treatments-xml/data/55/23/95/552395DB81535ABC987BEA547E83B8B0.xml

<document ID-CLB-Dataset="6744" ID-DOI="http://dx.doi.org/10.3897/dez.68.66200" ID-GBIF-Dataset="22670907-a451-4089-a3b0-e16edba09a28" ID-Pensoft-Pub="1860-1324-1-189" ID-Pensoft-UUID="84A6EF91377850DF803DE5E2A4062204" ID-ZooBank="F88F5F2BEC5A408E98190188E87DB024" ModsDocID="1860-1324-1-189" checkinTime="1621365470254" checkinUser="pensoft" docAuthor="Yang, Zhen-ming, Jia, Fenglong, Jiang, Lu &amp; Guo, Qiang" docDate="2021" docId="552395DB81535ABC987BEA547E83B8B0" docLanguage="en" docName="DeutEntomolZeit 68(1): 189-205" docOrigin="Deutsche Entomologische Zeitschrift 68 (1)" docPubDate="2021-05-18" docSource="http://dx.doi.org/10.3897/dez.68.66200" docTitle="Agraphydrus pseudoniger Yang &amp; Jia 2021, sp. nov." docType="treatment" docUuid="60AE015E-045B-493C-A251-4224BB6296B5" docUuidSource="ZooBank" docVersion="5" id="84A6EF91377850DF803DE5E2A4062204" lastPageNumber="189" masterDocId="84A6EF91377850DF803DE5E2A4062204" masterDocTitle="Four new species of Agraphydrus Regimbart, 1903 with additional faunastic record from China (Coleoptera, Hydrophilidae, Acidocerinae)" masterLastPageNumber="205" masterPageNumber="189" pageNumber="189" updateTime="1732802310598" updateUser="ExternalLinkService">
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<mods:title id="653FFAFADB22F75AD4B54A9DB69174E6">Four new species of Agraphydrus Regimbart, 1903 with additional faunastic record from China (Coleoptera, Hydrophilidae, Acidocerinae)</mods:title>
</mods:titleInfo>
<mods:name id="1CB72C1E4EED2D38E65393E636C1F20B" type="personal">
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<mods:namePart id="CF23417F17CC50F5E13018BF269D28F3">Yang, Zhen-ming</mods:namePart>
<mods:nameIdentifier id="3F56F0A8E87AD68CAAF311B9EAB8B188" type="ORCID">https://orcid.org/0000-0001-7838-8605</mods:nameIdentifier>
<mods:affiliation id="AB6221845C301D0CDE41EB075326D7D7">Institute of Entomology, Life Sciences School, Sun Yat-sen University, Guangzhou, 510275, Guangdong, China</mods:affiliation>
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<mods:namePart id="EA72A4784D439AD48FF746FE1DC80A65">Jia, Fenglong</mods:namePart>
<mods:affiliation id="C8B896F2578F13C183160EFF7CFD6D61">Institute of Entomology, Life Sciences School, Sun Yat-sen University, Guangzhou, 510275, Guangdong, China</mods:affiliation>
<mods:nameIdentifier id="A6055674DEA912699D5D9ED7CD3496CF" type="email">lssjfl@mail.sysu.edu.cn</mods:nameIdentifier>
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<mods:namePart id="53413D5AA8DD6A3405BBB62944F92BE9">Jiang, Lu</mods:namePart>
<mods:affiliation id="1E2F3DFD9216D503747DD0BC98050774">Shenzhen Wilde life Conservation Division, Shenzhen, China</mods:affiliation>
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<mods:namePart id="E7C53F14A50CF67056D24179D989E4F6">Guo, Qiang</mods:namePart>
<mods:affiliation id="6A3399304E521973082029C0CF9701EC">Shenzhen Wilde life Conservation Division, Shenzhen, China</mods:affiliation>
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<mods:title id="2CD043A1FB72EBA8DD1BBF7C7652C1B2">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date id="355260BE6CBAF575A6BB05BE5A916ADA">2021</mods:date>
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<mods:number id="22A6B9E9E1E78C1459588123ABB70370">68</mods:number>
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<mods:url id="D4C73A9584C5852EF98BED1220DCC580">http://dx.doi.org/10.3897/dez.68.66200</mods:url>
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<mods:classification id="671A84EABB2D077E56B9CF8CE7A24546">journal article</mods:classification>
<mods:identifier id="E601E27FF9FE4047881255B449830763" type="DOI">http://dx.doi.org/10.3897/dez.68.66200</mods:identifier>
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<treatment id="552395DB81535ABC987BEA547E83B8B0" ID-GBIF-Taxon="182223089" LSID="urn:lsid:zoobank.org:act:60AE015E-045B-493C-A251-4224BB6296B5" httpUri="http://treatment.plazi.org/id/552395DB81535ABC987BEA547E83B8B0" lastPageNumber="189" pageId="0" pageNumber="189" scope_family="Hydrophilidae" scope_order="Coleoptera" scope_subFamily="Acidocerinae">
<subSubSection id="745BA06C6DA4E3F0B6B5894397F08E53" pageId="0" pageNumber="189" type="nomenclature">
<paragraph id="6CDE4E5A470E652A20E3396313FDE959" pageId="0" pageNumber="189">
<taxonomicName id="3D66CB61346C273E58A36FE786AED167" ID-CoL="8XT9L" LSID="http://zoobank.org/60AE015E-045B-493C-A251-4224BB6296B5" authority="Yang &amp; Jia" authorityName="Yang &amp; Jia" authorityYear="2021" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agraphydrus pseudoniger" order="Coleoptera" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="pseudoniger" status="sp. nov.">Agraphydrus pseudoniger Yang &amp; Jia</taxonomicName>
<taxonomicNameLabel id="7262321C3E8FADBF642083C67BA80BF2" pageId="0" pageNumber="189">sp. nov.</taxonomicNameLabel>
<figureCitation id="887BE3F3F189464879C403466E8BC53A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus: A, B. Agraphydrus pseudoniger sp. nov. C, D. Agraphydrus sabulosus sp. nov." figureDoi="10.3897/dez.68.66200.figure2" httpUri="https://binary.pensoft.net/fig/544290" pageId="0" pageNumber="189">Figs 2A, B</figureCitation>
<figureCitation id="91395C6791BBCD280C0A67A1DE301BDD" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Agraphydrus pseudoniger sp. nov.: A. Clypeus; B. Clypeus; C. Antennae; D. Maxillary palpi; E. Mentum; F. Mesoventrite; G. Ventrite 5; H. Profemora; I. Mesofemora; J. Metafemora." figureDoi="10.3897/dez.68.66200.figure5" httpUri="https://binary.pensoft.net/fig/544293" pageId="0" pageNumber="189">; 5</figureCitation>
<figureCitation id="AC79778B235EBC9D97562E66400BC262" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Aedeagi: A. Agraphydrus dapengensis sp. nov.; B. Agraphydrus komareki sp. nov.; C. Agraphydrus pseudoniger sp. nov.; D. Agraphydrus sabulosus sp. nov." figureDoi="10.3897/dez.68.66200.figure7" httpUri="https://binary.pensoft.net/fig/544295" pageId="0" pageNumber="189">; 7C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="50C6992EC205D341960EA80F9CFD0359" pageId="0" pageNumber="189" type="type locality">
<paragraph id="25514E5F8135567AD46CDE90FDB8DEEA" pageId="0" pageNumber="189">Type locality.</paragraph>
<paragraph id="2D0DBB3BF3032AECEA98AD41319BF2D0" pageId="0" pageNumber="189">China, Jiangxi Province, Shangyou County, Guanggushan Nature Reserve.</paragraph>
</subSubSection>
<subSubSection id="7E45C2F6DD507BF73B7CACAF20ED3B44" pageId="0" pageNumber="189" type="materials_examined">
<paragraph id="1A36EB455892106F72F79C1A715434C1" pageId="0" pageNumber="189">Material examined.</paragraph>
<paragraph id="987F37934D3F380B6272A1A961ED47B1" pageId="0" pageNumber="189">
<materialsCitation id="A50BA27C9E57D043D37A80F3904125CF" collectingDate="2015-06-21" collectorName="Fenglong Jia, Yudan Tang" county="Shangyou County" elevation="846" latitude="25.919722" location="Jiangxi Province" longLatPrecision="20" longitude="114.05112" municipality="Guanggushan Nature Reserve" specimenCount="1" typeStatus="Holotype">
<emphasis id="427F9BA8073F724D46FC11A66A6CB067" bold="true" pageId="0" pageNumber="189">
<emphasis id="EAF230BC61138CCEF8676C46571446F3" italics="true" pageId="0" pageNumber="189">
<typeStatus id="01834C5F153EEFA5E5EED56ECF0D10B8">Holotype</typeStatus>
</emphasis>
:
</emphasis>
male (SYSU); 
<location id="623D07A2F63506E61C2B417CBD7CC54A" LSID="urn:lsid:plazi:treatment:552395DB81535ABC987BEA547E83B8B0:623D07A2F63506E61C2B417CBD7CC54A" county="Shangyou County" latitude="25.919722" longLatPrecision="20" longitude="114.05112" municipality="Guanggushan Nature Reserve" name="Jiangxi Province">Jiangxi Province</location>
, 
<collectingCounty id="6E8B33D7522A16C24411846A0FAC2D67">Shangyou County</collectingCounty>
, 
<collectingMunicipality id="7038B9D45BBE9197BEAB42AFD3A1C047">Guanggushan Nature Reserve</collectingMunicipality>
, ca 
<geoCoordinate id="356BF6F72208B98686A898F773A9F0E0" degrees="25" direction="north" minutes="55" orientation="latitude" precision="15" seconds="11" value="25.919722">25°55'11&quot;N</geoCoordinate>
, 
<geoCoordinate id="9A150656FC06B14C5CD809C36ABEC2FF" degrees="114" direction="east" minutes="03" orientation="longitude" precision="15" seconds="04" value="114.05112">114°03'04&quot;E</geoCoordinate>
, 
<elevation id="AEA0AE4A97E000FFB44FA2FC29E8CC36" metricMagnitude="2" metricUnit="m" metricValue="8.459999999999999" unit="m" value="846.0">
<quantity id="01E70D156A3CF9B8D843591F196379F8" metricMagnitude="2" metricUnit="m" metricValue="8.459999999999999" unit="m" value="846.0">846m</quantity>
a.s.l.
</elevation>
, 
<collectingDate id="54080EEB0258B976CBF34D0311FE8D0F" value="2015-06-21">21.vi.2015</collectingDate>
, 
<collectorName id="C7D0FBD2D09E86835E5207F8C6AF3136">Fenglong Jia</collectorName>
, 
<collectorName id="DAF2A26283AA879D90A4FF2997B95FE2">Yudan Tang</collectorName>
leg.
</materialsCitation>
<materialsCitation id="7DC11A5AF0F915C4E87FDAFD89F88C9C" collectingDate="2015-06-18" collectorName="Fenglong Jia, Yudan Tang" county="Suichuan County" elevation="816" latitude="26.284445" location="Jiangxi Province" longLatPrecision="20" longitude="114.064445" municipality="Nanfengmian Nature Reserve" specimenCount="47" typeStatus="Paratypes">
<emphasis id="C909693DC8F3A050826E33E184B11ECF" bold="true" italics="true" pageId="0" pageNumber="189">
<typeStatus id="EEAA39A0C2234B3B281F8B30593E2826">Paratypes</typeStatus>
</emphasis>
: (
<specimenCount id="66A7F56231AAF5FB2242D490DA72D439" type="generic">47exs.</specimenCount>
, IZCAS, SYSU): 
<specimenCount id="DA9AE16BDEA7346F06F321FF0C8E4F26" type="generic">21 exs.</specimenCount>
, same data as holotype; 
<specimenCount id="69CAEC4447E2438A516BE15861D284DD" type="generic">3 exs.</specimenCount>
, 
<location id="98D9FD35600BF72636F2F65E488DFBD4" LSID="urn:lsid:plazi:treatment:552395DB81535ABC987BEA547E83B8B0:98D9FD35600BF72636F2F65E488DFBD4" county="Suichuan County" latitude="26.284445" longLatPrecision="20" longitude="114.064445" municipality="Nanfengmian Nature Reserve" name="Jiangxi Province">Jiangxi Province</location>
, 
<collectingCounty id="C12E40611FA34ABF901A82ABE3872879">Suichuan County</collectingCounty>
, 
<collectingMunicipality id="4E7FBBFEE88766A276F8BFD4D314A857">Nanfengmian Nature Reserve</collectingMunicipality>
, 
<elevation id="A7E45FD539B6C28F3985C601A435B34F" metricMagnitude="2" metricUnit="m" metricValue="8.16" unit="m" value="816.0">
<quantity id="C92786FC8D7004AF2B992EC130E6869B" metricMagnitude="2" metricUnit="m" metricValue="8.16" unit="m" value="816.0">816m</quantity>
a.s.l.
</elevation>
, 
<geoCoordinate id="4FCF38C3D74A2A1397F23DE5276529C3" degrees="26" direction="north" minutes="17" orientation="latitude" precision="15" seconds="04" value="26.284445">26°17'04&quot;N</geoCoordinate>
, 
<geoCoordinate id="EEFAA01D6D3B1581A24E4A1C1BF6B740" degrees="114" direction="east" minutes="03" orientation="longitude" precision="15" seconds="52" value="114.064445">114°03'52&quot;E</geoCoordinate>
, 
<collectingDate id="FD89F6163CD52724533258A1FAFFAEEE" value="2015-06-18">18.vi.2015</collectingDate>
, 
<collectorName id="3E1BED5407679D4F528B3B6A7F9358F0">Fenglong Jia</collectorName>
, 
<collectorName id="2552F100B25D72844E2AA77108353AEB">Yudan Tang</collectorName>
leg
</materialsCitation>
; 
<materialsCitation id="F1383AD91BA0FAC187C3E51FD538701F" collectingDate="2013-06-19" collectingDateMax="2013-06-22" collectingDateMin="2013-06-19" collectorName="Fenglong Jia" location="Guangdong Province" municipality="Dadongshan Nature Reserve" specimenCount="3" typeStatus="paratype">
<specimenCount id="A3516B13EB0F1E277438691468D424E6" type="generic">3 exs.</specimenCount>
, 
<location id="978FA91A99E269A2759E12E819DE2E91" LSID="urn:lsid:plazi:treatment:552395DB81535ABC987BEA547E83B8B0:978FA91A99E269A2759E12E819DE2E91" municipality="Dadongshan Nature Reserve" name="Guangdong Province">Guangdong Province</location>
, 
<collectingMunicipality id="A5D0DA0657AD627D546067C871656DDC">Dadongshan Nature Reserve</collectingMunicipality>
, 
<collectingDate id="6310F2830DE07BCB3D622D30DD04E3B5" value="2013-06-19" valueMax="2013-06-22" valueMin="2013-06-19">19-22.vi.2013</collectingDate>
, 
<collectorName id="6ED192CA47C29F5A36D18FE1D0388849">Fenglong Jia</collectorName>
leg
</materialsCitation>
; 
<materialsCitation id="1808B0D8CFB791D4EC7AB80372C9F497" collectingDate="2009-06-22" collectingDateMax="2009-06-26" collectingDateMin="2009-06-22" collectorName="Fenglong Jia" county="Dadongshan" location="Guangdong Province" municipality="Nanling" specimenCount="24" typeStatus="paratype">
<specimenCount id="B78BBA72DC0D2E7344D99518368C7E66" type="generic">24 exs.</specimenCount>
, 
<location id="DBFCCF078F3912A3DE973B1711391A3D" LSID="urn:lsid:plazi:treatment:552395DB81535ABC987BEA547E83B8B0:DBFCCF078F3912A3DE973B1711391A3D" county="Dadongshan" municipality="Nanling" name="Guangdong Province">Guangdong Province</location>
, 
<collectingMunicipality id="D04F622F9F9375BB2EBD9C6142C97817">Nanling</collectingMunicipality>
, 
<collectingCounty id="068058BA6C95477BC25DCB9D6ED11790">Dadongshan</collectingCounty>
, 
<collectingDate id="42E6468CF4B61621615C3DC2B306B88F" value="2009-06-22" valueMax="2009-06-26" valueMin="2009-06-22">22-26.vi.2009</collectingDate>
, 
<collectorName id="ECA8BAC55E8CEEFC4D5959CBC0A7EB00">Fenglong Jia</collectorName>
leg
</materialsCitation>
; 
<materialsCitation id="7F7DD5F30AA56125784049A0C4F9722D" collectingDate="2008-09-23" collectorName="Yun Wang" county="Dadongshan" location="Guangdong Province" municipality="Lianzhou City" specimenCount="1" typeStatus="paratype">
5 exs, 
<location id="62A96E0A75E61866454EC591CE983D6D" LSID="urn:lsid:plazi:treatment:552395DB81535ABC987BEA547E83B8B0:62A96E0A75E61866454EC591CE983D6D" county="Dadongshan" municipality="Lianzhou City" name="Guangdong Province">Guangdong Province</location>
, 
<collectingMunicipality id="3706AA97451962F9014A0E1F6429C899">Lianzhou City</collectingMunicipality>
, 
<collectingCounty id="767BFC89884207F7CAEE1EC343096C9A">Dadongshan</collectingCounty>
, 
<collectingDate id="7629DB29D916B204512E49121EBE94D5" value="2008-09-23">23.ix.2008</collectingDate>
, 
<collectorName id="414A718F831728881AD9AAE94A2CEDAC">Yun Wang</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="CA7E7F0B8620F8D2BCFB2DC445A5A1E2" pageId="0" pageNumber="189" type="diagnosis">
<paragraph id="A37344EF2416F0D4F26AD2EA7BB04A6B" pageId="0" pageNumber="189">Diagnosis.</paragraph>
<paragraph id="EFEC0B7D16EA682F953DEC5ACA50EEA5" pageId="0" pageNumber="189">
This species is distinguished from other species of 
<taxonomicName id="5B64246A32E6854F8E6D50AE081D7E20" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agraphydrus" order="Coleoptera" pageId="0" pageNumber="189" phylum="Arthropoda" rank="genus">
<emphasis id="A39FE95D47C65D86B963B1643EDD3311" italics="true" pageId="0" pageNumber="189">Agraphydrus</emphasis>
</taxonomicName>
by the following combination of characters: labrum, clypeus and frons black; without preocular patches; maxillary palpi with apical palpomere slightly longer than penultimate, palpomere 4 without apical infuscation; clypeus without microsculpture, antennae with eight antennomeres; elytra with four rows of systematic punctures; meso- and metafemora pubescent in about basal 2/3; aedeagus with apex of the parameres distinct bending mesally, corona large, located at subapical position.
</paragraph>
</subSubSection>
<subSubSection id="641D56DADA4E6062583062D9375EE40C" pageId="0" pageNumber="189" type="description">
<paragraph id="0C8D451FEF8377520760CE7AF9BE1FAF" pageId="0" pageNumber="189">Description.</paragraph>
<paragraph id="00125E01BD90D3FCCE6CF907D214C89D" pageId="0" pageNumber="189">
<emphasis id="10DD971B7F4BBB3D6FFA70E37BC6A16E" bold="true" pageId="0" pageNumber="189">
<emphasis id="4FFD9A8FD8D1A28CB7049091CE4AD1CB" italics="true" pageId="0" pageNumber="189">Form and color</emphasis>
.
</emphasis>
Total length:1.9-2.3 mm; elytral width:1.0-1.2 mm; E.I.:1.1-1.2, P.I.:2.1-2.3, elytra 2.5-3 
<normalizedToken id="97C2B6458A0D7997B274A7DC70CD52AA" originalValue="×">x</normalizedToken>
as long as pronotum. Body oblong oval, distinct convex dorsally. Labrum, clypeus and frons black, preocular patches absent; maxillary palpi unicolored yellow; pronotum black with reddish-brown margin; elytral black; ventrites black; femora and tibia dark reddish-brown, tarsus yellow.
</paragraph>
<paragraph id="205D6A6027728DE3965F6293E16B322B" pageId="0" pageNumber="189">
<emphasis id="75AA40F22D4925E9AA1DD946709EABEC" bold="true" pageId="0" pageNumber="189">
<emphasis id="88035C222AA0CADCDEA934BE4F767FDF" italics="true" pageId="0" pageNumber="189">Head</emphasis>
.
</emphasis>
Labrum with dense, fine punctures and several coarse punctures. Clypeus (Fig. 
<figureCitation id="CD79E395D98F3DF9CDDDE7EA07E8AEA5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Agraphydrus pseudoniger sp. nov.: A. Clypeus; B. Clypeus; C. Antennae; D. Maxillary palpi; E. Mentum; F. Mesoventrite; G. Ventrite 5; H. Profemora; I. Mesofemora; J. Metafemora." figureDoi="10.3897/dez.68.66200.figure5" httpUri="https://binary.pensoft.net/fig/544293" pageId="0" pageNumber="189">5A</figureCitation>
) with angularly deep excised anterior margin and small median notch (Fig. 
<figureCitation id="5AAFC6372A633853767B6D9088FB0F75" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Agraphydrus pseudoniger sp. nov.: A. Clypeus; B. Clypeus; C. Antennae; D. Maxillary palpi; E. Mentum; F. Mesoventrite; G. Ventrite 5; H. Profemora; I. Mesofemora; J. Metafemora." figureDoi="10.3897/dez.68.66200.figure5" httpUri="https://binary.pensoft.net/fig/544293" pageId="0" pageNumber="189">5B</figureCitation>
), without microsculpture, with very sparse ground punctures, coarser than that on the labrum, interspaces 1-3 
<normalizedToken id="9D63458D61C2B717B1BB1B2E31AABF25" originalValue="×">x</normalizedToken>
as large as punctures. Systematic punctures distinct. Ground punctures on the frons as on the clypeus, systematic punctures distinct. Eyes small, not protruding. Antennae (Fig. 
<figureCitation id="E4383E0711C46B604EBC44E62303DB27" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Agraphydrus pseudoniger sp. nov.: A. Clypeus; B. Clypeus; C. Antennae; D. Maxillary palpi; E. Mentum; F. Mesoventrite; G. Ventrite 5; H. Profemora; I. Mesofemora; J. Metafemora." figureDoi="10.3897/dez.68.66200.figure5" httpUri="https://binary.pensoft.net/fig/544293" pageId="0" pageNumber="189">5C</figureCitation>
) with eight antennomeres. Maxillary palpi (Fig. 
<figureCitation id="7A7BA49E33AEF4A556EFBCD6EC969299" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Agraphydrus pseudoniger sp. nov.: A. Clypeus; B. Clypeus; C. Antennae; D. Maxillary palpi; E. Mentum; F. Mesoventrite; G. Ventrite 5; H. Profemora; I. Mesofemora; J. Metafemora." figureDoi="10.3897/dez.68.66200.figure5" httpUri="https://binary.pensoft.net/fig/544293" pageId="0" pageNumber="189">5D</figureCitation>
) 1.0-1.1 
<normalizedToken id="607248C1FB0F55076EF5568BA76C1B4C" originalValue="×">x</normalizedToken>
as long as pronotum in midline, 0.9-1.0 
<normalizedToken id="08612050A5F725BD0C1C44E79FFD5DD1" originalValue="×">x</normalizedToken>
as long as maximum width of clypeus; length ratio palpomere 4:3 =1.0-1.3, palpomere 4 asymmetrical. Mentum (Fig. 
<figureCitation id="C2316621B8A1434EBD4D4C3011C230CD" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Agraphydrus pseudoniger sp. nov.: A. Clypeus; B. Clypeus; C. Antennae; D. Maxillary palpi; E. Mentum; F. Mesoventrite; G. Ventrite 5; H. Profemora; I. Mesofemora; J. Metafemora." figureDoi="10.3897/dez.68.66200.figure5" httpUri="https://binary.pensoft.net/fig/544293" pageId="0" pageNumber="189">5E</figureCitation>
) with fine and sparse punctures on the lateral portions, without microsculpture.
</paragraph>
<caption id="6485C81B9A68A95CBA8F9368C1FF9502" doi="10.3897/dez.68.66200.figure5" httpUri="https://binary.pensoft.net/fig/544293" pageId="0" pageNumber="189" start="Figure 5" startId="F5">
<paragraph id="DBD32B7C24B6CB98D881F4627D4F8A59" pageId="0" pageNumber="189">
<emphasis id="FB5212878D3E80216208191D281F139A" bold="true" pageId="0" pageNumber="189">Figure 5.</emphasis>
<taxonomicName id="BF5BDAF06A3B57C1FBA721D3B719FA5E" authorityName="Yang &amp; Jia" authorityYear="2021" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agraphydrus pseudoniger" order="Coleoptera" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="pseudoniger">
<emphasis id="7DCA92918ABF7C7F9BC74B1F49D8CB81" italics="true" pageId="0" pageNumber="189">Agraphydrus pseudoniger</emphasis>
</taxonomicName>
sp. nov.: 
<emphasis id="E944E75339E3A9CD0815E36BC36181A3" bold="true" pageId="0" pageNumber="189">A.</emphasis>
Clypeus; 
<emphasis id="23AFE8694512CA2DDC39945335DB079D" bold="true" pageId="0" pageNumber="189">B.</emphasis>
Clypeus; 
<emphasis id="8B1D2A8A0DED4E3D02BD388B1283031B" bold="true" pageId="0" pageNumber="189">C.</emphasis>
Antennae; 
<emphasis id="A9066A824B40BD0370EF3DF5AAC4C2E0" bold="true" pageId="0" pageNumber="189">D.</emphasis>
Maxillary palpi; 
<emphasis id="C746FFD31AEBC3BCDCC6584B548F0D51" bold="true" pageId="0" pageNumber="189">E.</emphasis>
Mentum; 
<emphasis id="629C7CFCDD93B4C7235058BC43EA6B99" bold="true" pageId="0" pageNumber="189">F.</emphasis>
Mesoventrite; 
<emphasis id="390DA2B664327DE9975303BED41E30BE" bold="true" pageId="0" pageNumber="189">G.</emphasis>
Ventrite 5; 
<emphasis id="51E444871E94334D611BC02E7FC977A7" bold="true" pageId="0" pageNumber="189">H.</emphasis>
Profemora; 
<emphasis id="85756060FC39C20F84A19204AD4EDC84" bold="true" pageId="0" pageNumber="189">I.</emphasis>
Mesofemora; 
<emphasis id="FAE87625BB293A40D7821223E36C2720" bold="true" pageId="0" pageNumber="189">J.</emphasis>
Metafemora.
</paragraph>
</caption>
<paragraph id="CAAA1F5572AB659F2C76FC0AEFFA7500" pageId="0" pageNumber="189">
<emphasis id="7445023341F90D873E39797F090DB369" bold="true" pageId="0" pageNumber="189">
<emphasis id="1EC37F1B2D5A67C370188B71F70A62A5" italics="true" pageId="0" pageNumber="189">Thorax</emphasis>
.
</emphasis>
Pronotum ca. 1.10-1.25 
<normalizedToken id="1B6F66073E3ACA15A8BCE93DCDB9FB9F" originalValue="×">x</normalizedToken>
as wide as long, pronotal ground punctures sparse, as on frons, interspaces 1-3 
<normalizedToken id="EB8949B75740633F99B7CA4100FA1AC4" originalValue="×">x</normalizedToken>
as large as punctures, surface between ground punctures with microsculpture, systematic punctures distinct, located in the anterior and lateral middle position of the clypeus, each with a long seta on the middle. Elytral ground puncture slightly larger than that on head and pronotum, interspaces 1-2 
<normalizedToken id="9FD112D9F0A9F2AD8CA90C755BAD132F" originalValue="×">x</normalizedToken>
as large as punctures, surface between ground punctures as on the pronotum; with four distinct rows of systematic punctures, mesal rows reaching anterior margin. Prosternum slightly convex, without median carina. Mesoventrite (Fig. 
<figureCitation id="AAE4E1F25E389F464E92CE9AE462AD1D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Agraphydrus pseudoniger sp. nov.: A. Clypeus; B. Clypeus; C. Antennae; D. Maxillary palpi; E. Mentum; F. Mesoventrite; G. Ventrite 5; H. Profemora; I. Mesofemora; J. Metafemora." figureDoi="10.3897/dez.68.66200.figure5" httpUri="https://binary.pensoft.net/fig/544293" pageId="0" pageNumber="189">5F</figureCitation>
) with narrow, needle-like and strong median carina, extending to middle of mesofemora. Metaventrite moderately convex mesally, with small glabrous area posteromedially.
</paragraph>
<paragraph id="76BC74D62D225305D6C661B985F9D62E" pageId="0" pageNumber="189">
<emphasis id="52858C7C96D33267EF96E25AED8AE5C6" bold="true" pageId="0" pageNumber="189">
<emphasis id="8EC289DDFFA294072BB84C8E7D6E3F29" italics="true" pageId="0" pageNumber="189">Legs</emphasis>
.
</emphasis>
Pubescence present on proximal 2/3 of femora (Fig. 
<figureCitation id="3BFDD82E72FF00C3E025090423A28276" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Agraphydrus pseudoniger sp. nov.: A. Clypeus; B. Clypeus; C. Antennae; D. Maxillary palpi; E. Mentum; F. Mesoventrite; G. Ventrite 5; H. Profemora; I. Mesofemora; J. Metafemora." figureDoi="10.3897/dez.68.66200.figure5" httpUri="https://binary.pensoft.net/fig/544293" pageId="0" pageNumber="189">5H, I, J</figureCitation>
), hairline oblique on pro- and mesofemora, straight on metafemora.
</paragraph>
<paragraph id="ACB52139DEDF3251D69C9C5B3801B1CE" pageId="0" pageNumber="189">
<emphasis id="0A474D8B2532F1D7681B39C017F51424" bold="true" pageId="0" pageNumber="189">
<emphasis id="55F769243FD09836979EF203E026B92D" italics="true" pageId="0" pageNumber="189">Abdomen</emphasis>
.
</emphasis>
Ventrite 5 (Fig. 
<figureCitation id="7F4F308B1D78AF12D97457D372DED2E9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Agraphydrus pseudoniger sp. nov.: A. Clypeus; B. Clypeus; C. Antennae; D. Maxillary palpi; E. Mentum; F. Mesoventrite; G. Ventrite 5; H. Profemora; I. Mesofemora; J. Metafemora." figureDoi="10.3897/dez.68.66200.figure5" httpUri="https://binary.pensoft.net/fig/544293" pageId="0" pageNumber="189">5G</figureCitation>
) round, without apical emargination, with a row of strong setae.
</paragraph>
<paragraph id="5180DD7450C456C48FF338D2503652B7" pageId="0" pageNumber="189">
<emphasis id="D985C5642433A2B3EA7B7A8CA33E590A" bold="true" italics="true" pageId="0" pageNumber="189">Aedeagus</emphasis>
(Fig. 
<figureCitation id="FB3695C9DE3CAE3E31F848DFA9A47ADD" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Aedeagi: A. Agraphydrus dapengensis sp. nov.; B. Agraphydrus komareki sp. nov.; C. Agraphydrus pseudoniger sp. nov.; D. Agraphydrus sabulosus sp. nov." figureDoi="10.3897/dez.68.66200.figure7" httpUri="https://binary.pensoft.net/fig/544295" pageId="0" pageNumber="189">7C</figureCitation>
). Length: 0.31mm. Phallobase about as long as parameres, manubrium wide and triangular. Base of parameres wide, slightly narrowing apicad, apical 1/3 strongly narrowed, apex blunt, moderately bending mesad, lateral margin straight at basal 2/3, cambered in apical 1/3, basal protrusion extending into about half of phallobase. Median lobe slender, shaped like an index finger, margin cambered; apex bluntly rounded, slightly shorter than parameres, corona moderately larger, located at subapical position; basal apophyses long, reaching to half of phallobase.
</paragraph>
</subSubSection>
<subSubSection id="CD562BE0A87208216365B45730469C30" pageId="0" pageNumber="189" type="etymology">
<paragraph id="54A2BB91DB64A5824E67BDBF3011187B" pageId="0" pageNumber="189">Etymology.</paragraph>
<paragraph id="15BBA8E64EC8CF83D61E4C2046DC20FE" pageId="0" pageNumber="189">
Specific name derived from Latin 
<normalizedToken id="C7D1E3CCA3FBCDDB032637ECD39704A4" originalValue="“pseudo”">&quot;pseudo&quot;</normalizedToken>
, false, and &quot; 
<taxonomicName id="C9E075E39063DB378127EA255F600647" authorityName="Komarek &amp; Hebauer" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agraphydrus niger" order="Coleoptera" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="niger">Agraphydrus niger</taxonomicName>
&quot;, black, refers to the similarity with 
<taxonomicName id="28E14F3A0068360FB51C7F97CE60247B" authorityName="Komarek &amp; Hebauer" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agraphydrus niger" order="Coleoptera" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="3CEDEFC704DF2C02CBBA9B496C0EE1AB" italics="true" pageId="0" pageNumber="189">Agraphydrus niger</emphasis>
</taxonomicName>
Komarek &amp; Hebauer, 2018.
</paragraph>
</subSubSection>
<subSubSection id="984488D1B9B982260D836DDBD2842FCC" pageId="0" pageNumber="189" type="distribution">
<paragraph id="39C2DABB9C0AD1330A4DE5E35FD2110B" pageId="0" pageNumber="189">Distribution.</paragraph>
<paragraph id="0D16C34F8378B21975146E9CCD729038" pageId="0" pageNumber="189">CHINA (Jiangxi, Guangdong).</paragraph>
</subSubSection>
<subSubSection id="74B9CF244D173512B45C4FFD7C4B83CE" pageId="0" pageNumber="189" type="remarks">
<paragraph id="54CA8469CEB116CEE00A7FEB635075C0" pageId="0" pageNumber="189">Remarks.</paragraph>
<paragraph id="7396930DBB2FE5A8C43D224260CA719F" pageId="0" pageNumber="189">
This species shares four rows of distinct systematic punctures, absence of clypeal microsculpture, unicolorous maxillary palpomeres, antennae with eight antennomeres with 
<taxonomicName id="47E5429E41947DA05D1139C8719A937E" lsidName="A. fasciatus" pageId="0" pageNumber="189" rank="species" species="fasciatus">
<emphasis id="D510FE3A2325EEC213E5AC08A20141BC" italics="true" pageId="0" pageNumber="189">A. fasciatus</emphasis>
</taxonomicName>
Komarek &amp; Hebauer, 
<taxonomicName id="0785032470E78928947EA53DC8B30655" lsidName="A. niger" pageId="0" pageNumber="189" rank="species" species="niger">
<emphasis id="386CCA5FEB94A88FABD62049B3921D1F" italics="true" pageId="0" pageNumber="189">A. niger</emphasis>
</taxonomicName>
Komarek &amp; Hebauer, 
<taxonomicName id="75EEF0507A41436D95614848405DDC9C" lsidName="A. yunnanensis" pageId="0" pageNumber="189" rank="species" species="yunnanensis">
<emphasis id="8D6B6918F4A9955F8CEF6A8709583956" italics="true" pageId="0" pageNumber="189">A. yunnanensis</emphasis>
</taxonomicName>
Komarek &amp; Hebauer and 
<taxonomicName id="825EB8E9F2F1C0F72FD49302EFAB9A8F" lsidName="A. longipenis" pageId="0" pageNumber="189" rank="species" species="longipenis">
<emphasis id="EA5A760D08F2C9E78E70138BC8D83F7A" italics="true" pageId="0" pageNumber="189">A. longipenis</emphasis>
</taxonomicName>
Komarek &amp; Hebauer. It differs from 
<taxonomicName id="070EF4890FC469C7167D79F7E37C24B7" lsidName="A. yunnanensis" pageId="0" pageNumber="189" rank="species" species="yunnanensis">
<emphasis id="BB635D924CD06F33EFC3C0C6ED2DBDE3" italics="true" pageId="0" pageNumber="189">A. yunnanensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="F83F5A49A7680911ACC52D71A501D140" lsidName="A. longipenis" pageId="0" pageNumber="189" rank="species" species="longipenis">
<emphasis id="A95C501E1E91BD1A9A6CB45EC748103D" italics="true" pageId="0" pageNumber="189">A. longipenis</emphasis>
</taxonomicName>
by eyes small, mesoventrite carinate (
<taxonomicName id="8452946C0DD141F7D6053DC2293B2E56" lsidName="A. yunnanensis" pageId="0" pageNumber="189" rank="species" species="yunnanensis">
<emphasis id="51A032AAAF97B9BD44FD1A38C7345CA7" italics="true" pageId="0" pageNumber="189">A. yunnanensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="A4D7310A4894E94745B9AC91A97CFD82" lsidName="A. longipenis" pageId="0" pageNumber="189" rank="species" species="longipenis">
<emphasis id="4EDB0C22D6783DFEF707A1FF0E5625CA" italics="true" pageId="0" pageNumber="189">A. longipenis</emphasis>
</taxonomicName>
without mesoventral carina), parameres moderately curved inwards subapically, median lobe with corona located at subapical position (apex of parameres not curved inwards, median lobe with corona located at base in 
<taxonomicName id="A09F2FB51C50C42DD0B36B1FE4F0E3FC" lsidName="A. yunnanensis" pageId="0" pageNumber="189" rank="species" species="yunnanensis">
<emphasis id="12DE6FCD9A22BEFA6E0EE6304AB0F1F6" italics="true" pageId="0" pageNumber="189">A. yunnanensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="FECFAC28730C2BF43F4FC5944B17500B" lsidName="A. longipenis" pageId="0" pageNumber="189" rank="species" species="longipenis">
<emphasis id="117164B5F5FA5C86E37520265B228737" italics="true" pageId="0" pageNumber="189">A. longipenis</emphasis>
</taxonomicName>
). Shares small eyes, mesoventral carina with 
<taxonomicName id="6AF225EF9B9B16BC20A03B498F4B1360" lsidName="A. fasciatus" pageId="0" pageNumber="189" rank="species" species="fasciatus">
<emphasis id="BC274A145701B4A5B37AA9203CB03C75" italics="true" pageId="0" pageNumber="189">A. fasciatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="0A09E511E1FB05371EA744122543326B" lsidName="A. niger" pageId="0" pageNumber="189" rank="species" species="niger">
<emphasis id="CF2A38BC51F3FC03651DC84EC0BCDF89" italics="true" pageId="0" pageNumber="189">A. niger</emphasis>
</taxonomicName>
and 
<taxonomicName id="C0A0FBBB00BC555E53808285269168B8" lsidName="A. fujianensis" pageId="0" pageNumber="189" rank="species" species="fujianensis">
<emphasis id="6F156182126EDB73931A4DDA8C53B05A" italics="true" pageId="0" pageNumber="189">A. fujianensis</emphasis>
</taxonomicName>
Komarek &amp; Hebauer, similar aedeagus with 
<taxonomicName id="E9DE64E7EB5D7A331F9AE34A2128E248" lsidName="A. fasciatus" pageId="0" pageNumber="189" rank="species" species="fasciatus">
<emphasis id="97C0E8A9030C926335D97AF0E062F0F7" italics="true" pageId="0" pageNumber="189">A. fasciatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="FF97C00A71B2987200E2295E4EDCA9EE" lsidName="A. niger" pageId="0" pageNumber="189" rank="species" species="niger">
<emphasis id="5C14BE482E8DDE5ACE5CB6CDAB058AAF" italics="true" pageId="0" pageNumber="189">A. niger</emphasis>
</taxonomicName>
and 
<taxonomicName id="B36029BD3C420851EAFE6762B5E54CA5" lsidName="A. politus" pageId="0" pageNumber="189" rank="species" species="politus">
<emphasis id="76DEF3B9D9F42771F23890C461C0482B" italics="true" pageId="0" pageNumber="189">A. politus</emphasis>
</taxonomicName>
Komarek &amp; Hebauer, differs from 
<taxonomicName id="F89DB165A1AC3C3AA074E5E5B25F31EA" lsidName="A. fujianensis" pageId="0" pageNumber="189" rank="species" species="fujianensis">
<emphasis id="AC9F0DFFD7AC71CCB8818F4A24C3248B" italics="true" pageId="0" pageNumber="189">A. fujianensis</emphasis>
</taxonomicName>
by abdominal ventrite 5 with apical emargination, coloration of pronotum and elytra (dark yellowish to rufous brown in 
<taxonomicName id="7F98EBDDD50A8C818CE6F6A968CE291B" lsidName="A. fujianensis" pageId="0" pageNumber="189" rank="species" species="fujianensis">
<emphasis id="7E5FD53313A7941587BF4C00AB2F2F7E" italics="true" pageId="0" pageNumber="189">A. fujianensis</emphasis>
</taxonomicName>
, black in 
<taxonomicName id="21902323ED4E6821A03D34F4A1A0ABCA" lsidName="A. pseudoniger" pageId="0" pageNumber="189" rank="species" species="pseudoniger">
<emphasis id="AD8120EDD88432F6C4DE5CBF7855865A" italics="true" pageId="0" pageNumber="189">A. pseudoniger</emphasis>
</taxonomicName>
), parameres moderately curved inwards subapically and corona of median lobe locating at subapical position (apex of parameres not curved inwards, median lobe with corona basally in 
<taxonomicName id="BDC9A30145A96A76AAE1B2D48DAE551E" lsidName="A. fujianensis" pageId="0" pageNumber="189" rank="species" species="fujianensis">
<emphasis id="109021AC4DAC0194E324899DD9F9E23A" italics="true" pageId="0" pageNumber="189">A. fujianensis</emphasis>
</taxonomicName>
); from 
<taxonomicName id="07A5A2B3B6EBD5F5BAE4936CF3BD0E27" lsidName="A. politus" pageId="0" pageNumber="189" rank="species" species="politus">
<emphasis id="3916A743AE77C706551ECFFB2F933B16" italics="true" pageId="0" pageNumber="189">A. politus</emphasis>
</taxonomicName>
by small size, with four rows of punctures (ten in 
<taxonomicName id="C5AE4B9729E5F3E8FA39923E47056F3B" lsidName="A. politus" pageId="0" pageNumber="189" rank="species" species="politus">
<emphasis id="6A6CE441BA51A8D2DE95B28EFA94FFDD" italics="true" pageId="0" pageNumber="189">A. politus</emphasis>
</taxonomicName>
), phallobase about as long as parameres and apex of parameres moderately curved inwards (phallobase slightly longer than the parameres, parameres strongly curved inwards subapically in 
<taxonomicName id="01AC5CD402CAD9288DE8F77426ACD651" lsidName="A. politus" pageId="0" pageNumber="189" rank="species" species="politus">
<emphasis id="618B2E5DDDD43A65C77F566B37A3638D" italics="true" pageId="0" pageNumber="189">A. politus</emphasis>
</taxonomicName>
); from 
<taxonomicName id="A9B8EC02E5950FD6F638792BC64CB259" lsidName="A. fasciatus" pageId="0" pageNumber="189" rank="species" species="fasciatus">
<emphasis id="6DDF3BDD94F1D18E6FF1ED0B80E53F53" italics="true" pageId="0" pageNumber="189">A. fasciatus</emphasis>
</taxonomicName>
by clypeus without preocular patches, pronotum and elytra black and parameres slightly longer than median lobe (parameres about 2x as wide as median lobe in 
<taxonomicName id="224CC2D95365A6156FE789BCBF758395" lsidName="A. fasciatus" pageId="0" pageNumber="189" rank="species" species="fasciatus">
<emphasis id="5B1B5E0386109FF9C512EB5C7939099D" italics="true" pageId="0" pageNumber="189">A. fasciatus</emphasis>
</taxonomicName>
); from 
<taxonomicName id="4954D9D6A0E521C4D65E9306CC2F10F6" lsidName="A. niger" pageId="0" pageNumber="189" rank="species" species="niger">
<emphasis id="AC063874D63111A6170D8DC7007E623C" italics="true" pageId="0" pageNumber="189">A. niger</emphasis>
</taxonomicName>
by clypeus with a deep excision anteriorly and small median notch and mesal rows of systematic punctures reaching anterior margin.
</paragraph>
</subSubSection>
</treatment>
</document>