322DB7775E2D">journal article</mods:classification>
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<mods:identifier id="BBD1A52CBCC798FB879E6E771BC4882D" type="DOI">10.11646/zootaxa.4674.2.6</mods:identifier>
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<treatment id="5541526FFFC0FF9A9E84F8C5FAFEFA0A" ID-DOI="http://doi.org/10.5281/zenodo.5621550" ID-GBIF-Taxon="159398147" ID-Zenodo-Dep="5621550" LSID="urn:lsid:plazi:treatment:5541526FFFC0FF9A9E84F8C5FAFEFA0A" httpUri="http://treatment.plazi.org/id/5541526FFFC0FF9A9E84F8C5FAFEFA0A" lastPageId="13" lastPageNumber="256" pageId="1" pageNumber="244">
<subSubSection id="95F2B0F2FFC0FF969E84F8C5FD8AF8E0" box="[151,622,1844,1871]" pageId="1" pageNumber="244" type="nomenclature">
<paragraph id="DD57E379FFC0FF969E84F8C5FD8AF8E0" blockId="1.[151,995,1844,1936]" box="[151,622,1844,1871]" pageId="1" pageNumber="244">
<heading id="861F5415FFC0FF969E84F8C5FD8AF8E0" bold="true" box="[151,622,1844,1871]" fontSize="11" level="1" pageId="1" pageNumber="244" reason="1">
<emphasis id="EF9C3F6BFFC0FF969E84F8C5FD8AF8E0" bold="true" box="[151,622,1844,1871]" pageId="1" pageNumber="244">
<taxonomicName id="1AE898FAFFC0FF969E84F8C5FE2BF8E1" authority="Shibatta, 2019" authorityName="Shibatta" authorityYear="2019" box="[151,463,1844,1871]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="244" phylum="Chordata" rank="species" species="castaneus" status="sp. nov.">
<emphasis id="EF9C3F6BFFC0FF969E84F8C5FE2BF8E1" bold="true" box="[151,463,1844,1871]" italics="true" pageId="1" pageNumber="244">Batrochoglanis castaneus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="F4AF8210FFC0FF969FCDF8C5FD8AF8E0" box="[478,622,1844,1870]" pageId="1" pageNumber="244" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="95F2B0F2FFC0FF969E84F8A9FC07F83E" pageId="1" pageNumber="244" type="description">
<paragraph id="DD57E379FFC0FF969E84F8A9FE79F8DC" blockId="1.[151,995,1844,1936]" box="[151,413,1880,1906]" pageId="1" pageNumber="244">
(
<figureCitation id="45D3FFFCFFC0FF969E8CF8A9FED1F8DC" box="[159,309,1880,1906]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-10="FIGURE 11" captionStart-11="FIGURE 12" captionStart-12="FIGURE 13" captionStart-13="FIGURE 14" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStart-9="FIGURE 10" captionStartId-0="4.[151,250,1713,1738]" captionStartId-1="5.[151,250,1129,1154]" captionStartId-10="12.[151,250,1099,1124]" captionStartId-11="12.[151,250,1892,1917]" captionStartId-12="13.[151,250,1167,1192]" captionStartId-13="14.[151,248,1947,1972]" captionStartId-2="5.[151,250,1923,1948]" captionStartId-3="6.[151,250,930,955]" captionStartId-4="6.[151,250,1868,1893]" captionStartId-5="7.[151,250,1658,1683]" captionStartId-6="8.[151,250,1022,1047]" captionStartId-7="9.[151,250,1151,1176]" captionStartId-8="10.[151,250,1137,1162]" captionStartId-9="11.[151,250,1615,1640]" captionTargetBox-0="[151,1436,174,1693]" captionTargetBox-1="[162,1425,276,1102]" captionTargetBox-10="[200,1393,243,1062]" captionTargetBox-11="[191,1384,1184,1861]" captionTargetBox-12="[162,1417,190,1137]" captionTargetBox-13="[416,1166,216,1898]" captionTargetBox-2="[167,1417,1213,1896]" captionTargetBox-3="[163,1424,186,901]" captionTargetBox-4="[164,1421,1006,1838]" captionTargetBox-5="[162,1413,712,1624]" captionTargetBox-6="[161,1429,384,989]" captionTargetBox-7="[173,1414,194,1108]" captionTargetBox-8="[165,1421,187,1101]" captionTargetBox-9="[219,1366,181,1587]" captionTargetId-0="figure@4.[151,1436,174,1693]" captionTargetId-1="figure@5.[151,1436,265,1109]" captionTargetId-10="figure@12.[183,1405,229,1079]" captionTargetId-11="figure@12.[185,1402,1175,1873]" captionTargetId-12="figure@13.[151,1436,174,1148]" captionTargetId-13="figure@14.[373,1215,179,1920]" captionTargetId-2="figure@5.[151,1436,1204,1904]" captionTargetId-3="figure@6.[151,1436,174,910]" captionTargetId-4="figure@6.[151,1434,1006,1848]" captionTargetId-5="figure@7.[151,1436,697,1639]" captionTargetId-6="figure@8.[151,1435,373,1002]" captionTargetId-7="figure@9.[151,1436,174,1131]" captionTargetId-8="figure@10.[151,1435,174,1117]" captionTargetId-9="figure@11.[204,1382,174,1596]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-10="12" captionTargetPageId-11="12" captionTargetPageId-12="13" captionTargetPageId-13="14" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="8" captionTargetPageId-7="9" captionTargetPageId-8="10" captionTargetPageId-9="11" captionText-0="FIGURE 1. Holotype of Batrochoglanis castaneus, INPA 8059, 58,3 mm SL, Aripuanã River, Igarapé do Castanhal, MT." captionText-1="FIGURE 2. Cranium of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. A) dorsal view; b) ventral view. Light blue = cartilage; dark gray = fontanel or foramen. Scale bar = 5 mm." captionText-10="FIGURE 11. Dorsal fin of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. A) dorsal view of nuchal plate and first two rays; b) lateral view. Light blue = cartilage. Scale bar = 1 mm." captionText-11="FIGURE 12. Lateral view of anal-fin skeleton of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. Light blue = cartilage. Scale bar = 1 mm." captionText-12="FIGURE 13. Caudal skeleton of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. Light blue = cartilage. Scale bar = 1 mm." captionText-13="FIGURE 14. Laterosensory system of Batrochoglanis castaneus, INPA 8059, 58.3 mm SL, holotype. Scale bar = 5 mm. Open circle = laterosensory pore (i = infraorbital; ll = lateral line; pm = preoperculomandibular; po = postotic; s = supraorbital); dark dot = neuromast (al = anterior line; dtl = dorsal trunk line; ml = maxillary line; nl = nasal line; stal = supratemporal accessory line)." captionText-2="FIGURE 3. Autopalatine and maxilla of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. A) dorsal view; b) lateral view; c) ventral view; d) medial view. Light blue = cartilage. Scale bar = 1 mm." captionText-3="FIGURE 4. Lower jaw of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. Light blue = cartilage. Scale bar = 1 mm." captionText-4="FIGURE 5. Suspensorium of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. Light blue = cartilage; dark gray = foramen. Scale bar = 1 mm." captionText-5="FIGURE 6. Hyoid arch of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype.A) lateral view; b) dorsal view of the anterior region. Light blue = cartilage. Scale bar = 5 mm." captionText-6="FIGURE 7. Branchial arches of Batrochoglanis castaneus, a) dorsal view and b) ventral view of right epibranchials and pharyngobranchials, MZUEL 20113, 50.8 mm SL, paratype. Light blue = cartilage." captionText-7="FIGURE 8. Ventral view of the left pectoral girdle of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. Light blue = cartilage. Scale bar = 5 mm." captionText-8="FIGURE 9. Ventral view of right pelvic girdle of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. Light blue = cartilage. Scale bar = 1 mm." captionText-9="FIGURE 10. Complex vertebra of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. A) Ventral view; b) ventral-lateral view of the anterior region. Scale bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3458975" figureDoi-1="http://doi.org/10.5281/zenodo.3458977" figureDoi-10="http://doi.org/10.5281/zenodo.3458997" figureDoi-11="http://doi.org/10.5281/zenodo.3458999" figureDoi-12="http://doi.org/10.5281/zenodo.3459001" figureDoi-13="http://doi.org/10.5281/zenodo.3459003" figureDoi-2="http://doi.org/10.5281/zenodo.3458979" figureDoi-3="http://doi.org/10.5281/zenodo.3458981" figureDoi-4="http://doi.org/10.5281/zenodo.3458983" figureDoi-5="http://doi.org/10.5281/zenodo.3458985" figureDoi-6="http://doi.org/10.5281/zenodo.3458987" figureDoi-7="http://doi.org/10.5281/zenodo.3458989" figureDoi-8="http://doi.org/10.5281/zenodo.3458993" figureDoi-9="http://doi.org/10.5281/zenodo.3458995" httpUri-0="https://zenodo.org/record/3458975/files/figure.png" httpUri-1="https://zenodo.org/record/3458977/files/figure.png" httpUri-10="https://zenodo.org/record/3458997/files/figure.png" httpUri-11="https://zenodo.org/record/3458999/files/figure.png" httpUri-12="https://zenodo.org/record/3459001/files/figure.png" httpUri-13="https://zenodo.org/record/3459003/files/figure.png" httpUri-2="https://zenodo.org/record/3458979/files/figure.png" httpUri-3="https://zenodo.org/record/3458981/files/figure.png" httpUri-4="https://zenodo.org/record/3458983/files/figure.png" httpUri-5="https://zenodo.org/record/3458985/files/figure.png" httpUri-6="https://zenodo.org/record/3458987/files/figure.png" httpUri-7="https://zenodo.org/record/3458989/files/figure.png" httpUri-8="https://zenodo.org/record/3458993/files/figure.png" httpUri-9="https://zenodo.org/record/3458995/files/figure.png" pageId="1" pageNumber="244">Figures 1–14</figureCitation>
,
<tableCitation id="906AD6C2FFC0FF969F52F8A9FE76F8DC" box="[321,402,1880,1906]" captionStart="TABLE 1" captionStartId="3.[151,239,658,682]" captionTargetPageId="3" captionText="TABLE 1. Morphometric data of Batrochoglanis castaneus (n = 9). Min = minimum, max = maximum, SD = standard deviation. The holotype is included in the statistics." httpUri="http://table.plazi.org/id/8997B3F1FFC2FF949E84FD63FD58FD61" pageId="1" pageNumber="244" tableUuid="8997B3F1FFC2FF949E84FD63FD58FD61">Table 1</tableCitation>
)
</paragraph>
<paragraph id="DD57E379FFC0FF969E84F884FC07F83E" blockId="1.[151,995,1844,1936]" box="[151,995,1909,1936]" pageId="1" pageNumber="244">
<uri id="A979EF7BFFC0FF969E84F884FC07F83E" box="[151,995,1909,1936]" pageId="1" pageNumber="244">
urn:lsid:zoobank.org:act:
<uuid id="A94ED9ACFFC0FF969FB9F884FC07F83E" box="[426,995,1909,1936]" pageId="1" pageNumber="244">0BF7C8B5-F053-4D05-AADC-75DADC1FFFE8</uuid>
</uri>
</paragraph>
</subSubSection>
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<paragraph id="DD57E379FFC0FF969E84F849FEF7F858" blockId="1.[151,1436,1975,2039]" pageId="1" pageNumber="244">
<materialsCitation id="6D80E924FFC0FF969E84F849FEF7F858" ID-GBIF-Occurrence="2417904855" collectingDate="1976-11-22" collectionCode="INPA" collectorName="de Ictiologia" county="Equipe de Ictiologia" location="Igarape do Castanhal" municipality="Aripuana River" pageId="1" pageNumber="244" specimenCode="INPA 8059" specimenCount="1" typeStatus="holotype">
<emphasis id="EF9C3F6BFFC0FF969E84F849FEEFF87C" bold="true" box="[151,267,1976,2002]" pageId="1" pageNumber="244">
<typeStatus id="02535DDBFFC0FF969E84F849FEE0F87C" box="[151,260,1976,2002]" pageId="1" pageNumber="244" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="8D4E4B02FFC0FF969F01F849FE75F87C" box="[274,401,1975,2002]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="1" pageNumber="244">INPA 8059</specimenCode>
,
<quantity id="1A104E9CFFC0FF969F88F846FE1BF87D" box="[411,511,1975,2003]" metricMagnitude="-2" metricUnit="m" metricValue="5.83" pageId="1" pageNumber="244" unit="mm" value="58.3">58.3 mm</quantity>
SL,
<location id="D837B5A2FFC0FF969C27F849FCC6F87C" LSID="urn:lsid:plazi:treatment:5541526FFFC0FF9A9E84F8C5FAFEFA0A:D837B5A2FFC0FF969C27F849FCC6F87C" box="[564,802,1976,2002]" county="Equipe de Ictiologia" municipality="Aripuana River" name="Igarape do Castanhal" pageId="1" pageNumber="244">Igarapé do Castanhal</location>
,
<collectingMunicipality id="3D337903FFC0FF969D38F849FC3CF87C" box="[811,984,1976,2002]" pageId="1" pageNumber="244">Aripuanã River</collectingMunicipality>
basin,
<collectingCounty id="34369BF5FFC0FF969A3BF849FAEEF87C" box="[1064,1290,1976,2002]" pageId="1" pageNumber="244">
Equipe
<collectorName id="701D86AFFFC0FF969A6DF849FAEEF87C" box="[1150,1290,1976,2002]" pageId="1" pageNumber="244">de Ictiologia</collectorName>
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do
<collectionCode id="BBF97BBCFFC0FF969B27F849FA92F87C" box="[1332,1398,1976,2002]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="1" pageNumber="244">INPA</collectionCode>
,
<date id="A956C5B9FFC0FF969B93F849FEEBF858" pageId="1" pageNumber="244" value="1976-11-22">
<collectingDate id="B9123C51FFC0FF969B93F849FEEBF858" pageId="1" pageNumber="244" value="1976-11-22">22 Nov. 1976</collectingDate>
</date>
.
</materialsCitation>
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<paragraph id="DD57E379FFC3FF959ED4FF68FBF5FF55" blockId="2.[151,1437,152,2015]" pageId="2" pageNumber="245">
<materialsCitation id="6D80E924FFC3FF959ED4FF68FE7BFF79" ID-GBIF-Occurrence="2417904859" collectionCode="INPA" country="Brazil" county="Igarape do Castanhal" location="Equipe de Ictiologia" municipality="Aripuana River" pageId="2" pageNumber="245" specimenCode="INPA 8054, 1" specimenCount="1" stateProvince="Mato Grosso" typeStatus="paratype">
<emphasis id="EF9C3F6BFFC3FF959ED4FF68FEA2FF1D" bold="true" box="[199,326,153,179]" pageId="2" pageNumber="245">
<typeStatus id="02535DDBFFC3FF959ED4FF68FEDBFF1D" box="[199,319,153,179]" pageId="2" pageNumber="245" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="A5FFA3E9FFC3FF959F43FF68FE7CFF1D" box="[336,408,153,179]" name="Brazil" pageId="2" pageNumber="245">Brazil</collectingCountry>
.
<collectingRegion id="1F2C2D9BFFC3FF959FB5FF68FDDAFF1D" box="[422,574,153,179]" country="Brazil" name="Mato Grosso" pageId="2" pageNumber="245">Mato Grosso</collectingRegion>
.
<specimenCode id="8D4E4B02FFC3FF959C5FFF68FD2AFF1D" box="[588,718,152,179]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="245">INPA 8054</specimenCode>
,
<specimenCode id="8D4E4B02FFC3FF959CCFFF68FD0EFF1D" box="[732,746,153,179]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="245">1</specimenCode>
,
<quantity id="1A104E9CFFC3FF959CEFFF69FC87FF1D" box="[764,867,152,179]" metricMagnitude="-2" metricUnit="m" metricValue="5.43" pageId="2" pageNumber="245" unit="mm" value="54.3">54.3 mm</quantity>
CP,
<collectingCounty id="34369BF5FFC3FF959D8EFF68FB76FF1D" box="[925,1170,153,179]" pageId="2" pageNumber="245">Igarapé do Castanhal</collectingCounty>
,
<collectingMunicipality id="3D337903FFC3FF959A8CFF68FAB4FF1D" box="[1183,1360,153,179]" pageId="2" pageNumber="245">Aripuanã River</collectingMunicipality>
basin,
<location id="D837B5A2FFC3FF959E84FF4CFE9FFF79" LSID="urn:lsid:plazi:treatment:5541526FFFC0FF9A9E84F8C5FAFEFA0A:D837B5A2FFC3FF959E84FF4CFE9FFF79" box="[151,379,189,215]" country="Brazil" county="Igarape do Castanhal" municipality="Aripuana River" name="Equipe de Ictiologia" pageId="2" pageNumber="245" stateProvince="Mato Grosso">Equipe de Ictiologia</location>
do
</materialsCitation>
<materialsCitation id="6D80E924FFC3FF959FB4FF4CFD74FF79" ID-GBIF-Occurrence="2417904858" box="[423,656,189,215]" collectingDate="1976-08-22" collectionCode="INPA" country="Brazil" pageId="2" pageNumber="245" specimenCount="1" stateProvince="Mato Grosso" typeStatus="paratype">
<collectionCode id="BBF97BBCFFC3FF959FB4FF4CFE0DFF79" box="[423,489,189,215]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="245">INPA</collectionCode>
,
<date id="A956C5B9FFC3FF959FE7FF4CFD74FF79" box="[500,656,189,215]" pageId="2" pageNumber="245" value="1976-08-22">
<collectingDate id="B9123C51FFC3FF959FE7FF4CFD74FF79" box="[500,656,189,215]" pageId="2" pageNumber="245" value="1976-08-22">22 Aug. 1976</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="6D80E924FFC3FF959C8FFF4CFF2FFF55" ID-GBIF-Occurrence="2417904856" collectingDate="1976-11-22" collectionCode="INPA" collectorName="de Ictiologia" country="Brazil" county="Equipe de Ictiologia" location="Igarape do Castanhal" municipality="Aripuana River" pageId="2" pageNumber="245" specimenCode="INPA 58786, 13" specimenCount="1" stateProvince="Mato Grosso" typeStatus="paratype">
<specimenCode id="8D4E4B02FFC3FF959C8FFF4CFCCEFF79" box="[668,810,188,215]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="245">INPA 58786</specimenCode>
,
<specimenCode id="8D4E4B02FFC3FF959D26FF4CFCB5FF79" box="[821,849,189,215]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="245">13</specimenCode>
(1 C&amp;S),
<quantity id="1A104E9CFFC3FF959DDBFF4DFB88FF79" box="[968,1132,188,215]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="5.07" metricValueMin="2.55" pageId="2" pageNumber="245" unit="mm" value="38.1" valueMax="50.7" valueMin="25.5">25.5–50.7 mm</quantity>
CP, same data as the holotype
</materialsCitation>
.
<materialsCitation id="6D80E924FFC3FF959EC6FF10FBE9FF55" ID-GBIF-Occurrence="2417904857" box="[213,1037,224,251]" collectingDate="1976-11-22" collectionCode="MZUEL" collectorName="de Ictiologia" country="Brazil" county="Equipe de Ictiologia" location="Igarape do Castanhal" municipality="Aripuana River" pageId="2" pageNumber="245" specimenCode="MZUEL 20113, 3" specimenCount="1" stateProvince="Mato Grosso" typeStatus="paratype">
<specimenCode id="8D4E4B02FFC3FF959EC6FF10FE60FF55" box="[213,388,225,251]" collectionCode="MZUEL" pageId="2" pageNumber="245">MZUEL 20113</specimenCode>
,
<specimenCode id="8D4E4B02FFC3FF959F9CFF10FE79FF55" box="[399,413,225,251]" collectionCode="MZUEL" pageId="2" pageNumber="245">3</specimenCode>
(1 C&amp;S),
<quantity id="1A104E9CFFC3FF959C02FF11FD50FF55" box="[529,692,224,251]" metricMagnitude="-2" metricUnit="m" metricValue="5.33" metricValueMax="5.58" metricValueMin="5.08" pageId="2" pageNumber="245" unit="mm" value="53.3" valueMax="55.8" valueMin="50.8">50.8–55.8 mm</quantity>
SL, same data as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95F2B0F2FFC3FF959ED4FEF5FCB2FD61" pageId="2" pageNumber="245" type="diagnosis">
<paragraph id="DD57E379FFC3FF959ED4FEF5FCB2FD61" blockId="2.[151,1437,152,2015]" pageId="2" pageNumber="245">
<emphasis id="EF9C3F6BFFC3FF959ED4FEF5FEA6FEB1" bold="true" box="[199,322,260,287]" pageId="2" pageNumber="245">Diagnosis.</emphasis>
<taxonomicName id="1AE898FAFFC3FF959F59FEF4FD88FEB1" authorityName="Shibatta" authorityYear="2019" box="[330,620,261,287]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="245" phylum="Chordata" rank="species" species="castaneus">
<emphasis id="EF9C3F6BFFC3FF959F59FEF4FD88FEB1" box="[330,620,261,287]" italics="true" pageId="2" pageNumber="245">Batrochoglanis castaneus</emphasis>
</taxonomicName>
differs from its congeners, except
<taxonomicName id="1AE898FAFFC3FF959DE9FEF7FB95FEB1" baseAuthorityName="Valenciennes" baseAuthorityYear="1840" box="[1018,1137,262,287]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="245" phylum="Chordata" rank="species" species="raninus">
<emphasis id="EF9C3F6BFFC3FF959DE9FEF7FB95FEB1" box="[1018,1137,262,287]" italics="true" pageId="2" pageNumber="245">B. raninus</emphasis>
</taxonomicName>
and
<taxonomicName id="1AE898FAFFC3FF959AB9FEF7FA94FEB1" baseAuthorityName="Regan" baseAuthorityYear="1913" box="[1194,1392,262,287]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="245" phylum="Chordata" rank="species" species="transmontanus">
<emphasis id="EF9C3F6BFFC3FF959AB9FEF7FA94FEB1" box="[1194,1392,262,287]" italics="true" pageId="2" pageNumber="245">B. transmontanus</emphasis>
</taxonomicName>
, by the dark brown color of the body, with a light brown band on the nape, a light blotch immediately posterior to the vertical through the terminus of the dorsal-fin base, and on the terminus of the adipose fin (vs. body mottled with dark brown spots in
<taxonomicName id="1AE898FAFFC3FF959F6BFE83FE08FE25" baseAuthorityName="Eigenmann" baseAuthorityYear="1912" box="[376,492,369,395]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="245" phylum="Chordata" rank="species" species="villosus">
<emphasis id="EF9C3F6BFFC3FF959F6BFE83FE08FE25" box="[376,492,369,395]" italics="true" pageId="2" pageNumber="245">B. villosus</emphasis>
</taxonomicName>
and
<taxonomicName id="1AE898FAFFC3FF959C33FE83FD1BFE25" baseAuthorityName="Güntert" baseAuthorityYear="1942" box="[544,767,369,395]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="245" phylum="Chordata" rank="species" species="acanthochiroides">
<emphasis id="EF9C3F6BFFC3FF959C33FE83FD1BFE25" box="[544,767,369,395]" italics="true" pageId="2" pageNumber="245">B. acanthochiroides</emphasis>
</taxonomicName>
), and caudal fin hyaline with a dark brown stripe (vs. caudal fin completely dark in
<taxonomicName id="1AE898FAFFC3FF959F8CFE67FDD3FE01" authorityName="Shibatta &amp; Pavanelli" authorityYear="2005" box="[415,567,405,431]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="245" phylum="Chordata" rank="species" species="melanurus">
<emphasis id="EF9C3F6BFFC3FF959F8CFE67FDD3FE01" box="[415,567,405,431]" italics="true" pageId="2" pageNumber="245">B. melanurus</emphasis>
</taxonomicName>
).
<taxonomicName id="1AE898FAFFC3FF959C42FE64FC90FE01" authorityName="Shibatta" authorityYear="2019" box="[593,884,405,431]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="245" phylum="Chordata" rank="species" species="castaneus">
<emphasis id="EF9C3F6BFFC3FF959C42FE64FC90FE01" box="[593,884,405,431]" italics="true" pageId="2" pageNumber="245">Batrochoglanis castaneus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="1AE898FAFFC3FF959A1CFE67FB62FE01" baseAuthorityName="Valenciennes" baseAuthorityYear="1840" box="[1039,1158,406,431]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="245" phylum="Chordata" rank="species" species="raninus">
<emphasis id="EF9C3F6BFFC3FF959A1CFE67FB62FE01" box="[1039,1158,406,431]" italics="true" pageId="2" pageNumber="245">B. raninus</emphasis>
</taxonomicName>
by the longer maxillary barbel (131.8%±10.1 of HL
<emphasis id="EF9C3F6BFFC3FF959FF3FE4BFE1BFE7D" box="[480,511,442,467]" italics="true" pageId="2" pageNumber="245">vs.</emphasis>
87.7%±9.3, p &lt;0.01), longer prepelvic length (57.4%±0.7 of SL
<emphasis id="EF9C3F6BFFC3FF959AE1FE4BFAF5FE7D" box="[1266,1297,442,467]" italics="true" pageId="2" pageNumber="245">vs.</emphasis>
55.9%±1.4, p&lt;0.01), greater prepectoral length (31.5%±1.6 of SL
<emphasis id="EF9C3F6BFFC3FF959CEFFE2FFCFFFE59" box="[764,795,478,503]" italics="true" pageId="2" pageNumber="245">vs.</emphasis>
29.5%±1.04, p&lt;0.01), anal-fin base length (12.0±0.5
<emphasis id="EF9C3F6BFFC3FF959B6DFE2FFA78FE59" box="[1406,1436,478,503]" italics="true" pageId="2" pageNumber="245">vs.</emphasis>
13.4±0.92, p&lt;0.01), and head depth (31.1±2.5
<emphasis id="EF9C3F6BFFC3FF959C8AFDF3FD53FDB5" box="[665,695,514,539]" italics="true" pageId="2" pageNumber="245">vs.</emphasis>
34.6±2.2, p&lt;0.01). The new species differs from
<taxonomicName id="1AE898FAFFC3FF959ACBFDF3FA78FDB5" baseAuthorityName="Regan" baseAuthorityYear="1913" box="[1240,1436,514,539]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="245" phylum="Chordata" rank="species" species="transmontanus">
<emphasis id="EF9C3F6BFFC3FF959ACBFDF3FA78FDB5" box="[1240,1436,514,539]" italics="true" pageId="2" pageNumber="245">B. transmontanus</emphasis>
</taxonomicName>
by the longer caudal-fin dorsal lobe, presence of translucent spot or stripe on pelvic, dorsal, and anal fins, longer maxillary barbel length (131.8%±10.1 of HL
<emphasis id="EF9C3F6BFFC3FF959C8BFDBBFD53FDCD" box="[664,695,586,611]" italics="true" pageId="2" pageNumber="245">vs.</emphasis>
87.7%±9.3, p&lt;0.01), longer pectoral-fin spine length (21.0%±1.7 of SL
<emphasis id="EF9C3F6BFFC3FF959ECEFD9FFF1FFD29" box="[221,251,622,647]" italics="true" pageId="2" pageNumber="245">vs.</emphasis>
16.1%±0.85, p&lt;0.01), longer dorsal-fin spine length (16.5%±1.1 of SL
<emphasis id="EF9C3F6BFFC3FF959A30FD9FFBA5FD29" box="[1059,1089,622,647]" italics="true" pageId="2" pageNumber="245">vs.</emphasis>
12.1±1.6, p&lt;0.01), greater eye diameter (12.2%±1.3 of HL
<emphasis id="EF9C3F6BFFC3FF959FC3FD63FE0AFD05" box="[464,494,658,683]" italics="true" pageId="2" pageNumber="245">vs.</emphasis>
7.7%±1.0, p&lt;0.01), smaller head depth (31.1%±2.5 of HL
<emphasis id="EF9C3F6BFFC3FF959A68FD63FB7DFD05" box="[1147,1177,658,683]" italics="true" pageId="2" pageNumber="245">vs.</emphasis>
37.4±3.8, p&lt;0.01), and larger mouth width (71.2%±3.9 of HL
<emphasis id="EF9C3F6BFFC3FF959C5AFD47FD83FD61" box="[585,615,694,719]" italics="true" pageId="2" pageNumber="245">vs.</emphasis>
63.2%±6.0, p&lt;0.01).
</paragraph>
</subSubSection>
<subSubSection id="95F2B0F2FFC3FF9D9ED4FD29FD4AF908" lastPageId="10" lastPageNumber="253" pageId="2" pageNumber="245" type="description">
<paragraph id="DD57E379FFC3FF959ED4FD29FB62FB45" blockId="2.[151,1437,152,2015]" pageId="2" pageNumber="245">
<emphasis id="EF9C3F6BFFC3FF959ED4FD29FEBDFD5D" bold="true" box="[199,345,728,755]" pageId="2" pageNumber="245">Description.</emphasis>
Body proportions presented in
<tableCitation id="906AD6C2FFC3FF959CAAFD28FCE9FD5D" box="[697,781,729,755]" captionStart="TABLE 1" captionStartId="3.[151,239,658,682]" captionTargetPageId="3" captionText="TABLE 1. Morphometric data of Batrochoglanis castaneus (n = 9). Min = minimum, max = maximum, SD = standard deviation. The holotype is included in the statistics." httpUri="http://table.plazi.org/id/8997B3F1FFC2FF949E84FD63FD58FD61" pageId="2" pageNumber="245" tableUuid="8997B3F1FFC2FF949E84FD63FD58FD61">Table 1</tableCitation>
. Dorsal profile slightly convex, almost straight from tip of snout to terminus of adipose fin. Ventral profile slightly convex from edge of mandible to beginning of caudal fin. Head larger than wide. Profile of head trapezoidal in dorsal view, greater width at cleithrum bulge. Gill opening wide. Gill rakers simple, without branching or bifurcations; epibranchials 2(3), 3(6*) rakers; epibranchial-ceratobranchial cartilage 1(9*) raker, ceratobranchial 6(5), 7(7*) rakers. Maxillary barbel surpassing the opercular border, but not reaching vertical through beginning of dorsal fin. External mental barbel surpassing base of pectoral fin. Dorsal fin trapezoidal, when adpressed not reaching adipose fin; I+6(9*) rays. Dorsal-fin spine sharp-pointed, anterior and posterior margins smooth, without serrae or denticuli. Adipose fin elongated with posterior end free and angled. Pectoral fin triangular, when adpressed not reaching ventral fin, I+6(9*) rays. Pectoral-fin spine covered by thin skin, serrated on both sides, antrorse and retrorse serrae on anterior margin and retrorse serrae on posterior margin; anterior serrae a little shorter than posterior ones. Pelvic fin triangular, with rounded border, terminus at vertical through base of fifth branched ray of dorsal fin, reaching origin of anal fin when adpressed; i+5(9*) rays. Anal fin semi-circular, its base length similar to adipose-fin base length; ii(3*), iii(6) + 6(1), 7(6), 8(2*) rays. Caudal fin with upper rays longer; 8(8*) rays in upper lobe, 6(4*), 7(4) rays in ventral lobe. Postcleithrum process elongated with little blunted point. Lateral line incomplete, reaching vertical through anterior third of adipose fin. Axillary pore absent.
</paragraph>
<paragraph id="DD57E379FFC3FF959ED4FB05FBC7F8A9" blockId="2.[151,1437,152,2015]" pageId="2" pageNumber="245">
<emphasis id="EF9C3F6BFFC3FF959ED4FB05FE2EFAA1" bold="true" box="[199,458,1268,1295]" pageId="2" pageNumber="245">Osteological features.</emphasis>
Cranial roof approximately triangular (
<figureCitation id="45D3FFFCFFC3FF959D82FB04FC02FAA1" box="[913,998,1269,1295]" captionStart="FIGURE 2" captionStartId="5.[151,250,1129,1154]" captionTargetBox="[162,1425,276,1102]" captionTargetId="figure@5.[151,1436,265,1109]" captionTargetPageId="5" captionText="FIGURE 2. Cranium of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. A) dorsal view; b) ventral view. Light blue = cartilage; dark gray = fontanel or foramen. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3458977" httpUri="https://zenodo.org/record/3458977/files/figure.png" pageId="2" pageNumber="245">Fig. 2a</figureCitation>
). Surface irregular, with several small rounded depressions. A single and large fontanel between mesethmoid and frontals; epiphyseal bar absent. Mesethmoid in the anterodorsal region of neurocranium, bifurcated anteriorly, with two cornua, ventrally attached by ligaments to the premaxilla. Lateral ethmoid broad, irregular shaped, medially articulated to mesethmoid, laterally articulated to autopalatine condyle. Ethmoid cartilage broad, located on the anterior region of lateral ethmoid and laterally to mesethmoid. Frontal bone anteriorly narrow and posteriorly expanded, sutured to sphenotic and parietosupraoccipital bones; frontals separated by large fontanel anteriorly. Sphenotic bone longer than wide, sutured to frontal, parieto-supraoccipital and pterotic bones. Pterotic approximately same size as sphenotic; sutured to sphenotic, parieto-supraoccipital and epiotic bones. Sphenotic and pterotic articulated to hyomandibula on their ventral surfaces. Extrascapula approximately half size of pterotic, located on posterior region of neurocranium, articulated to pterotic and parieto-supraoccipital. Sphenotic, pterotic and epiotic forming the posterolateral roof of skull. Parieto-supraoccipital a little wider than long, forming the posteromedian surface of cranial roof, sutured to frontals, sphenotics, pterotics, and epiotics. A single parieto-supraoccipital fontanel, posteriorly placed, ellipsoid shaped. Parieto-supraoccipital crest extending posteromedially, triangular shaped, posteriorly pointed, not touching anterior nuchal plate. Nasal bone tubular, slightly curved, laterally placed to mesethmoid.
</paragraph>
<paragraph id="DD57E379FFC3FF949ED4F8E0FCEEFEED" blockId="2.[151,1437,152,2015]" lastBlockId="3.[151,1437,153,611]" lastPageId="3" lastPageNumber="246" pageId="2" pageNumber="245">
Cranial floor surface less ornamented with rounded depressions than roof (
<figureCitation id="45D3FFFCFFC3FF959A16F8E0FBBCF882" box="[1029,1112,1809,1836]" captionStart="FIGURE 2" captionStartId="5.[151,250,1129,1154]" captionTargetBox="[162,1425,276,1102]" captionTargetId="figure@5.[151,1436,265,1109]" captionTargetPageId="5" captionText="FIGURE 2. Cranium of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. A) dorsal view; b) ventral view. Light blue = cartilage; dark gray = fontanel or foramen. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3458977" httpUri="https://zenodo.org/record/3458977/files/figure.png" pageId="2" pageNumber="245">Fig. 2b</figureCitation>
). Vomer absent. Mesethmoid articulated medially to lateral ethmoid. Parasphenoid narrow anteriorly and wider posteriorly; anteriorly sutured with mesethmoid and lateral ethmoid; laterally articulated to orbitosphenoid and pterosphenoid, and posteriorly with prootic and basioccipital. Orbitosphenoid large, lateral wall dorsally sutured to frontals and anteriorly with synchondral joint to lateral ethmoid. Pterosphenoid irregular shaped, posterior to orbitosphenoid, lateral to parasphenoid, and medially to sphenotic. Prootic large, latero-posteriorly sutured with pterotic; posteriorly with exoccipital. Wide foramen for trigeminofacialis nerve between pterosphenoid anteriorly, prootic posteriorly, and parasphenoid medially. Exoccipital forming posterior corner of neurocranium lateral to basioccipital. Basioccipital forming the posterior region of cranial floor, posteriorly articulated to first vertebra. Epiotic (visible only in posterior view) triangular-shaped, posteroventral to extrascapula. Supraoccipital process ventral region with a
<emphasis id="EF9C3F6BFFC2FF949A80FEF7FB40FEB1" box="[1171,1188,262,287]" italics="true" pageId="3" pageNumber="246">V</emphasis>
-shaped gutter. Lateral line with one ossified anteriormost canal near first pore.
</paragraph>
<paragraph id="DD57E379FFC2FF949ED4FEBCFEA1FE7D" blockId="3.[151,1437,153,611]" pageId="3" pageNumber="246">
First infraorbital irregular shaped, a thin curved point anteriorly, posterolateral and medial processes bearing a canal and pore of lateral line, and a posterior elongated process. Second infraorbital small, elongated, slightly curved. Third infraorbital straight, elongated, lateral to eye. Fourth infraorbital larger, angular, almost 90 degrees curved (
<figureCitation id="45D3FFFCFFC2FF949EE0FE48FEDCFE7D" box="[243,312,441,467]" captionStart="FIGURE 2" captionStartId="5.[151,250,1129,1154]" captionTargetBox="[162,1425,276,1102]" captionTargetId="figure@5.[151,1436,265,1109]" captionTargetPageId="5" captionText="FIGURE 2. Cranium of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. A) dorsal view; b) ventral view. Light blue = cartilage; dark gray = fontanel or foramen. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3458977" httpUri="https://zenodo.org/record/3458977/files/figure.png" pageId="3" pageNumber="246">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="DD57E379FFC2FF949ED4FE2CFEEEFDCD" blockId="3.[151,1437,153,611]" pageId="3" pageNumber="246">
Posttemporal-supracleithrum (
<figureCitation id="45D3FFFCFFC2FF949C0EFE2CFD86FE59" box="[541,610,477,503]" captionStart="FIGURE 2" captionStartId="5.[151,250,1129,1154]" captionTargetBox="[162,1425,276,1102]" captionTargetId="figure@5.[151,1436,265,1109]" captionTargetPageId="5" captionText="FIGURE 2. Cranium of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. A) dorsal view; b) ventral view. Light blue = cartilage; dark gray = fontanel or foramen. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3458977" httpUri="https://zenodo.org/record/3458977/files/figure.png" pageId="3" pageNumber="246">Fig. 2</figureCitation>
) well-developed, elongated, attached to pterotic and extrascapula anteriorly, and posteriorly to anterior process of parapophysis of fourth vertebra. Ventrally, an elongated cylindrical ossified transcapular ligament attached to basioccipital by a ligament. Ventrolateral limb attached to the upper limb of cleithrum.
</paragraph>
<caption id="8997B3F1FFC2FF949E84FD63FD58FD61" ID-Table-UUID="8997B3F1FFC2FF949E84FD63FD58FD61" httpUri="http://table.plazi.org/id/8997B3F1FFC2FF949E84FD63FD58FD61" pageId="3" pageNumber="246" targetBox="[161,1371,730,2003]" targetIsTable="true" targetPageId="3">
<paragraph id="DD57E379FFC2FF949E84FD63FD58FD61" blockId="3.[151,1436,658,719]" pageId="3" pageNumber="246">
<emphasis id="EF9C3F6BFFC2FF949E84FD63FEEFFD04" bold="true" box="[151,267,658,682]" pageId="3" pageNumber="246">TABLE 1.</emphasis>
Morphometric data of
<taxonomicName id="1AE898FAFFC2FF949C15FD62FCFEFD04" authorityName="Shibatta" authorityYear="2019" box="[518,794,659,683]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="246" phylum="Chordata" rank="species" species="castaneus">
<emphasis id="EF9C3F6BFFC2FF949C15FD62FCFEFD04" box="[518,794,659,683]" italics="true" pageId="3" pageNumber="246">Batrochoglanis castaneus</emphasis>
</taxonomicName>
(n = 9). Min = minimum, max = maximum, SD = standard deviation. The holotype is included in the statistics.
</paragraph>
</caption>
<paragraph id="DD57E379FFC2FF949EB2FD2BFABFF87C" pageId="3" pageNumber="246">
<table id="AFE811D9FFC200689EB2FD2BFABFF87D" box="[161,1371,730,2003]" gridcols="4" gridrows="34" pageId="3" pageNumber="246">
<tr id="63D8E13BFFC200689EB2FD2BFABFFD5D" box="[161,1371,730,755]" gridrow="0" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2FD2BFDBDFD5D" box="[161,601,730,755]" gridcol="0" gridrow="0" pageId="3" pageNumber="246">Character</th>
<th id="20098847FFC200689D65FD2BFC30FD5D" box="[886,980,730,755]" gridcol="1" gridrow="0" pageId="3" pageNumber="246">Holotype</th>
<th id="20098847FFC200689A06FD2BFB76FD5D" box="[1045,1170,730,755]" gridcol="2" gridrow="0" pageId="3" pageNumber="246">Min-max</th>
<th id="20098847FFC200689AFBFD2BFABFFD5D" box="[1256,1371,730,755]" gridcol="3" gridrow="0" pageId="3" pageNumber="246">Mean±SD</th>
</tr>
<tr id="63D8E13BFFC200689EB2FCF1FABFFCB7" box="[161,1371,768,793]" gridrow="1" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2FCF1FDBDFCB7" box="[161,601,768,793]" gridcol="0" gridrow="1" pageId="3" pageNumber="246">Standard length (mm)</th>
<td id="20098847FFC200689D65FCF1FC30FCB7" box="[886,980,768,793]" gridcol="1" gridrow="1" pageId="3" pageNumber="246">58.3</td>
<td id="20098847FFC200689A06FCF1FB76FCB7" box="[1045,1170,768,793]" gridcol="2" gridrow="1" pageId="3" pageNumber="246">49.7–58.3</td>
<td id="20098847FFC200689AFBFCF1FABFFCB7" box="[1256,1371,768,793]" gridcol="3" gridrow="1" pageId="3" pageNumber="246">52.6±3.1</td>
</tr>
<tr id="63D8E13BFFC200689EB2FCD4FABFFC90" box="[161,1371,805,830]" gridrow="2" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2FCD4FABFFC90" box="[161,1371,805,830]" colspan="4" colspanRight="3" gridcol="0" gridrow="2" pageId="3" pageNumber="246">
<emphasis id="EF9C3F6BFFC2FF949EB2FCD7FE0BFC90" bold="true" box="[161,495,805,830]" pageId="3" pageNumber="246">Percentages of standard length</emphasis>
</th>
</tr>
<tr id="63D8E13BFFC200689EB2FCBDFABFFCCB" box="[161,1371,844,869]" gridrow="3" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2FCBDFDBDFCCB" box="[161,601,844,869]" gridcol="0" gridrow="3" pageId="3" pageNumber="246">Head length</th>
<td id="20098847FFC200689D65FCBDFC30FCCB" box="[886,980,844,869]" gridcol="1" gridrow="3" pageId="3" pageNumber="246">33.0</td>
<td id="20098847FFC200689A06FCBDFB76FCCB" box="[1045,1170,844,869]" gridcol="2" gridrow="3" pageId="3" pageNumber="246">31.8–34.0</td>
<td id="20098847FFC200689AFBFCBDFABFFCCB" box="[1256,1371,844,869]" gridcol="3" gridrow="3" pageId="3" pageNumber="246">32.9±0.7</td>
</tr>
<tr id="63D8E13BFFC200689EB2FC80FABFFC24" box="[161,1371,881,906]" gridrow="4" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2FC80FDBDFC24" box="[161,601,881,906]" gridcol="0" gridrow="4" pageId="3" pageNumber="246">Predorsal length</th>
<td id="20098847FFC200689D65FC80FC30FC24" box="[886,980,881,906]" gridcol="1" gridrow="4" pageId="3" pageNumber="246">41.3</td>
<td id="20098847FFC200689A06FC80FB76FC24" box="[1045,1170,881,906]" gridcol="2" gridrow="4" pageId="3" pageNumber="246">41.3–44.1</td>
<td id="20098847FFC200689AFBFC80FABFFC24" box="[1256,1371,881,906]" gridcol="3" gridrow="4" pageId="3" pageNumber="246">43.1±0.8</td>
</tr>
<tr id="63D8E13BFFC200689EB2FC66FABFFC1E" box="[161,1371,919,944]" gridrow="5" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2FC66FDBDFC1E" box="[161,601,919,944]" gridcol="0" gridrow="5" pageId="3" pageNumber="246">Dorsal-fin base length</th>
<td id="20098847FFC200689D65FC66FC30FC1E" box="[886,980,919,944]" gridcol="1" gridrow="5" pageId="3" pageNumber="246">14.8</td>
<td id="20098847FFC200689A06FC66FB76FC1E" box="[1045,1170,919,944]" gridcol="2" gridrow="5" pageId="3" pageNumber="246">14.0–16.4</td>
<td id="20098847FFC200689AFBFC66FABFFC1E" box="[1256,1371,919,944]" gridcol="3" gridrow="5" pageId="3" pageNumber="246">15.3±0.7</td>
</tr>
<tr id="63D8E13BFFC200689EB2FC4CFABFFC78" box="[161,1371,957,982]" gridrow="6" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2FC4CFDBDFC78" box="[161,601,957,982]" gridcol="0" gridrow="6" pageId="3" pageNumber="246">Adipose-fin base length</th>
<td id="20098847FFC200689D65FC4CFC30FC78" box="[886,980,957,982]" gridcol="1" gridrow="6" pageId="3" pageNumber="246">13.0</td>
<td id="20098847FFC200689A06FC4CFB76FC78" box="[1045,1170,957,982]" gridcol="2" gridrow="6" pageId="3" pageNumber="246">11.3–17.0</td>
<td id="20098847FFC200689AFBFC4CFABFFC78" box="[1256,1371,957,982]" gridcol="3" gridrow="6" pageId="3" pageNumber="246">14.2±1.6</td>
</tr>
<tr id="63D8E13BFFC200689EB2FC12FABFFC52" box="[161,1371,995,1020]" gridrow="7" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2FC12FDBDFC52" box="[161,601,995,1020]" gridcol="0" gridrow="7" pageId="3" pageNumber="246">Prepelvic length</th>
<td id="20098847FFC200689D65FC12FC30FC52" box="[886,980,995,1020]" gridcol="1" gridrow="7" pageId="3" pageNumber="246">57.2</td>
<td id="20098847FFC200689A06FC12FB76FC52" box="[1045,1170,995,1020]" gridcol="2" gridrow="7" pageId="3" pageNumber="246">56.3–58.8</td>
<td id="20098847FFC200689AFBFC12FABFFC52" box="[1256,1371,995,1020]" gridcol="3" gridrow="7" pageId="3" pageNumber="246">57.4±0.7</td>
</tr>
<tr id="63D8E13BFFC200689EB2FBF8FABFFB8C" box="[161,1371,1033,1058]" gridrow="8" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2FBF8FDBDFB8C" box="[161,601,1033,1058]" gridcol="0" gridrow="8" pageId="3" pageNumber="246">Pelvic fin to anal fin distance</th>
<td id="20098847FFC200689D65FBF8FC30FB8C" box="[886,980,1033,1058]" gridcol="1" gridrow="8" pageId="3" pageNumber="246">40.3</td>
<td id="20098847FFC200689A06FBF8FB76FB8C" box="[1045,1170,1033,1058]" gridcol="2" gridrow="8" pageId="3" pageNumber="246">19.0–22.4</td>
<td id="20098847FFC200689AFBFBF8FABFFB8C" box="[1256,1371,1033,1058]" gridcol="3" gridrow="8" pageId="3" pageNumber="246">20.8±1.3</td>
</tr>
<tr id="63D8E13BFFC200689EB2FBDFFABFFBE9" box="[161,1371,1070,1095]" gridrow="9" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2FBDFFDBDFBE9" box="[161,601,1070,1095]" gridcol="0" gridrow="9" pageId="3" pageNumber="246">Anal-fin base length</th>
<td id="20098847FFC200689D65FBDFFC30FBE9" box="[886,980,1070,1095]" gridcol="1" gridrow="9" pageId="3" pageNumber="246">12.3</td>
<td id="20098847FFC200689A06FBDFFB76FBE9" box="[1045,1170,1070,1095]" gridcol="2" gridrow="9" pageId="3" pageNumber="246">11.2–12.6</td>
<td id="20098847FFC200689AFBFBDFFABFFBE9" box="[1256,1371,1070,1095]" gridcol="3" gridrow="9" pageId="3" pageNumber="246">12.0±0.5</td>
</tr>
<tr id="63D8E13BFFC200689EB2FBA5FABFFBC3" box="[161,1371,1108,1133]" gridrow="10" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2FBA5FDBDFBC3" box="[161,601,1108,1133]" gridcol="0" gridrow="10" pageId="3" pageNumber="246">Body depth</th>
<td id="20098847FFC200689D65FBA5FC30FBC3" box="[886,980,1108,1133]" gridcol="1" gridrow="10" pageId="3" pageNumber="246">26.8</td>
<td id="20098847FFC200689A06FBA5FB76FBC3" box="[1045,1170,1108,1133]" gridcol="2" gridrow="10" pageId="3" pageNumber="246">21.4–28.0</td>
<td id="20098847FFC200689AFBFBA5FABFFBC3" box="[1256,1371,1108,1133]" gridcol="3" gridrow="10" pageId="3" pageNumber="246">25.3±2.2</td>
</tr>
<tr id="63D8E13BFFC200689EB2FB8BFABFFB3D" box="[161,1371,1146,1171]" gridrow="11" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2FB8BFDBDFB3D" box="[161,601,1146,1171]" gridcol="0" gridrow="11" pageId="3" pageNumber="246">Caudal peduncle depth</th>
<td id="20098847FFC200689D65FB8BFC30FB3D" box="[886,980,1146,1171]" gridcol="1" gridrow="11" pageId="3" pageNumber="246">11.2</td>
<td id="20098847FFC200689A06FB8BFB76FB3D" box="[1045,1170,1146,1171]" gridcol="2" gridrow="11" pageId="3" pageNumber="246">10.2–11.8</td>
<td id="20098847FFC200689AFBFB8BFABFFB3D" box="[1256,1371,1146,1171]" gridcol="3" gridrow="11" pageId="3" pageNumber="246">10.9±0.5</td>
</tr>
<tr id="63D8E13BFFC200689EB2FB51FABFFB17" box="[161,1371,1184,1209]" gridrow="12" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2FB51FDBDFB17" box="[161,601,1184,1209]" gridcol="0" gridrow="12" pageId="3" pageNumber="246">Pectoral girdle width</th>
<td id="20098847FFC200689D65FB51FC30FB17" box="[886,980,1184,1209]" gridcol="1" gridrow="12" pageId="3" pageNumber="246">34.1</td>
<td id="20098847FFC200689A06FB51FB76FB17" box="[1045,1170,1184,1209]" gridcol="2" gridrow="12" pageId="3" pageNumber="246">33.1–34.3</td>
<td id="20098847FFC200689AFBFB51FABFFB17" box="[1256,1371,1184,1209]" gridcol="3" gridrow="12" pageId="3" pageNumber="246">33.8±0.4</td>
</tr>
<tr id="63D8E13BFFC200689EB2FB37FABFFB71" box="[161,1371,1222,1247]" gridrow="13" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2FB37FDBDFB71" box="[161,601,1222,1247]" gridcol="0" gridrow="13" pageId="3" pageNumber="246">Pectoral-fin spine length</th>
<td id="20098847FFC200689D65FB37FC30FB71" box="[886,980,1222,1247]" gridcol="1" gridrow="13" pageId="3" pageNumber="246">20.0</td>
<td id="20098847FFC200689A06FB37FB76FB71" box="[1045,1170,1222,1247]" gridcol="2" gridrow="13" pageId="3" pageNumber="246">17.7–23.3</td>
<td id="20098847FFC200689AFBFB37FABFFB71" box="[1256,1371,1222,1247]" gridcol="3" gridrow="13" pageId="3" pageNumber="246">21.0±1.6</td>
</tr>
<tr id="63D8E13BFFC200689EB2FB1AFABFFAAA" box="[161,1371,1259,1284]" gridrow="14" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2FB1AFDBDFAAA" box="[161,601,1259,1284]" gridcol="0" gridrow="14" pageId="3" pageNumber="246">Dorsal-fin spine length</th>
<td id="20098847FFC200689D65FB1AFC30FAAA" box="[886,980,1259,1284]" gridcol="1" gridrow="14" pageId="3" pageNumber="246">16.0</td>
<td id="20098847FFC200689A06FB1AFB76FAAA" box="[1045,1170,1259,1284]" gridcol="2" gridrow="14" pageId="3" pageNumber="246">14.8–18.4</td>
<td id="20098847FFC200689AFBFB1AFABFFAAA" box="[1256,1371,1259,1284]" gridcol="3" gridrow="14" pageId="3" pageNumber="246">16.5±1.0</td>
</tr>
<tr id="63D8E13BFFC200689EB2FAE0FABFFA84" box="[161,1371,1297,1322]" gridrow="15" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2FAE0FDBDFA84" box="[161,601,1297,1322]" gridcol="0" gridrow="15" pageId="3" pageNumber="246">Prepectoral length</th>
<td id="20098847FFC200689D65FAE0FC30FA84" box="[886,980,1297,1322]" gridcol="1" gridrow="15" pageId="3" pageNumber="246">31.3</td>
<td id="20098847FFC200689A06FAE0FB76FA84" box="[1045,1170,1297,1322]" gridcol="2" gridrow="15" pageId="3" pageNumber="246">29.0–33.5</td>
<td id="20098847FFC200689AFBFAE0FABFFA84" box="[1256,1371,1297,1322]" gridcol="3" gridrow="15" pageId="3" pageNumber="246">31.5±1.6</td>
</tr>
<tr id="63D8E13BFFC200689EB2FAC6FABFFAFE" box="[161,1371,1335,1360]" gridrow="16" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2FAC6FDBDFAFE" box="[161,601,1335,1360]" gridcol="0" gridrow="16" pageId="3" pageNumber="246">Predorsal length</th>
<td id="20098847FFC200689D65FAC6FC30FAFE" box="[886,980,1335,1360]" gridcol="1" gridrow="16" pageId="3" pageNumber="246">41.3</td>
<td id="20098847FFC200689A06FAC6FB76FAFE" box="[1045,1170,1335,1360]" gridcol="2" gridrow="16" pageId="3" pageNumber="246">41.3–44.1</td>
<td id="20098847FFC200689AFBFAC6FABFFAFE" box="[1256,1371,1335,1360]" gridcol="3" gridrow="16" pageId="3" pageNumber="246">43.1±0.8</td>
</tr>
<tr id="63D8E13BFFC200689EB2FAACFABFFAD8" box="[161,1371,1373,1398]" gridrow="17" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2FAACFDBDFAD8" box="[161,601,1373,1398]" gridcol="0" gridrow="17" pageId="3" pageNumber="246">Pectoral fin to dorsal fin</th>
<td id="20098847FFC200689D65FAACFC30FAD8" box="[886,980,1373,1398]" gridcol="1" gridrow="17" pageId="3" pageNumber="246">26.2</td>
<td id="20098847FFC200689A06FAACFB76FAD8" box="[1045,1170,1373,1398]" gridcol="2" gridrow="17" pageId="3" pageNumber="246">24.8—27.9</td>
<td id="20098847FFC200689AFBFAACFABFFAD8" box="[1256,1371,1373,1398]" gridcol="3" gridrow="17" pageId="3" pageNumber="246">26.8±1.0</td>
</tr>
<tr id="63D8E13BFFC200689EB2FA72FABFFA32" box="[161,1371,1411,1436]" gridrow="18" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2FA72FDBDFA32" box="[161,601,1411,1436]" gridcol="0" gridrow="18" pageId="3" pageNumber="246">Pectoral fin to pelvic fin</th>
<td id="20098847FFC200689D65FA72FC30FA32" box="[886,980,1411,1436]" gridcol="1" gridrow="18" pageId="3" pageNumber="246">32.6</td>
<td id="20098847FFC200689A06FA72FB76FA32" box="[1045,1170,1411,1436]" gridcol="2" gridrow="18" pageId="3" pageNumber="246">28.5–35.5</td>
<td id="20098847FFC200689AFBFA72FABFFA32" box="[1256,1371,1411,1436]" gridcol="3" gridrow="18" pageId="3" pageNumber="246">31.7±2.0</td>
</tr>
<tr id="63D8E13BFFC200689EB2FA59FABFFA6F" box="[161,1371,1448,1473]" gridrow="19" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2FA59FDBDFA6F" box="[161,601,1448,1473]" gridcol="0" gridrow="19" pageId="3" pageNumber="246">Dorsal-fin origin to pelvic-fin origin</th>
<td id="20098847FFC200689D65FA59FC30FA6F" box="[886,980,1448,1473]" gridcol="1" gridrow="19" pageId="3" pageNumber="246">27.1</td>
<td id="20098847FFC200689A06FA59FB76FA6F" box="[1045,1170,1448,1473]" gridcol="2" gridrow="19" pageId="3" pageNumber="246">25.9–29.1</td>
<td id="20098847FFC200689AFBFA59FABFFA6F" box="[1256,1371,1448,1473]" gridcol="3" gridrow="19" pageId="3" pageNumber="246">27.3±1.1</td>
</tr>
<tr id="63D8E13BFFC200689EB2FA3FFABFFA49" box="[161,1371,1486,1511]" gridrow="20" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2FA3FFDBDFA49" box="[161,601,1486,1511]" gridcol="0" gridrow="20" pageId="3" pageNumber="246">Dorsal-fin origin to anal-fin origin</th>
<td id="20098847FFC200689D65FA3FFC30FA49" box="[886,980,1486,1511]" gridcol="1" gridrow="20" pageId="3" pageNumber="246">40.6</td>
<td id="20098847FFC200689A06FA3FFB76FA49" box="[1045,1170,1486,1511]" gridcol="2" gridrow="20" pageId="3" pageNumber="246">38.3–40.5</td>
<td id="20098847FFC200689AFBFA3FFABFFA49" box="[1256,1371,1486,1511]" gridcol="3" gridrow="20" pageId="3" pageNumber="246">39.3±0.9</td>
</tr>
<tr id="63D8E13BFFC200689EB2FA05FABFF9A3" box="[161,1371,1524,1549]" gridrow="21" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2FA05FDBDF9A3" box="[161,601,1524,1549]" gridcol="0" gridrow="21" pageId="3" pageNumber="246">Dorsal-fin terminus to pelvic-fin origin</th>
<td id="20098847FFC200689D65FA05FC30F9A3" box="[886,980,1524,1549]" gridcol="1" gridrow="21" pageId="3" pageNumber="246">22.1</td>
<td id="20098847FFC200689A06FA05FB76F9A3" box="[1045,1170,1524,1549]" gridcol="2" gridrow="21" pageId="3" pageNumber="246">19.7–22.5</td>
<td id="20098847FFC200689AFBFA05FABFF9A3" box="[1256,1371,1524,1549]" gridcol="3" gridrow="21" pageId="3" pageNumber="246">21.1±0.9</td>
</tr>
<tr id="63D8E13BFFC200689EB2F9EBFABFF99D" box="[161,1371,1562,1587]" gridrow="22" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2F9EBFDBDF99D" box="[161,601,1562,1587]" gridcol="0" gridrow="22" pageId="3" pageNumber="246">Dorsal-fin terminus to anal-fin origin</th>
<td id="20098847FFC200689D65F9EBFC30F99D" box="[886,980,1562,1587]" gridcol="1" gridrow="22" pageId="3" pageNumber="246">27.2</td>
<td id="20098847FFC200689A06F9EBFB76F99D" box="[1045,1170,1562,1587]" gridcol="2" gridrow="22" pageId="3" pageNumber="246">23.7–28.1</td>
<td id="20098847FFC200689AFBF9EBFABFF99D" box="[1256,1371,1562,1587]" gridcol="3" gridrow="22" pageId="3" pageNumber="246">26.1±1.3</td>
</tr>
<tr id="63D8E13BFFC200689EB2F9B1FABFF9F7" box="[161,1371,1600,1625]" gridrow="23" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2F9B1FDBDF9F7" box="[161,601,1600,1625]" gridcol="0" gridrow="23" pageId="3" pageNumber="246">Dorsal-fin terminus to adipose-fin terminus</th>
<td id="20098847FFC200689D65F9B1FC30F9F7" box="[886,980,1600,1625]" gridcol="1" gridrow="23" pageId="3" pageNumber="246">33.0</td>
<td id="20098847FFC200689A06F9B1FB76F9F7" box="[1045,1170,1600,1625]" gridcol="2" gridrow="23" pageId="3" pageNumber="246">28.8–34.8</td>
<td id="20098847FFC200689AFBF9B1FABFF9F7" box="[1256,1371,1600,1625]" gridcol="3" gridrow="23" pageId="3" pageNumber="246">31.4±1.8</td>
</tr>
<tr id="63D8E13BFFC200689EB2F994FABFF9D0" box="[161,1371,1637,1662]" gridrow="24" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2F994FDBDF9D0" box="[161,601,1637,1662]" gridcol="0" gridrow="24" pageId="3" pageNumber="246">Dorsal-fin terminus to anal-fin terminus</th>
<td id="20098847FFC200689D65F994FC30F9D0" box="[886,980,1637,1662]" gridcol="1" gridrow="24" pageId="3" pageNumber="246">34.3</td>
<td id="20098847FFC200689A06F994FB76F9D0" box="[1045,1170,1637,1662]" gridcol="2" gridrow="24" pageId="3" pageNumber="246">31.5–34.8</td>
<td id="20098847FFC200689AFBF994FABFF9D0" box="[1256,1371,1637,1662]" gridcol="3" gridrow="24" pageId="3" pageNumber="246">33.4±0.9</td>
</tr>
<tr id="63D8E13BFFC200689EB2F97AFABFF90A" box="[161,1371,1675,1700]" gridrow="25" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2F97AFDBDF90A" box="[161,601,1675,1700]" gridcol="0" gridrow="25" pageId="3" pageNumber="246">Anal-fin origin to adipose-fin terminus</th>
<td id="20098847FFC200689D65F97AFC30F90A" box="[886,980,1675,1700]" gridcol="1" gridrow="25" pageId="3" pageNumber="246">19.6</td>
<td id="20098847FFC200689A06F97AFB76F90A" box="[1045,1170,1675,1700]" gridcol="2" gridrow="25" pageId="3" pageNumber="246">17.8–19.9</td>
<td id="20098847FFC200689AFBF97AFABFF90A" box="[1256,1371,1675,1700]" gridcol="3" gridrow="25" pageId="3" pageNumber="246">18.8±0.7</td>
</tr>
<tr id="63D8E13BFFC200689EB2F940FABFF964" box="[161,1371,1713,1738]" gridrow="26" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2F940FDBDF964" box="[161,601,1713,1738]" gridcol="0" gridrow="26" pageId="3" pageNumber="246">Adipose-fin terminus to anal-fin terminus</th>
<td id="20098847FFC200689D65F940FC30F964" box="[886,980,1713,1738]" gridcol="1" gridrow="26" pageId="3" pageNumber="246">11.2</td>
<td id="20098847FFC200689A06F940FB76F964" box="[1045,1170,1713,1738]" gridcol="2" gridrow="26" pageId="3" pageNumber="246">10.3–12.5</td>
<td id="20098847FFC200689AFBF940FABFF964" box="[1256,1371,1713,1738]" gridcol="3" gridrow="26" pageId="3" pageNumber="246">11.2±0.7</td>
</tr>
<tr id="63D8E13BFFC200689EB2F927FABFF941" box="[161,1371,1750,1775]" gridrow="27" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2F927FABFF941" box="[161,1371,1750,1775]" colspan="4" colspanRight="3" gridcol="0" gridrow="27" pageId="3" pageNumber="246">
<emphasis id="EF9C3F6BFFC2FF949EB2F926FE26F941" bold="true" box="[161,450,1750,1775]" pageId="3" pageNumber="246">Percentages of head length</emphasis>
</th>
</tr>
<tr id="63D8E13BFFC200689EB2F90CFABFF8B8" box="[161,1371,1789,1814]" gridrow="28" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2F90CFDBDF8B8" box="[161,601,1789,1814]" gridcol="0" gridrow="28" pageId="3" pageNumber="246">Eye diameter</th>
<td id="20098847FFC200689D65F90CFC30F8B8" box="[886,980,1789,1814]" gridcol="1" gridrow="28" pageId="3" pageNumber="246">9.9</td>
<td id="20098847FFC200689A06F90CFB76F8B8" box="[1045,1170,1789,1814]" gridcol="2" gridrow="28" pageId="3" pageNumber="246">9.87–14.1</td>
<td id="20098847FFC200689AFBF90CFABFF8B8" box="[1256,1371,1789,1814]" gridcol="3" gridrow="28" pageId="3" pageNumber="246">12.1±1.2</td>
</tr>
<tr id="63D8E13BFFC200689EB2F8D3FABFF895" box="[161,1371,1826,1851]" gridrow="29" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2F8D3FDBDF895" box="[161,601,1826,1851]" gridcol="0" gridrow="29" pageId="3" pageNumber="246">Interorbital width</th>
<td id="20098847FFC200689D65F8D3FC30F895" box="[886,980,1826,1851]" gridcol="1" gridrow="29" pageId="3" pageNumber="246">43.1</td>
<td id="20098847FFC200689A06F8D3FB76F895" box="[1045,1170,1826,1851]" gridcol="2" gridrow="29" pageId="3" pageNumber="246">39.7–48.1</td>
<td id="20098847FFC200689AFBF8D3FABFF895" box="[1256,1371,1826,1851]" gridcol="3" gridrow="29" pageId="3" pageNumber="246">43.7±2.1</td>
</tr>
<tr id="63D8E13BFFC200689EB2F8B9FABFF8CF" box="[161,1371,1864,1889]" gridrow="30" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2F8B9FDBDF8CF" box="[161,601,1864,1889]" gridcol="0" gridrow="30" pageId="3" pageNumber="246">Snout length</th>
<td id="20098847FFC200689D65F8B9FC30F8CF" box="[886,980,1864,1889]" gridcol="1" gridrow="30" pageId="3" pageNumber="246">36.4</td>
<td id="20098847FFC200689A06F8B9FB76F8CF" box="[1045,1170,1864,1889]" gridcol="2" gridrow="30" pageId="3" pageNumber="246">32.3–36.3</td>
<td id="20098847FFC200689AFBF8B9FABFF8CF" box="[1256,1371,1864,1889]" gridcol="3" gridrow="30" pageId="3" pageNumber="246">34.2±1.1</td>
</tr>
<tr id="63D8E13BFFC200689EB2F89FFABFF829" box="[161,1371,1902,1927]" gridrow="31" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2F89FFDBDF829" box="[161,601,1902,1927]" gridcol="0" gridrow="31" pageId="3" pageNumber="246">Mouth width</th>
<td id="20098847FFC200689D65F89FFC30F829" box="[886,980,1902,1927]" gridcol="1" gridrow="31" pageId="3" pageNumber="246">74.0</td>
<td id="20098847FFC200689A06F89FFB76F829" box="[1045,1170,1902,1927]" gridcol="2" gridrow="31" pageId="3" pageNumber="246">65.2–78.7</td>
<td id="20098847FFC200689AFBF89FFABFF829" box="[1256,1371,1902,1927]" gridcol="3" gridrow="31" pageId="3" pageNumber="246">71.2±3.7</td>
</tr>
<tr id="63D8E13BFFC200689EB2F865FABFF803" box="[161,1371,1940,1965]" gridrow="32" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2F865FDBDF803" box="[161,601,1940,1965]" gridcol="0" gridrow="32" pageId="3" pageNumber="246">Head depth</th>
<td id="20098847FFC200689D65F865FC30F803" box="[886,980,1940,1965]" gridcol="1" gridrow="32" pageId="3" pageNumber="246">35.3</td>
<td id="20098847FFC200689A06F865FB76F803" box="[1045,1170,1940,1965]" gridcol="2" gridrow="32" pageId="3" pageNumber="246">27.0–35.3</td>
<td id="20098847FFC200689AFBF865FABFF803" box="[1256,1371,1940,1965]" gridcol="3" gridrow="32" pageId="3" pageNumber="246">31.1±2.3</td>
</tr>
<tr id="63D8E13BFFC200689EB2F84BFABFF87D" box="[161,1371,1978,2003]" gridrow="33" pageId="3" pageNumber="246">
<th id="20098847FFC200689EB2F84BFDBDF87D" box="[161,601,1978,2003]" gridcol="0" gridrow="33" pageId="3" pageNumber="246">Maxillary barbel length</th>
<td id="20098847FFC200689D65F84BFC30F87D" box="[886,980,1978,2003]" gridcol="1" gridrow="33" pageId="3" pageNumber="246">129.9</td>
<td id="20098847FFC200689A06F84BFB76F87D" box="[1045,1170,1978,2003]" gridcol="2" gridrow="33" pageId="3" pageNumber="246">115.6–150.2</td>
<td id="20098847FFC200689AFBF84BFABFF87D" box="[1256,1371,1978,2003]" gridcol="3" gridrow="33" pageId="3" pageNumber="246">131.8±10.1</td>
</tr>
</table>
</paragraph>
<caption id="8997B3F1FFC5FF939E84F940FA8CF967" ID-DOI="http://doi.org/10.5281/zenodo.3458975" ID-Zenodo-Dep="3458975" box="[151,1384,1713,1738]" httpUri="https://zenodo.org/record/3458975/files/figure.png" pageId="4" pageNumber="247" startId="4.[151,250,1713,1738]" targetBox="[151,1436,174,1693]" targetPageId="4">
<paragraph id="DD57E379FFC5FF939E84F940FA8CF967" blockId="4.[151,1384,1713,1738]" box="[151,1384,1713,1738]" pageId="4" pageNumber="247">
<emphasis id="EF9C3F6BFFC5FF939E84F940FEE9F964" bold="true" box="[151,269,1713,1738]" pageId="4" pageNumber="247">FIGURE 1</emphasis>
. Holotype of
<taxonomicName id="1AE898FAFFC5FF939F8AF943FD7BF967" authorityName="Shibatta" authorityYear="2019" box="[409,671,1714,1738]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="247" phylum="Chordata" rank="species" species="castaneus">
<emphasis id="EF9C3F6BFFC5FF939F8AF943FD7BF967" box="[409,671,1714,1738]" italics="true" pageId="4" pageNumber="247">Batrochoglanis castaneus</emphasis>
</taxonomicName>
, INPA 8059, 58,3 mm SL, Aripuanã River, Igarapé do Castanhal, MT.
</paragraph>
</caption>
<paragraph id="DD57E379FFC5FF939ED4F909FB81F80C" blockId="4.[151,1437,1784,2027]" pageId="4" pageNumber="247">
Premaxilla longer than wide, with lateral point projecting posteriorly; ventrally bearing tooth plate with four irregular rows of small conical teeth turned slightly backward (
<figureCitation id="45D3FFFCFFC5FF939D49F8EDFC44F898" box="[858,928,1820,1846]" captionStart="FIGURE 2" captionStartId="5.[151,250,1129,1154]" captionTargetBox="[162,1425,276,1102]" captionTargetId="figure@5.[151,1436,265,1109]" captionTargetPageId="5" captionText="FIGURE 2. Cranium of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. A) dorsal view; b) ventral view. Light blue = cartilage; dark gray = fontanel or foramen. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3458977" httpUri="https://zenodo.org/record/3458977/files/figure.png" pageId="4" pageNumber="247">Fig. 2</figureCitation>
). Maxilla small, approximately wing-shaped, with two proximal rounded heads (dorsal and ventral condyle), medially articulated to autopalatine. Autopalatine (
<figureCitation id="45D3FFFCFFC5FF939EB3F895FF07F8D0" box="[160,227,1892,1918]" captionStart="FIGURE 3" captionStartId="5.[151,250,1923,1948]" captionTargetBox="[167,1417,1213,1896]" captionTargetId="figure@5.[151,1436,1204,1904]" captionTargetPageId="5" captionText="FIGURE 3. Autopalatine and maxilla of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. A) dorsal view; b) lateral view; c) ventral view; d) medial view. Light blue = cartilage. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3458979" httpUri="https://zenodo.org/record/3458979/files/figure.png" pageId="4" pageNumber="247">Fig. 3</figureCitation>
) elongated anteroposteriorly, spoon-shaped, with a dorsal keel in the posterior tip; latero-anteriorly articulated to the maxillary bone, medially to lateral ethmoid by a rounded articulation facet.
</paragraph>
<paragraph id="DD57E379FFC5FF929ED4F85CFAB1FF55" blockId="4.[151,1437,1784,2027]" lastBlockId="5.[151,1436,153,251]" lastPageId="5" lastPageNumber="248" pageId="4" pageNumber="247">
Dentary (
<figureCitation id="45D3FFFCFFC5FF939F23F85DFE97F868" box="[304,371,1964,1990]" captionStart="FIGURE 4" captionStartId="6.[151,250,930,955]" captionTargetBox="[163,1424,186,901]" captionTargetId="figure@6.[151,1436,174,910]" captionTargetPageId="6" captionText="FIGURE 4. Lower jaw of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. Light blue = cartilage. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3458981" httpUri="https://zenodo.org/record/3458981/files/figure.png" pageId="4" pageNumber="247">Fig. 4</figureCitation>
) long and straight, slightly deep posteriorly bearing the dorsal coronoid process; three irregular series of small conical teeth on dorsal surface. Six preoperculomandibular laterosensory branches (pm1-pm6) along lateroventral surface of dentary. Angular, articular, and retroarticular bones fused, horizontally elongated, articulated with dentary anteriorly and posteriorly attached to quadrate by ligaments. Coronomeckelian bone on medial surface, small, irregular shaped, posterodorsally articulated with horizontally elongated Meckelian cartilage.
</paragraph>
<caption id="8997B3F1FFC4FF929E84FB98FC6FFB0B" ID-DOI="http://doi.org/10.5281/zenodo.3458977" ID-Zenodo-Dep="3458977" httpUri="https://zenodo.org/record/3458977/files/figure.png" pageId="5" pageNumber="248" startId="5.[151,250,1129,1154]" targetBox="[162,1425,276,1102]" targetPageId="5">
<paragraph id="DD57E379FFC4FF929E84FB98FC6FFB0B" blockId="5.[151,1437,1129,1190]" pageId="5" pageNumber="248">
<emphasis id="EF9C3F6BFFC4FF929E84FB98FEF1FB2C" bold="true" box="[151,277,1129,1154]" pageId="5" pageNumber="248">FIGURE 2.</emphasis>
Cranium of
<taxonomicName id="1AE898FAFFC4FF929F88FB98FD47FB2F" authorityName="Shibatta" authorityYear="2019" box="[411,675,1129,1153]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="248" phylum="Chordata" rank="species" species="castaneus">
<emphasis id="EF9C3F6BFFC4FF929F88FB98FD47FB2F" box="[411,675,1129,1153]" italics="true" pageId="5" pageNumber="248">Batrochoglanis castaneus</emphasis>
</taxonomicName>
, MZUEL 20113, 50.8 mm SL, paratype. A) dorsal view; b) ventral view. Light blue = cartilage; dark gray = fontanel or foramen. Scale bar = 5 mm.
</paragraph>
</caption>
<caption id="8997B3F1FFC4FF929E84F872FBE2F86E" ID-DOI="http://doi.org/10.5281/zenodo.3458979" ID-Zenodo-Dep="3458979" httpUri="https://zenodo.org/record/3458979/files/figure.png" pageId="5" pageNumber="248" startId="5.[151,250,1923,1948]" targetBox="[167,1417,1213,1896]" targetPageId="5">
<paragraph id="DD57E379FFC4FF929E84F872FBE2F86E" blockId="5.[151,1436,1923,1984]" pageId="5" pageNumber="248">
<emphasis id="EF9C3F6BFFC4FF929E84F872FEF7F832" bold="true" box="[151,275,1923,1948]" pageId="5" pageNumber="248">FIGURE 3.</emphasis>
Autopalatine and maxilla of
<taxonomicName id="1AE898FAFFC4FF929C28F875FCA4F832" authorityName="Shibatta" authorityYear="2019" box="[571,832,1924,1948]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="248" phylum="Chordata" rank="species" species="castaneus">
<emphasis id="EF9C3F6BFFC4FF929C28F875FCA4F832" box="[571,832,1924,1948]" italics="true" pageId="5" pageNumber="248">Batrochoglanis castaneus</emphasis>
</taxonomicName>
, MZUEL 20113, 50.8 mm SL, paratype. A) dorsal view; b) lateral view; c) ventral view; d) medial view. Light blue = cartilage. Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="8997B3F1FFC7FF919E84FC53FF09FC70" ID-DOI="http://doi.org/10.5281/zenodo.3458981" ID-Zenodo-Dep="3458981" httpUri="https://zenodo.org/record/3458981/files/figure.png" pageId="6" pageNumber="249" startId="6.[151,250,930,955]" targetBox="[163,1424,186,901]" targetPageId="6">
<paragraph id="DD57E379FFC7FF919E84FC53FF09FC70" blockId="6.[151,1436,930,991]" pageId="6" pageNumber="249">
<emphasis id="EF9C3F6BFFC7FF919E84FC53FEF7FC15" bold="true" box="[151,275,930,955]" pageId="6" pageNumber="249">FIGURE 4.</emphasis>
Lower jaw of
<taxonomicName id="1AE898FAFFC7FF919FB9FC52FD54FC15" authorityName="Shibatta" authorityYear="2019" box="[426,688,931,955]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="249" phylum="Chordata" rank="species" species="castaneus">
<emphasis id="EF9C3F6BFFC7FF919FB9FC52FD54FC15" box="[426,688,931,955]" italics="true" pageId="6" pageNumber="249">Batrochoglanis castaneus</emphasis>
</taxonomicName>
, MZUEL 20113, 50.8 mm SL, paratype. Light blue = cartilage. Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="8997B3F1FFC7FF919E84F8BDFE1EF826" ID-DOI="http://doi.org/10.5281/zenodo.3458983" ID-Zenodo-Dep="3458983" httpUri="https://zenodo.org/record/3458983/files/figure.png" pageId="6" pageNumber="249" startId="6.[151,250,1868,1893]" targetBox="[164,1421,1006,1838]" targetPageId="6">
<paragraph id="DD57E379FFC7FF919E84F8BDFE1EF826" blockId="6.[151,1436,1868,1929]" pageId="6" pageNumber="249">
<emphasis id="EF9C3F6BFFC7FF919E84F8BDFEF0F8CB" bold="true" box="[151,276,1868,1893]" pageId="6" pageNumber="249">FIGURE 5.</emphasis>
Suspensorium of
<taxonomicName id="1AE898FAFFC7FF919FC3F8BCFD32F8CA" authorityName="Shibatta" authorityYear="2019" box="[464,726,1869,1893]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="249" phylum="Chordata" rank="species" species="castaneus">
<emphasis id="EF9C3F6BFFC7FF919FC3F8BCFD32F8CA" box="[464,726,1869,1893]" italics="true" pageId="6" pageNumber="249">Batrochoglanis castaneus</emphasis>
</taxonomicName>
, MZUEL 20113, 50.8 mm SL, paratype. Light blue = cartilage; dark gray = foramen. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="DD57E379FFC7FF909ED4F846FE5BFEC9" blockId="6.[151,1436,1975,2038]" lastBlockId="7.[151,1437,153,683]" lastPageId="7" lastPageNumber="250" pageId="6" pageNumber="249">
Suspensorium with broad hyomandibula anteriorly sutured to irregularly shaped metapterygoid, and ventrally to quadrate (
<figureCitation id="45D3FFFCFFC7FF919F15F82AFEADF85B" box="[262,329,2010,2037]" captionStart="FIGURE 5" captionStartId="6.[151,250,1868,1893]" captionTargetBox="[164,1421,1006,1838]" captionTargetId="figure@6.[151,1434,1006,1848]" captionTargetPageId="6" captionText="FIGURE 5. Suspensorium of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. Light blue = cartilage; dark gray = foramen. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3458983" httpUri="https://zenodo.org/record/3458983/files/figure.png" pageId="6" pageNumber="249">Fig. 5</figureCitation>
); dorsally articulated to pterotic and sphenotic, and posteriorly with opercle. Metapterygoid anterior region wider and with three pointed processes (basal, anterior, and ectopterygoid processes). Quadrate somewhat triangular, articulated by cartilage to hyomandibula and metapterygoid. Quadrate with a fenestra in ventral region to articulate with posterior dorsal projection of anguloarticular. Entopterygoid (
<figureCitation id="45D3FFFCFFC6FF909DE3FF10FBA6FF52" box="[1008,1090,225,252]" captionStart="FIGURE 2" captionStartId="5.[151,250,1129,1154]" captionTargetBox="[162,1425,276,1102]" captionTargetId="figure@5.[151,1436,265,1109]" captionTargetPageId="5" captionText="FIGURE 2. Cranium of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. A) dorsal view; b) ventral view. Light blue = cartilage; dark gray = fontanel or foramen. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3458977" httpUri="https://zenodo.org/record/3458977/files/figure.png" pageId="7" pageNumber="250">Fig. 2b</figureCitation>
) irregular shaped, with a sharp anterolateral process, articulated to ectopterygoid anteriorly and attached to metapterygoid by ligaments posteriorly. Ectopterygoid (
<figureCitation id="45D3FFFCFFC6FF909F55FED8FE7CFEEA" box="[326,408,297,324]" captionStart="FIGURE 2" captionStartId="5.[151,250,1129,1154]" captionTargetBox="[162,1425,276,1102]" captionTargetId="figure@5.[151,1436,265,1109]" captionTargetPageId="5" captionText="FIGURE 2. Cranium of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. A) dorsal view; b) ventral view. Light blue = cartilage; dark gray = fontanel or foramen. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3458977" httpUri="https://zenodo.org/record/3458977/files/figure.png" pageId="7" pageNumber="250">Fig. 2b</figureCitation>
) comma-shaped, attached to posteroventral surface of autopalatine and posteromedially articulated to entopterygoid.
</paragraph>
<paragraph id="DD57E379FFC6FF909ED4FE80FF2EFE59" blockId="7.[151,1437,153,683]" pageId="7" pageNumber="250">
Opercle (
<figureCitation id="45D3FFFCFFC6FF909F22FE80FE91FE25" box="[305,373,368,395]" captionStart="FIGURE 5" captionStartId="6.[151,250,1868,1893]" captionTargetBox="[164,1421,1006,1838]" captionTargetId="figure@6.[151,1434,1006,1848]" captionTargetPageId="6" captionText="FIGURE 5. Suspensorium of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. Light blue = cartilage; dark gray = foramen. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3458983" httpUri="https://zenodo.org/record/3458983/files/figure.png" pageId="7" pageNumber="250">Fig. 5</figureCitation>
) large, somewhat triangular, posteriorly rounded, anteriorly articulated to hyomandibula, and anteroventrally to interopercle. Preopercle long and thin, sutured along posterior margin of hyomandibula and quadrate. Interopercle triangular, slightly concave anteriorly, articulated to opercle posteriorly and preopercle anteriorly.
</paragraph>
<paragraph id="DD57E379FFC6FF909ED4FDF0FAF7FD05" blockId="7.[151,1437,153,683]" pageId="7" pageNumber="250">
Hyoid arch (
<figureCitation id="45D3FFFCFFC6FF909F45FDF0FE7FFDB2" box="[342,411,513,540]" captionStart="FIGURE 6" captionStartId="7.[151,250,1658,1683]" captionTargetBox="[162,1413,712,1624]" captionTargetId="figure@7.[151,1436,697,1639]" captionTargetPageId="7" captionText="FIGURE 6. Hyoid arch of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype.A) lateral view; b) dorsal view of the anterior region. Light blue = cartilage. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3458985" httpUri="https://zenodo.org/record/3458985/files/figure.png" pageId="7" pageNumber="250">Fig. 6</figureCitation>
) with small triangular ventral hypohyal, well developed anterior ceratohyal and triangular posterior ceratohyal. Dorsal hypohyal absent. Nine branchiostegal rays articulated with hyoid arch; six with anterior ceratohyal, one with interceratohyal cartilage, and two with posterior ceratohyal. These two posteriormost branchiostegal rays largest and posteriorly expanded. Parurohyal located between ventral hypohyals; anterior portion blunt, lateral process like triangular wing, medially with laminar process and short pointed posterior process.
</paragraph>
<caption id="8997B3F1FFC6FF909E84F98BFCE5F919" ID-DOI="http://doi.org/10.5281/zenodo.3458985" ID-Zenodo-Dep="3458985" httpUri="https://zenodo.org/record/3458985/files/figure.png" pageId="7" pageNumber="250" startId="7.[151,250,1658,1683]" targetBox="[162,1413,712,1624]" targetPageId="7">
<paragraph id="DD57E379FFC6FF909E84F98BFCE5F919" blockId="7.[151,1436,1658,1719]" pageId="7" pageNumber="250">
<emphasis id="EF9C3F6BFFC6FF909E84F98BFEF6F93D" bold="true" box="[151,274,1658,1683]" pageId="7" pageNumber="250">FIGURE 6.</emphasis>
Hyoid arch of
<taxonomicName id="1AE898FAFFC6FF909FB9F98AFD4BF93D" authorityName="Shibatta" authorityYear="2019" box="[426,687,1659,1683]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="250" phylum="Chordata" rank="species" species="castaneus">
<emphasis id="EF9C3F6BFFC6FF909FB9F98AFD4BF93D" box="[426,687,1659,1683]" italics="true" pageId="7" pageNumber="250">Batrochoglanis castaneus</emphasis>
</taxonomicName>
, MZUEL 20113, 50.8 mm SL, paratype. A) lateral view; b) dorsal view of the anterior region. Light blue = cartilage. Scale bar = 5 mm.
</paragraph>
</caption>
<paragraph id="DD57E379FFC6FF9F9ED4F914FB0FFEC9" blockId="7.[151,1437,1765,2043]" lastBlockId="8.[151,1437,153,359]" lastPageId="8" lastPageNumber="251" pageId="7" pageNumber="250">
Pharyngobranchial 1 absent. Pharyngobranchial 2 cartilaginous, elongated, articulated with epibranchials 1 and 2, and pharyngobranchial 3. Pharyngobranchial 3 elongated, ossified, somewhat triangular, with wider posterior region. Pharyngobranchial 4 rounded, ossified, with a tooth plate with irregular five rows of conical teeth. Epibranchial 1 rod-like, with a posterodorsal uncinate process, and a row of five elongated rakers. Epibranchial 2 slightly curved, with one row of elongated rakers. Epibranchial 3 rod-like with a posterior uncinate process, with two rows of small conical rakers. Epibranchial 4 broad, slightly convex anteriorly and posteriorly; toothless. First and second ceratobranchials supporting a single row of elongated rakers. Third and fourth ceratobranchial with two rows of rakers. Fifth ceratobranchial medially expanded, supporting tooth plate dorsally, with conic teeth in approximately six irregular rows. Hypobranchial 1 ossified, laminar, trapezoid, with medial margin cartilaginous; laterally to basibranchial 2. Hypobranchial 2 ossified, laminar, ovoid, with posterior margin cartilaginous; laterally to basibranchial 3. Hypobranchial 3 entirely cartilaginous, trapezoid, laterally to posterior cartilaginous tip of basibranchial 3. Hypobranchial 4 absent. Basibranchial 1 absent. Basibranchial 2 ossified, nearly spoon-shaped, with anterior cartilaginous tip, and posteriorly sharing cartilage with basibranchial 3. Basibranchial 3 small, elongated, ossified, with a small posterior cartilaginous tip. Basibranchial 4 pentagonal shaped, entirely cartilaginous (
<figureCitation id="45D3FFFCFFC9FF9F9A8AFEBCFB39FEC9" box="[1177,1245,333,359]" captionStart="FIGURE 7" captionStartId="8.[151,250,1022,1047]" captionTargetBox="[161,1429,384,989]" captionTargetId="figure@8.[151,1435,373,1002]" captionTargetPageId="8" captionText="FIGURE 7. Branchial arches of Batrochoglanis castaneus, a) dorsal view and b) ventral view of right epibranchials and pharyngobranchials, MZUEL 20113, 50.8 mm SL, paratype. Light blue = cartilage." figureDoi="http://doi.org/10.5281/zenodo.3458987" httpUri="https://zenodo.org/record/3458987/files/figure.png" pageId="8" pageNumber="251">Fig. 7</figureCitation>
).
</paragraph>
<caption id="8997B3F1FFC9FF9F9E84FC0FFC24FB95" ID-DOI="http://doi.org/10.5281/zenodo.3458987" ID-Zenodo-Dep="3458987" httpUri="https://zenodo.org/record/3458987/files/figure.png" pageId="8" pageNumber="251" startId="8.[151,250,1022,1047]" targetBox="[161,1429,384,989]" targetPageId="8">
<paragraph id="DD57E379FFC9FF9F9E84FC0FFC24FB95" blockId="8.[151,1436,1022,1083]" pageId="8" pageNumber="251">
<emphasis id="EF9C3F6BFFC9FF9F9E84FC0FFEF0FBB8" bold="true" box="[151,276,1022,1047]" pageId="8" pageNumber="251">FIGURE 7.</emphasis>
Branchial arches of
<taxonomicName id="1AE898FAFFC9FF9F9FFAFC0FFD0BFBB8" authorityName="Shibatta" authorityYear="2019" box="[489,751,1022,1046]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="251" phylum="Chordata" rank="species" species="castaneus">
<emphasis id="EF9C3F6BFFC9FF9F9FFAFC0FFD0BFBB8" box="[489,751,1022,1046]" italics="true" pageId="8" pageNumber="251">Batrochoglanis castaneus</emphasis>
</taxonomicName>
, a) dorsal view and b) ventral view of right epibranchials and pharyngobranchials, MZUEL 20113, 50.8 mm SL, paratype. Light blue = cartilage.
</paragraph>
</caption>
<paragraph id="DD57E379FFC9FF9F9ED4FB98FCEBFA0D" blockId="8.[151,1437,1129,1983]" pageId="8" pageNumber="251">
Cleithrum with a dorsoventral laminar bone forming a cylindrical crest anteriorly; right and left sides a little separated medially; posteriorly articulated with coracoid (
<figureCitation id="45D3FFFCFFC9FF9F9D0DFB7CFC87FB09" box="[798,867,1165,1191]" captionStart="FIGURE 8" captionStartId="9.[151,250,1151,1176]" captionTargetBox="[173,1414,194,1108]" captionTargetId="figure@9.[151,1436,174,1131]" captionTargetPageId="9" captionText="FIGURE 8. Ventral view of the left pectoral girdle of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. Light blue = cartilage. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3458989" httpUri="https://zenodo.org/record/3458989/files/figure.png" pageId="8" pageNumber="251">Fig. 8</figureCitation>
). Posterior cleithral process elongated, with a little blunt point, not surpassing anterior dorsal process of cleithrum posteriorly. Anterior and posterior dorsal process of vertical limb of cleithrum pointed; anterior larger than posterior. Coracoid wide, laminar; sutured to cleithrum anterolaterally; right and left coracoid interdigitated medially with strong serrations. Coracoid process triangular, articulated with pectoral-fin spine. Mesocoracoid arch thin. Radials elongated; first cartilaginous, posterior three ossified, articulated with six divided rays. Pectoral-fin spine serrated, anterior serrae antrorses (0(1*), 2(3), 3(1), 4(4), 5(1) serrae) and retrorses (2(1), 5(2), 6(2), 8(2), 10(1), 12(1), 13(1) serrae), total of 7(1), 9(3), 10(2), 12(2), 13(1*), 14(1); posterior serrae retrorse, 6(1), 8(4), 10(2), 11(3*).
</paragraph>
<paragraph id="DD57E379FFC9FF9F9ED4FA5CFAECF9A1" blockId="8.[151,1437,1129,1983]" pageId="8" pageNumber="251">
Pelvic girdle composed of a wide basipterygium, with two elongated anterior processes, the medial one (internal) wider and shorter than the lateral (external) (
<figureCitation id="45D3FFFCFFC9FF9F9CAEFA20FCE6FA45" box="[701,770,1489,1515]" captionStart="FIGURE 9" captionStartId="10.[151,250,1137,1162]" captionTargetBox="[165,1421,187,1101]" captionTargetId="figure@10.[151,1435,174,1117]" captionTargetPageId="10" captionText="FIGURE 9. Ventral view of right pelvic girdle of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. Light blue = cartilage. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3458993" httpUri="https://zenodo.org/record/3458993/files/figure.png" pageId="8" pageNumber="251">Fig. 9</figureCitation>
). Cartilage in lateral (including the lateral process), medial, and posterior sides, and on the tip of external anterior process. Six rays articulated to lateral cartilage.
</paragraph>
<paragraph id="DD57E379FFC9FF9F9ED4F9E8FB5DF881" blockId="8.[151,1437,1129,1983]" pageId="8" pageNumber="251">
Axial skeleton composed of three bones of Weberian apparatus, complex vertebra composed of vertebrae 1, 4 and 5; 28(6) or 29(3) free vertebrae; one compound caudal vertebra; and 5(1), 6(2), 7(6) or 8(1) ribs, with first rib attached to sixth centrum.
<emphasis id="EF9C3F6BFFC9FF9F9FB4F993FE0BF9D5" box="[423,495,1634,1659]" italics="true" pageId="8" pageNumber="251">Tripus</emphasis>
triangular, with the longest tip pointed anteriorly, medial-posteriorly articulated with
<emphasis id="EF9C3F6BFFC9FF9F9E84F977FEB5F931" box="[151,337,1670,1695]" italics="true" pageId="8" pageNumber="251">os suspensorium</emphasis>
.
<emphasis id="EF9C3F6BFFC9FF9F9F4DF974FE1DF931" box="[350,505,1669,1695]" italics="true" pageId="8" pageNumber="251">Intercalarium</emphasis>
slightly fusiform, little wrapped.
<taxonomicName id="1AE898FAFFC9FF9F9D63F974FC3AF931" box="[880,990,1669,1695]" class="Magnoliopsida" family="Malvaceae" genus="Scaphium" kingdom="Plantae" order="Malvales" pageId="8" pageNumber="251" phylum="Tracheophyta" rank="genus">
<emphasis id="EF9C3F6BFFC9FF9F9D63F974FC3AF931" box="[880,990,1669,1695]" italics="true" pageId="8" pageNumber="251">Scaphium</emphasis>
</taxonomicName>
curved spoon-shaped, with the concave region dorsal.
<emphasis id="EF9C3F6BFFC9FF9F9F2BF958FE4AF96D" box="[312,430,1705,1731]" italics="true" pageId="8" pageNumber="251">Claustrum</emphasis>
small, concave. Complex vertebra with anterior and posterior parapophysis of fourth vertebra and parapophysis of fifth vertebra (
<figureCitation id="45D3FFFCFFC9FF9F9C70F93CFD53F949" box="[611,695,1741,1767]" captionStart="FIGURE 10" captionStartId="11.[151,250,1615,1640]" captionTargetBox="[219,1366,181,1587]" captionTargetId="figure@11.[204,1382,174,1596]" captionTargetPageId="11" captionText="FIGURE 10. Complex vertebra of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. A) Ventral view; b) ventral-lateral view of the anterior region. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3458995" httpUri="https://zenodo.org/record/3458995/files/figure.png" pageId="8" pageNumber="251">Fig. 10</figureCitation>
).
<emphasis id="EF9C3F6BFFC9FF9F9CDFF93CFC69F949" box="[716,909,1741,1767]" italics="true" pageId="8" pageNumber="251">Os suspensorium</emphasis>
in the ventral region of parapophysis of fourth vertebra, lateral to centrum of complex vertebra, articulated with
<emphasis id="EF9C3F6BFFC9FF9F9D96F903FC22F8A5" box="[901,966,1778,1803]" italics="true" pageId="8" pageNumber="251">tripus</emphasis>
. The anterior process of parapophysis of fourth vertebra lateroposteriorly curved, anteriorly articulated to posttemporal-supracleithrum.
</paragraph>
<paragraph id="DD57E379FFC9FF9F9ED4F8C8FAEAF811" blockId="8.[151,1437,1129,1983]" pageId="8" pageNumber="251">
Dorsal fin with spinelet, one spine, and six divided rays supported by five wide proximal radials (
<figureCitation id="45D3FFFCFFC9FF9F9B11F8C8FAB7F8FD" box="[1282,1363,1849,1875]" captionStart="FIGURE 11" captionStartId="12.[151,250,1099,1124]" captionTargetBox="[200,1393,243,1062]" captionTargetId="figure@12.[183,1405,229,1079]" captionTargetPageId="12" captionText="FIGURE 11. Dorsal fin of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. A) dorsal view of nuchal plate and first two rays; b) lateral view. Light blue = cartilage. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3458997" httpUri="https://zenodo.org/record/3458997/files/figure.png" pageId="8" pageNumber="251">Fig. 11</figureCitation>
). Dorsal-fin spine smooth anteriorly and posteriorly, with no serrae or denticuli. Anterior nuchal plate triangular, anteriorly pointed. Middle nuchal plate trapezoidal, and posterior third paired and rectangular. Dorsal-fin rays articulated to wide pterygiophores by cartilages. Third to seventh pterygiophores broad, only narrowly separated.
</paragraph>
<caption id="8997B3F1FFC8FF9E9E84FB8EFE18FB15" ID-DOI="http://doi.org/10.5281/zenodo.3458989" ID-Zenodo-Dep="3458989" httpUri="https://zenodo.org/record/3458989/files/figure.png" pageId="9" pageNumber="252" startId="9.[151,250,1151,1176]" targetBox="[173,1414,194,1108]" targetPageId="9">
<paragraph id="DD57E379FFC8FF9E9E84FB8EFE18FB15" blockId="9.[151,1436,1151,1212]" pageId="9" pageNumber="252">
<emphasis id="EF9C3F6BFFC8FF9E9E84FB8EFEF7FB36" bold="true" box="[151,275,1151,1176]" pageId="9" pageNumber="252">FIGURE 8.</emphasis>
Ventral view of the left pectoral girdle of
<taxonomicName id="1AE898FAFFC8FF9E9CAEFB71FC27FB36" authorityName="Shibatta" authorityYear="2019" box="[701,963,1152,1176]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="252" phylum="Chordata" rank="species" species="castaneus">
<emphasis id="EF9C3F6BFFC8FF9E9CAEFB71FC27FB36" box="[701,963,1152,1176]" italics="true" pageId="9" pageNumber="252">Batrochoglanis castaneus</emphasis>
</taxonomicName>
, MZUEL 20113, 50.8 mm SL, paratype. Light blue = cartilage. Scale bar = 5 mm.
</paragraph>
</caption>
<paragraph id="DD57E379FFC8FF9E9ED4FB1BFEFEFADE" blockId="9.[151,1437,1258,2040]" pageId="9" pageNumber="252">
Anal fin composed of proximal radials, distal radials, and ii(3*), iii(6) + 6(1), 7(6), 8(2*) rays (
<figureCitation id="45D3FFFCFFC8FF9E9AFCFB1BFAA6FAAA" box="[1263,1346,1258,1284]" captionStart="FIGURE 12" captionStartId="12.[151,250,1892,1917]" captionTargetBox="[191,1384,1184,1861]" captionTargetId="figure@12.[185,1402,1175,1873]" captionTargetPageId="12" captionText="FIGURE 12. Lateral view of anal-fin skeleton of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. Light blue = cartilage. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3458999" httpUri="https://zenodo.org/record/3458999/files/figure.png" pageId="9" pageNumber="252">Fig. 12</figureCitation>
). Proximal radials elongated, pointed, with laminar process; the last one rectangular shaped. Distal radials rounded, first two and last cartilaginous, others ossified and anteriorly cartilaginous. First anal-fin ray rudimentary, usually hidden by the skin.
</paragraph>
<paragraph id="DD57E379FFC8FF9E9ED4FA8BFC50F98B" blockId="9.[151,1437,1258,2040]" pageId="9" pageNumber="252">
Caudal fin skeleton (
<figureCitation id="45D3FFFCFFC8FF9E9FA9FA8BFDE9FA3A" box="[442,525,1402,1428]" captionStart="FIGURE 13" captionStartId="13.[151,250,1167,1192]" captionTargetBox="[162,1417,190,1137]" captionTargetId="figure@13.[151,1436,174,1148]" captionTargetPageId="13" captionText="FIGURE 13. Caudal skeleton of Batrochoglanis castaneus, MZUEL 20113, 50.8 mm SL, paratype. Light blue = cartilage. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3459001" httpUri="https://zenodo.org/record/3459001/files/figure.png" pageId="9" pageNumber="252">Fig. 13</figureCitation>
) composed of a plate formed by parhypural + hypurals 1 and 2, another plate formed by hypurals 3 and 4, hypural 5 free, and a pleurostile fused to compound centrum. A free elongated epural, slightly curved, with distal portion wider. Opisthural cartilage slightly rounded. A laminar secondary hypurapophysis fused to the anterodorsal region of hypural 2 (
<typeStatus id="02535DDBFFC8FF9E9CA6FA17FD03F9AE" box="[693,743,1510,1536]" pageId="9" pageNumber="252">type</typeStatus>
C hypurapophysis
<emphasis id="EF9C3F6BFFC8FF9E9DAEFA16FC1FF9AE" box="[957,1019,1511,1536]" italics="true" pageId="9" pageNumber="252">sensu</emphasis>
<bibRefCitation id="B9799E88FFC8FF9E9A12FA17FAC2F9AF" author="Lundberg, J. G. &amp; Baskin, J. N." box="[1025,1318,1510,1537]" pageId="9" pageNumber="252" pagination="1 - 49" refId="ref9632" refString="Lundberg, J. G. &amp; Baskin, J. N. (1969) The caudal skeleton of the catfishes, order Siluriformes. American Museum Novitates, 2398, 1 - 49." type="journal article" year="1969">Lundberg &amp; Baskin, 1969</bibRefCitation>
). Eighteen dorsal procurrent rays; i,7,6,i principal rays; 11 ventral procurrent rays.
</paragraph>
<paragraph id="DD57E379FFC8FF9E9ED4F9DFFF28F856" blockId="9.[151,1437,1258,2040]" pageId="9" pageNumber="252">
<emphasis id="EF9C3F6BFFC8FF9E9ED4F9DFFDA6F9E6" bold="true" box="[199,578,1581,1608]" pageId="9" pageNumber="252">
Laterosensory system (
<figureCitation id="45D3FFFCFFC8FF9E9FCEF9DCFDD7F9E6" box="[477,563,1581,1608]" captionStart="FIGURE 14" captionStartId="14.[151,248,1947,1972]" captionTargetBox="[416,1166,216,1898]" captionTargetId="figure@14.[373,1215,179,1920]" captionTargetPageId="14" captionText="FIGURE 14. Laterosensory system of Batrochoglanis castaneus, INPA 8059, 58.3 mm SL, holotype. Scale bar = 5 mm. Open circle = laterosensory pore (i = infraorbital; ll = lateral line; pm = preoperculomandibular; po = postotic; s = supraorbital); dark dot = neuromast (al = anterior line; dtl = dorsal trunk line; ml = maxillary line; nl = nasal line; stal = supratemporal accessory line)." figureDoi="http://doi.org/10.5281/zenodo.3459003" httpUri="https://zenodo.org/record/3459003/files/figure.png" pageId="9" pageNumber="252">Fig. 14</figureCitation>
).
</emphasis>
Head sensory canals composed of single pores. Supraorbital sensory canal continuous with five pores (s1, s2, s3, s6 (epiphyseal branch), and s8 (parietal branch)) and connected to otic and infraorbital sensory canals posteriorly. Left and right epiphyseal branches (s6) never fused medially. S4 and s5 branch and pore absent. Presence of s7 pore (postorbital) in two of eight examined specimens. Otic sensory canal short, without pores, anteriorly continuous with supra- and infraorbital sensory canals, and posteriorly with postotic sensory canal. Postotic (or temporal) sensory canal located between posterior limit of otic sensory canal and anterior limit of lateral line, with two branches (po1, and po2). First postotic branch (po1) fused to a posteriormost branch of preoperculomandibular sensory canal (pm11), forming po1+pm11 complex pore. Infraorbital sensory canal with six branches (
<date id="A956C5B9FFC8FF9E9F20F8BFFE8EF8C7" box="[307,362,1870,1897]" pageId="9" pageNumber="252">i1-i6</date>
). Preoperculomandibular sensory canal with 11 pores; pm1 not fused to its opposite counterpart branch. Lateral-line sensory canal continuous with postotic sensory canal and extending to vertical through origin of anal fin. First lateral-line pore immediately posterior to vertical through posterior margin of opercular bone. Lateral line with 18(1), 20(3), 21(2*) pores on the left side, and 18(1), 19(2), 20(1), 22(1), and 24(1*) pores on the right side.
</paragraph>
<caption id="8997B3F1FFCBFF9D9E84FB80FE2EFB03" ID-DOI="http://doi.org/10.5281/zenodo.3458993" ID-Zenodo-Dep="3458993" httpUri="https://zenodo.org/record/3458993/files/figure.png" pageId="10" pageNumber="253" startId="10.[151,250,1137,1162]" targetBox="[165,1421,187,1101]" targetPageId="10">
<paragraph id="DD57E379FFCBFF9D9E84FB80FE2EFB03" blockId="10.[151,1437,1137,1198]" pageId="10" pageNumber="253">
<emphasis id="EF9C3F6BFFCBFF9D9E84FB80FEF6FB24" bold="true" box="[151,274,1137,1162]" pageId="10" pageNumber="253">FIGURE 9.</emphasis>
Ventral view of right pelvic girdle of
<taxonomicName id="1AE898FAFFCBFF9D9C9CFB80FC70FB27" authorityName="Shibatta" authorityYear="2019" box="[655,916,1137,1161]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="253" phylum="Chordata" rank="species" species="castaneus">
<emphasis id="EF9C3F6BFFCBFF9D9C9CFB80FC70FB27" box="[655,916,1137,1161]" italics="true" pageId="10" pageNumber="253">Batrochoglanis castaneus</emphasis>
</taxonomicName>
, MZUEL 20113, 50.8 mm SL, paratype. Light blue = cartilage. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="DD57E379FFCBFF9D9ED4FB2DFEFAFA28" blockId="10.[151,1437,1244,1918]" pageId="10" pageNumber="253">
Rostral line with one neuromast located between anterior nostrils; nasal line with four neuromasts, in front of anterior nostril; anterior line in the occipital region, after pore S8, with two neuromasts; dorsal trunk line anterior to dorsal fin, with two neuromasts; mandibular line with four neuromasts in the ventral region of head, forming an arch between
<date id="A956C5B9FFCBFF9D9F24FAB9FE79FACC" box="[311,413,1351,1378]" pageId="10" pageNumber="253">i4 and i5</date>
pores and pm6 and pm7; supratemporal accessory line located posteriorly to po2, with 2 neuromasts.
</paragraph>
<paragraph id="DD57E379FFCBFF9D9ED4FA7EFD4AF908" blockId="10.[151,1437,1244,1918]" pageId="10" pageNumber="253">
<emphasis id="EF9C3F6BFFCBFF9D9ED4FA7EFE69FA04" bold="true" box="[199,397,1423,1450]" pageId="10" pageNumber="253">Color in alcohol.</emphasis>
Head and trunk dark brown. Light brown stripe along nape, running from right to left pectoral fins. Light brown spots above lateral line soon after vertical through dorsal fin terminus, and light brown stripe crossing caudal peduncle transversely, between adipose and anal fins. Light brown saddle-shaped spot sometimes between adipose and caudal fins. Adipose fin dark brown with light brown spot on the anterior and posterior region. Dorsal fin dark brown, with hyaline stripe on top and medially on five last rays. Pectoral and pelvic fins dark brown with hyaline spot on base and margin. Anal fin with dark stripe on base and middle. Posterior cleithral process light brown. Caudal fin hyaline, with dark brown spots scattered on first two thirds, and broad dark brown band on posterior third. Ventral region light brown. (
<figureCitation id="45D3FFFCFFCBFF9D9C4FF97DFD79F908" box="[604,669,1676,1702]" captionStart="FIGURE 1" captionStartId="4.[151,250,1713,1738]" captionTargetBox="[151,1436,174,1693]" captionTargetId="figure@4.[151,1436,174,1693]" captionTargetPageId="4" captionText="FIGURE 1. Holotype of Batrochoglanis castaneus, INPA 8059, 58,3 mm SL, Aripuanã River, Igarapé do Castanhal, MT." figureDoi="http://doi.org/10.5281/zenodo.3458975" httpUri="https://zenodo.org/record/3458975/files/figure.png" pageId="10" pageNumber="253">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="95F2B0F2FFCBFF9D9ED4F95EFAC3F8D0" pageId="10" pageNumber="253" type="distribution">
<paragraph id="DD57E379FFCBFF9D9ED4F95EFAC3F8D0" blockId="10.[151,1437,1244,1918]" pageId="10" pageNumber="253">
<emphasis id="EF9C3F6BFFCBFF9D9ED4F95EFE84F964" bold="true" box="[199,352,1711,1738]" pageId="10" pageNumber="253">Distribution.</emphasis>
Known only from the type-locality, Igarapé do Castanhal, Aripuanã River basin, state of
<collectingRegion id="1F2C2D9BFFCBFF9D9B72F941FF0DF940" country="Brazil" name="Mato Grosso" pageId="10" pageNumber="253">Mato Grosso</collectingRegion>
,
<collectingCountry id="A5FFA3E9FFCBFF9D9EE5F925FEDAF940" box="[246,318,1748,1774]" name="Brazil" pageId="10" pageNumber="253">Brazil</collectingCountry>
. No coordinate or municipality was recovered from the labels, which did not enable determination of a specific site of the type-locality. The collecting site is tentatively estimated at
<geoCoordinate id="B8DC85BEFFCBFF9D9A4CF906FB0DF8BC" box="[1119,1257,1783,1810]" degrees="59" direction="west" minutes="27" orientation="longitude" pageId="10" pageNumber="253" precision="15" seconds="34" value="-59.459446">
59
<superScript id="2A9D4E31FFCBFF9D9A68F906FB67F8AB" attach="none" box="[1147,1155,1783,1797]" fontSize="6" pageId="10" pageNumber="253">o</superScript>
27’34”W
</geoCoordinate>
,
<geoCoordinate id="B8DC85BEFFCBFF9D9AE8F909FA98F8BC" box="[1275,1404,1783,1810]" degrees="10" direction="south" minutes="10" orientation="latitude" pageId="10" pageNumber="253" precision="15" seconds="00" value="-10.166667">
10
<superScript id="2A9D4E31FFCBFF9D9B04F906FAFBF8AB" attach="none" box="[1303,1311,1783,1797]" fontSize="6" pageId="10" pageNumber="253">o</superScript>
10’00”S
</geoCoordinate>
in SpeciesLink (http://splink.cria.org.br/), which is near Aripuanã City. However, in the Aripuanã River basin, a river called Igarapé Castanhal occurs in the Colniza municipality, state of
<collectingRegion id="1F2C2D9BFFCBFF9D9D8EF8B1FBCAF8F4" box="[925,1070,1856,1882]" country="Brazil" name="Mato Grosso" pageId="10" pageNumber="253">Mato Grosso</collectingRegion>
(e.g.,
<geoCoordinate id="B8DC85BEFFCBFF9D9A65F8B1FB16F8F4" box="[1142,1266,1855,1882]" degrees="9" direction="south" minutes="41" orientation="latitude" pageId="10" pageNumber="253" precision="15" seconds="10" value="-9.686111">
9
<superScript id="2A9D4E31FFCBFF9D9A97F8CEFB68F8E3" attach="none" box="[1156,1164,1855,1869]" fontSize="6" pageId="10" pageNumber="253">o</superScript>
41’10” S
</geoCoordinate>
<geoCoordinate id="B8DC85BEFFCBFF9D9AEAF8CEFA6FF8F4" box="[1273,1419,1855,1882]" degrees="59" direction="west" minutes="35" orientation="longitude" pageId="10" pageNumber="253" precision="15" seconds="19" value="-59.58861">
59
<superScript id="2A9D4E31FFCBFF9D9B06F8CEFAF9F8E3" attach="none" box="[1301,1309,1855,1869]" fontSize="6" pageId="10" pageNumber="253">o</superScript>
35’19” W
</geoCoordinate>
). Therefore, it is very imprecise to determine the exact point of type-locality with the current information.
</paragraph>
</subSubSection>
<caption id="8997B3F1FFCAFF9C9E84F9BEFD38F925" ID-DOI="http://doi.org/10.5281/zenodo.3458995" ID-Zenodo-Dep="3458995" httpUri="https://zenodo.org/record/3458995/files/figure.png" pageId="11" pageNumber="254" startId="11.[151,250,1615,1640]" targetBox="[219,1366,181,1587]" targetPageId="11">
<paragraph id="DD57E379FFCAFF9C9E84F9BEFD38F925" blockId="11.[151,1436,1615,1676]" pageId="11" pageNumber="254">
<emphasis id="EF9C3F6BFFCAFF9C9E84F9BEFEC4F9C6" bold="true" box="[151,288,1615,1640]" pageId="11" pageNumber="254">FIGURE 10.</emphasis>
Complex vertebra of
<taxonomicName id="1AE898FAFFCAFF9C9FEDF9BEFCE0F9C9" authorityName="Shibatta" authorityYear="2019" box="[510,772,1615,1639]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="254" phylum="Chordata" rank="species" species="castaneus">
<emphasis id="EF9C3F6BFFCAFF9C9FEDF9BEFCE0F9C9" box="[510,772,1615,1639]" italics="true" pageId="11" pageNumber="254">Batrochoglanis castaneus</emphasis>
</taxonomicName>
, MZUEL 20113, 50.8 mm SL, paratype. A) Ventral view; b) ventral-lateral view of the anterior region. Scale bar = 1 mm.
</paragraph>
</caption>
<subSubSection id="95F2B0F2FFCAFF9A9ED4F94BFAFEFA0A" lastPageId="13" lastPageNumber="256" pageId="11" pageNumber="254" type="etymology">
<paragraph id="DD57E379FFCAFF9C9ED4F94BFA92F957" blockId="11.[151,1437,1722,2036]" pageId="11" pageNumber="254">
<emphasis id="EF9C3F6BFFCAFF9C9ED4F94BFEAAF97A" bold="true" box="[199,334,1722,1748]" pageId="11" pageNumber="254">Etymology.</emphasis>
The species name,
<taxonomicName id="1AE898FAFFCAFF9C9C37F94AFD76F97A" authorityName="Shibatta" authorityYear="2019" box="[548,658,1723,1748]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="254" phylum="Chordata" rank="species" species="castaneus">
<emphasis id="EF9C3F6BFFCAFF9C9C37F94AFD76F97A" box="[548,658,1723,1748]" italics="true" pageId="11" pageNumber="254">castaneus</emphasis>
</taxonomicName>
is a Latin word that means the color of chestnuts, related to the brownish body color of analyzed specimens and to the
<collectingCountry id="A5FFA3E9FFCAFF9C9CABF92FFD18F956" box="[696,764,1758,1784]" name="Brazil" pageId="11" pageNumber="254">Brazil</collectingCountry>
nut that named the type-locality (Igarapé do Castanhal).
</paragraph>
<paragraph id="DD57E379FFCAFF9B9ED4F8F3FDC1FF79" blockId="11.[151,1437,1722,2036]" lastBlockId="12.[151,1436,153,215]" lastPageId="12" lastPageNumber="255" pageId="11" pageNumber="254">
<emphasis id="EF9C3F6BFFCAFF9C9ED4F8F3FE92F8B2" bold="true" box="[199,374,1794,1820]" pageId="11" pageNumber="254">Morphometry.</emphasis>
The first Principal Component retained 96.3% of the original data variance (eigenvalue = 0.591616), all variable loadings were positive, which allows identification of this axis as the representative size. The morphometric discrimination of the new species concerning the comparative congeners occurred in the second axis, which retained 1.17% of the variance (eigenvalue = 0.00720634). Also, in this axis, it is observed that
<taxonomicName id="1AE898FAFFCAFF9C9B35F89EFA79F826" baseAuthorityName="Valenciennes" baseAuthorityYear="1840" box="[1318,1437,1903,1928]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="254" phylum="Chordata" rank="species" species="raninus">
<emphasis id="EF9C3F6BFFCAFF9C9B35F89EFA79F826" box="[1318,1437,1903,1928]" italics="true" pageId="11" pageNumber="254">B. raninus</emphasis>
</taxonomicName>
<typeStatus id="02535DDBFFCAFF9C9E84F863FF0AF802" box="[151,238,1938,1964]" pageId="11" pageNumber="254" type="syntype">syntype</typeStatus>
was closer to the group of
<taxonomicName id="1AE898FAFFCAFF9C9C33F862FD71F802" baseAuthorityName="Valenciennes" baseAuthorityYear="1840" box="[544,661,1939,1964]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="254" phylum="Chordata" rank="species" species="raninus">
<emphasis id="EF9C3F6BFFCAFF9C9C33F862FD71F802" box="[544,661,1939,1964]" italics="true" pageId="11" pageNumber="254">B. raninus</emphasis>
</taxonomicName>
of Amazonia. In the third axis, which retained 0.60% of the variance (eigenvalue = 0.00367162), intraspecific variation was observed in
<taxonomicName id="1AE898FAFFCAFF9C9D80F846FBBDF87E" baseAuthorityName="Regan" baseAuthorityYear="1913" box="[915,1113,1975,2000]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="254" phylum="Chordata" rank="species" species="transmontanus">
<emphasis id="EF9C3F6BFFCAFF9C9D80F846FBBDF87E" box="[915,1113,1975,2000]" italics="true" pageId="11" pageNumber="254">B. transmontanus</emphasis>
</taxonomicName>
(
<figureCitation id="45D3FFFCFFCAFF9C9A79F847FB5AF87E" box="[1130,1214,1973,2000]" captionStart="FIGURE 15" captionStartId="16.[151,250,1166,1191]" captionTargetBox="[180,1410,190,1136]" captionTargetId="figure@16.[164,1424,174,1147]" captionTargetPageId="16" captionText="FIGURE 15. Scatterplot of individual scores of combined samples of Batrochoglanis castaneus (filled vine square, n=9), B. raninus from Amazon basin (green X, n=30), B. raninus syntype (red cross, n=1), B. transmontanus (open blue square, n=5) on the second and third axis of Principal Components." figureDoi="http://doi.org/10.5281/zenodo.3459005" httpUri="https://zenodo.org/record/3459005/files/figure.png" pageId="11" pageNumber="254">Fig. 15</figureCitation>
). From the analysis of individual scores distribution in PC1 and PC2, it was observed that the smaller individuals of
<taxonomicName id="1AE898FAFFCAFF9C9AC5F82AFA78F85A" baseAuthorityName="Regan" baseAuthorityYear="1913" box="[1238,1436,2011,2036]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="254" phylum="Chordata" rank="species" species="transmontanus">
<emphasis id="EF9C3F6BFFCAFF9C9AC5F82AFA78F85A" box="[1238,1436,2011,2036]" italics="true" pageId="11" pageNumber="254">B. transmontanus</emphasis>
</taxonomicName>
differ morphometrically from the larger individuals. In turn, the largest individuals tended to have similar morphometric characteristics to
<taxonomicName id="1AE898FAFFCDFF9B9FB9FF4FFDFBFF79" baseAuthorityName="Valenciennes" baseAuthorityYear="1840" box="[426,543,190,215]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="255" phylum="Chordata" rank="species" species="raninus">
<emphasis id="EF9C3F6BFFCDFF9B9FB9FF4FFDFBFF79" box="[426,543,190,215]" italics="true" pageId="12" pageNumber="255">B. raninus</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="8997B3F1FFCDFF9B9E84FBBAFC70FB29" ID-DOI="http://doi.org/10.5281/zenodo.3458997" ID-Zenodo-Dep="3458997" httpUri="https://zenodo.org/record/3458997/files/figure.png" pageId="12" pageNumber="255" startId="12.[151,250,1099,1124]" targetBox="[200,1393,243,1062]" targetPageId="12">
<paragraph id="DD57E379FFCDFF9B9E84FBBAFC70FB29" blockId="12.[151,1436,1099,1160]" pageId="12" pageNumber="255">
<emphasis id="EF9C3F6BFFCDFF9B9E84FBBAFEC4FBCA" bold="true" box="[151,288,1099,1124]" pageId="12" pageNumber="255">FIGURE 11.</emphasis>
Dorsal fin of
<taxonomicName id="1AE898FAFFCDFF9B9FA1FBBDFD5DFBCA" authorityName="Shibatta" authorityYear="2019" box="[434,697,1100,1124]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="255" phylum="Chordata" rank="species" species="castaneus">
<emphasis id="EF9C3F6BFFCDFF9B9FA1FBBDFD5DFBCA" box="[434,697,1100,1124]" italics="true" pageId="12" pageNumber="255">Batrochoglanis castaneus</emphasis>
</taxonomicName>
, MZUEL 20113, 50.8 mm SL, paratype. A) dorsal view of nuchal plate and first two rays; b) lateral view. Light blue = cartilage. Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="8997B3F1FFCDFF9B9E84F895FE2EF80E" ID-DOI="http://doi.org/10.5281/zenodo.3458999" ID-Zenodo-Dep="3458999" httpUri="https://zenodo.org/record/3458999/files/figure.png" pageId="12" pageNumber="255" startId="12.[151,250,1892,1917]" targetBox="[191,1384,1184,1861]" targetPageId="12">
<paragraph id="DD57E379FFCDFF9B9E84F895FE2EF80E" blockId="12.[151,1437,1892,1953]" pageId="12" pageNumber="255">
<emphasis id="EF9C3F6BFFCDFF9B9E84F895FEFBF8D3" bold="true" box="[151,287,1892,1917]" pageId="12" pageNumber="255">FIGURE 12.</emphasis>
Lateral view of anal-fin skeleton of
<taxonomicName id="1AE898FAFFCDFF9B9C9CF894FC70F8D3" authorityName="Shibatta" authorityYear="2019" box="[655,916,1893,1917]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="255" phylum="Chordata" rank="species" species="castaneus">
<emphasis id="EF9C3F6BFFCDFF9B9C9CF894FC70F8D3" box="[655,916,1893,1917]" italics="true" pageId="12" pageNumber="255">Batrochoglanis castaneus</emphasis>
</taxonomicName>
, MZUEL 20113, 50.8 mm SL, paratype. Light blue = cartilage. Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="8997B3F1FFCCFF9A9E84FB7EFEF0FB65" ID-DOI="http://doi.org/10.5281/zenodo.3459001" ID-Zenodo-Dep="3459001" httpUri="https://zenodo.org/record/3459001/files/figure.png" pageId="13" pageNumber="256" startId="13.[151,250,1167,1192]" targetBox="[162,1417,190,1137]" targetPageId="13">
<paragraph id="DD57E379FFCCFF9A9E84FB7EFEF0FB65" blockId="13.[151,1436,1167,1228]" pageId="13" pageNumber="256">
<emphasis id="EF9C3F6BFFCCFF9A9E84FB7EFEFAFB06" bold="true" box="[151,286,1167,1192]" pageId="13" pageNumber="256">FIGURE 13.</emphasis>
Caudal skeleton of
<taxonomicName id="1AE898FAFFCCFF9A9FF7FB61FD0DFB06" authorityName="Shibatta" authorityYear="2019" box="[484,745,1168,1192]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="256" phylum="Chordata" rank="species" species="castaneus">
<emphasis id="EF9C3F6BFFCCFF9A9FF7FB61FD0DFB06" box="[484,745,1168,1192]" italics="true" pageId="13" pageNumber="256">Batrochoglanis castaneus</emphasis>
</taxonomicName>
, MZUEL 20113, 50.8 mm SL, paratype. Light blue = cartilage. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="DD57E379FFCCFF9A9ED4FAEFFAFEFA0A" blockId="13.[151,1436,1310,1445]" pageId="13" pageNumber="256">
The morphometric distinction of
<taxonomicName id="1AE898FAFFCCFF9A9C53FAEEFD2AFA96" authorityName="Shibatta" authorityYear="2019" box="[576,718,1311,1336]" class="Actinopterygii" family="Pseudopimelodidae" genus="Batrochoglanis" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="256" phylum="Chordata" rank="species" species="castaneus">
<emphasis id="EF9C3F6BFFCCFF9A9C53FAEEFD2AFA96" box="[576,718,1311,1336]" italics="true" pageId="13" pageNumber="256">B. castaneus</emphasis>
</taxonomicName>
in relation to the other species analyzed was mainly due to the following characteristics: greater maxillary barbel length, larger eye diameter, greater dorsal-fin spine length, and larger mouth width (positive loadings on PC2). In turn, the new species had the smallest lengths of the adipose-fin base, anal-fin origin to the adipose-fin terminus and anal-fin base (negative loading on PC2). (
<tableCitation id="906AD6C2FFCCFF9A9AA6FA7BFAE9FA0A" box="[1205,1293,1418,1444]" captionStart="TABLE 2" captionStartId="15.[151,239,154,178]" captionTargetPageId="15" captionText="TABLE 2. Variable loadings on PC1 to PC3 of combined samples of B. cataneus, B. raninus from Amazon, B. raninus syntype and B. transmontanus." httpUri="http://table.plazi.org/id/8997B3F1FFCEFF989E84FF6BFE05FF7B" pageId="13" pageNumber="256" tableUuid="8997B3F1FFCEFF989E84FF6BFE05FF7B">Table 2</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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