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<mods:title id="EC5DA26BAD326039069998252570446F">Replacement names and nomenclatural comments for problematic species-group names in Europe's Neogene freshwater Gastropoda. Part 2</mods:title>
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<mods:namePart id="6F972ACA51B390E3754F981A109A22EC">Neubauer, Thomas A.</mods:namePart>
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<mods:namePart id="4C620514FDD433DA11D0F83702A675B4">Elisavet, Georgopoulou</mods:namePart>
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<mods:namePart id="F134A41F87B0315C6BBCA774DDFB7AB0">Mandic, Oleg</mods:namePart>
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<treatment id="5532FA2DACA3343E2B5E8629F6F4EAEF" ID-GBIF-Taxon="152054656" LSID="urn:lsid:plazi:treatment:5532FA2DACA3343E2B5E8629F6F4EAEF" httpUri="http://treatment.plazi.org/id/5532FA2DACA3343E2B5E8629F6F4EAEF" lastPageNumber="27" pageId="14" pageNumber="27">
<subSubSection id="18D179A7646F7AA57273F36F926F0F45" pageId="14" pageNumber="27" type="multiple">
<paragraph id="1904E641ECD74B1E0161C0B7169921D8" pageId="14" pageNumber="27">Taxon classification Animalia ORDO FAMILIA</paragraph>
</subSubSection>
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<paragraph id="1B5317B5F012CF3CF02CC5BE1C811825" pageId="14" pageNumber="27">
<taxonomicName id="DA3667BCC3A4E58FFB7FE1C8C812AC81" ID-CoL="5JLKP" authority="Sauerzopf, 1952 non Handmann, 1882" authorityName="Sauerzopf, 1952 non Handmann" authorityYear="1882" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanopsis pygmaea" order="Neogastropoda" pageId="14" pageNumber="27" phylum="Mollusca" rank="subSpecies" species="pygmaea" subSpecies="inflata">Melanopsis pygmaea inflata Sauerzopf, 1952 non Handmann, 1882</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="11A9480DBB203D04ABE0CE1A8C657775" pageId="14" pageNumber="27" type="reference_group">
<paragraph id="1DD45C86EEE8DE2F6D13519D7DF6ED61" pageId="14" pageNumber="27">
<taxonomicName id="6452367A5573D74C47E03F8FD50B73FB" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanopsis pygmaea subsp. inflata" order="Neogastropoda" pageId="14" pageNumber="27" phylum="Mollusca" rank="subSpecies" species="pygmaea" subSpecies="inflata">Melanopsis pygmaea inflata</taxonomicName>
Sauerzopf, 1952: 13, pl. 2, fig. 4 [non
<taxonomicName id="A53E5F729A08EB768FA95BB8142B6DEB" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanopsis pygmaea subsp. inflata" order="Neogastropoda" pageId="14" pageNumber="27" phylum="Mollusca" rank="subSpecies" species="pygmaea" subSpecies="inflata">Melanopsis pygmaea inflata</taxonomicName>
Handmann, 1882].
</paragraph>
</subSubSection>
<subSubSection id="4A98B4FD7D48828EBA847C73867A131F" pageId="14" pageNumber="27" type="type locality">
<paragraph id="F374629BC11B5F60E42963E8A50170E2" pageId="14" pageNumber="27">Type locality.</paragraph>
<paragraph id="6BE1798918837CEF200D9D988A5CDF8B" pageId="14" pageNumber="27">No locus typicus given; occurs in Stegersbach, Litzelsdorf, Olbendorf, and Oberdorf in the Styrian Basin, Austria.</paragraph>
</subSubSection>
<subSubSection id="A67C4355006CFFC2CFB91D45F9E47ABB" pageId="14" pageNumber="27" type="age">
<paragraph id="8CC2AC4DA66E685472499C26BF486931" pageId="14" pageNumber="27">Age.</paragraph>
<paragraph id="BB1062BF427F738EA22FF16F6B9E0132" pageId="14" pageNumber="27">
Late Miocene (Pannonian, Serbian, biozones
<normalizedToken id="E8B72981FAAB358A653FBBC93A6DDF5C" originalValue="EF">E-F</normalizedToken>
).
</paragraph>
</subSubSection>
<subSubSection id="7D51880C88A0565A361CE337B4E61682" pageId="14" pageNumber="27" type="type material">
<paragraph id="A16E1094177B414BBA212CA3FD00FEFA" pageId="14" pageNumber="27">Type material.</paragraph>
<paragraph id="F4662F7308EE5951B60AB673C5191AA7" pageId="14" pageNumber="27">No storage or types indicated (material derived from Sauerzopf's private collections).</paragraph>
</subSubSection>
<subSubSection id="7F206B0D0998929075E63ED8128AEC82" pageId="14" pageNumber="27" type="discussion">
<paragraph id="D7ED377A9C4D1A3749C4EDA60A2A10BE" pageId="14" pageNumber="27">Discussion.</paragraph>
<paragraph id="1DC333B76E873C7E71A991D647199DA5" pageId="14" pageNumber="27">
There are several issues with the name
<taxonomicName id="722A66B9E2C2F3AE2AB0855DB1F5BC10" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanopsis inflata" order="Neogastropoda" pageId="14" pageNumber="27" phylum="Mollusca" rank="species" species="inflata">Melanopsis inflata</taxonomicName>
. First, the name introduced by Sauerzopf definitely constitutes a primary homonym of
<taxonomicName id="06D7194C35D98D3B926C7455DD90846A" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanopsis pygmaea subsp. inflata" order="Neogastropoda" pageId="14" pageNumber="27" phylum="Mollusca" rank="subSpecies" species="pygmaea" subSpecies="inflata">Melanopsis pygmaea inflata</taxonomicName>
Handmann, 1882.
<bibRefCitation id="140629C5E44911FC5571847F2E0E6D50" author="Sauerzopf, F" journalOrPublisher="Burgenlaendische Heimatblaetter" pageId="28" pageNumber="41" pagination="1 - 16" title="Beitrag zur Entwicklungsgeschichte des suedburgenlaendischen Pannons." volume="14" year="1952">Sauerzopf (1952)</bibRefCitation>
explicitly introduced it as new taxon, although the combination is identical to that established by Handmann. Both taxa were obviously erected for different morphologies: while Sauerzopf's form is elongated conical, Handmann's subspecies is rather globular. Handmann's taxon is meanwhile considered as junior synonym of
<taxonomicName id="4C81F68F34705E695E179A124F80D8BD" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanopsis pygmaea" order="Neogastropoda" pageId="14" pageNumber="27" phylum="Mollusca" rank="species" species="pygmaea">Melanopsis pygmaea</taxonomicName>
<normalizedToken id="BED45F997187114852C0BD3239FBB378" originalValue="Hörnes">Hoernes</normalizedToken>
, 1856 (
<bibRefCitation id="A45A3C860ABF17FFF874B6E7C0ED8CD1" author="Wenz, W" journalOrPublisher="Senckenbergiana" pageId="29" pageNumber="42" url="http://www.biodiversitylibrary.org/bibliography/61839/summary" year="1923 - 1930">Wenz 1929a</bibRefCitation>
, p. 2813).
<taxonomicName id="7F5E86286CE0ECA32080B0ED78D32527" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanopsis pygmaea subsp. inflata" order="Neogastropoda" pageId="14" pageNumber="27" phylum="Mollusca" rank="subSpecies" species="pygmaea" subSpecies="inflata">Melanopsis pygmaea inflata</taxonomicName>
Sauerzopf, 1952, in turn, highly resembles
<taxonomicName id="9C36089758CEC92CEF79BCFEE819C9C2" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanopsis fuchsi" order="Neogastropoda" pageId="14" pageNumber="27" phylum="Mollusca" rank="species" species="fuchsi">Melanopsis fuchsi</taxonomicName>
Handmann, 1882 concerning its size, the regular conical outline and the slightly inflated last whorl. Exactly these last two criteria were for both authors the reason to separate their forms from
<taxonomicName id="51440B78D4E1F3F55D738AFFE077B88A" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanopsis pygmaea" order="Neogastropoda" pageId="14" pageNumber="27" phylum="Mollusca" rank="species" species="pygmaea">Melanopsis pygmaea</taxonomicName>
(see
<bibRefCitation id="F02C389A5DD43886CC1DF05623516E66" author="Handmann, R" journalOrPublisher="Aschendorff, Muenster" pageId="23" pageNumber="36" title="Die fossile Conchylienfauna von Leobersdorf im Tertiaerbecken von Wien." url="http://sammlungen.ulb.uni-muenster.de/hd/content/pageview/2023144" year="1887">Handmann 1887</bibRefCitation>
, p. 13;
<bibRefCitation id="6515707917B491560FBF5DB89E067CDB" author="Sauerzopf, F" journalOrPublisher="Burgenlaendische Heimatblaetter" pageId="28" pageNumber="41" pagination="1 - 16" title="Beitrag zur Entwicklungsgeschichte des suedburgenlaendischen Pannons." volume="14" year="1952">Sauerzopf 1952</bibRefCitation>
, p. 13). Therefore we consider both synonymous and refrain from introducing a replacement name.
</paragraph>
<paragraph id="669C009566D0FC99EB659D4298B65950" pageId="14" pageNumber="27">
The second problem regards the availability of
<taxonomicName id="CB805896D5906926354195F45C74B2D9" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanopsis inflata" order="Neogastropoda" pageId="14" pageNumber="27" phylum="Mollusca" rank="species" species="inflata">Melanopsis inflata</taxonomicName>
Handmann, 1882. This name was already introduced as subordinate taxon by
<bibRefCitation id="DAFB9B4C6C4BE8741844117115F61B9C" author="Ferussac, AEJPJF d'Audebard de" journalOrPublisher="Memoires de la Societe d'Histoire Naturelle de Paris" pageId="21" pageNumber="34" pagination="132 - 164" title="Monographie des especes vivantes et fossiles du genre melanopside, Melanopsis, et observations geologiques a leur sujet." url="http://www.biodiversitylibrary.org/item/23445page/148/mode/1up" volume="1" year="1823">
<normalizedToken id="30C5EE1E30F45E036A6AFD54A123DEA9" originalValue="Férussac">Ferussac</normalizedToken>
(1823)
</bibRefCitation>
within
<taxonomicName id="ACD4A881E00A92A227E8C11DF9B212CB" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanopsis buccinoidea" order="Neogastropoda" pageId="14" pageNumber="27" phylum="Mollusca" rank="species" species="buccinoidea">Melanopsis buccinoidea</taxonomicName>
. Whether it is available as species-group name, however, cannot easily be determined, given the chaotic system in
<normalizedToken id="075399CDA429F3C445764843C4733CD0" originalValue="Férussac's">Ferussac's</normalizedToken>
work (see also discussion of
<taxonomicName id="EE207C9B3D41C9A13B2BFF89EE74BC2C" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanopsis elongata" order="Neogastropoda" pageId="14" pageNumber="27" phylum="Mollusca" rank="species" species="elongata">Melanopsis elongata</taxonomicName>
below) and the fact that it is not found to be used as species-group name attributed to
<normalizedToken id="1B2B62532C35364CE8681A45996B53FC" originalValue="Férussac">Ferussac</normalizedToken>
in the literature, which would have made it available via
<bibRefCitation id="9FC10315F37FC86243C50B3B4360D23A" author="ICZN," journalOrPublisher="International Trust for Zoological Nomenclature, London" pageId="23" pageNumber="36" title="International Code of Zoological Nomenclature." url="http://www.nhm.ac.uk/hosted-sites/iczn/code/index.jsp" year="1999">ICZN 1999</bibRefCitation>
, Article 45.6.4.1. If
<normalizedToken id="E3B88C23A20FF1A816A49DF14EE5DFEF" originalValue="Férussac's">Ferussac's</normalizedToken>
name is accepted as species-group name, Handmann's taxon would become a primary homonym. Since this is apparently not the case and Handmann's subspecies was synonymized anyway, the introduction of a replacement name would be inexpedient.
</paragraph>
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