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<document id="6E3362E7133A1F9B0059619943F16F02" ID-DOI="10.11646/zootaxa.3934.1.1" ID-GBIF-Dataset="0abffe3e-a2c6-4623-b7c6-c8b26cee4a2b" ID-ISSN="1175-5326" ID-Zenodo-Dep="236066" ID-ZooBank="11C2A2CB-30B3-4694-B379-AE9D47332F0C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461012229557" checkinUser="plazi" docAuthor="Winterbottom, Richard &amp; Hoese, Douglass F." docDate="2015" docId="5519879AB849F308FF1FFF6E68DEEA64" docLanguage="en" docName="zt03934p102.pdf" docOrigin="Zootaxa 3934 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Trimma readerae Winterbottom &amp; Hoese, 2015, sp. nov." docType="treatment" docUuid="2069462A-E421-4BFA-8E28-681D3729D090" docUuidSource="ZooBank" docVersion="11" lastPageNumber="78" masterDocId="A920FFE2B803F345FF88FFF66B4DEB78" masterDocTitle="A revision of the Australian species of Trimma (Actinopterygii, Gobiidae), with descriptions of six new species and redescriptions of twenty-three valid species" masterLastPageNumber="102" masterPageNumber="1" pageNumber="75" updateTime="1698567702668" updateUser="plazi">
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<mods:title id="7478594B487E98EBDE8C58BFE94875F0">A revision of the Australian species of Trimma (Actinopterygii, Gobiidae), with descriptions of six new species and redescriptions of twenty-three valid species</mods:title>
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<mods:namePart id="184DB4640D43B5F624FB3A345ED77FFE">Winterbottom, Richard</mods:namePart>
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<mods:namePart id="3542CA07206F61CF43DCC51BB9B7CC34">Hoese, Douglass F.</mods:namePart>
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<mods:date id="51592A52D4E5209685C4A3BBDA761A42">2015</mods:date>
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<treatment id="5519879AB849F308FF1FFF6E68DEEA64" ID-DOI="http://doi.org/10.5281/zenodo.5621538" ID-GBIF-Taxon="119600466" ID-Zenodo-Dep="5621538" LSID="urn:lsid:zoobank.org:act:2069462A-E421-4BFA-8E28-681D3729D090" httpUri="http://treatment.plazi.org/id/5519879AB849F308FF1FFF6E68DEEA64" lastPageId="77" lastPageNumber="78" pageId="74" pageNumber="75">
<subSubSection id="95AA6507B849F30FFF1FFF6E6ADAEB8E" pageId="74" pageNumber="75" type="nomenclature">
<paragraph id="DD0F368CB849F30FFF1FFF6E6A83EBCA" blockId="74.[151,462,152,247]" box="[151,462,152,178]" pageId="74" pageNumber="75">
<heading id="864781E0B849F30FFF1FFF6E6A83EBCA" bold="true" box="[151,462,152,178]" fontSize="11" level="1" pageId="74" pageNumber="75" reason="1">
<emphasis id="EFC4EA9EB849F30FFF1FFF6E6A83EBCA" bold="true" box="[151,462,152,178]" pageId="74" pageNumber="75">
<taxonomicName id="1AB04D0FB849F30FFF1FFF6E6A24EBCA" ID-CoL="7CTHR" box="[151,361,152,178]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="74" pageNumber="75" phylum="Chordata" rank="species" species="readerae" status="sp. nov.">
<emphasis id="EFC4EA9EB849F30FFF1FFF6E6A24EBCA" bold="true" box="[151,361,152,178]" italics="true" pageId="74" pageNumber="75">Trimma readerae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4F757E5B849F30FFEF8FF6E6A83EBCA" box="[368,462,152,178]" pageId="74" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="DD0F368CB849F30FFF1FFF4C6ADAEB8E" blockId="74.[151,462,152,247]" pageId="74" pageNumber="75">
Red-spotted Pygmygoby
<figureCitation id="458B2A09B849F30FFF1FFF286A53EB8F" box="[151,286,222,247]" captionStart-0="FIGURE 42" captionStart-1="FIGURE 43" captionStartId-0="76.[151,250,656,678]" captionStartId-1="77.[151,250,1207,1229]" captionTargetBox-0="[152,1435,193,634]" captionTargetBox-1="[158,1425,351,1171]" captionTargetId-0="figure@76.[152,1435,193,635]" captionTargetId-1="figure@77.[151,1436,339,1185]" captionTargetPageId-0="76" captionTargetPageId-1="77" captionText-0="FIGURE 42. Trimma readerae, 22.5 mm SL holotype, AMS I. 20784 - 020, Yonge Reef, Queensland. Photo by D. F. Hoese." captionText-1="FIGURE 43. Distribution map for Trimma readerae (solid diamonds)." httpUri-0="https://zenodo.org/record/236110/files/figure.png" httpUri-1="https://zenodo.org/record/236111/files/figure.png" pageId="74" pageNumber="75">Figs. 4243</figureCitation>
; Pl. 2 D, E
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B849F30FFF1FFEDE69D6EA02" pageId="74" pageNumber="75" type="reference_group">
<paragraph id="DD0F368CB849F30FFF1FFEDE69BAEA23" blockId="74.[151,1436,294,378]" pageId="74" pageNumber="75">
<treatmentCitationGroup id="FDA011A2B849F30FFF1FFEDE69BAEA23" pageId="74" pageNumber="75">
<taxonomicName id="1AB04D0FB849F30FFF1FFEDE6A02EA44" ID-CoL="58V73" box="[151,335,294,317]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="74" pageNumber="75" phylum="Chordata" rank="species" species="okinawae">
<emphasis id="EFC4EA9EB849F30FFF1FFEDE6A02EA44" box="[151,335,294,317]" italics="true" pageId="74" pageNumber="75">Trimma okinawae</emphasis>
</taxonomicName>
(
<emphasis id="EFC4EA9EB849F30FFEE9FEDE6ACAEA45" box="[353,391,296,317]" italics="true" pageId="74" pageNumber="75">non</emphasis>
<bibRefCitation id="B9214B7DB849F30FFE18FED0696CEA44" author="Aoyagi" box="[400,545,294,316]" pageId="74" pageNumber="75" refString="Aoyagi, H. (1949) Studies on the coral reef fishes of the Riu-Kiu Islands. V. The Zoological Magazine, The Zoological Society of Japan (Tokyo), 58 (9), 171 - 173." type="journal article" year="1949">Aoyagi, 1949</bibRefCitation>
): Randall, 1990:408 (Great Barrier Reef, but photo is of BPBM 8012, from Enewetak);
<treatmentCitation id="5C11109DB849F30FFF4FFEB06A90EA23" author="Hoese" box="[199,477,325,347]" page="1690" pageId="74" pageNumber="75" year="2006">
<bibRefCitation id="B9214B7DB849F30FFF4FFEB06AE5EA23" author="Hoese" box="[199,424,325,347]" pageId="74" pageNumber="75" refString="Hoese, D. F. &amp; Larson, H. K. (2006) Family Gobiidae. In: Beesley, P. L. &amp; Wells, A. (Eds.), Zoological Catalogue of Australia. Vol. 35. Australia. ABRS &amp; CSIRO Publishing Parts 1 - 3, pp. 1612 - 1697." type="book chapter" year="2006">Hoese &amp; Larson, 2006</bibRefCitation>
:1690
</treatmentCitation>
(in part Great Barrier Reef).
</treatmentCitationGroup>
</paragraph>
<paragraph id="DD0F368CB849F30FFF1FFE9269D6EA02" blockId="74.[151,1436,294,378]" box="[151,667,356,378]" pageId="74" pageNumber="75">
<treatmentCitationGroup id="FDA011A2B849F30FFF1FFE9269D6EA02" box="[151,667,356,378]" pageId="74" pageNumber="75">
<taxonomicName id="1AB04D0FB849F30FFF1FFE926BA5EA02" ID-CoL="827C" box="[151,232,356,378]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="74" pageNumber="75" phylum="Chordata" rank="genus">Trimma</taxonomicName>
sp.:
<treatmentCitation id="5C11109DB849F30FFE90FE926909EA02" author="Hayashi" box="[280,580,356,378]" page="44" pageId="74" pageNumber="75" year="2003">
<bibRefCitation id="B9214B7DB849F30FFE90FE92696BEA02" author="Hayashi" box="[280,550,356,378]" pageId="74" pageNumber="75" refString="Hayashi, M. &amp; Shiratori, T. (2003) Gobies of Japanese Waters. Hankyu Books, Osaka, 224 pp. [in Japanese]" type="book" year="2003">Hayashi &amp; Shiratori, 2003</bibRefCitation>
:44
</treatmentCitation>
(
<collectingCountry id="A5A7761CB849F30FFDD9FE9269DDEA02" box="[593,656,356,378]" name="Japan" pageId="74" pageNumber="75">Japan</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B849F30FFF1FFE506E4DEED7" pageId="74" pageNumber="75" type="materials_examined">
<paragraph id="DD0F368CB849F30FFF1FFE50697BEA9B" blockId="74.[151,1437,422,2031]" pageId="74" pageNumber="75">
<emphasis id="EFC4EA9EB849F30FFF1FFE506AC9EAC7" bold="true" box="[151,388,422,447]" pageId="74" pageNumber="75">
Material.
<typeStatus id="020B882EB849F30FFE98FE506A32EAC7" box="[272,383,422,447]" pageId="74" pageNumber="75" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="BBA1AE49B849F30FFE06FE516A86EAC7" box="[398,459,423,447]" pageId="74" pageNumber="75">AMS</collectionCode>
I.20784-020,
<quantity id="1A489B69B849F30FFDF8FE50699AEAC6" box="[624,727,422,447]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" pageId="74" pageNumber="75" unit="mm" value="22.5">22.5 mm</quantity>
SL female, inner side of Yonge Reef, Queensland, (
<geoCoordinate id="B884504BB849F30FFAB5FE516ED5EAC7" box="[1341,1432,423,447]" direction="south" orientation="latitude" pageId="74" pageNumber="75" precision="925" value="-14.616667">14°37'S</geoCoordinate>
,
<geoCoordinate id="B884504BB849F30FFF1FFE3C6A4FEA9B" box="[151,258,458,483]" direction="east" orientation="longitude" pageId="74" pageNumber="75" precision="925" value="145.63333">145°38'E</geoCoordinate>
),
<date id="A90E104CB849F30FFE9DFE3D6A2CEA9A" box="[277,353,459,483]" pageId="74" pageNumber="75">1 Dec.</date>
, 1978,
<collectionCode id="BBA1AE49B849F30FFE3BFE3D6ABDEA9B" box="[435,496,459,483]" pageId="74" pageNumber="75">AMS</collectionCode>
party.
</paragraph>
<paragraph id="DD0F368CB849F30FFF4FFE186F28E85F" blockId="74.[151,1437,422,2031]" pageId="74" pageNumber="75">
<emphasis id="EFC4EA9EB849F30FFF4FFE186A0BE97F" bold="true" box="[199,326,494,519]" pageId="74" pageNumber="75">
<typeStatus id="020B882EB849F30FFF4FFE186A0CE97F" box="[199,321,494,519]" pageId="74" pageNumber="75" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Outer Barrier Reef:
<collectionCode id="BBA1AE49B849F30FFDA4FE196924E97F" box="[556,617,495,519]" pageId="74" pageNumber="75">AMS</collectionCode>
I.20774-113, 45(1122), off Cape Melville (
<geoCoordinate id="B884504BB849F30FFBD4FE186FF8E97F" box="[1116,1205,494,519]" direction="south" orientation="latitude" pageId="74" pageNumber="75" precision="925" value="-13.933333">13°56'S</geoCoordinate>
<geoCoordinate id="B884504BB849F30FFB34FE186E6BE97F" box="[1212,1318,494,519]" direction="east" orientation="longitude" pageId="74" pageNumber="75" precision="925" value="144.6">144°36'E</geoCoordinate>
),
<quantity id="1A489B69B849F30FFAB2FE196ECEE97E" box="[1338,1411,495,519]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="8.0" metricValueMin="0.0" pageId="74" pageNumber="75" unit="m" value="4.0" valueMax="8.0" valueMin="0.0">08 m</quantity>
,
<date id="A90E104CB849F30FFA06FE196A47E953" pageId="74" pageNumber="75" value="1979-02-09">9 Feb. 1979</date>
,
<collectionCode id="BBA1AE49B849F30FFE9DFDE56A1FE953" box="[277,338,531,555]" pageId="74" pageNumber="75">AMS</collectionCode>
party. Yonge Reef:
<collectionCode id="BBA1AE49B849F30FFDBBFDE5693DE953" box="[563,624,531,555]" pageId="74" pageNumber="75">AMS</collectionCode>
I.19456-076, 18(1121) inner side of reef (
<geoCoordinate id="B884504BB849F30FFBD2FDE46FF8E953" box="[1114,1205,530,555]" direction="south" orientation="latitude" pageId="74" pageNumber="75" precision="925" value="-14.583333">14°35'S</geoCoordinate>
,
<geoCoordinate id="B884504BB849F30FFB49FDE46E61E953" box="[1217,1324,530,555]" direction="east" orientation="longitude" pageId="74" pageNumber="75" precision="925" value="145.61667">145°37'E</geoCoordinate>
),
<quantity id="1A489B69B849F30FFAC8FDE46EDAE952" box="[1344,1431,530,555]" metricMagnitude="1" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="74" pageNumber="75" unit="m" value="10.0" valueMax="15.0" valueMin="5.0">515 m</quantity>
,
<date id="A90E104CB849F30FFF1FFDC16A71E937" box="[151,316,566,591]" pageId="74" pageNumber="75" value="1975-11-12">12 Nov. 1975</date>
,
<collectionCode id="BBA1AE49B849F30FFEC5FDC16AC7E937" box="[333,394,567,591]" pageId="74" pageNumber="75">AMS</collectionCode>
Party;
<collectionCode id="BBA1AE49B849F30FFE6DFDC1696FE937" box="[485,546,567,591]" pageId="74" pageNumber="75">AMS</collectionCode>
I.20784-087, 13(1622) taken with
<typeStatus id="020B882EB849F30FFC5BFDC06F74E937" box="[979,1081,566,591]" pageId="74" pageNumber="75" type="holotype">holotype</typeStatus>
;
<collectionCode id="BBA1AE49B849F30FFBC1FDC16FCBE937" box="[1097,1158,567,591]" pageId="74" pageNumber="75">AMS</collectionCode>
I.25065-011, 5(1320), channel between Yonge and Carter Reefs (
<geoCoordinate id="B884504BB849F30FFDFAFDAC6980E90B" box="[626,717,602,627]" direction="south" orientation="latitude" pageId="74" pageNumber="75" precision="925" value="-14.583333">14°35'S</geoCoordinate>
,
<geoCoordinate id="B884504BB849F30FFD50FDAC680FE90B" box="[728,834,602,627]" direction="east" orientation="longitude" pageId="74" pageNumber="75" precision="925" value="145.6">145°36'E</geoCoordinate>
),
<quantity id="1A489B69B849F30FFCDEFDAD68CEE90A" box="[854,899,603,627]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="74" pageNumber="75" unit="m" value="9.0">9 m</quantity>
,
<date id="A90E104CB849F30FFC06FDAD6F5BE90B" box="[910,1046,602,627]" pageId="74" pageNumber="75" value="1977-02-07">7 Feb. 1977</date>
, H. Larson and B. Goldman;
<collectionCode id="BBA1AE49B849F30FFAD5FDAD6ED0E90B" LSID="urn:lsid:biocol.org:col:34778" box="[1373,1437,603,627]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="74" pageNumber="75">ROM</collectionCode>
39329, 2(1923) inner side of reef (
<geoCoordinate id="B884504BB849F30FFDA7FD8869C4E9EF" box="[559,649,638,663]" direction="south" orientation="latitude" pageId="74" pageNumber="75" precision="925" value="-14.583333">14°35'S</geoCoordinate>
,
<geoCoordinate id="B884504BB849F30FFD1EFD88684CE9EF" box="[662,769,638,663]" direction="east" orientation="longitude" pageId="74" pageNumber="75" precision="925" value="145.6">145°36'E</geoCoordinate>
),
<quantity id="1A489B69B849F30FFC9EFD896822E9EE" box="[790,879,639,663]" metricMagnitude="1" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="74" pageNumber="75" unit="m" value="10.5" valueMax="13.0" valueMin="8.0">813 m</quantity>
,
<date id="A90E104CB849F30FFCF4FD896F56E9EF" box="[892,1051,639,663]" pageId="74" pageNumber="75" value="1981-09-18">18 Sept. 1981</date>
, R. Winterbottom. Lizard Id: QM I.39131, 11(1721), North Point, (
<geoCoordinate id="B884504BB849F30FFD97FD556934E9C3" box="[543,633,675,699]" direction="south" orientation="latitude" pageId="74" pageNumber="75" precision="925" value="-14.666667">14°40'S</geoCoordinate>
,
<geoCoordinate id="B884504BB849F30FFD00FD5469BFE9C3" box="[648,754,674,699]" direction="east" orientation="longitude" pageId="74" pageNumber="75" precision="925" value="145.45">145°27'E</geoCoordinate>
),
<quantity id="1A489B69B849F30FFC80FD546819E9C2" box="[776,852,674,699]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="74" pageNumber="75" unit="m" value="5.5" valueMax="6.0" valueMin="5.0">56 m</quantity>
,
<date id="A90E104CB849F30FFCE9FD5568BAE9C3" box="[865,1015,674,699]" pageId="74" pageNumber="75" value="1975-01-30">30 Jan. 1975</date>
, D. Hoese
<emphasis id="EFC4EA9EB849F30FFBFFFD526FFCE9C3" box="[1143,1201,674,699]" italics="true" pageId="74" pageNumber="75">et al.</emphasis>
, Escape Reef:
<collectionCode id="BBA1AE49B849F30FFAD7FD556ED6E9C3" box="[1375,1435,675,699]" pageId="74" pageNumber="75">AMS</collectionCode>
I.22578-064, 14(1821), (
<geoCoordinate id="B884504BB849F30FFE49FD306951E9A7" box="[449,540,710,735]" direction="south" orientation="latitude" pageId="74" pageNumber="75" precision="925" value="-15.816667">15°49'S</geoCoordinate>
,
<geoCoordinate id="B884504BB849F30FFDA3FD3069D8E9A7" box="[555,661,710,735]" direction="east" orientation="longitude" pageId="74" pageNumber="75" precision="925" value="145.83333">145°50'E</geoCoordinate>
),
<quantity id="1A489B69B849F30FFD25FD30684AE9A6" box="[685,775,710,735]" metricMagnitude="0" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="74" pageNumber="75" unit="m" value="8.0" valueMax="10.0" valueMin="6.0">610 m</quantity>
,
<date id="A90E104CB849F30FFC9EFD3168E2E9A7" box="[790,943,711,735]" pageId="74" pageNumber="75" value="1981-10-28">28 Oct. 1981</date>
<collectionCode id="BBA1AE49B849F30FFC31FD3168BBE9A7" box="[953,1014,711,735]" pageId="74" pageNumber="75">AMS</collectionCode>
party;
<collectionCode id="BBA1AE49B849F30FFBC4FD316FC4E9A7" box="[1100,1161,711,735]" pageId="74" pageNumber="75">AMS</collectionCode>
I.22621-037, 9(1522), inner side of reef (
<geoCoordinate id="B884504BB849F30FFEE6FD1C6A84E87B" box="[366,457,746,771]" direction="south" orientation="latitude" pageId="74" pageNumber="75" precision="925" value="-15.816667">15°49'S</geoCoordinate>
,
<geoCoordinate id="B884504BB849F30FFE5FFD1C690CE87B" box="[471,577,746,771]" direction="east" orientation="longitude" pageId="74" pageNumber="75" precision="925" value="145.83333">145°50'E</geoCoordinate>
),
<quantity id="1A489B69B849F30FFDD0FD1C69EEE87A" box="[600,675,746,771]" metricMagnitude="0" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="74" pageNumber="75" unit="m" value="6.5" valueMax="8.0" valueMin="5.0">58 m</quantity>
,
<date id="A90E104CB849F30FFD39FD1D681DE87B" box="[689,848,747,771]" pageId="74" pageNumber="75" value="1980-11-27">27 Nov. 1980</date>
, W. and J. Starck. One Tree Id:
<collectionCode id="BBA1AE49B849F30FFB4EFD1D6E4EE87B" box="[1222,1283,747,771]" pageId="74" pageNumber="75">AMS</collectionCode>
I.15682-061, 8(2024) outside of lagoon (
<geoCoordinate id="B884504BB849F30FFE5FFCF9697CE85F" box="[471,561,783,807]" direction="south" orientation="latitude" pageId="74" pageNumber="75" precision="925" value="-23.5">23°30'S</geoCoordinate>
,
<geoCoordinate id="B884504BB849F30FFDB5FCF869EAE85F" box="[573,679,782,807]" direction="east" orientation="longitude" pageId="74" pageNumber="75" precision="925" value="152.08333">152°05'E</geoCoordinate>
),
<quantity id="1A489B69B849F30FFD34FCF969BAE85E" box="[700,759,783,807]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="74" pageNumber="75" unit="m" value="30.0">30 m</quantity>
,
<date id="A90E104CB849F30FFC8AFCF968D0E85F" box="[770,925,782,807]" pageId="74" pageNumber="75" value="1969-11-27">27 Nov. 1969</date>
, F.H. Talbot
<emphasis id="EFC4EA9EB849F30FFBA4FCF96F28E85F" box="[1068,1125,782,807]" italics="true" pageId="74" pageNumber="75">et al.</emphasis>
</paragraph>
<paragraph id="DD0F368CB849F30FFF4FFCC46F21EFCA" blockId="74.[151,1437,422,2031]" pageId="74" pageNumber="75">
<emphasis id="EFC4EA9EB849F30FFF4FFCC46906E833" bold="true" box="[199,587,818,843]" pageId="74" pageNumber="75">Non-type material. Queensland:</emphasis>
Tijou Reef:
<collectionCode id="BBA1AE49B849F30FFD63FCC5686AE833" box="[747,807,819,843]" pageId="74" pageNumber="75">AMS</collectionCode>
I.20956-021, 11(1522),
<quantity id="1A489B69B849F30FFBDDFCC56FFFE832" box="[1109,1202,819,843]" metricMagnitude="0" metricUnit="m" metricValue="7.5" metricValueMax="12.0" metricValueMin="3.0" pageId="74" pageNumber="75" unit="m" value="7.5" valueMax="12.0" valueMin="3.0">312 m</quantity>
;
<collectionCode id="BBA1AE49B849F30FFB4DFCC56E4CE833" box="[1221,1281,819,843]" pageId="74" pageNumber="75">AMS</collectionCode>
I.20779-149, 13(1222). Outer Barrier Reef off Cape Melville:
<collectionCode id="BBA1AE49B849F30FFD40FCA16849E817" box="[712,772,855,879]" pageId="74" pageNumber="75">AMS</collectionCode>
I.20774-084, (19),
<quantity id="1A489B69B849F30FFC57FCA16F67E816" box="[991,1066,855,879]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="8.0" metricValueMin="0.0" pageId="74" pageNumber="75" unit="m" value="4.0" valueMax="8.0" valueMin="0.0">08 m</quantity>
;
<collectionCode id="BBA1AE49B849F30FFBBFFCA16F3EE817" box="[1079,1139,855,879]" pageId="74" pageNumber="75">AMS</collectionCode>
I.20774-257, (11),
<quantity id="1A489B69B849F30FFAC6FCA16ED5E816" box="[1358,1432,855,879]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="8.0" metricValueMin="0.0" pageId="74" pageNumber="75" unit="m" value="4.0" valueMax="8.0" valueMin="0.0">08 m</quantity>
, Yonge Reef:
<collectionCode id="BBA1AE49B849F30FFEBBFC8D6A22E8EB" box="[307,367,891,915]" pageId="74" pageNumber="75">AMS</collectionCode>
I.18740-077, 12(720),
<quantity id="1A489B69B849F30FFD06FC8D69A7E8EA" box="[654,746,891,915]" metricMagnitude="1" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="74" pageNumber="75" unit="m" value="10.5" valueMax="12.0" valueMin="9.0">912 m</quantity>
;
<collectionCode id="BBA1AE49B849F30FFD74FC8D6875E8EB" box="[764,824,891,915]" pageId="74" pageNumber="75">AMS</collectionCode>
I.19454-092, 2(1718),
<quantity id="1A489B69B849F30FFBD0FC8D6FF9E8EA" box="[1112,1204,891,915]" metricMagnitude="0" metricUnit="m" metricValue="9.5" metricValueMax="18.0" metricValueMin="1.0" pageId="74" pageNumber="75" unit="m" value="9.5" valueMax="18.0" valueMin="1.0">118 m</quantity>
;
<collectionCode id="BBA1AE49B849F30FFB4EFC8D6E4FE8EB" box="[1222,1282,891,915]" pageId="74" pageNumber="75">AMS</collectionCode>
I.19472-091, 28(921),
<quantity id="1A489B69B849F30FFE81FC686A2DE8CE" box="[265,352,926,951]" metricMagnitude="1" metricUnit="m" metricValue="1.1" metricValueMax="1.5" metricValueMin="0.7" pageId="74" pageNumber="75" unit="m" value="11.0" valueMax="15.0" valueMin="7.0">
7
<specimenCount id="CBB6FD05B849F30FFEAEFC686A2DE8CE" box="[294,352,926,951]" pageId="74" pageNumber="75" type="generic">15 m</specimenCount>
</quantity>
. McGillivray Reef:
<collectionCode id="BBA1AE49B849F30FFDB3FC69693AE8CF" box="[571,631,927,951]" pageId="74" pageNumber="75">AMS</collectionCode>
I.19482-135, 23(925),
<quantity id="1A489B69B849F30FFC0CFC686891E8CE" box="[900,988,926,951]" metricMagnitude="1" metricUnit="m" metricValue="1.4" metricValueMax="2.5" metricValueMin="0.3" pageId="74" pageNumber="75" unit="m" value="14.0" valueMax="25.0" valueMin="3.0">
3
<specimenCount id="CBB6FD05B849F30FFC17FC686891E8CE" box="[927,988,926,951]" pageId="74" pageNumber="75" type="generic">25 m</specimenCount>
</quantity>
. Lizard Id:
<collectionCode id="BBA1AE49B849F30FFBD2FC696FDBE8CF" box="[1114,1174,927,951]" pageId="74" pageNumber="75">AMS</collectionCode>
I.19607-024, 2(1416),
<quantity id="1A489B69B849F30FFF1FFC346BAFE8A2" box="[151,226,962,987]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="74" pageNumber="75" unit="m" value="5.5" valueMax="6.0" valueMin="5.0">56 m</quantity>
;
<collectionCode id="BBA1AE49B849F30FFF79FC356A60E8A3" box="[241,301,963,987]" pageId="74" pageNumber="75">AMS</collectionCode>
I.21578-004, (22);
<collectionCode id="BBA1AE49B849F30FFD84FC356905E8A3" box="[524,584,963,987]" pageId="74" pageNumber="75">AMS</collectionCode>
I.21894-011, (16). North Direction Id:
<collectionCode id="BBA1AE49B849F30FFB81FC356F08E8A3" box="[1033,1093,963,987]" pageId="74" pageNumber="75">AMS</collectionCode>
I.19442-079, 3(1422),
<specimenCount id="CBB6FD05B849F30FFAD2FC356ED1E8A2" box="[1370,1436,963,987]" pageId="74" pageNumber="75" type="generic">
<quantity id="1A489B69B849F30FFAD2FC356ED4E8A2" box="[1370,1433,963,987]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="74" pageNumber="75" unit="m" value="12.0">12 m</quantity>
.
</specimenCount>
Outer Barrier Reef off Townsville:
<collectionCode id="BBA1AE49B849F30FFD9CFC11691DE887" box="[532,592,999,1023]" pageId="74" pageNumber="75">AMS</collectionCode>
I.20965-015, 7(1025),
<quantity id="1A489B69B849F30FFCD2FC1168F3E886" box="[858,958,999,1023]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="74" pageNumber="75" unit="m" value="15.0" valueMax="20.0" valueMin="10.0">
10
<specimenCount id="CBB6FD05B849F30FFC0AFC1168F3E886" box="[898,958,999,1023]" pageId="74" pageNumber="75" type="generic">20 m</specimenCount>
</quantity>
. Escape Reef:
<collectionCode id="BBA1AE49B849F30FFBD4FC116FD5E887" box="[1116,1176,999,1023]" pageId="74" pageNumber="75">AMS</collectionCode>
I.22573-036, 3(1619),
<quantity id="1A489B69B849F30FFF1FFBFD6BB6EF5A" box="[151,251,1035,1059]" metricMagnitude="1" metricUnit="m" metricValue="1.55" metricValueMax="1.7" metricValueMin="1.4" pageId="74" pageNumber="75" unit="m" value="15.5" valueMax="17.0" valueMin="14.0">1417 m</quantity>
;
<collectionCode id="BBA1AE49B849F30FFE80FBFD6A09EF5B" box="[264,324,1035,1059]" pageId="74" pageNumber="75">AMS</collectionCode>
I.22574-031, 7(1223),
<quantity id="1A489B69B849F30FFDDBFBFC69D0EF5A" box="[595,669,1034,1059]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="74" pageNumber="75" unit="m" value="4.5" valueMax="6.0" valueMin="3.0">36 m</quantity>
;
<collectionCode id="BBA1AE49B849F30FFD22FBFD69ABEF5B" box="[682,742,1035,1059]" pageId="74" pageNumber="75">AMS</collectionCode>
I.22579-079, 4(1720),
<quantity id="1A489B69B849F30FFC7DFBFD6F73EF5A" box="[1013,1086,1035,1059]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="74" pageNumber="75" unit="m" value="2.5" valueMax="4.0" valueMin="1.0">14 m</quantity>
;
<collectionCode id="BBA1AE49B849F30FFBC3FBFD6FCAEF5B" box="[1099,1159,1035,1059]" pageId="74" pageNumber="75">AMS</collectionCode>
I.22582-052, 19(1022),
<quantity id="1A489B69B849F30FFF1FFBD96BB6EF3E" box="[151,251,1071,1095]" metricMagnitude="1" metricUnit="m" metricValue="1.55" metricValueMax="1.7" metricValueMin="1.4" pageId="74" pageNumber="75" unit="m" value="15.5" valueMax="17.0" valueMin="14.0">1417 m</quantity>
;
<collectionCode id="BBA1AE49B849F30FFE8EFBD96A0FEF3F" box="[262,322,1071,1095]" pageId="74" pageNumber="75">AMS</collectionCode>
I.22583-039, 2(1516),
<quantity id="1A489B69B849F30FFDC5FBD969C5EF3E" box="[589,648,1071,1095]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="74" pageNumber="75" unit="m" value="29.0">29 m</quantity>
;
<collectionCode id="BBA1AE49B849F30FFD1BFBD96982EF3F" box="[659,719,1071,1095]" pageId="74" pageNumber="75">AMS</collectionCode>
I.22584-029, (20);
<collectionCode id="BBA1AE49B849F30FFC2DFBD968ACEF3F" box="[933,993,1071,1095]" pageId="74" pageNumber="75">AMS</collectionCode>
I.22587-020, (18),
<quantity id="1A489B69B849F30FFB3EFBD96FBCEF3E" box="[1206,1265,1071,1095]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="74" pageNumber="75" unit="m" value="40.0">40 m</quantity>
;
<collectionCode id="BBA1AE49B849F30FFB74FBD96E75EF3F" box="[1276,1336,1071,1095]" pageId="74" pageNumber="75">AMS</collectionCode>
I.22612- 0 32, 8(1121),
<quantity id="1A489B69B849F30FFEB6FBA56A34EF12" box="[318,377,1107,1131]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="74" pageNumber="75" unit="m" value="20.0">20 m</quantity>
;
<collectionCode id="BBA1AE49B849F30FFE0CFBA56A8DEF13" box="[388,448,1107,1131]" pageId="74" pageNumber="75">AMS</collectionCode>
I.22616-047, 14(1019),
<quantity id="1A489B69B849F30FFD51FBA46863EF12" box="[729,814,1106,1131]" metricMagnitude="1" metricUnit="m" metricValue="1.15" metricValueMax="1.8" metricValueMin="0.5" pageId="74" pageNumber="75" unit="m" value="11.5" valueMax="18.0" valueMin="5.0">518 m</quantity>
;
<collectionCode id="BBA1AE49B849F30FFCB2FBA5683BEF13" box="[826,886,1107,1131]" pageId="74" pageNumber="75">AMS</collectionCode>
I.22619-042, 16(1120),
<quantity id="1A489B69B849F30FFB06FBA56FBFEF12" box="[1166,1266,1107,1131]" metricMagnitude="1" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="74" pageNumber="75" unit="m" value="20.5" valueMax="22.0" valueMin="19.0">1922 m</quantity>
;
<collectionCode id="BBA1AE49B849F30FFB74FBA56E75EF13" box="[1276,1336,1107,1131]" pageId="74" pageNumber="75">AMS</collectionCode>
I.22634- 0 13, 7(920),
<specimenCount id="CBB6FD05B849F30FFEBBFB816A3EEFF6" box="[307,371,1143,1167]" pageId="74" pageNumber="75" type="generic">
<quantity id="1A489B69B849F30FFEBBFB816A22EFF6" box="[307,367,1143,1167]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="74" pageNumber="75" unit="m" value="18.0">18 m</quantity>
.
</specimenCount>
<emphasis id="EFC4EA9EB849F30FFEF2FB806914EFF7" bold="true" box="[378,601,1142,1167]" pageId="74" pageNumber="75">
Western
<collectingCountry id="A5A7761CB849F30FFE6BFB806919EFF7" box="[483,596,1142,1167]" name="Australia" pageId="74" pageNumber="75">Australia</collectingCountry>
:
</emphasis>
<collectingCountry id="A5A7761CB849F30FFDE9FB8069E3EFF7" box="[609,686,1142,1167]" name="Ashmore and Cartier Islands" pageId="74" pageNumber="75">Cartier</collectingCountry>
Id, Timor Sea:
<collectionCode id="BBA1AE49B849F30FFCEBFB8168EDEFF7" box="[867,928,1143,1167]" pageId="74" pageNumber="75">AMS</collectionCode>
I.26752-024, 7(1922), reef front off N side
<specimenCount id="CBB6FD05B849F30FFF1FFB6D6B9AEFCA" box="[151,215,1179,1203]" pageId="74" pageNumber="75" type="generic">
<quantity id="1A489B69B849F30FFF1FFB6D6B9EEFCA" box="[151,211,1179,1203]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="74" pageNumber="75" unit="m" value="12.0">12 m</quantity>
.
</specimenCount>
North West Cape:
<collectionCode id="BBA1AE49B849F30FFE25FB6D6ABFEFCB" LSID="urn:lsid:biocol.org:col:34981" box="[429,498,1179,1203]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="74" pageNumber="75">WAM</collectionCode>
P.26631-015, 19(1626), N of Mangrove Bay,
<quantity id="1A489B69B849F30FFB8AFB6D6F25EFCA" box="[1026,1128,1179,1203]" metricMagnitude="1" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="74" pageNumber="75" unit="m" value="13.0" valueMax="14.0" valueMin="12.0">
12
<specimenCount id="CBB6FD05B849F30FFBA5FB6D6F25EFCA" box="[1069,1128,1179,1203]" pageId="74" pageNumber="75" type="generic">14 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="DD0F368CB849F30FFF4FFB486830EEF2" blockId="74.[151,1437,422,2031]" pageId="74" pageNumber="75">
<emphasis id="EFC4EA9EB849F30FFF4FFB48694AEFAF" bold="true" box="[199,519,1214,1239]" pageId="74" pageNumber="75">
Other material.
<collectingCountry id="A5A7761CB849F30FFE01FB48694FEFAF" box="[393,514,1214,1239]" name="Indonesia" pageId="74" pageNumber="75">Indonesia</collectingCountry>
:
</emphasis>
N E side of
<collectingCountry id="A5A7761CB849F30FFD1DFB4869A4EFAF" box="[661,745,1214,1239]" name="Indonesia" pageId="74" pageNumber="75">Ambon</collectingCountry>
Id about
<quantity id="1A489B69B849F30FFCDDFB4968C3EFAF" box="[853,910,1214,1239]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="74" pageNumber="75" unit="km" value="2.0">2 km</quantity>
E of Sawa Telu: ex-USNM
<date id="A90E104CB849F30FFB45FB496E69EFAF" box="[1229,1316,1215,1239]" pageId="74" pageNumber="75" value="30-02-21">210230</date>
, 3(1121),
<quantity id="1A489B69B849F30FFF1FFB156BA4EF82" box="[151,233,1251,1275]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="8.0" metricValueMin="0.0" pageId="74" pageNumber="75" unit="m" value="4.0" valueMax="8.0" valueMin="0.0">
0
<specimenCount id="CBB6FD05B849F30FFF3CFB156BA4EF82" box="[180,233,1251,1275]" pageId="74" pageNumber="75" type="generic">8 m</specimenCount>
</quantity>
. Saparua:
<collectionCode id="BBA1AE49B849F30FFEE4FB156AF0EF83" LSID="urn:lsid:biocol.org:col:34871" box="[364,445,1251,1275]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="74" pageNumber="75">USNM</collectionCode>
209963, 20(
<quantity id="1A489B69B849F30FFDD3FB1569F6EF83" box="[603,699,1251,1275]" metricMagnitude="1" metricUnit="m" metricValue="1.45" metricValueMax="2.0" metricValueMin="0.9" pageId="74" pageNumber="75" unit="m" value="14.5" valueMax="20.0" valueMin="9.0">920 m</quantity>
),
<quantity id="1A489B69B849F30FFD5FFB156864EF82" box="[727,809,1251,1275]" metricMagnitude="0" metricUnit="m" metricValue="6.5" metricValueMax="9.0" metricValueMin="4.0" pageId="74" pageNumber="75" unit="m" value="6.5" valueMax="9.0" valueMin="4.0">
4
<specimenCount id="CBB6FD05B849F30FFD7CFB156864EF82" box="[756,809,1251,1275]" pageId="74" pageNumber="75" type="generic">9 m</specimenCount>
</quantity>
.
<emphasis id="EFC4EA9EB849F30FFCB4FB1468ABEF83" bold="true" box="[828,998,1250,1275]" pageId="74" pageNumber="75">
<collectingCountry id="A5A7761CB849F30FFCB4FB1468E4EF83" box="[828,937,1250,1275]" name="Marshall Islands" pageId="74" pageNumber="75">Marshall</collectingCountry>
Ids:
</emphasis>
Eniwetok Atoll:
<collectionCode id="BBA1AE49B849F30FFB37FB156FB1EF83" box="[1215,1276,1251,1275]" pageId="74" pageNumber="75">AMS</collectionCode>
I.37693-014, 17(1721);
<collectionCode id="BBA1AE49B849F30FFE96FAF16A16EE67" box="[286,347,1287,1311]" pageId="74" pageNumber="75">AMS</collectionCode>
I.37687-004, 5(1320),
<quantity id="1A489B69B849F30FFDF4FAF169E3EE66" box="[636,686,1287,1310]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="74" pageNumber="75" unit="m" value="7.0">7 m</quantity>
;
<collectionCode id="BBA1AE49B849F30FFD36FAF169B6EE67" box="[702,763,1287,1311]" pageId="74" pageNumber="75">AMS</collectionCode>
I.37695-002, (14);
<collectionCode id="BBA1AE49B849F30FFC6BFAF16F6DEE67" box="[995,1056,1287,1311]" pageId="74" pageNumber="75">AMS</collectionCode>
I.37709-007, 20(1724),
<collectionCode id="BBA1AE49B849F30FFAC6FAF16ED1EE66" LSID="urn:lsid:biocol.org:col:34697" box="[1358,1436,1287,1310]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="74" pageNumber="75">BPBM</collectionCode>
29222, 2(1316),
<specimenCount id="CBB6FD05B849F30FFEEDFADD6AD4EE3A" box="[357,409,1323,1346]" pageId="74" pageNumber="75" type="generic">
<quantity id="1A489B69B849F30FFEEDFADD6AD8EE3A" box="[357,405,1323,1346]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="74" pageNumber="75" unit="m" value="7.0">7 m</quantity>
.
</specimenCount>
<collectingCountry id="A5A7761CB849F30FFE2AFADC694FEE3B" box="[418,514,1322,1347]" name="Kiribati" pageId="74" pageNumber="75">
<emphasis id="EFC4EA9EB849F30FFE2AFADC694FEE3B" bold="true" box="[418,514,1322,1347]" pageId="74" pageNumber="75">Kiribati</emphasis>
</collectingCountry>
: Abaiang Atoll:
<collectionCode id="BBA1AE49B849F30FFD4FFADD6849EE3B" box="[711,772,1323,1347]" pageId="74" pageNumber="75">AMS</collectionCode>
I.18043-064, 6(1221),
<quantity id="1A489B69B849F30FFB97FADC6F26EE3A" box="[1055,1131,1322,1347]" metricMagnitude="0" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="74" pageNumber="75" unit="m" value="6.5" valueMax="8.0" valueMin="5.0">
5
<specimenCount id="CBB6FD05B849F30FFBB4FADC6F26EE3A" box="[1084,1131,1322,1347]" pageId="74" pageNumber="75" type="generic">8 m</specimenCount>
</quantity>
.
<emphasis id="EFC4EA9EB849F30FFBF0FADC6E55EE3B" bold="true" box="[1144,1304,1322,1347]" pageId="74" pageNumber="75">Solomon Ids:</emphasis>
Florida Id:
<collectionCode id="BBA1AE49B849F30FFF1FFAB96B99EE1F" box="[151,212,1359,1383]" pageId="74" pageNumber="75">AMS</collectionCode>
I.17497-016, 3(1618),
<quantity id="1A489B69B849F30FFE64FAB8697AEE1E" box="[492,567,1358,1383]" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="74" pageNumber="75" unit="m" value="3.5" valueMax="5.0" valueMin="2.0">25 m</quantity>
; Florida Id:
<collectionCode id="BBA1AE49B849F30FFD4EFAB9684EEE1F" box="[710,771,1359,1383]" pageId="74" pageNumber="75">AMS</collectionCode>
I.17500-028, 2(1921),
<quantity id="1A489B69B849F30FFB93FAB96FCFEE1E" box="[1051,1154,1359,1383]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="74" pageNumber="75" unit="m" value="25.0" valueMax="30.0" valueMin="20.0">
20
<specimenCount id="CBB6FD05B849F30FFBCDFAB96FCFEE1E" box="[1093,1154,1359,1383]" pageId="74" pageNumber="75" type="generic">30 m</specimenCount>
</quantity>
. Savo Id:
<collectionCode id="BBA1AE49B849F30FFB7FFAB96E79EE1F" box="[1271,1332,1359,1383]" pageId="74" pageNumber="75">AMS</collectionCode>
I.17491- 0 39, 3(1719),
<specimenCount id="CBB6FD05B849F30FFECCFA856AC9EEF2" box="[324,388,1395,1419]" pageId="74" pageNumber="75" type="generic">
<quantity id="1A489B69B849F30FFECCFA856A32EEF2" box="[324,383,1395,1419]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="74" pageNumber="75" unit="m" value="10.0">10 m</quantity>
.
</specimenCount>
Guadalcanal:
<collectionCode id="BBA1AE49B849F30FFDADFA85692FEEF3" box="[549,610,1395,1419]" pageId="74" pageNumber="75">AMS</collectionCode>
I.27020-003, (19),
<specimenCount id="CBB6FD05B849F30FFCB5FA856830EEF2" box="[829,893,1395,1419]" pageId="74" pageNumber="75" type="generic">
<quantity id="1A489B69B849F30FFCB5FA856834EEF2" box="[829,889,1395,1419]" metricMagnitude="1" metricUnit="m" metricValue="3.2" pageId="74" pageNumber="75" unit="m" value="32.0">32 m</quantity>
.
</specimenCount>
</paragraph>
<paragraph id="DD0F368CB849F30FFF4FFA606E4DEED7" blockId="74.[151,1437,422,2031]" box="[199,1280,1430,1455]" pageId="74" pageNumber="75">
<emphasis id="EFC4EA9EB849F30FFF4FFA606A52EED7" bold="true" box="[199,287,1430,1455]" pageId="74" pageNumber="75">Photos.</emphasis>
KPM-NR0034522,
<collectingCountry id="A5A7761CB849F30FFD8BFA606920EED7" box="[515,621,1430,1455]" name="Malaysia" pageId="74" pageNumber="75">Malaysia</collectingCountry>
; KPM-NR0052877, Borneo; KPM-NR0052878, Okinawa.
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B849F30FFF4FFA4C6A19EC97" pageId="74" pageNumber="75" type="diagnosis">
<paragraph id="DD0F368CB849F30FFF4FFA4C6A19EC97" blockId="74.[151,1437,422,2031]" pageId="74" pageNumber="75">
<emphasis id="EFC4EA9EB849F30FFF4FFA4C6A0CEEAB" bold="true" box="[199,321,1466,1491]" pageId="74" pageNumber="75">Diagnosis.</emphasis>
A species of
<taxonomicName id="1AB04D0FB849F30FFE53FA4A697FEEAB" box="[475,562,1468,1491]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="74" pageNumber="75" phylum="Chordata" rank="genus">
<emphasis id="EFC4EA9EB849F30FFE53FA4A697FEEAB" box="[475,562,1468,1491]" italics="true" pageId="74" pageNumber="75">Trimma</emphasis>
</taxonomicName>
with a moderately deep interorbital trench in adults, a shallow groove behind the upper half of the eye in adults, largely disappearing below the papilla at the posterodorsal margin of the eye; predorsal almost completely covered with ctenoid scales, with 36 rows crossing the midline anteriorly to just behind the interorbital trench, anterior-most scale on the midline just behind a small naked patch posterior to the eyes, scale not larger than the second scale on the midline; opercle and preopercle naked; pectoral-fin base covered with 56 large cycloid scales, lowermost 34 scales smaller than upper two scales; prepelvic area with a small scale covering basal membrane between the two fins, followed anteriorly by a large scale between the bases of fins, and 3 rows of large cycloid scales anteriorly; central pectoral rays branched; pelvic fins widely separate, connected only at their bases, the distance between the bases of the fins about threequarters to equal to the base of each pelvic fin; fifth pelvic ray usually unbranched, rarely with a single branch (2 terminal tips); fifth ray 6080% the length of the fourth; second dorsal spine usually longest, elongated into a long filament in adult males, reaching to beyond the end of the second dorsal fin, females sometimes with a short filament reaching to above the anterior rays of the second dorsal fin; third spine sometimes a short filament; second dorsal usually
<date id="A90E104CB849F30FFB06F89D6FE1ECFB" box="[1166,1196,1899,1923]" pageId="74" pageNumber="75">I 9</date>
and anal rays usually
<date id="A90E104CB849F30FFF1FF8796BF4ECDF" box="[151,185,1935,1959]" pageId="74" pageNumber="75">I 8</date>
; two distinct round red to reddish-orange spots below the eye; branchiostegal membranes with distinct bright reddish-orange spots; body with small yellow to red spots in 45 irregular rows anteriorly and three rows on the caudal peduncle.
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B848F309FF4FFF616FFDE899" lastPageId="76" lastPageNumber="77" pageId="75" pageNumber="76" type="description">
<paragraph id="DD0F368CB848F30EFF4FFF616988EE2C" blockId="75.[151,1437,151,1977]" pageId="75" pageNumber="76">
<emphasis id="EFC4EA9EB848F30EFF4FFF616A14EBC8" bold="true" box="[199,345,151,176]" pageId="75" pageNumber="76">Description.</emphasis>
The description is based on specimens from the Timor Sea, Great Barrier Reef, Coral Sea and
<collectingCountry id="A5A7761CB848F30EFF1FFF4A6A1BEBAD" box="[151,342,188,213]" name="Solomon Islands" pageId="75" pageNumber="76">Solomon Islands</collectingCountry>
. Dorsal fin usually VI +
<date id="A90E104CB848F30EFDE3FF4B69C7EBAD" box="[619,650,188,213]" pageId="75" pageNumber="76">
I
<emphasis id="EFC4EA9EB848F30EFDF4FF4A69C7EBAD" bold="true" box="[636,650,188,213]" pageId="75" pageNumber="76">9</emphasis>
</date>
(
<date id="A90E104CB848F30EFD11FF4A69F5EBAC" box="[665,696,188,212]" pageId="75" pageNumber="76">I 8</date>
or
<date id="A90E104CB848F30EFD56FF4B6847EBAC" box="[734,778,188,212]" pageId="75" pageNumber="76" value="2010-01">
I
<quantity id="1A489B69B848F30EFD66FF4A6847EBAC" box="[750,778,188,212]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="75" pageNumber="76" unit="in" value="10.0">10</quantity>
</date>
in 10% of individuals), mean = I 9.0, n = 60, second spine longest, often elongated into filament (reaching first segmented rays in juveniles and well beyond end of second dorsal fin in some large males, in some females first spine also filamentous and longest; first ray of second dorsal fin usually branched in specimens above
<quantity id="1A489B69B848F30EFDEEFEDE69FBEA47" box="[614,694,296,320]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="75" pageNumber="76" unit="mm" value="14.0">14 mm</quantity>
SL and unbranched in smaller specimens; anterior element of last ray branched; anal fin
<date id="A90E104CB848F30EFE1DFEBB6AF9EA1D" box="[405,436,332,357]" pageId="75" pageNumber="76">
I
<emphasis id="EFC4EA9EB848F30EFE2EFEBA6AF9EA1D" bold="true" box="[422,436,332,357]" pageId="75" pageNumber="76">8</emphasis>
</date>
(rarely
<date id="A90E104CB848F30EFD86FEBB6960EA1C" box="[526,557,333,356]" pageId="75" pageNumber="76">I 7</date>
or
<date id="A90E104CB848F30EFDDCFEBB6938EA1C" box="[596,629,332,356]" pageId="75" pageNumber="76">I 9</date>
, in 6% of individuals), mean = I 8.0, n = 60, first ray usually branched, anterior element of last ray branched; pectoral-fin rays usually 18
<emphasis id="EFC4EA9EB848F30EFC03FE9968EAEAF0" bold="true" box="[907,935,367,392]" pageId="75" pageNumber="76">19</emphasis>
, (Queensland mean = 18.1, n = 37, Western
<collectingCountry id="A5A7761CB848F30EFF1FFE626BB3EAD5" box="[151,254,404,429]" name="Australia" pageId="75" pageNumber="76">Australia</collectingCountry>
mean = 18.5, n = 15,
<collectingCountry id="A5A7761CB848F30EFE7BFE6269E3EAD5" box="[499,686,404,429]" name="Solomon Islands" pageId="75" pageNumber="76">Solomon Islands</collectingCountry>
mean = 17.8, n = 9), upper 58 rays unbranched, central 512 rays branched and lower 36 rays unbranched; pectoral fin reaching just posteriorly to a vertical in line from urogenital papilla to about second segmented ray of anal fin; pelvic fin
<date id="A90E104CB848F30EFCEFFE2B68C6EA8D" box="[871,907,476,501]" pageId="75" pageNumber="76">I 5</date>
, fins widely separate with very low interradial connecting membrane, covered by a median scale, distance between bases of fins slightly less than base of each pelvic fin; first four rays with one sequential branch, fifth ray usually unbranched, 5080% length of fourth ray, which reaches posteriorly to below point between first and third anal rays, segmented caudal rays usually
<emphasis id="EFC4EA9EB848F30EFA84FDB16E6AE918" bold="true" box="[1292,1319,583,608]" pageId="75" pageNumber="76">17</emphasis>
; branched caudal rays usually 6 upper + 5 lower (rarely
<emphasis id="EFC4EA9EB848F30EFD12FD9A6995E9FD" bold="true" box="[666,728,620,645]" pageId="75" pageNumber="76">
6 +
<quantity id="1A489B69B848F30EFD42FD9A6995E9FD" box="[714,728,620,645]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="75" pageNumber="76" unit="in" value="6.0">6</quantity>
</emphasis>
in &lt;3% of specimens); predorsal scales crossing midline 3
<emphasis id="EFC4EA9EB848F30EFA0FFD9A6ED8E9FD" bold="true" box="[1415,1429,620,645]" pageId="75" pageNumber="76">6</emphasis>
, anterior two scales on midline, posterior scales overlapping on midline and no scales on midline covering nuchal crest (Queensland mean 3.6; n = 19, Western
<collectingCountry id="A5A7761CB848F30EFD21FD42685DE9B5" box="[681,784,692,717]" name="Australia" pageId="75" pageNumber="76">Australia</collectingCountry>
mean = 4.9, n = 14, mean
<collectingCountry id="A5A7761CB848F30EFBD9FD426E42E9B5" box="[1105,1295,692,717]" name="Solomon Islands" pageId="75" pageNumber="76">Solomon Islands</collectingCountry>
5.3, n = 8); lateral scales 2225, usually 23 or
<emphasis id="EFC4EA9EB848F30EFD90FD216979E988" bold="true" box="[536,564,727,752]" pageId="75" pageNumber="76">24</emphasis>
(Queensland mean 23.8; n = 36, Western
<collectingCountry id="A5A7761CB848F30EFB8EFD216F20E988" box="[1030,1133,727,752]" name="Australia" pageId="75" pageNumber="76">Australia</collectingCountry>
mean = 23.9, n = 14, mean
<collectingCountry id="A5A7761CB848F30EFF1FFD0A6A1AE86D" box="[151,343,764,789]" name="Solomon Islands" pageId="75" pageNumber="76">Solomon Islands</collectingCountry>
23.3, n = 8); transverse scale count 68, usually
<emphasis id="EFC4EA9EB848F30EFC23FD0A68F4E86D" bold="true" box="[939,953,764,789]" pageId="75" pageNumber="76">7</emphasis>
(Queensland mean 7.1; n = 31, Western
<collectingCountry id="A5A7761CB848F30EFF1FFCE96BB3E840" box="[151,254,799,824]" name="Australia" pageId="75" pageNumber="76">Australia</collectingCountry>
mean = 7.0, n = 14,
<collectingCountry id="A5A7761CB848F30EFE6DFCE969EDE840" box="[485,672,799,824]" name="Solomon Islands" pageId="75" pageNumber="76">Solomon Islands</collectingCountry>
mean 7.0, n = 8); preopercle and opercle scaleless; scales extending to edge of postorbital trench above posterior part eye; scales just below dorsal fin with small accessory scales; pectoral-fin base covered with 56 large cycloid scales, lowermost 34 scales smaller than other scales; prepelvic area fully scaled, with small scale covering basal membrane between fins, followed anteriorly by large scale between bases of fins and 3 rows of large of cycloid scales anteriorly. Nape crest low and indistinct, extending to just before dorsal origin and above posterior opercular margin. Teeth in lower jaw consist of enlarged outer row of curved, slightly enlarged wide-set canines ending near bend in dentary and inner row of similar, but smaller canines extending full length of dentary, with one or two irregular rows of smaller conical teeth in between; outer row of teeth in upper jaw wide-set and extending full length of premaxilla, similar to those of lower jaw, with 12 small irregular inner rows of teeth, innermost row not enlarged. Tongue tip slightly concave. Gill opening extending anteroventrally to below posterior margin of eye. Outer gill rakers on first arch elongate, almost equal to filament length at angle, 34 + 1113 = 1518 (mean = 16.3, n = 20). Anterior nares at end of narrow tube, just above upper lip, posterior nares small pore with raised rim about two nostril diameters from anterior nares and two nostril diameters from eye; nasal sac elevated as raised oval sac located just above upper lip. Interorbital very narrow, with eyes almost touching, trench typically present in well preserved adults, but often obscured in poorly preserved specimens and not well developed in juveniles below
<quantity id="1A489B69B848F30EFDCFFACA69DBEE2C" box="[583,662,1340,1364]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="75" pageNumber="76" unit="mm" value="18.0">18 mm</quantity>
SL.
</paragraph>
<paragraph id="DD0F368CB848F30EFF4FFAA96AF5ECC1" blockId="75.[151,1437,151,1977]" pageId="75" pageNumber="76">
<emphasis id="EFC4EA9EB848F30EFF4FFAA9691EEE00" box="[199,595,1375,1400]" italics="true" pageId="75" pageNumber="76">Colour pattern. Freshly collected</emphasis>
specimens from Great Barrier Reef (from slides of freshly collected specimens from Yonge Reef and Escape Reef, Queensland,
<figureCitation id="458B2A09B848F30EFCD9FA7268E5EEE4" box="[849,936,1412,1437]" captionStart="FIGURE 42" captionStartId="76.[151,250,656,678]" captionTargetBox="[152,1435,193,634]" captionTargetId="figure@76.[152,1435,193,635]" captionTargetPageId="76" captionText="FIGURE 42. Trimma readerae, 22.5 mm SL holotype, AMS I. 20784 - 020, Yonge Reef, Queensland. Photo by D. F. Hoese." httpUri="https://zenodo.org/record/236110/files/figure.png" pageId="75" pageNumber="76">Fig. 42</figureCitation>
). Head and body with numerous orange to yellow-orange or reddish-orange spots with interspaces grey to bluish grey, except ventrally on body where interspaces whitish. Short orange bar extending from anterior margin of eye below nostrils to upper lip and along dorsal margin of upper lip; followed by elongate orange spot or bar extending to just behind posterior end of jaws; two round orange spots below middle of eye; followed by similar sized spots near end of preopercle; upper preopercle with oval orange spot extending onto upper opercle, followed posteriorly by oval spots extending from upper part of opercle onto nape; pale orange vertically elongate spot ventrally on opercle; two large round reddish spots ventrally on branchiostegal membranes below opercle and preopercle, spot below opercle with thin dorsal projection onto posterior end of opercle; pectoral-fin base with two large round orange spots; nape with large orange spots, some connected across nape forming bars; orange to reddish-orange spots on body round anteriorly, but becoming smaller and vertically elongate posteriorly and ventrally, spots forming 35 horizontal rows on caudal peduncle; dorsal and anal fins with large spots of similar colouration to body spots, with largest spots near base of fin, followed by 34 rows of smaller spots distally; pectoral fin whitish; pelvic fin clear to grey, with orange pigment near base of fins; caudal fin with 46 rows of smaller spots of similar colouration to body spots; pectoralfin base with upper and lower orange to yellow-orange or reddish-orange spot, with central spot extending onto bases of pectoral-fin rays.
</paragraph>
<caption id="89CF6604B84FF309FF1FFD666E35E9DE" box="[151,1400,656,678]" httpUri="https://zenodo.org/record/236110/files/figure.png" pageId="76" pageNumber="77" targetBox="[152,1435,193,634]" targetPageId="76">
<paragraph id="DD0F368CB84FF309FF1FFD666E35E9DE" blockId="76.[151,1400,656,678]" box="[151,1400,656,678]" pageId="76" pageNumber="77">
<emphasis id="EFC4EA9EB84FF309FF1FFD666A52E9DE" bold="true" box="[151,287,656,678]" pageId="76" pageNumber="77">FIGURE 42.</emphasis>
<taxonomicName id="1AB04D0FB84FF309FEADFD676A9BE9DE" box="[293,470,656,678]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="readerae">
<emphasis id="EFC4EA9EB84FF309FEADFD676A9BE9DE" box="[293,470,656,678]" italics="true" pageId="76" pageNumber="77">Trimma readerae</emphasis>
</taxonomicName>
, 22.5 mm SL holotype, AMS I.20784-020, Yonge Reef, Queensland. Photo by D. F. Hoese.
</paragraph>
</caption>
<paragraph id="DD0F368CB84FF309FF4FFD3868C0E856" blockId="76.[151,1437,717,2038]" pageId="76" pageNumber="77">
<emphasis id="EFC4EA9EB84FF309FF4FFD386BBAE99D" box="[199,247,718,741]" italics="true" pageId="76" pageNumber="77">Live</emphasis>
. (from two images from Great Barrier Reef, Pl 3 D and E): similar to above, but body somewhat translucent; white spot present on top of pectoral-fin base, better developed in light phase than in dark phase; iris red with some darker pigment near outer rim dorsally and ventrally.
</paragraph>
<paragraph id="DD0F368CB84FF309FF4FFCCF6FFDE899" blockId="76.[151,1437,717,2038]" pageId="76" pageNumber="77">
<emphasis id="EFC4EA9EB84FF309FF4FFCCF6A7AE82A" box="[199,311,825,850]" italics="true" pageId="76" pageNumber="77">Preserved</emphasis>
. Head and body light brown, Grey interspaces dark grey to brown; orange to yellow-orange or reddish-orange spots becoming light brown; dark brown bar connecting vertical bars under eye between spots faint and sometimes absent, with margins of bands concave above and below indicating position of spots; fins often clear, but in freshly collected specimens dorsal, anal and caudal fins with melanophores surrounding clear spots; spots on branchiostegal membranes in life not visible; pectoral-fin base with three light spots.
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B84FF309FF4FFC1B692AEF35" pageId="76" pageNumber="77" type="etymology">
<paragraph id="DD0F368CB84FF309FF4FFC1B692AEF35" blockId="76.[151,1437,717,2038]" pageId="76" pageNumber="77">
<emphasis id="EFC4EA9EB84FF309FF4FFC1B6A03EF7E" bold="true" box="[199,334,1005,1030]" pageId="76" pageNumber="77">Etymology.</emphasis>
Named for Sally Reader, of the Australian Museum, in appreciation of the many years she has spent painstakingly gathering, analyzing and documenting data on
<taxonomicName id="1AB04D0FB84FF309FC18FBE468ABEF51" box="[912,998,1042,1065]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="76" pageNumber="77" phylum="Chordata" rank="genus">
<emphasis id="EFC4EA9EB84FF309FC18FBE468ABEF51" box="[912,998,1042,1065]" italics="true" pageId="76" pageNumber="77">Trimma</emphasis>
</taxonomicName>
for DFHs research program, some of which has been used in the present paper.
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B84FF308FF4FFBAF6984EB80" lastPageId="77" lastPageNumber="78" pageId="76" pageNumber="77" type="distribution">
<paragraph id="DD0F368CB84FF309FF4FFBAF6980ED5A" blockId="76.[151,1437,717,2038]" pageId="76" pageNumber="77">
<emphasis id="EFC4EA9EB84FF309FF4FFBAF6A2DEF0A" bold="true" box="[199,352,1113,1138]" pageId="76" pageNumber="77">Distribution.</emphasis>
<taxonomicName id="1AB04D0FB84FF309FEE0FBAC6967EF0A" box="[360,554,1113,1138]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="readerae">
<emphasis id="EFC4EA9EB84FF309FEE0FBAC6967EF0A" box="[360,554,1113,1138]" italics="true" pageId="76" pageNumber="77">Trimma readerae</emphasis>
</taxonomicName>
is known from mid to outer reefs on the Great Barrier Reef, and inshore reefs of Western
<collectingCountry id="A5A7761CB84FF309FE93FB8B6AC8EFEE" box="[283,389,1149,1174]" name="Australia" pageId="76" pageNumber="77">Australia</collectingCountry>
, but rarely from the Coral Sea and Timor Sea. It sometimes occurs with
<taxonomicName id="1AB04D0FB84FF309FB42FB886ED8EFEE" box="[1226,1429,1149,1174]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="okinawae">
<emphasis id="EFC4EA9EB84FF309FB42FB886ED8EFEE" box="[1226,1429,1149,1174]" italics="true" pageId="76" pageNumber="77">Trimma okinawae</emphasis>
</taxonomicName>
, which is typically more common on offshore reefs of the Coral and Timor seas. On the outer reefs
<taxonomicName id="1AB04D0FB84FF309FACDFB546BB6EFA6" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="readerae">
<emphasis id="EFC4EA9EB84FF309FACDFB546BB6EFA6" italics="true" pageId="76" pageNumber="77">Trimma readerae</emphasis>
</taxonomicName>
is usually found in back reef leeward areas. The apparent scarcity of the species north of Tijou Reef is probably an artefact relating to collecting effort. For example, samples from Raine
<collectingCountry id="A5A7761CB84FF309FBDFFB1F6FD6EE7A" box="[1111,1179,1257,1282]" name="Iceland" pageId="76" pageNumber="77">Island</collectingCountry>
and other reefs of the northern Great Barrier Reef were generally from the seaward face of the reef. The species is known from a wide range of depths from
<quantity id="1A489B69B84FF309FE16FAC46AB6EE31" box="[414,507,1330,1354]" metricMagnitude="1" metricUnit="m" metricValue="2.15" metricValueMax="4.0" metricValueMin="0.3" pageId="76" pageNumber="77" unit="m" value="21.5" valueMax="40.0" valueMin="3.0">
3
<specimenCount id="CBB6FD05B84FF309FE32FAC46AB6EE31" box="[442,507,1330,1354]" pageId="76" pageNumber="77" type="generic">40 m</specimenCount>
</quantity>
. We tentatively include here, in what appears to represent a species complex, specimens and images from the
<collectingCountry id="A5A7761CB84FF309FE76FAA369F1EE16" box="[510,700,1365,1390]" name="Solomon Islands" pageId="76" pageNumber="77">Solomon Islands</collectingCountry>
,
<collectingCountry id="A5A7761CB84FF309FD4FFAA3687AEE16" box="[711,823,1365,1390]" name="Indonesia" pageId="76" pageNumber="77">Indonesia</collectingCountry>
, Borneo, New
<collectingCountry id="A5A7761CB84FF309FC56FAA36F7FEE16" box="[990,1074,1365,1390]" name="Guinea-Bissau" pageId="76" pageNumber="77">Guinea</collectingCountry>
,
<collectingCountry id="A5A7761CB84FF309FBB5FAA06F30EE15" box="[1085,1149,1366,1389]" name="Japan" pageId="76" pageNumber="77">Japan</collectingCountry>
and the
<collectingCountry id="A5A7761CB84FF309FB53FAA36ED5EE16" box="[1243,1432,1365,1390]" name="Marshall Islands" pageId="76" pageNumber="77">Marshall Islands</collectingCountry>
, based on the spotting pattern on the cheek. However,
<bibRefCitation id="B9214B7DB84FF309FCB1FA8F6F32EEEA" box="[825,1151,1401,1426]" pageId="76" pageNumber="77" refString="Winterbottom, R., Hanner, R. H., Burridge, M. &amp; Zur, M. (2014) A cornucopia of cryptic species-a DNA barcode analysis of the gobiid fish genus Trimma (Percomorpha, Gobiiformes). ZooKeys, 381, 79 - 111. http: // dx. doi. org / 10.3897 / zookeys. 381.6445" type="journal article">
Winterbottom
<emphasis id="EFC4EA9EB84FF309FC6CFA8C6F68EEEA" box="[996,1061,1401,1426]" italics="true" pageId="76" pageNumber="77">et al.</emphasis>
(2014)
</bibRefCitation>
noted 8 distinct forms (haplogroups) of what they identified as
<taxonomicName id="1AB04D0FB84FF309FDEFFA6869A2EECE" box="[615,751,1437,1462]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="okinawae">
<emphasis id="EFC4EA9EB84FF309FDEFFA6869A2EECE" box="[615,751,1437,1462]" italics="true" pageId="76" pageNumber="77">T. okinawae</emphasis>
</taxonomicName>
(see below), with all groups except 5 and 6 from
<collectingCountry id="A5A7761CB84FF309FAA3FA686E26EECD" box="[1323,1387,1438,1461]" name="Japan" pageId="76" pageNumber="77">Japan</collectingCountry>
and
<collectingCountry id="A5A7761CB84FF309FF1FFA376BA4EEA2" box="[151,233,1473,1498]" name="Taiwan" pageId="76" pageNumber="77">Taiwan</collectingCountry>
having spots under the eye. Specimens from Raja Ampat in
<collectingCountry id="A5A7761CB84FF309FC54FA376F04EEA2" box="[988,1097,1473,1498]" name="Indonesia" pageId="76" pageNumber="77">Indonesia</collectingCountry>
closely resemble Australian specimens in live colouration, but differ significantly genetically, suggesting multiple species. Consequently we restrict the
<typeStatus id="020B882EB84FF309FE9DF9FC6A08ED5A" box="[277,325,1546,1570]" pageId="76" pageNumber="77">type</typeStatus>
material to the Great Barrier Reef.
</paragraph>
<paragraph id="DD0F368CB84FF309FF4FF9DB68C2ECF1" blockId="76.[151,1437,717,2038]" pageId="76" pageNumber="77">
<emphasis id="EFC4EA9EB84FF309FF4FF9DB6A21ED3E" bold="true" box="[199,364,1581,1606]" pageId="76" pageNumber="77">Comparisons.</emphasis>
The species is most similar to the species treated here as
<taxonomicName id="1AB04D0FB84FF309FB9DF9D86FAEED3E" box="[1045,1251,1581,1606]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="okinawae">
<emphasis id="EFC4EA9EB84FF309FB9DF9D86FAEED3E" box="[1045,1251,1581,1606]" italics="true" pageId="76" pageNumber="77">Trimma okinawae</emphasis>
</taxonomicName>
(q.v.). It differs primarily in having light spots under the eye, rather than light vertical bars, and in having filamentous dorsal spines in males and frequently females. In addition, the spot posteriorly on the middle of the preopercle is much smaller than other spots on the head (versus subequal in size to, or continuous with, the spot above in
<taxonomicName id="1AB04D0FB84FF309FB49F96C6EC6EDCA" box="[1217,1419,1689,1714]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="okinawae">
<emphasis id="EFC4EA9EB84FF309FB49F96C6EC6EDCA" box="[1217,1419,1689,1714]" italics="true" pageId="76" pageNumber="77">Trimma okinawae</emphasis>
</taxonomicName>
). The second dorsal spine is usually filamentous reaching to or beyond the middle of the second dorsal fin. In females the second spine is longest and sometimes filamentous, reaching to above the first few rays of the second dorsal fin.
<taxonomicName id="1AB04D0FB84FF309FE86F8F06A95EC66" box="[270,472,1797,1822]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="okinawae">
<emphasis id="EFC4EA9EB84FF309FE86F8F06A95EC66" box="[270,472,1797,1822]" italics="true" pageId="76" pageNumber="77">Trimma okinawae</emphasis>
</taxonomicName>
normally lacks filamentous dorsal spines, but also has the second dorsal spine longest. In other populations the spines is frequently filamentous. The pelvic fin varies considerably in length, but normally does not extend beyond the second segmented anal ray, versus fourth or fifth segmented ray in
<taxonomicName id="1AB04D0FB84FF309FB43F8B86ED8EC1E" box="[1227,1429,1869,1894]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="okinawae">
<emphasis id="EFC4EA9EB84FF309FB43F8B86ED8EC1E" box="[1227,1429,1869,1894]" italics="true" pageId="76" pageNumber="77">Trimma okinawae</emphasis>
</taxonomicName>
. Faded preserved specimens can be difficult to consistently separate.
</paragraph>
<paragraph id="DD0F368CB84FF308FF4FF8636984EB80" blockId="76.[151,1437,717,2038]" lastBlockId="77.[151,1436,151,285]" lastPageId="77" lastPageNumber="78" pageId="76" pageNumber="77">
<bibRefCitation id="B9214B7DB84FF309FF4FF8636AB7ECD6" box="[199,506,1941,1966]" pageId="76" pageNumber="77" refString="Winterbottom, R., Hanner, R. H., Burridge, M. &amp; Zur, M. (2014) A cornucopia of cryptic species-a DNA barcode analysis of the gobiid fish genus Trimma (Percomorpha, Gobiiformes). ZooKeys, 381, 79 - 111. http: // dx. doi. org / 10.3897 / zookeys. 381.6445" type="journal article">
Winterbottom
<emphasis id="EFC4EA9EB84FF309FEE4F8606AEBECD6" box="[364,422,1941,1966]" italics="true" pageId="76" pageNumber="77">et al.</emphasis>
(2014)
</bibRefCitation>
provided a DNA barcode analysis of
<specimenCount id="CBB6FD05B84FF309FC3AF8606F01ECD6" box="[946,1100,1941,1966]" pageId="76" pageNumber="77" type="generic">28 specimens</specimenCount>
provisionally identified as
<taxonomicName id="1AB04D0FB84FF309FA0FF8606A4FECAA" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="okinawae">
<emphasis id="EFC4EA9EB84FF309FA0FF8606A4FECAA" italics="true" pageId="76" pageNumber="77">T. okinawae</emphasis>
</taxonomicName>
. They recovered eight haplogroups, all but one confined to a single geographic area, suggesting the possibility of multiple cryptic species. The two specimens from the Great Barrier Reef in their analysis (Group 8) were phenetically closest to three specimens from Raja Ampat,
<collectingCountry id="A5A7761CB84EF308FCFDFF6168AFEBC8" box="[885,994,151,176]" name="Indonesia" pageId="77" pageNumber="78">Indonesia</collectingCountry>
plus one specimen from Rabaul, New
<collectingCountry id="A5A7761CB84EF308FF1FFF4A6BA8EBAD" box="[151,229,188,213]" name="United Kingdom" pageId="77" pageNumber="78">Britain</collectingCountry>
(Group 7), but separated from them by 7.1% of the CO1 base pairs. The other specimens were from
<collectingCountry id="A5A7761CB84EF308FAC1FF4A6ED5EBAD" box="[1353,1432,188,213]" name="Brunei" pageId="77" pageNumber="78">Brunei</collectingCountry>
,
<collectingCountry id="A5A7761CB84EF308FF1FFF296B8CEB80" box="[151,193,223,248]" name="Fiji" pageId="77" pageNumber="78">Fiji</collectingCountry>
,
<collectingCountry id="A5A7761CB84EF308FF44FF166A42EB8F" box="[204,271,224,247]" name="Japan" pageId="77" pageNumber="78">Japan</collectingCountry>
,
<collectingCountry id="A5A7761CB84EF308FE92FF166A87EB80" box="[282,458,223,248]" name="New Caledonia" pageId="77" pageNumber="78">New Caledonia</collectingCountry>
,
<collectingCountry id="A5A7761CB84EF308FE5DFF29696AEB80" box="[469,551,223,248]" name="Taiwan" pageId="77" pageNumber="78">Taiwan</collectingCountry>
and
<collectingCountry id="A5A7761CB84EF308FDD5FF296989EB80" box="[605,708,223,248]" name="Thailand" pageId="77" pageNumber="78">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B84EF308FF4FFEF268DEEA64" box="[199,915,260,285]" pageId="77" pageNumber="78" type="discussion">
<paragraph id="DD0F368CB84EF308FF4FFEF268DEEA64" blockId="77.[151,1436,151,285]" box="[199,915,260,285]" pageId="77" pageNumber="78">
<emphasis id="EFC4EA9EB84EF308FF4FFEF26A01EA65" bold="true" box="[199,332,260,285]" pageId="77" pageNumber="78">Discussion.</emphasis>
The species reaches a maximum size of
<quantity id="1A489B69B84EF308FC9DFEF26829EA64" box="[789,868,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="77" pageNumber="78" unit="mm" value="28.0">28 mm</quantity>
SL.
</paragraph>
</subSubSection>
</treatment>
</document>