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<document id="ECC91A3D2880B843522C192E6E422E92" ID-DOI="10.11646/zootaxa.3934.1.1" ID-GBIF-Dataset="0abffe3e-a2c6-4623-b7c6-c8b26cee4a2b" ID-ISSN="1175-5326" ID-Zenodo-Dep="236066" ID-ZooBank="11C2A2CB-30B3-4694-B379-AE9D47332F0C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461012229557" checkinUser="plazi" docAuthor="Winterbottom, Richard &amp; Hoese, Douglass F." docDate="2015" docId="5519879AB84BF30CFF1FFD2E6E3EEC8A" docLanguage="en" docName="zt03934p102.pdf" docOrigin="Zootaxa 3934 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Trimma preclarum Winterbottom 2006" docType="treatment" docVersion="11" lastPageNumber="74" masterDocId="A920FFE2B803F345FF88FFF66B4DEB78" masterDocTitle="A revision of the Australian species of Trimma (Actinopterygii, Gobiidae), with descriptions of six new species and redescriptions of twenty-three valid species" masterLastPageNumber="102" masterPageNumber="1" pageNumber="73" updateTime="1698567702668" updateUser="plazi">
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<mods:title id="8B3DD059809FD96CC13A0AC575003B8C">A revision of the Australian species of Trimma (Actinopterygii, Gobiidae), with descriptions of six new species and redescriptions of twenty-three valid species</mods:title>
</mods:titleInfo>
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<mods:namePart id="BB6C43341E78400E9B2CA8C364641EE8">Winterbottom, Richard</mods:namePart>
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<mods:namePart id="D0B9AC59A087CE79F0E449F3BD19CD0A">Hoese, Douglass F.</mods:namePart>
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<mods:typeOfResource id="A6927903F3DD9D274A628E34780929EF">text</mods:typeOfResource>
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<mods:title id="9B25CA2072B273F2230B4867A7A079E0">Zootaxa</mods:title>
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<mods:date id="3010BF52B032F23E5B0622DF3845C272">2015</mods:date>
<mods:detail id="F123A1901E92CDC62419137C9DF0C76D" type="volume">
<mods:number id="3220445015E4A61AF65C5E54AA9BAEF8">3934</mods:number>
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<mods:identifier id="2288BD2794F943532CB2D0007717F50F" type="DOI">10.11646/zootaxa.3934.1.1</mods:identifier>
<mods:identifier id="D31D9AF6A110C0967E0F59B041B08CBA" type="GBIF-Dataset">0abffe3e-a2c6-4623-b7c6-c8b26cee4a2b</mods:identifier>
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<treatment id="5519879AB84BF30CFF1FFD2E6E3EEC8A" ID-DOI="http://doi.org/10.5281/zenodo.3508956" ID-GBIF-Taxon="119600467" ID-Zenodo-Dep="3508956" LSID="urn:lsid:plazi:treatment:5519879AB84BF30CFF1FFD2E6E3EEC8A" httpUri="http://treatment.plazi.org/id/5519879AB84BF30CFF1FFD2E6E3EEC8A" lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73">
<subSubSection id="95AA6507B84BF30DFF1FFD2E6A3BE84F" pageId="72" pageNumber="73" type="nomenclature">
<paragraph id="DD0F368CB84BF30DFF1FFD2E6932E98A" blockId="72.[151,639,728,823]" box="[151,639,728,754]" pageId="72" pageNumber="73">
<heading id="864781E0B84BF30DFF1FFD2E6932E98A" bold="true" box="[151,639,728,754]" fontSize="11" level="1" pageId="72" pageNumber="73" reason="1">
<taxonomicName id="1AB04D0FB84BF30DFF1FFD2E6932E98A" ID-CoL="58V78" authority="Winterbottom, 2006" authorityName="Winterbottom" authorityYear="2006" box="[151,639,728,754]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="72" pageNumber="73" phylum="Chordata" rank="species" species="preclarum">
<emphasis id="EFC4EA9EB84BF30DFF1FFD2E6932E98A" bold="true" box="[151,639,728,754]" pageId="72" pageNumber="73">
<emphasis id="EFC4EA9EB84BF30DFF1FFD2E6A30E98A" bold="true" box="[151,381,728,754]" italics="true" pageId="72" pageNumber="73">Trimma preclarum</emphasis>
<bibRefCitation id="B9214B7DB84BF30DFE0BFD2E6932E98A" author="Winterbottom" box="[387,639,728,754]" pageId="72" pageNumber="73" refString="Winterbottom, R. (2006) Two new species of the gobiid fish Trimma from the coral reefs of the western Pacific Ocean (Pisces; Perciformes; Gobioidei). Zootaxa, 1331, 55 - 68." type="journal article" year="2006">Winterbottom, 2006</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="DD0F368CB84BF30DFF1FFD0C6AC2E86B" blockId="72.[151,639,728,823]" box="[151,399,762,787]" pageId="72" pageNumber="73">Exquisite Pygmygoby</paragraph>
<paragraph id="DD0F368CB84BF30DFF1FFCE86A3BE84F" blockId="72.[151,639,728,823]" box="[151,374,798,823]" pageId="72" pageNumber="73">
<figureCitation id="458B2A09B84BF30DFF1FFCE86A50E84F" box="[151,285,798,823]" captionStart-0="FIGURE 40" captionStart-1="FIGURE 41" captionStartId-0="71.[151,250,1899,1921]" captionStartId-1="73.[151,250,1030,1052]" captionTargetBox-0="[161,1425,1049,1867]" captionTargetBox-1="[180,1385,678,978]" captionTargetId-0="figure@71.[151,1436,1043,1877]" captionTargetId-1="figure@73.[151,1435,654,1009]" captionTargetPageId-0="71" captionTargetPageId-1="73" captionText-0="FIGURE 40. Distribution map for Trimma pentherum (open triangles) and T. preclarum (solid squares). A solid star indicates localities at which both species occur." captionText-1="FIGURE 41. Trimma preclarum, 21.3 mm SL female, ROM 83037, Merir Id, Palau. Photo by R. Winterbottom." httpUri-0="https://zenodo.org/record/236107/files/figure.png" httpUri-1="https://zenodo.org/record/236108/files/figure.png" pageId="72" pageNumber="73">Figs. 4041</figureCitation>
, Pl. 3 C
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B84BF30DFF1FFC9E69D7E8E3" pageId="72" pageNumber="73" type="reference_group">
<paragraph id="DD0F368CB84BF30DFF1FFC9E69D7E8E3" blockId="72.[151,1257,870,923]" pageId="72" pageNumber="73">
<treatmentCitationGroup id="FDA011A2B84BF30DFF1FFC9E69D7E8E3" pageId="72" pageNumber="73">
<taxonomicName id="1AB04D0FB84BF30DFF1FFC9E690AE804" ID-CoL="58V78" authority="Winterbottom, 2006:63" authorityName="Winterbottom" authorityPageNumber="63" authorityYear="2006" box="[151,583,870,893]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="72" pageNumber="73" phylum="Chordata" rank="species" species="preclarum">
<emphasis id="EFC4EA9EB84BF30DFF1FFC9E6A18E804" box="[151,341,870,893]" italics="true" pageId="72" pageNumber="73">Trimma preclarum</emphasis>
<treatmentCitation id="5C11109DB84BF30DFED3FC90690AE804" ID-DOI="http://doi.org/10.5281/zenodo.3501549" ID-Zenodo-Dep="3501549" LSID="urn:lsid:zoobank.org:act:15E8AE99-6636-4C83-920E-99FB98C86E75" author="Winterbottom" box="[347,583,870,892]" httpUri="http://treatment.plazi.org/id/C28D1C3C5B20E96B60E21100A3F2AFB3" page="63" pageId="72" pageNumber="73" year="2006">
<bibRefCitation id="B9214B7DB84BF30DFED3FC906966E804" author="Winterbottom" box="[347,555,870,892]" pageId="72" pageNumber="73" refString="Winterbottom, R. (2006) Two new species of the gobiid fish Trimma from the coral reefs of the western Pacific Ocean (Pisces; Perciformes; Gobioidei). Zootaxa, 1331, 55 - 68." type="journal article" year="2006">Winterbottom, 2006</bibRefCitation>
:63
</treatmentCitation>
</taxonomicName>
(
<collectingCountry id="A5A7761CB84BF30DFDDDFC906931E804" box="[597,636,870,892]" name="Fiji" pageId="72" pageNumber="73">Fiji</collectingCountry>
, also
<collectingCountry id="A5A7761CB84BF30DFD3DFC9069BFE804" box="[693,754,870,892]" name="Palau" pageId="72" pageNumber="73">Palau</collectingCountry>
);
<treatmentCitation id="5C11109DB84BF30DFC8BFC906F69E804" author="Allen" box="[771,1060,870,892]" page="944" pageId="72" pageNumber="73" year="2012">
<bibRefCitation id="B9214B7DB84BF30DFC8BFC9068B7E804" author="Allen" box="[771,1018,870,892]" pageId="72" pageNumber="73" refString="Allen, G. R. &amp; Erdmann, M. V. (2012) Reef fishes of the East Indies. Vol. III. Tropical Reef Research, Perth, pp. 857 - 1292." type="book chapter" year="2012">Allen &amp; Erdmann, 2012</bibRefCitation>
:944
</treatmentCitation>
(
<collectingCountry id="A5A7761CB84BF30DFBBAFC906FDBE804" box="[1074,1174,870,892]" name="Indonesia" pageId="72" pageNumber="73">Indonesia</collectingCountry>
to
<collectingCountry id="A5A7761CB84BF30DFB3EFC906F93E804" box="[1206,1246,870,892]" name="Fiji" pageId="72" pageNumber="73">Fiji</collectingCountry>
).
<taxonomicName id="1AB04D0FB84BF30DFF1FFC706BABE8E3" ID-CoL="827C" box="[151,230,902,923]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="72" pageNumber="73" phylum="Chordata" rank="genus">
<emphasis id="EFC4EA9EB84BF30DFF1FFC706BABE8E3" box="[151,230,902,923]" italics="true" pageId="72" pageNumber="73">Trimma</emphasis>
</taxonomicName>
sp.:
<treatmentCitation id="5C11109DB84BF30DFE9EFC736909E8E3" author="Hayashi" box="[278,580,901,923]" page="44" pageId="72" pageNumber="73" year="2003">
<bibRefCitation id="B9214B7DB84BF30DFE9EFC73696BE8E3" author="Hayashi" box="[278,550,901,923]" pageId="72" pageNumber="73" refString="Hayashi, M. &amp; Shiratori, T. (2003) Gobies of Japanese Waters. Hankyu Books, Osaka, 224 pp. [in Japanese]" type="book" year="2003">Hayashi &amp; Shiratori, 2003</bibRefCitation>
:44
</treatmentCitation>
, Saipan.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B84BF30DFF1FFC316809EF87" pageId="72" pageNumber="73" type="materials_examined">
<paragraph id="DD0F368CB84BF30DFF1FFC3168C2EF08" blockId="72.[151,1437,967,2029]" pageId="72" pageNumber="73">
<emphasis id="EFC4EA9EB84BF30DFF1FFC316924E898" bold="true" box="[151,617,967,992]" pageId="72" pageNumber="73">
Australian material. New South
<collectingCountry id="A5A7761CB84BF30DFD90FC316929E898" box="[536,612,967,992]" name="United Kingdom" pageId="72" pageNumber="73">Wales</collectingCountry>
:
</emphasis>
Coffs Harbour:
<collectionCode id="BBA1AE49B84BF30DFCA1FC3E682BE898" box="[809,870,968,992]" pageId="72" pageNumber="73">AMS</collectionCode>
I.30456-003, (22.8).
<emphasis id="EFC4EA9EB84BF30DFBDCFC316FA6E898" bold="true" box="[1108,1259,967,992]" pageId="72" pageNumber="73">Queensland:</emphasis>
<collectingCountry id="A5A7761CB84BF30DFB7AFC316E14E898" box="[1266,1369,967,992]" name="Ashmore and Cartier Islands" pageId="72" pageNumber="73">Ashmore</collectingCountry>
Reef:
<collectionCode id="BBA1AE49B84BF30DFF1FFC1A6B99EF7C" box="[151,212,1004,1028]" pageId="72" pageNumber="73">AMS</collectionCode>
I.33717-083, 4(9.816.0),
<quantity id="1A489B69B84BF30DFD81FC1A6929EF7C" box="[521,612,1004,1028]" metricMagnitude="1" metricUnit="m" metricValue="1.65" metricValueMax="3.2" metricValueMin="0.1" pageId="72" pageNumber="73" unit="m" value="16.5" valueMax="32.0" valueMin="1.0">
1
<specimenCount id="CBB6FD05B84BF30DFDABFC1A6929EF7C" box="[547,612,1004,1028]" pageId="72" pageNumber="73" type="generic">32 m</specimenCount>
</quantity>
.
<collectionCode id="BBA1AE49B84BF30DFDF8FC1A69E0EF7C" box="[624,685,1004,1028]" pageId="72" pageNumber="73">AMS</collectionCode>
I.33720-063, 10(9.617.0),
<quantity id="1A489B69B84BF30DFC79FC1A6F06EF7C" box="[1009,1099,1004,1028]" metricMagnitude="1" metricUnit="m" metricValue="1.7" metricValueMax="3.2" metricValueMin="0.2" pageId="72" pageNumber="73" unit="m" value="17.0" valueMax="32.0" valueMin="2.0">
2
<specimenCount id="CBB6FD05B84BF30DFB85FC1A6F06EF7C" box="[1037,1099,1004,1028]" pageId="72" pageNumber="73" type="generic">32 m</specimenCount>
</quantity>
. Escape Reef:
<collectionCode id="BBA1AE49B84BF30DFB7FFC1A6E79EF7C" box="[1271,1332,1004,1028]" pageId="72" pageNumber="73">AMS</collectionCode>
I.22627- 0 17, 2(1821). Herald Cays:
<collectionCode id="BBA1AE49B84BF30DFE57FBE6696EEF50" LSID="urn:lsid:biocol.org:col:34981" box="[479,547,1040,1064]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="72" pageNumber="73">WAM</collectionCode>
P.28537-004, 2(13.216.2);
<collectionCode id="BBA1AE49B84BF30DFCEEFBE668E6EF50" LSID="urn:lsid:biocol.org:col:34981" box="[870,939,1040,1064]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="72" pageNumber="73">WAM</collectionCode>
P.28537-006, 5(12.518.7);
<collectionCode id="BBA1AE49B84BF30DFB65FBE66E7FEF50" LSID="urn:lsid:biocol.org:col:34981" box="[1261,1330,1040,1064]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="72" pageNumber="73">WAM</collectionCode>
P.29630- 0 0 9, 2(17.417.7). Holmes Reef:
<collectionCode id="BBA1AE49B84BF30DFDAAFBC2692AEF34" LSID="urn:lsid:biocol.org:col:34981" box="[546,615,1076,1100]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="72" pageNumber="73">WAM</collectionCode>
P.28541-046, 15(10.618.2);
<collectionCode id="BBA1AE49B84BF30DFC4CFBC26F45EF34" LSID="urn:lsid:biocol.org:col:34981" box="[964,1032,1076,1100]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="72" pageNumber="73">WAM</collectionCode>
P.28541-047, 3(15.517.6);
<collectionCode id="BBA1AE49B84BF30DFAD0FBC26ED0EF34" LSID="urn:lsid:biocol.org:col:34981" box="[1368,1437,1076,1100]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="72" pageNumber="73">WAM</collectionCode>
P.29627-040, 3(19.020.0). Lihou Reef:
<collectionCode id="BBA1AE49B84BF30DFDD6FBAE69EEEF08" LSID="urn:lsid:biocol.org:col:34981" box="[606,675,1112,1136]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="72" pageNumber="73">WAM</collectionCode>
P.29641-014, (20.1).
</paragraph>
<paragraph id="DD0F368CB84BF30DFF4FFB8A6809EF87" blockId="72.[151,1437,967,2029]" pageId="72" pageNumber="73">
<emphasis id="EFC4EA9EB84BF30DFF4FFB8A6ACFEFED" bold="true" box="[199,386,1148,1173]" pageId="72" pageNumber="73">Other material.</emphasis>
In addition to the
<typeStatus id="020B882EB84BF30DFDDDFB8B69C8EFED" box="[597,645,1149,1173]" pageId="72" pageNumber="73">type</typeStatus>
material from
<collectingCountry id="A5A7761CB84BF30DFCA6FB8A6818EFED" box="[814,853,1148,1173]" name="Fiji" pageId="72" pageNumber="73">Fiji</collectingCountry>
and
<collectingCountry id="A5A7761CB84BF30DFC05FB8A6883EFED" box="[909,974,1148,1173]" name="Palau" pageId="72" pageNumber="73">Palau</collectingCountry>
, we have also examined specimens from the following localities:
<emphasis id="EFC4EA9EB84BF30DFE20FB696932EFC0" bold="true" box="[424,639,1183,1208]" pageId="72" pageNumber="73">
<collectingCountry id="A5A7761CB84BF30DFE20FB696937EFC0" box="[424,634,1183,1208]" name="American Samoa" pageId="72" pageNumber="73">American Samoa</collectingCountry>
:
</emphasis>
<collectionCode id="BBA1AE49B84BF30DFD0EFB56698BEFC0" box="[646,710,1184,1208]" pageId="72" pageNumber="73">AMS</collectionCode>
.
<emphasis id="EFC4EA9EB84BF30DFD59FB696876EFC0" bold="true" box="[721,827,1183,1208]" pageId="72" pageNumber="73">
<collectingCountry id="A5A7761CB84BF30DFD59FB69687BEFC0" box="[721,822,1183,1208]" name="Kiribati" pageId="72" pageNumber="73">Kiribati</collectingCountry>
:
</emphasis>
Abaiang Atoll:
<collectionCode id="BBA1AE49B84BF30DFC78FB566F7DEFC0" box="[1008,1072,1184,1208]" pageId="72" pageNumber="73">AMS</collectionCode>
.
<emphasis id="EFC4EA9EB84BF30DFBB3FB696FF4EFC0" bold="true" box="[1083,1209,1183,1208]" pageId="72" pageNumber="73">
<collectingCountry id="A5A7761CB84BF30DFBB3FB696FF9EFC0" box="[1083,1204,1183,1208]" name="Indonesia" pageId="72" pageNumber="73">Indonesia</collectingCountry>
:
</emphasis>
Raja Ampat:
<collectionCode id="BBA1AE49B84BF30DFADDFB566ED5EFC0" LSID="urn:lsid:biocol.org:col:34778" box="[1365,1432,1184,1208]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="72" pageNumber="73">ROM</collectionCode>
.
<emphasis id="EFC4EA9EB84BF30DFF1FFB326AC3EFA5" bold="true" box="[151,398,1220,1245]" pageId="72" pageNumber="73">
<collectingCountry id="A5A7761CB84BF30DFF1FFB326AC4EFA5" box="[151,393,1220,1245]" name="Papua New Guinea" pageId="72" pageNumber="73">Papua New Guinea</collectingCountry>
:
</emphasis>
Ninigo Ids:
<collectionCode id="BBA1AE49B84BF30DFDAFFB326936EFA4" LSID="urn:lsid:biocol.org:col:34871" box="[551,635,1220,1244]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="72" pageNumber="73">USNM</collectionCode>
. Hermit Ids:
<collectionCode id="BBA1AE49B84BF30DFC92FB326823EFA4" LSID="urn:lsid:biocol.org:col:34871" box="[794,878,1220,1244]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="72" pageNumber="73">USNM</collectionCode>
.
<emphasis id="EFC4EA9EB84BF30DFCF4FB3268A4EFA5" bold="true" box="[892,1001,1220,1245]" pageId="72" pageNumber="73">
<collectingCountry id="A5A7761CB84BF30DFCF4FB326892EFA5" box="[892,991,1220,1245]" name="Vanuatu" pageId="72" pageNumber="73">Vanuatu</collectingCountry>
:
</emphasis>
<collectionCode id="BBA1AE49B84BF30DFC7EFB326F7BEFA4" box="[1014,1078,1220,1244]" pageId="72" pageNumber="73">AMS</collectionCode>
.
<emphasis id="EFC4EA9EB84BF30DFBCDFB326F99EFA5" bold="true" box="[1093,1236,1220,1245]" pageId="72" pageNumber="73">
<collectingCountry id="A5A7761CB84BF30DFBCDFB326F82EFA5" box="[1093,1231,1220,1245]" name="Micronesia" pageId="72" pageNumber="73">Micronesia</collectingCountry>
:
</emphasis>
<collectingCountry id="A5A7761CB84BF30DFB57FB336E7BEFA4" box="[1247,1334,1221,1244]" name="Micronesia" pageId="72" pageNumber="73">Ponape</collectingCountry>
:
<collectionCode id="BBA1AE49B84BF30DFACCFB326ED4EFA4" LSID="urn:lsid:biocol.org:col:34871" box="[1348,1433,1220,1244]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="72" pageNumber="73">USNM</collectionCode>
. Kapingamarangi Atoll:
<collectionCode id="BBA1AE49B84BF30DFE29FB1E6A96EE78" LSID="urn:lsid:biocol.org:col:34700" box="[417,475,1256,1280]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="72" pageNumber="73">CAS</collectionCode>
.
<emphasis id="EFC4EA9EB84BF30DFE6DFB11692BEE78" bold="true" box="[485,614,1255,1280]" pageId="72" pageNumber="73">Marshalls:</emphasis>
Kwajalein:
<collectionCode id="BBA1AE49B84BF30DFD67FB1E680DEF87" LSID="urn:lsid:biocol.org:col:34697" box="[751,832,1256,1279]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="72" pageNumber="73">BPBM</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B84BF30DFF4FFAFA6EDCEEAF" pageId="72" pageNumber="73" type="diagnosis">
<paragraph id="DD0F368CB84BF30DFF4FFAFA6EDCEEAF" blockId="72.[151,1437,967,2029]" pageId="72" pageNumber="73">
<emphasis id="EFC4EA9EB84BF30DFF4FFAFA6A0CEE5D" bold="true" box="[199,321,1292,1317]" pageId="72" pageNumber="73">Diagnosis.</emphasis>
<taxonomicName id="1AB04D0FB84BF30DFEC1FAFB6951EE5D" box="[329,540,1292,1317]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="72" pageNumber="73" phylum="Chordata" rank="species" species="preclarum">
<emphasis id="EFC4EA9EB84BF30DFEC1FAFB6951EE5D" box="[329,540,1292,1317]" italics="true" pageId="72" pageNumber="73">Trimma preclarum</emphasis>
</taxonomicName>
is characterized by the presence of scales in the predorsal midline, second and third spines of the first dorsal fin usually elongated, moderate interorbital and postorbital trenches, no opercular scales, and scales pockets outlined with pigment, especially across the dorsal half of the body. The fifth pelvic fin ray is branched sequentially once or unbranched depending on the locality. When alive
<taxonomicName id="1AB04D0FB84BF30DFB1FFA8F6E65EEE8" box="[1175,1320,1399,1424]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="72" pageNumber="73" phylum="Chordata" rank="species" species="preclarum">
<emphasis id="EFC4EA9EB84BF30DFB1FFA8F6E65EEE8" box="[1175,1320,1399,1424]" italics="true" pageId="72" pageNumber="73">T. preclarum</emphasis>
</taxonomicName>
has three yellow stripes on a grey background which extend along the body, two stripes in the dorsal and anal fins (one dark basal stripe and a more distal yellow one), and a red or cerise iris with four large, irregularly-spaced yellow spots.
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B84BF30CFF4FFA1269E6EEE1" lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73" type="description">
<paragraph id="DD0F368CB84BF30CFF4FFA1268D6EAD0" blockId="72.[151,1437,967,2029]" lastBlockId="73.[151,1437,151,602]" lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73">
<emphasis id="EFC4EA9EB84BF30DFF4FFA126A14EE85" bold="true" box="[199,345,1508,1533]" pageId="72" pageNumber="73">Description.</emphasis>
The description is based on
<specimenCount id="CBB6FD05B84BF30DFD2FFA12680FEE85" box="[679,834,1508,1533]" pageId="72" pageNumber="73" type="generic">10 specimens</specimenCount>
from the
<collectingCountry id="A5A7761CB84BF30DFC3FFA126F50EE85" box="[951,1053,1508,1533]" name="Australia" pageId="72" pageNumber="73">Australia</collectingCountry>
for comparison with the original description. Dorsal fins VI
<date id="A90E104CB84BF30DFE5CF9FE6AB8ED58" box="[468,501,1544,1568]" pageId="72" pageNumber="73">I 9</date>
(one specimen deformed, lacking sixth dorsal spine and anal spine), second and sometimes third spine longest, with second spine reaching between spine and third ray of second dorsal fin, all rays branched except posterior element of last fin-ray, and sometimes first ray, and once first and seventh to ninth rays; anal fin
<date id="A90E104CB84BF30DFF73F9846A74EDF2" box="[251,313,1650,1674]" pageId="72" pageNumber="73">I 89</date>
(mean = 8.7); pectoral fin 1719 (mean = 18.0), with 36 (mean = 4.5) and 48 (mean = 6.1) unbranched dorsal and ventral unbranched rays respectively, reaching posteriorly to vertical line with first few elements of anal fin; pelvic fin
<date id="A90E104CB84BF30DFE7BF94F695EEDA9" box="[499,531,1720,1745]" pageId="72" pageNumber="73">I 5</date>
, no fraenum, basal membrane vestigial, with small membrane from inner base of fifth ray that does not reach midline, first four rays with one sequential branch, fifth ray unbranched and 4856% (mean = 51.6) length of fourth, fourth ray reaching posteriorly to a vertical line with second to fifth element of anal fin. Lateral scales 2223 (mean = 22.7), transverse scales 68, (mean = 6.8); midline predorsal scales 78 (mean = 7.9), pectoral-fin base with 23 vertical rows of cycloid scales, with
<quantity id="1A489B69B84BF30DFC19F8B0689AEC27" box="[913,983,1862,1887]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.27" metricValueMin="0.762" pageId="72" pageNumber="73" unit="in" value="4.0" valueMax="5.0" valueMin="3.0">35 in</quantity>
outer row,
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middle row and
<quantity id="1A489B69B84BF30DFADEF8B16ED1EC27" box="[1366,1436,1862,1887]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="7.62" metricValueMin="5.08" pageId="72" pageNumber="73" unit="in" value="2.5" valueMax="3.0" valueMin="2.0">23 in</quantity>
inner row (if present); 57 (mean = 5.4) cycloid scales in midline anterior to pelvic fin; scales on head extend anteriorly to above vertical with posterior half of iris; cheek and opercle scaleless. Gill opening extending anteroventrally to below mid to posterior margin of pupil. Upper jaw with outer row of widely spaced enlarged canines extending to about bend of jaw, with few irregular inner rows of small conical teeth that grade to single row posteriorly. Lower jaw with outer row of enlarged widely spaced canines across front of jaw, one or two middle rows of small closely spaced canines, and inner row of widely spaced medium-sized canines. Inner rows of teeth on lower jaw extend around whole jaw as far as coronoid process, where they may be reduced to single row. Tongue round, or rounded with central tip. Gill rakers on first arch 34 + 14 = 1718 (mean = 17.9). Anterior nares short tube adjacent to upper lip, posterior nares pore with raised rim, both nares on raised oval sac confined to anterior half of snout. Bony interorbital 1/3 to 1/2 pupil width, with moderately developed interorbital and postorbital trenches (both wider than deep), but with postorbital trench poorly developed (especially posterolaterally) in smaller specimens. Abdominal/caudal vertebral transition of
<typeStatus id="020B882EB84AF30CFCCAFE676837EAD1" box="[834,890,401,425]" pageId="73" pageNumber="74">Type</typeStatus>
B.
</paragraph>
<paragraph id="DD0F368CB84AF30CFF4FFE456FF3E922" blockId="73.[151,1437,151,602]" pageId="73" pageNumber="74">
<emphasis id="EFC4EA9EB84AF30CFF4FFE45691BEAB4" box="[199,598,435,460]" italics="true" pageId="73" pageNumber="74">Colour Pattern. Freshly collected.</emphasis>
Helen Larson (NTM) provided the following notes made from freshly collected specimens from
<collectingCountry id="A5A7761CB84AF30CFE34FE21696EEA88" box="[444,547,471,496]" name="Ashmore and Cartier Islands" pageId="73" pageNumber="74">Ashmore</collectingCountry>
Reef (AMS I.33717-083): “Dusky grey with pink heads. Three yellow lines on body converge on caudal base. Bluish lines on bases of second dorsal and anal”. This description essentially agrees with that provided in the original description (
<figureCitation id="458B2A09B84AF30CFD37FDEB685AE94E" box="[703,791,541,566]" captionStart="FIGURE 41" captionStartId="73.[151,250,1030,1052]" captionTargetBox="[180,1385,678,978]" captionTargetId="figure@73.[151,1435,654,1009]" captionTargetPageId="73" captionText="FIGURE 41. Trimma preclarum, 21.3 mm SL female, ROM 83037, Merir Id, Palau. Photo by R. Winterbottom." httpUri="https://zenodo.org/record/236108/files/figure.png" pageId="73" pageNumber="74">Fig. 41</figureCitation>
; Pl. 3 C), and rules out the possibility that Australian specimens are
<taxonomicName id="1AB04D0FB84AF30CFEB4FDB4695FE922" box="[316,530,577,602]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="73" pageNumber="74" phylum="Chordata" rank="species" species="anthrenum">
<emphasis id="EFC4EA9EB84AF30CFEB4FDB4695FE922" box="[316,530,577,602]" italics="true" pageId="73" pageNumber="74">Trimma anthrenum</emphasis>
</taxonomicName>
(these two species are difficult to tell apart when preserved).
</paragraph>
<caption id="89CF6604B84AF30CFF1FFBF06E46EF64" box="[151,1291,1030,1052]" httpUri="https://zenodo.org/record/236108/files/figure.png" pageId="73" pageNumber="74" targetBox="[180,1385,678,978]" targetPageId="73">
<paragraph id="DD0F368CB84AF30CFF1FFBF06E46EF64" blockId="73.[151,1291,1030,1052]" box="[151,1291,1030,1052]" pageId="73" pageNumber="74">
<emphasis id="EFC4EA9EB84AF30CFF1FFBF06A52EF64" bold="true" box="[151,287,1030,1052]" pageId="73" pageNumber="74">FIGURE 41.</emphasis>
<taxonomicName id="1AB04D0FB84AF30CFEADFBF16AA9EF64" box="[293,484,1030,1052]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="73" pageNumber="74" phylum="Chordata" rank="species" species="preclarum">
<emphasis id="EFC4EA9EB84AF30CFEADFBF16AA9EF64" box="[293,484,1030,1052]" italics="true" pageId="73" pageNumber="74">Trimma preclarum</emphasis>
</taxonomicName>
, 21.3 mm SL female, ROM 83037, Merir Id, Palau. Photo by R. Winterbottom.
</paragraph>
</caption>
<paragraph id="DD0F368CB84AF30CFF4FFBB569E6EEE1" blockId="73.[151,1437,1091,2034]" pageId="73" pageNumber="74">
<emphasis id="EFC4EA9EB84AF30CFF4FFBB56A7AEF24" box="[199,311,1091,1116]" italics="true" pageId="73" pageNumber="74">Preserved</emphasis>
. A
<quantity id="1A489B69B84AF30CFEECFBB26A84EF23" box="[356,457,1092,1116]" metricMagnitude="-2" metricUnit="m" metricValue="2.28" pageId="73" pageNumber="74" unit="mm" value="22.8">22.8 mm</quantity>
SL female with yellow-brown body, scale pockets strongly outlined with dark brown chromatophores (pigmentation about twice width of dorsal spines); small brown chromatophores on posterior cheek, snout, opercle and pectoral base; body with larger, pale brown chromatophores mostly below scale pockets and more concentrated over abdomen and above anal fin, but no discrete, wedge-shaped pattern as in original description of Palauan specimens; one-third pupil-diameter dark stripe of chromatophores at bases of second dorsal and anal fins, with few scattered chromatophores in interradial membranes. Female (
<quantity id="1A489B69B84AF30CFBFBFB056F96EE73" box="[1139,1243,1267,1292]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="73" pageNumber="74" unit="mm" value="16.3">16.3 mm</quantity>
SL) similar, but whole cheek pigmented, melanophores on body scales, and subdermal chromatophores virtually confined to abdomen and lower body above anal fin. This specimen similar to 17.0 mm SL female, but melanophores widely distributed over body, especially outlining scale pockets, and subdermal chromatophores present over whole body (although still more concentrated on abdomen).
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B84AF30CFF4FFA556820EEA7" pageId="73" pageNumber="74" type="etymology">
<paragraph id="DD0F368CB84AF30CFF4FFA556820EEA7" blockId="73.[151,1437,1091,2034]" pageId="73" pageNumber="74">
<emphasis id="EFC4EA9EB84AF30CFF4FFA556A03EEC4" bold="true" box="[199,334,1443,1468]" pageId="73" pageNumber="74">Etymology.</emphasis>
From the Latin word “preclarus”, meaning splendid or very beautiful, in reference to the wonderful colouration when alive, especially the multi-hued iris.
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B84AF30CFF4FFA1C6E3EEC8A" pageId="73" pageNumber="74" type="distribution">
<paragraph id="DD0F368CB84AF30CFF4FFA1C69F9ED30" blockId="73.[151,1437,1091,2034]" pageId="73" pageNumber="74">
<emphasis id="EFC4EA9EB84AF30CFF4FFA1C6A2DED7B" bold="true" box="[199,352,1514,1539]" pageId="73" pageNumber="74">Distribution.</emphasis>
Found though the western Pacific from
<collectingCountry id="A5A7761CB84AF30CFCA5FA1C6826ED7B" box="[813,875,1514,1539]" name="Palau" pageId="73" pageNumber="74">Palau</collectingCountry>
and
<collectingCountry id="A5A7761CB84AF30CFC2CFA1C68A5ED7A" box="[932,1000,1514,1538]" name="Guam" pageId="73" pageNumber="74">Guam</collectingCountry>
in the north to
<collectingCountry id="A5A7761CB84AF30CFB13FA1C6FAAED7A" box="[1179,1255,1514,1538]" name="Samoa" pageId="73" pageNumber="74">Samoa</collectingCountry>
and west to the eastern coast of
<collectingCountry id="A5A7761CB84AF30CFED8F9FB6AFAED5E" box="[336,439,1549,1574]" name="Australia" pageId="73" pageNumber="74">Australia</collectingCountry>
(
<figureCitation id="458B2A09B84AF30CFE40F9FB6953ED5E" box="[456,542,1549,1574]" captionStart="FIGURE 40" captionStartId="71.[151,250,1899,1921]" captionTargetBox="[161,1425,1049,1867]" captionTargetId="figure@71.[151,1436,1043,1877]" captionTargetPageId="71" captionText="FIGURE 40. Distribution map for Trimma pentherum (open triangles) and T. preclarum (solid squares). A solid star indicates localities at which both species occur." httpUri="https://zenodo.org/record/236107/files/figure.png" pageId="73" pageNumber="74">Fig. 40</figureCitation>
). In
<collectingCountry id="A5A7761CB84AF30CFDDBF9FB69F4ED5E" box="[595,697,1549,1574]" name="Australia" pageId="73" pageNumber="74">Australia</collectingCountry>
it is known from outer barrier reef islands in depths of
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and reefs in the Coral Sea in depths of
<quantity id="1A489B69B84AF30CFDC1F9C669E2ED30" box="[585,687,1584,1609]" metricMagnitude="1" metricUnit="m" metricValue="1.85" metricValueMax="2.5" metricValueMin="1.2" pageId="73" pageNumber="74" unit="m" value="18.5" valueMax="25.0" valueMin="12.0">
12
<specimenCount id="CBB6FD05B84AF30CFDFBF9C669E2ED30" box="[627,687,1584,1609]" pageId="73" pageNumber="74" type="generic">25 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="DD0F368CB84AF30CFF4FF9A56E3EEC8A" blockId="73.[151,1437,1091,2034]" pageId="73" pageNumber="74">
<emphasis id="EFC4EA9EB84AF30CFF4FF9A56A21ED14" bold="true" box="[199,364,1619,1644]" pageId="73" pageNumber="74">Comparisons.</emphasis>
This species is most easily confused, when preserved, with
<taxonomicName id="1AB04D0FB84AF30CFBF4F9A36E10ED14" box="[1148,1373,1619,1644]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="73" pageNumber="74" phylum="Chordata" rank="species" species="anthrenum">
<emphasis id="EFC4EA9EB84AF30CFBF4F9A36E10ED14" box="[1148,1373,1619,1644]" italics="true" pageId="73" pageNumber="74">Trimma anthrenum</emphasis>
</taxonomicName>
(see
<bibRefCitation id="B9214B7DB84AF30CFF1FF9816A3AEDE8" author="Winterbottom" box="[151,375,1655,1680]" pageId="73" pageNumber="74" refString="Winterbottom, R. (2006) Two new species of the gobiid fish Trimma from the coral reefs of the western Pacific Ocean (Pisces; Perciformes; Gobioidei). Zootaxa, 1331, 55 - 68." type="journal article" year="2006">Winterbottom 2006</bibRefCitation>
). The specimens from
<collectingCountry id="A5A7761CB84AF30CFDFCF9816996EDE8" box="[628,731,1655,1680]" name="Australia" pageId="73" pageNumber="74">Australia</collectingCountry>
differ from the original description in that none of them has an elongated second dorsal spine, which is often, but not always, present in the Fijian and Palauan
<typeStatus id="020B882EB84AF30CFB45F96D6FB0EDCB" box="[1229,1277,1691,1715]" pageId="73" pageNumber="74">type</typeStatus>
material. (We note that only 3 of
<specimenCount id="CBB6FD05B84AF30CFEE7F9496945EDAF" box="[367,520,1726,1751]" pageId="73" pageNumber="74" type="generic">44 specimens</specimenCount>
identified as this species from the
<collectingCountry id="A5A7761CB84AF30CFC1AF94868A4EDAF" box="[914,1001,1726,1751]" name="Kiribati" pageId="73" pageNumber="74">Kiribati</collectingCountry>
had the second dorsal spine elongated beyond the base of the fifth dorsal fin ray). In addition, the wedge of chromatophores above the anal-fin origin is not as apparent in these specimens as the illustration provided in the original description (
<bibRefCitation id="B9214B7DB84AF30CFB0CF8F36BF7EC39" author="Winterbottom" pageId="73" pageNumber="74" refString="Winterbottom, R. (2006) Two new species of the gobiid fish Trimma from the coral reefs of the western Pacific Ocean (Pisces; Perciformes; Gobioidei). Zootaxa, 1331, 55 - 68." type="journal article" year="2006">Winterbottom, 2006, fig. 4B</bibRefCitation>
), although it is still discernible. The well developed outlining of the scale pockets and the relatively poorly developed posterior interorbital trench are, however, consistent with
<taxonomicName id="1AB04D0FB84AF30CFC1EF8BB6F6BEC1D" box="[918,1062,1868,1893]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="73" pageNumber="74" phylum="Chordata" rank="species" species="preclarum">
<emphasis id="EFC4EA9EB84AF30CFC1EF8BB6F6BEC1D" box="[918,1062,1868,1893]" italics="true" pageId="73" pageNumber="74">T. preclarum</emphasis>
</taxonomicName>
to the exclusion of
<taxonomicName id="1AB04D0FB84AF30CFA89F8BB6ED9EC1D" box="[1281,1428,1868,1893]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="73" pageNumber="74" phylum="Chordata" rank="species" species="anthrenum">
<emphasis id="EFC4EA9EB84AF30CFA89F8BB6ED9EC1D" box="[1281,1428,1868,1893]" italics="true" pageId="73" pageNumber="74">T. anthrenum</emphasis>
</taxonomicName>
, as is the colour pattern of freshly collected specimens. Although it is certainly possible that the Australian and, perhaps, the
<collectingCountry id="A5A7761CB84AF30CFEADF8656A31ECD4" box="[293,380,1939,1964]" name="Kiribati" pageId="73" pageNumber="74">Kiribati</collectingCountry>
material represent an undescribed species allied to
<taxonomicName id="1AB04D0FB84AF30CFC36F8626F00ECD4" box="[958,1101,1939,1964]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="73" pageNumber="74" phylum="Chordata" rank="species" species="preclarum">
<emphasis id="EFC4EA9EB84AF30CFC36F8626F00ECD4" box="[958,1101,1939,1964]" italics="true" pageId="73" pageNumber="74">T. preclarum</emphasis>
</taxonomicName>
, we prefer to await collection of fresh material and subsequent genetic analyses to resolve this issue. Two tissue samples of
<taxonomicName id="1AB04D0FB84AF30CFB42F8416E14ECB7" box="[1226,1369,1974,1999]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="73" pageNumber="74" phylum="Chordata" rank="species" species="preclarum">
<emphasis id="EFC4EA9EB84AF30CFB42F8416E14ECB7" box="[1226,1369,1974,1999]" italics="true" pageId="73" pageNumber="74">T. preclarum</emphasis>
</taxonomicName>
, both from
<collectingCountry id="A5A7761CB84AF30CFF5CF82F6A58EC8A" box="[212,277,2009,2034]" name="Palau" pageId="73" pageNumber="74">Palau</collectingCountry>
, were available for genetic analysis, and exhibited no variation in CO I (
<bibRefCitation id="B9214B7DB84AF30CFBC8F82F6E25EC8A" author="Winterbottom" box="[1088,1384,2009,2034]" pageId="73" pageNumber="74" refString="Winterbottom, R., Hanner, R. H., Burridge, M. &amp; Zur, M. (2014) A cornucopia of cryptic species-a DNA barcode analysis of the gobiid fish genus Trimma (Percomorpha, Gobiiformes). ZooKeys, 381, 79 - 111. http: // dx. doi. org / 10.3897 / zookeys. 381.6445" type="journal article" year="2014">
Winterbottom
<emphasis id="EFC4EA9EB84AF30CFB6DF82D6E50EC8A" box="[1253,1309,2009,2034]" italics="true" pageId="73" pageNumber="74">et al.</emphasis>
, 2014
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>