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<mods:title id="0FBB3302F74B6FC4536761B549C36F1A">Revision of the troglomorphic genus Troglostygnopsis Šilhavý, 1974 (Opiliones, Laniatores, Stygnopsidae)</mods:title>
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<mods:affiliation id="3746800F67F850E2933F0C9B18997C20">Colección Nacional de Arácnidos, Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México. 3 er. Circuito exterior s / n. Apartado Postal 70 - 153, C. P. 04510, Ciudad Universitaria, Coyoacán, Mexico City, Mexico. &amp; Instituto Nacional de Investigaciones Forestales, Agrícolas y Pecuarias (INIFAP), Campo Experimental Valles Centrales, Oaxaca, Melchor Ocampo No. 7, C. P. 68200, Santo Domingo Barrio Bajo, Etla, Oaxaca, Mexico.</mods:affiliation>
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<taxonomicName id="1A944D75034CC31BBC65825DFBBA302F" ID-CoL="7Z6HK" authority="Silhavy, 1974" authorityName="Silhavy" authorityYear="1974" box="[522,1066,264,294]" class="Arachnida" family="Stygnopsidae" genus="Troglostygnopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="78" phylum="Arthropoda" rank="species" species="anophthalma">
<emphasis id="EFE0EAE4034CC31BBC65825DFCEF302F" bold="true" box="[522,895,264,291]" italics="true" pageId="4" pageNumber="78">Troglostygnopsis anophthalma</emphasis>
<bibRefCitation id="B9054B07034CC31BBDE9825FFBBA302F" author="Silhavy V." box="[902,1066,265,294]" pageId="4" pageNumber="78" pagination="175 - 194" refId="ref6023" refString="Silhavy V. 1974. Cavernicolous opilionids from Mexico. Subterranean fauna of Mexico. Part. II. Quaderno Accademia Nazionale dei Lincei 171: 175 - 194." type="journal article" year="1974">Šilhavý, 1974</bibRefCitation>
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<figureCitation id="45AF2A73034CC31BBC888279FCDD304A" box="[743,845,300,326]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="3.[189,232,1963,1989]" captionStartId-1="4.[189,232,1963,1989]" captionStartId-2="5.[189,232,1963,1989]" captionTargetBox-0="[198,1389,272,1927]" captionTargetBox-1="[189,1398,1162,1927]" captionTargetBox-2="[189,1398,1251,1921]" captionTargetId-0="figure-15@3.[198,1389,265,1927]" captionTargetId-1="figure-156@4.[189,1398,1162,1927]" captionTargetId-2="figure-283@5.[189,1398,1251,1921]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Fig. 1. Troglostygnopsis anophthalma Šilhavý, 1974, ♂, topotype (AMNH). A. Chelicera, mesal view. B. Chelicera, frontal view. C. Pedipalpal femur, mesal view. D. Pedipalpal tibia and tarsus, ventral view." captionText-1="Fig. 2. Troglostygnopsis anophthalma Šilhavý, 1974, ♂, topotype (AMNH). A. Apical femur, patella and basal tibia IV. B. Patella IV." captionText-2="Fig. 3. Troglostygnopsis anophthalma Šilhavý, 1974, ♂, topotype (AMNH), penis. A. Dorsal view. B. Lateral view. C. Ventral view. Scale bar in C applicable to all figures." figureDoi-0="http://doi.org/10.5281/zenodo.5708142" figureDoi-1="http://doi.org/10.5281/zenodo.5708144" figureDoi-2="http://doi.org/10.5281/zenodo.5708146" httpUri-0="https://zenodo.org/record/5708142/files/figure.png" httpUri-1="https://zenodo.org/record/5708144/files/figure.png" httpUri-2="https://zenodo.org/record/5708146/files/figure.png" pageId="4" pageNumber="78">Figs 13</figureCitation>
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<emphasis id="EFE0EAE4034CC31BBED2823BFDB73084" box="[189,551,366,392]" italics="true" pageId="4" pageNumber="78">Troglostygnopsis anophthalma</emphasis>
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<taxonomicName id="1A944D75034CC31BBED282FAFDB930C5" ID-CoL="7Z6HK" authorityName="Silhavy" authorityYear="1974" box="[189,553,431,457]" class="Arachnida" family="Stygnopsidae" genus="Troglostygnopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="78" phylum="Arthropoda" rank="species" species="anophthalma">Troglostygnopsis anophthalma</taxonomicName>
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<treatmentCitation id="5C3510E7034CC31BBC2482FAFCFD30C5" author="Sbordoni V. &amp; Argano R. &amp; Zullini A." box="[587,877,431,457]" page="32" pageId="4" pageNumber="78" year="1974">
<bibRefCitation id="B9054B07034CC31BBC2482FAFCFD30C5" author="Sbordoni V. &amp; Argano R. &amp; Zullini A." box="[587,877,431,457]" pageId="4" pageNumber="78" pagination="5 - 45" refId="ref5983" refString="Sbordoni V., Argano R. &amp; Zullini A. 1974. Biological investigations on the caves of Chiapas (Mexico) and adjacent countries: Introduction. Quaderno Accademia Nazionale dei Lincei 171: 5 - 45." type="journal article" year="1974">
Sbordoni
<emphasis id="EFE0EAE4034CC31BBCD182E5FD6B30C5" box="[702,763,431,457]" italics="true" pageId="4" pageNumber="78">et al.</emphasis>
1974: 32
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(nomen nudum). —
<treatmentCitation id="5C3510E7034CC31BBA0382FAFADC30C6" author="Reddell J. R." box="[1132,1356,431,458]" page="165" pageId="4" pageNumber="78" year="1981">
<bibRefCitation id="B9054B07034CC31BBA0382FAFADC30C6" author="Reddell J. R." box="[1132,1356,431,458]" pageId="4" pageNumber="78" pagination="69 - 257" refId="ref5951" refString="Reddell J. R. 1981. A review of the cavernicole fauna of Mexico, Guatemala and Belize. Texas Memorial Museum, Speleological Monographs 27: 69 - 257." type="journal article" year="1981">Reddell 1981: 165</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="5C3510E7034CC31BBE858287FDC930E0" author="Rambla M. &amp; Juberthie C." box="[234,601,466,492]" page="221" pageId="4" pageNumber="78" year="1994">
<bibRefCitation id="B9054B07034CC31BBE858287FDC930E0" author="Rambla M. &amp; Juberthie C." box="[234,601,466,492]" pageId="4" pageNumber="78" pagination="215 - 230" refId="ref5915" refString="Rambla M. &amp; Juberthie C. 1994. Opiliones. In: Juberthie C. &amp; Decu V. (eds) Encyclopaedia Biospeologica: 215 - 230. Societe de Biospeologie, Netherlands." type="book chapter" year="1994">Rambla &amp; Juberthie 1994: 221</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="5C3510E7034CC31BBCE58286FB8430E0" author="Kury A. B. &amp; Cokendolpher J. C." box="[650,1044,466,493]" page="149" pageId="4" pageNumber="78" year="2000">
<bibRefCitation id="B9054B07034CC31BBCE58286FB8430E0" author="Kury A. B. &amp; Cokendolpher J. C." box="[650,1044,466,493]" pageId="4" pageNumber="78" pagination="137 - 157" refId="ref5688" refString="Kury A. B. &amp; Cokendolpher J. C. 2000. Opiliones. In: Llorente-Bousquets J. E., Gonzalez-Soriano E. &amp; Papavero N. (eds) Biodiversidad, taxonomia y biogeografia de artropodos de Mexico: hacia una sintesis de su conocimiento. Vol. II: 137 - 157. CONABIO, Mexico City." type="book chapter" year="2000">Kury &amp; Cokendolpher 2000: 149</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="5C3510E7034CC31BBA2A8286FA9430E0" author="Kury A. B." box="[1093,1284,466,493]" page="240" pageId="4" pageNumber="78" year="2003">
<bibRefCitation id="B9054B07034CC31BBA2A8286FA9430E0" author="Kury A. B." box="[1093,1284,466,493]" pageId="4" pageNumber="78" refId="ref5653" refString="Kury A. B. 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia. Vol. Especial monografico 1. Zaragoza." type="book" year="2003">Kury 2003: 240</bibRefCitation>
</treatmentCitation>
. — Cruz- López &amp; Francke 2013b: 303; 2015: 829; 2016: 328; 2020: 435. —
<treatmentCitation id="5C3510E7034CC31BBA6782A0FE9B333F" author="Aguinaga M. A. &amp; Cruz-Lopez J. A." page="10" pageId="4" pageNumber="78" year="2019">
<bibRefCitation id="B9054B07034CC31BBA6782A0FE9B333F" author="Aguinaga M. A. &amp; Cruz-Lopez J. A." pageId="4" pageNumber="78" pagination="902907" refId="ref4759" refString="Aguinaga M. A. &amp; Cruz-Lopez J. A. 2019. Dos nuevas especies cavernicolas del genero Serrobunus (Opiliones: Stygnopsidae: Stygnopsinae) del noreste de Mexico. Revista Mexicana de Biodiversidad 90: e 902907. https: // doi. org / 10.22201 / ib. 20078706 e. 2019.90.2907" type="journal article" year="2019">Aguiñaga &amp; Cruz-López 2019: 10</bibRefCitation>
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.
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</paragraph>
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<emphasis id="EFE0EAE4034CC31BBED2810FFEA63378" bold="true" box="[189,310,602,628]" pageId="4" pageNumber="78">Diagnosis</emphasis>
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<paragraph id="DD2B36F6034CC31BBED281DCFE0F33E5" blockId="4.[189,1399,649,745]" pageId="4" pageNumber="78">
Movable finger of chelicera with three teeth (
<figureCitation id="45AF2A73034CC31BBCB581DCFCA633AF" box="[730,822,649,675]" captionStart="Fig" captionStartId="3.[189,232,1963,1989]" captionTargetBox="[198,1389,272,1927]" captionTargetId="figure-15@3.[198,1389,265,1927]" captionTargetPageId="3" captionText="Fig. 1. Troglostygnopsis anophthalma Šilhavý, 1974, ♂, topotype (AMNH). A. Chelicera, mesal view. B. Chelicera, frontal view. C. Pedipalpal femur, mesal view. D. Pedipalpal tibia and tarsus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5708142" httpUri="https://zenodo.org/record/5708142/files/figure.png" pageId="4" pageNumber="78">Fig. 1B</figureCitation>
), small and wide. Tubercles of leg IV small and scattered; patella with few, small tubercles (
<figureCitation id="45AF2A73034CC31BBCA781F9FCB633CA" box="[712,806,684,710]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[189,1398,1162,1927]" captionTargetId="figure-156@4.[189,1398,1162,1927]" captionTargetPageId="4" captionText="Fig. 2. Troglostygnopsis anophthalma Šilhavý, 1974, ♂, topotype (AMNH). A. Apical femur, patella and basal tibia IV. B. Patella IV." figureDoi="http://doi.org/10.5281/zenodo.5708144" httpUri="https://zenodo.org/record/5708144/files/figure.png" pageId="4" pageNumber="78">Fig. 2B</figureCitation>
). Penis with one pair of MS D, MS E1 below the level of E2 (
<figureCitation id="45AF2A73034CC31BBF20819AFE1F33E5" box="[335,399,719,745]" captionStart="Fig" captionStartId="5.[189,232,1963,1989]" captionTargetBox="[189,1398,1251,1921]" captionTargetId="figure-283@5.[189,1398,1251,1921]" captionTargetPageId="5" captionText="Fig. 3. Troglostygnopsis anophthalma Šilhavý, 1974, ♂, topotype (AMNH), penis. A. Dorsal view. B. Lateral view. C. Ventral view. Scale bar in C applicable to all figures." figureDoi="http://doi.org/10.5281/zenodo.5708146" httpUri="https://zenodo.org/record/5708146/files/figure.png" pageId="4" pageNumber="78">Fig 3</figureCitation>
).
</paragraph>
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<emphasis id="EFE0EAE4034CC31BBED28044FEFC3226" bold="true" box="[189,364,784,811]" pageId="4" pageNumber="78">Type material</emphasis>
(not examined)
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<typeStatus id="022F8854034CC31BBE85806AFECC3255" box="[234,348,831,857]" pageId="4" pageNumber="78" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingRegion id="1F50F814034CC31BBED28037FEA13270" box="[189,305,866,892]" country="Mexico" name="Mexico" pageId="4" pageNumber="78">MEXICO</collectingRegion>
<specimenCount id="CB92FD7F034CC31BBF228036FEF53271" box="[333,357,867,893]" pageId="4" pageNumber="78" type="male"></specimenCount>
;
<collectingRegion id="1F50F814034CC31BBF1B8037FE453270" box="[372,469,866,892]" country="Mexico" name="Chiapas" pageId="4" pageNumber="78">Chiapas</collectingRegion>
,
<location id="D84B602D034CC31BBF8D8037FCB03270" LSID="urn:lsid:plazi:treatment:553D87E0034CC319BC65825DFDC935A5:D84B602D034CC31BBF8D8037FCB03270" box="[482,800,866,892]" country="Mexico" county="San Cristobal" municipality="Casas" name="San Cristobal de las Casas" pageId="4" pageNumber="78" stateProvince="MEXICO">San Cristóbal de las Casas</location>
,
<location id="D84B602D034CC31BBD418037FBCE3271" LSID="urn:lsid:plazi:treatment:553D87E0034CC319BC65825DFDC935A5:D84B602D034CC31BBD418037FBCE3271" box="[814,1118,866,893]" country="Mexico" county="San Cristobal" municipality="Casas" name="Grutas de Rancho Nuevo" pageId="4" pageNumber="78" stateProvince="MEXICO">Grutas de Rancho Nuevo</location>
;
<elevation id="56B9D1C5034CC31BBA048037FA9E3270" box="[1131,1294,866,893]" metricMagnitude="3" metricUnit="m" metricValue="2.275" pageId="4" pageNumber="78" unit="m" value="2275.0">
<quantity id="1A6C9B13034CC31BBA048037FB5D3271" box="[1131,1229,866,893]" metricMagnitude="3" metricUnit="m" metricValue="2.275" pageId="4" pageNumber="78" unit="m" value="2275.0">2,275 m</quantity>
a.s.l.
</elevation>
;
<date id="A92A1036034CC31BBB728037FF6B32AC" pageId="4" pageNumber="78" value="1971-02-25">
<collectingDate id="B96EE9DE034CC31BBB728037FF6B32AC" pageId="4" pageNumber="78" value="1971-02-25">25 Feb. 1971</collectingDate>
</date>
;
<collectorName id="70615320034CC31BBF6880D3FEEF32AC" box="[263,383,902,928]" pageId="4" pageNumber="78">R. Argano</collectorName>
leg.;
<collectionCode id="BB85AE33034CC31BBFAE80D3FD8832AC" box="[449,536,902,928]" collectionName="Italy, Roma, Museo Civico di Zoologia" country="Italy" httpUri="http://grbio.org/cool/mmbj-pfva" name="Museo Civico di Zoologia" pageId="4" pageNumber="78">MCZR</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="DD2B36F6034CC31BBE858092FC103508" blockId="4.[189,896,967,1029]" pageId="4" pageNumber="78">
<materialsCitation id="6DFC3CAB034CC31BBE858092FC103508" ID-GBIF-Occurrence="3407626301" collectingDate="1971-02-25" collectionCode="MCZR" collectorName="R. Argano" country="Mexico" county="San Cristobal" elevation="2275" location="MEXICO" municipality="Casas" pageId="4" pageNumber="78" specimenCount="1" specimenCount-female="1" stateProvince="MEXICO" typeStatus="allotype">
<emphasis id="EFE0EAE4034CC31BBE858092FEC432ED" bold="true" box="[234,340,967,993]" pageId="4" pageNumber="78">
<typeStatus id="022F8854034CC31BBE858092FEC432ED" box="[234,340,967,993]" pageId="4" pageNumber="78" type="allotype">Allotype</typeStatus>
</emphasis>
<collectingRegion id="1F50F814034CC31BBED280BFFEA13508" box="[189,305,1002,1028]" country="Mexico" name="Mexico" pageId="4" pageNumber="78">MEXICO</collectingRegion>
<specimenCount id="CB92FD7F034CC31BBF2580BEFEF13509" box="[330,353,1003,1029]" pageId="4" pageNumber="78" type="female"></specimenCount>
; same collection data as for holotype;
<collectionCode id="BB85AE33034CC31BBD4A80BFFCEC3508" box="[805,892,1002,1028]" collectionName="Italy, Roma, Museo Civico di Zoologia" country="Italy" httpUri="http://grbio.org/cool/mmbj-pfva" name="Museo Civico di Zoologia" pageId="4" pageNumber="78">MCZR</collectionCode>
.
</materialsCitation>
</paragraph>
<caption id="89EB667E034CC31BBED284FEFDA436E5" ID-DOI="http://doi.org/10.5281/zenodo.5708144" ID-Zenodo-Dep="5708144" httpUri="https://zenodo.org/record/5708144/files/figure.png" pageId="4" pageNumber="78" startId="4.[189,232,1963,1989]" targetBox="[189,1398,1162,1927]" targetPageId="4">
<paragraph id="DD2B36F6034CC31BBED284FEFDA436E5" blockId="4.[189,1398,1963,2025]" pageId="4" pageNumber="78">
<emphasis id="EFE0EAE4034CC31BBED284FEFE9D36C9" bold="true" box="[189,269,1963,1989]" pageId="4" pageNumber="78">Fig. 2.</emphasis>
<taxonomicName id="1A944D75034CC31BBF7984FEFCA336C9" authority="Silhavy, 1974" authorityName="Silhavy" authorityYear="1974" box="[278,819,1963,1992]" class="Arachnida" family="Stygnopsidae" genus="Troglostygnopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="78" phylum="Arthropoda" rank="species" species="anophthalma">
<emphasis id="EFE0EAE4034CC31BBF7984FEFD1236C9" box="[278,642,1963,1989]" italics="true" pageId="4" pageNumber="78">Troglostygnopsis anophthalma</emphasis>
<bibRefCitation id="B9054B07034CC31BBCE484F9FCA336C9" author="Silhavy V." box="[651,819,1963,1992]" pageId="4" pageNumber="78" pagination="175 - 194" refId="ref6023" refString="Silhavy V. 1974. Cavernicolous opilionids from Mexico. Subterranean fauna of Mexico. Part. II. Quaderno Accademia Nazionale dei Lincei 171: 175 - 194." type="journal article" year="1974">Šilhavý, 1974</bibRefCitation>
</taxonomicName>
, ♂, topotype (AMNH).
<emphasis id="EFE0EAE4034CC31BBA3684FEFBFC36C9" bold="true" box="[1113,1132,1963,1989]" pageId="4" pageNumber="78">A</emphasis>
. Apical femur, patella and basal tibia IV.
<emphasis id="EFE0EAE4034CC31BBFF7849BFE3C36E4" bold="true" box="[408,428,1998,2024]" pageId="4" pageNumber="78">B</emphasis>
. Patella IV.
</paragraph>
</caption>
<paragraph id="DD2B36F6034DC31ABED2825DFE3A302E" blockId="5.[189,426,264,290]" box="[189,426,264,290]" pageId="5" pageNumber="79">
<heading id="8663819A034DC31ABED2825DFE3A302E" bold="true" box="[189,426,264,290]" fontSize="11" level="3" pageId="5" pageNumber="79" reason="3">
<emphasis id="EFE0EAE4034DC31ABED2825DFE3A302E" bold="true" box="[189,426,264,290]" pageId="5" pageNumber="79">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DD2B36F6034DC31ABE858262FB973094" blockId="5.[189,1399,311,408]" pageId="5" pageNumber="79">
<materialsCitation id="6DFC3CAB034DC31ABE858262FB933094" ID-GBIF-Occurrence="3407626305" collectingDate="1980-01-20" collectionCode="AMNH" collectorName="V. Roth" country="Mexico" county="San Cristobal" latitude="16.736668" location="San Cristobal de las Casas" longLatPrecision="21" longitude="-92.63833" municipality="Casas" pageId="5" pageNumber="79" specimenCount="1" stateProvince="Chiapas" typeStatus="topotype">
<emphasis id="EFE0EAE4034DC31ABE858262FECE305D" bold="true" box="[234,350,311,337]" pageId="5" pageNumber="79">
<typeStatus id="022F8854034DC31ABE858262FECE305D" box="[234,350,311,337]" pageId="5" pageNumber="79" type="topotype">Topotype</typeStatus>
</emphasis>
<collectingCountry id="A5837666034DC31ABED2820FFEA13078" box="[189,305,346,372]" name="Mexico" pageId="5" pageNumber="79">MEXICO</collectingCountry>
<specimenCount id="CB92FD7F034DC31ABF20820FFEEF3079" box="[335,383,346,373]" pageId="5" pageNumber="79">1 ♂</specimenCount>
,
<collectingRegion id="1F50F814034DC31ABFE0820FFE603078" box="[399,496,346,372]" country="Mexico" name="Chiapas" pageId="5" pageNumber="79">Chiapas</collectingRegion>
,
<location id="D84B602D034DC31ABF90820FFCD23078" LSID="urn:lsid:plazi:treatment:553D87E0034CC319BC65825DFDC935A5:D84B602D034DC31ABF90820FFCD23078" box="[511,834,346,372]" country="Mexico" county="San Cristobal" latitude="16.736668" longLatPrecision="21" longitude="-92.63833" municipality="Casas" name="San Cristobal de las Casas" pageId="5" pageNumber="79" stateProvince="Chiapas">San Cristóbal de las Casas</location>
,
<location id="D84B602D034DC31ABD3E820FFBE13078" LSID="urn:lsid:plazi:treatment:553D87E0034CC319BC65825DFDC935A5:D84B602D034DC31ABD3E820FFBE13078" box="[849,1137,346,372]" country="Mexico" county="San Cristobal" latitude="16.736668" longLatPrecision="21" longitude="-92.63833" municipality="Casas" name="Grutas de San Cristobal" pageId="5" pageNumber="79" stateProvince="Chiapas">Grutas de San Cristóbal</location>
[= Grutas de Rancho Nuevo];
<geoCoordinate id="B8A05031034DC31ABF4B8228FE273094" box="[292,439,380,408]" degrees="16" direction="north" minutes="44" orientation="latitude" pageId="5" pageNumber="79" precision="15" seconds="12" value="16.736668">16º4412″ N</geoCoordinate>
,
<geoCoordinate id="B8A05031034DC31ABFAC822BFDCC3094" box="[451,604,380,408]" degrees="92" direction="west" minutes="38" orientation="longitude" pageId="5" pageNumber="79" precision="15" seconds="18" value="-92.63833">92º3818″ W</geoCoordinate>
;
<date id="A92A1036034DC31ABC07822BFC903094" box="[616,768,382,408]" pageId="5" pageNumber="79" value="1980-01-20">
<collectingDate id="B96EE9DE034DC31ABC07822BFC903094" box="[616,768,382,408]" pageId="5" pageNumber="79" value="1980-01-20">20 Jan. 1980</collectingDate>
</date>
;
<collectorName id="70615320034DC31ABD63822BFCF5309B" box="[780,869,381,408]" pageId="5" pageNumber="79">V. Roth</collectorName>
leg.;
<collectionCode id="BB85AE33034DC31ABDCA822BFB933094" box="[933,1027,382,408]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="5" pageNumber="79">AMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="958E657D034DC319BED282EAFEFA3535" lastPageId="6" lastPageNumber="80" pageId="5" pageNumber="79" type="description">
<paragraph id="DD2B36F6034DC31ABED282EAFD8030D5" blockId="5.[189,528,447,473]" box="[189,528,447,473]" pageId="5" pageNumber="79">
<heading id="8663819A034DC31ABED282EAFD8030D5" bold="true" box="[189,528,447,473]" fontSize="11" level="3" pageId="5" pageNumber="79" reason="3">
<emphasis id="EFE0EAE4034DC31ABED282EAFD8030D5" bold="true" box="[189,528,447,473]" pageId="5" pageNumber="79">Preliminary considerations</emphasis>
</heading>
</paragraph>
<paragraph id="DD2B36F6034DC31ABED282B8FB623287" blockId="5.[187,1401,493,907]" pageId="5" pageNumber="79">
The
<typeStatus id="022F8854034DC31ABE9D82B8FEC9330B" box="[242,345,493,519]" pageId="5" pageNumber="79" type="holotype">holotype</typeStatus>
was not examined; however, according to several caves studies and maps collected by the Association for Mexican Cave Studies (AMCS, information available at:http://www.mexicancaves.org/) we know that Grutas de Rancho Nuevo are the same as Grutas de
<collectingCountry id="A5837666034DC31ABD8D8161FB173342" box="[994,1159,564,590]" name="Saint Kitts and Nevis" pageId="5" pageNumber="79">San Cristobal</collectingCountry>
; therefore, the male examined here is considered a topotype. This specimen is poorly preserved, without almost all legs, incomplete scutum and broken body. In the present work, we could only illustrate the chelicera, pedipalp, ornamentation of femur and patella IV, and male genitalia in detail. Fortunately, these structures are enough to recognize and diagnose this species. Additionally,
<bibRefCitation id="B9054B07034DC31ABDEE8194FBBF33D7" author="Silhavy V." box="[897,1071,705,733]" pageId="5" pageNumber="79" pagination="175 - 194" refId="ref6023" refString="Silhavy V. 1974. Cavernicolous opilionids from Mexico. Subterranean fauna of Mexico. Part. II. Quaderno Accademia Nazionale dei Lincei 171: 175 - 194." type="journal article" year="1974">Šilhavý (1974)</bibRefCitation>
made excellent drawings of dorsal and lateral view of the
<typeStatus id="022F8854034DC31ABC4F81B1FD1A33F2" box="[544,650,740,766]" pageId="5" pageNumber="79" type="holotype">holotype</typeStatus>
, which are very clear and informative for taxonomic purposes. Finally,
<bibRefCitation id="B9054B07034DC31ABF72805DFE5D322D" author="Silhavy V." box="[285,461,775,804]" pageId="5" pageNumber="79" pagination="175 - 194" refId="ref6023" refString="Silhavy V. 1974. Cavernicolous opilionids from Mexico. Subterranean fauna of Mexico. Part. II. Quaderno Accademia Nazionale dei Lincei 171: 175 - 194." type="journal article" year="1974">Šilhavý (1974)</bibRefCitation>
, based on immature stages, mentioned two additional records for
<taxonomicName id="1A944D75034DC31ABAD0805DFAE5322D" authorityName="Silhavy" authorityYear="1974" box="[1215,1397,775,801]" class="Arachnida" family="Stygnopsidae" genus="Troglostygnopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="79" phylum="Arthropoda" rank="species" species="anophthalma">
<emphasis id="EFE0EAE4034DC31ABAD0805DFAE5322D" box="[1215,1397,775,801]" italics="true" pageId="5" pageNumber="79">T. anophthalma</emphasis>
</taxonomicName>
from Cueva de la Golondrina and Cueva del Nacimiento del Río San Antonio, both in Bochil, Chiapas. Due to the specificity in cave habitats in these species, it is necessary to collect adults from those caves to corroborate if they are conspecific with
<taxonomicName id="1A944D75034DC31ABCC08024FCF63287" authorityName="Silhavy" authorityYear="1974" box="[687,870,881,907]" class="Arachnida" family="Stygnopsidae" genus="Troglostygnopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="79" phylum="Arthropoda" rank="species" species="anophthalma">
<emphasis id="EFE0EAE4034DC31ABCC08024FCF63287" box="[687,870,881,907]" italics="true" pageId="5" pageNumber="79">T. anophthalma</emphasis>
</taxonomicName>
or represent undescribed species.
</paragraph>
<paragraph id="DD2B36F6034DC31ABED280E7FEFB32C0" blockId="5.[189,363,946,972]" box="[189,363,946,972]" pageId="5" pageNumber="79">
<heading id="8663819A034DC31ABED280E7FEFB32C0" bold="true" box="[189,363,946,972]" fontSize="11" level="3" pageId="5" pageNumber="79" reason="3">
<emphasis id="EFE0EAE4034DC31ABED280E7FEFB32C0" bold="true" box="[189,363,946,972]" pageId="5" pageNumber="79">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="DD2B36F6034DC31ABE8580B4FEBA32F7" blockId="5.[189,835,993,1054]" box="[234,298,993,1019]" pageId="5" pageNumber="79">
<emphasis id="EFE0EAE4034DC31ABE8580B4FEBA32F7" bold="true" box="[234,298,993,1019]" pageId="5" pageNumber="79">Male</emphasis>
</paragraph>
<paragraph id="DD2B36F6034DC31ABED28751FCD33512" blockId="5.[189,835,993,1054]" box="[189,835,1028,1054]" pageId="5" pageNumber="79">MEASUREMENTS. Scutum length: 4.9, scutum width: 4.1.</paragraph>
<paragraph id="DD2B36F6034DC31ABED28710FAE6358F" blockId="5.[189,1399,1093,1155]" pageId="5" pageNumber="79">
CHELICERA (
<figureCitation id="45AF2A73034DC31ABF258710FE1F3553" box="[330,399,1093,1119]" captionStart="Fig" captionStartId="3.[189,232,1963,1989]" captionTargetBox="[198,1389,272,1927]" captionTargetId="figure-15@3.[198,1389,265,1927]" captionTargetPageId="3" captionText="Fig. 1. Troglostygnopsis anophthalma Šilhavý, 1974, ♂, topotype (AMNH). A. Chelicera, mesal view. B. Chelicera, frontal view. C. Pedipalpal femur, mesal view. D. Pedipalpal tibia and tarsus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5708142" httpUri="https://zenodo.org/record/5708142/files/figure.png" pageId="5" pageNumber="79">Fig. 1</figureCitation>
AB). Scutum/cheliceral hand ratio: 1.37. Basichelicerite long, with diffuse bulla. Cheliceral hand slightly swollen, fixed finger with six teeth distributed evenly throughout, middle one
</paragraph>
<caption id="89EB667E034DC31ABED284FEFB9C36E4" ID-DOI="http://doi.org/10.5281/zenodo.5708146" ID-Zenodo-Dep="5708146" httpUri="https://zenodo.org/record/5708146/files/figure.png" pageId="5" pageNumber="79" startId="5.[189,232,1963,1989]" targetBox="[189,1398,1251,1921]" targetPageId="5">
<paragraph id="DD2B36F6034DC31ABED284FEFB9C36E4" blockId="5.[189,1399,1963,2025]" pageId="5" pageNumber="79">
<emphasis id="EFE0EAE4034DC31ABED284FEFE9E36C9" bold="true" box="[189,270,1963,1989]" pageId="5" pageNumber="79">Fig. 3.</emphasis>
<taxonomicName id="1A944D75034DC31ABF7584FEFCAC36C9" authority="Silhavy, 1974" authorityName="Silhavy" authorityYear="1974" box="[282,828,1963,1992]" class="Arachnida" family="Stygnopsidae" genus="Troglostygnopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="79" phylum="Arthropoda" rank="species" species="anophthalma">
<emphasis id="EFE0EAE4034DC31ABF7584FEFD1736C9" box="[282,647,1963,1989]" italics="true" pageId="5" pageNumber="79">Troglostygnopsis anophthalma</emphasis>
<bibRefCitation id="B9054B07034DC31ABCFD84F9FCAC36C9" author="Silhavy V." box="[658,828,1963,1992]" pageId="5" pageNumber="79" pagination="175 - 194" refId="ref6023" refString="Silhavy V. 1974. Cavernicolous opilionids from Mexico. Subterranean fauna of Mexico. Part. II. Quaderno Accademia Nazionale dei Lincei 171: 175 - 194." type="journal article" year="1974">Šilhavý, 1974</bibRefCitation>
</taxonomicName>
, ♂, topotype (AMNH), penis.
<emphasis id="EFE0EAE4034DC31ABAD884FEFB5C36C9" bold="true" box="[1207,1228,1963,1989]" pageId="5" pageNumber="79">A</emphasis>
. Dorsal view.
<emphasis id="EFE0EAE4034DC31ABED2849BFF4136E4" bold="true" box="[189,209,1998,2024]" pageId="5" pageNumber="79">B</emphasis>
. Lateral view.
<emphasis id="EFE0EAE4034DC31ABF10849BFE0436E4" bold="true" box="[383,404,1998,2024]" pageId="5" pageNumber="79">C</emphasis>
. Ventral view. Scale bar in C applicable to all figures.
</paragraph>
</caption>
<paragraph id="DD2B36F6034EC319BED2825CFED8304A" blockId="6.[189,1399,265,327]" pageId="6" pageNumber="80">biggest; movable finger with three scattered teeth, basalmost and middle ones small and triangular, distal one diffuse.</paragraph>
<paragraph id="DD2B36F6034EC319BED2823BFC723373" blockId="6.[188,1400,365,639]" pageId="6" pageNumber="80">
PEDIPALPS (
<figureCitation id="45AF2A73034EC319BF29823BFE013084" box="[326,401,366,392]" captionStart="Fig" captionStartId="3.[189,232,1963,1989]" captionTargetBox="[198,1389,272,1927]" captionTargetId="figure-15@3.[198,1389,265,1927]" captionTargetPageId="3" captionText="Fig. 1. Troglostygnopsis anophthalma Šilhavý, 1974, ♂, topotype (AMNH). A. Chelicera, mesal view. B. Chelicera, frontal view. C. Pedipalpal femur, mesal view. D. Pedipalpal tibia and tarsus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5708142" httpUri="https://zenodo.org/record/5708142/files/figure.png" pageId="6" pageNumber="80">Fig. 1</figureCitation>
CD). Measurements: 2.30/1.00/1.75/1.80/1.00. Trochanter globose, with three ventral spiniform tubercles, apical one very large, dorsally with one spiniform tubercle. Femur slightly compressed laterally, with one ventral and one dorsal row of large spiniform tubercles pointing distally, these tubercles uniformly distributed throughout femur, ventral basalmost one most prominent. Patella with few small spiniform tubercles. Tibia with long armature, with relative sizes of three major tubercles (SST) on mesal side 1 = 3&gt; 2, and on ectal margin 3 = 2&gt; 1, on this segment two small setiferous tubercles between SST 1 and 2, and at apical end of the margin. Tarsus with six major tubercles on mesal side (2&gt; 4 = 5&gt; 6&gt; 1 = 3) and five on ectal side (1&gt; 4&gt; 2 = 3 = 5).
</paragraph>
<paragraph id="DD2B36F6034EC319BED281F3FE3B320A" blockId="6.[189,1398,677,774]" pageId="6" pageNumber="80">
LEGS (
<figureCitation id="45AF2A73034EC319BF6681F0FEC533CC" box="[265,341,677,704]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[189,1398,1162,1927]" captionTargetId="figure-156@4.[189,1398,1162,1927]" captionTargetPageId="4" captionText="Fig. 2. Troglostygnopsis anophthalma Šilhavý, 1974, ♂, topotype (AMNH). A. Apical femur, patella and basal tibia IV. B. Patella IV." figureDoi="http://doi.org/10.5281/zenodo.5708144" httpUri="https://zenodo.org/record/5708144/files/figure.png" pageId="6" pageNumber="80">Fig. 2</figureCitation>
). All segments very long and slender, femur IV longer than scutum. Ornamentation of femur IV composed of small and scattered tubercles forming longitudinal rows, patella covered with few small tubercles.
</paragraph>
<paragraph id="DD2B36F6034EC319BED2807BFABA32D8" blockId="6.[189,1401,813,981]" pageId="6" pageNumber="80">
PENIS (
<figureCitation id="45AF2A73034EC319BF628078FEC6324B" box="[269,342,813,839]" captionStart="Fig" captionStartId="5.[189,232,1963,1989]" captionTargetBox="[189,1398,1251,1921]" captionTargetId="figure-283@5.[189,1398,1251,1921]" captionTargetPageId="5" captionText="Fig. 3. Troglostygnopsis anophthalma Šilhavý, 1974, ♂, topotype (AMNH), penis. A. Dorsal view. B. Lateral view. C. Ventral view. Scale bar in C applicable to all figures." figureDoi="http://doi.org/10.5281/zenodo.5708146" httpUri="https://zenodo.org/record/5708146/files/figure.png" pageId="6" pageNumber="80">Fig. 3</figureCitation>
). Pars distalis with an apical concavity, lateral and apical margins softly rolled dorsally. Follis rugose, apices of dorsal bilobular projection long and pointed. One pair of small MS D lateral to base of follis, near to lateral margins of pars distalis. Four pairs of acute MS C forming two irregular rows on lateral sides. MS A+B formed by many spatulate setae, on latero-basal portion of pars distalis, some of them on ventral side. MS E composed of two pairs, MS E1 markedly below level of E2.
</paragraph>
<paragraph id="DD2B36F6034EC319BE8580A9FEFA3535" blockId="6.[189,362,1020,1081]" pageId="6" pageNumber="80">
<emphasis id="EFE0EAE4034EC319BE8580A9FED5351A" bold="true" box="[234,325,1020,1046]" pageId="6" pageNumber="80">Female</emphasis>
Not examined.
</paragraph>
</subSubSection>
<subSubSection id="958E657D034EC319BED28735FDC935A5" box="[189,601,1120,1193]" pageId="6" pageNumber="80" type="distribution">
<paragraph id="DD2B36F6034EC319BED28735FEC63576" blockId="6.[189,342,1120,1146]" box="[189,342,1120,1146]" pageId="6" pageNumber="80">
<heading id="8663819A034EC319BED28735FEC63576" bold="true" box="[189,342,1120,1146]" fontSize="11" level="3" pageId="6" pageNumber="80" reason="3">
<emphasis id="EFE0EAE4034EC319BED28735FEC63576" bold="true" box="[189,342,1120,1146]" pageId="6" pageNumber="80">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD2B36F6034EC319BED287DAFDC935A5" blockId="6.[189,601,1167,1193]" box="[189,601,1167,1193]" pageId="6" pageNumber="80">
Known only from the
<typeStatus id="022F8854034EC319BFAD87DAFE6535A5" box="[450,501,1167,1193]" pageId="6" pageNumber="80">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
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