303 lines
34 KiB
XML
303 lines
34 KiB
XML
<document id="FA6BF2AC8B2E66CCB96426FE69FAA2D9" ID-DOI="10.11646/zootaxa.3934.1.1" ID-GBIF-Dataset="0abffe3e-a2c6-4623-b7c6-c8b26cee4a2b" ID-ISSN="1175-5326" ID-Zenodo-Dep="236066" ID-ZooBank="11C2A2CB-30B3-4694-B379-AE9D47332F0C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461012229557" checkinUser="plazi" docAuthor="Winterbottom, Richard & Hoese, Douglass F." docDate="2015" docId="5519879AB837F373FF1FFB6C6F73EE99" docLanguage="en" docName="zt03934p102.pdf" docOrigin="Zootaxa 3934 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Trimma meristum Winterbottom & Hoese, 2015, sp. nov." docType="treatment" docUuid="F3B46252-CC3D-4082-9E35-08A403DAC110" docUuidSource="ZooBank" docVersion="11" lastPageNumber="55" masterDocId="A920FFE2B803F345FF88FFF66B4DEB78" masterDocTitle="A revision of the Australian species of Trimma (Actinopterygii, Gobiidae), with descriptions of six new species and redescriptions of twenty-three valid species" masterLastPageNumber="102" masterPageNumber="1" pageNumber="53" updateTime="1698567702668" updateUser="plazi">
|
||
<mods:mods id="97C5720C965E38BFA8614E3BF2A06BF7" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="FB2695437F057E11094D530E572AA82D">
|
||
<mods:title id="C7F1343462C7D43754EF31CFF0A05B14">A revision of the Australian species of Trimma (Actinopterygii, Gobiidae), with descriptions of six new species and redescriptions of twenty-three valid species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="ECE641DAA0ADF7BD880C0EEEC7DE55EB" type="personal">
|
||
<mods:role id="E9A7177FB18DD92C5A9D812F41F02778">
|
||
<mods:roleTerm id="A5AE7AD9C6F4A50223422E1C930DE5F3">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="CE151940FD166AAC08E5B2ECD24A0BF9">Winterbottom, Richard</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="32DA944F4177DAAF9BF91A6EF153F1EF" type="personal">
|
||
<mods:role id="478DA69DCD45A8830FC6EEA12CD68558">
|
||
<mods:roleTerm id="BE8F596437A278B513914B1DC372BFEE">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="314BBE4DB159E866BAB6336DE238089E">Hoese, Douglass F.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="EB1B22A4F0B32737F9E9D6F4BF3018B2">text</mods:typeOfResource>
|
||
<mods:relatedItem id="37B23BB9B164D127FF3F5A15CB6B37AE" type="host">
|
||
<mods:titleInfo id="094B5E3271D9321911B732A2BF7BA0D1">
|
||
<mods:title id="65F76C82444DE0E0B45523D0067BAD24">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="CEB5CAE21F6AAC1980196C39B4EBED90">
|
||
<mods:date id="BE4A516917712733875BC6AACA0DB873">2015</mods:date>
|
||
<mods:detail id="70F3E7304E21C2671C5474C1F393CE89" type="volume">
|
||
<mods:number id="61692B9A34D03C8183E2A8966CBAA34E">3934</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="4AD65523F18069C174CE48FFDB2A850E" type="issue">
|
||
<mods:number id="0AAE376254D3BF2B431B3A203D40594E">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="708D340ED437CE163E350A571685E254" unit="page">
|
||
<mods:start id="E58DB53F68A2DFD417F621C108898ADE">1</mods:start>
|
||
<mods:end id="D7B9B13707AD2DAE21E058D3EF648564">102</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="0187A25FAA0EC5ACCD0128DA6A83A46C">journal article</mods:classification>
|
||
<mods:identifier id="F8815061778E08CF205F7F4F7397B1CB" type="DOI">10.11646/zootaxa.3934.1.1</mods:identifier>
|
||
<mods:identifier id="E991C174BE17D3B8C8EC9B5446C932DF" type="GBIF-Dataset">0abffe3e-a2c6-4623-b7c6-c8b26cee4a2b</mods:identifier>
|
||
<mods:identifier id="0C8DAD40CE7ACEEBFD4542CC68767814" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="23F2D500B6FA1822AED40ADF062B8E4D" type="Zenodo-Dep">236066</mods:identifier>
|
||
<mods:identifier id="5100B6C518C27257691C65DF49A628FD" type="ZooBank">11C2A2CB-30B3-4694-B379-AE9D47332F0C</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="5519879AB837F373FF1FFB6C6F73EE99" ID-DOI="http://doi.org/10.5281/zenodo.5621524" ID-GBIF-Taxon="119600472" ID-Zenodo-Dep="5621524" LSID="urn:lsid:zoobank.org:act:F3B46252-CC3D-4082-9E35-08A403DAC110" httpUri="http://treatment.plazi.org/id/5519879AB837F373FF1FFB6C6F73EE99" lastPageId="54" lastPageNumber="55" pageId="52" pageNumber="53">
|
||
<subSubSection id="95AA6507B837F371FF1FFB6C6903EE47" pageId="52" pageNumber="53" type="nomenclature">
|
||
<paragraph id="DD0F368CB837F371FF1FFB6C6A9BEFCC" blockId="52.[151,470,1178,1273]" box="[151,470,1178,1204]" pageId="52" pageNumber="53">
|
||
<heading id="864781E0B837F371FF1FFB6C6A9BEFCC" bold="true" box="[151,470,1178,1204]" fontSize="11" level="1" pageId="52" pageNumber="53" reason="1">
|
||
<emphasis id="EFC4EA9EB837F371FF1FFB6C6A9BEFCC" bold="true" box="[151,470,1178,1204]" pageId="52" pageNumber="53">
|
||
<taxonomicName id="1AB04D0FB837F371FF1FFB6C6A3CEFCC" ID-CoL="58V6T" box="[151,369,1178,1204]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="meristum" status="sp. nov.">
|
||
<emphasis id="EFC4EA9EB837F371FF1FFB6C6A3CEFCC" bold="true" box="[151,369,1178,1204]" italics="true" pageId="52" pageNumber="53">Trimma meristum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F4F757E5B837F371FEF0FB6C6A9BEFCC" box="[376,470,1178,1204]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="DD0F368CB837F371FF1FFB4A6A56EF81" blockId="52.[151,470,1178,1273]" pageId="52" pageNumber="53">
|
||
Split-ray Pygmygoby
|
||
<figureCitation id="458B2A09B837F371FF1FFB166A56EF81" box="[151,283,1248,1273]" captionStart-0="FIGURE 27" captionStart-1="FIGURE 28" captionStart-2="FIGURE 29" captionStartId-0="52.[151,250,1055,1077]" captionStartId-1="53.[151,250,1553,1575]" captionStartId-2="54.[151,250,632,654]" captionTargetBox-0="[163,1427,203,1022]" captionTargetBox-1="[162,1435,601,1516]" captionTargetBox-2="[151,1435,193,610]" captionTargetId-0="figure@52.[151,1436,193,1034]" captionTargetId-1="figure@53.[151,1435,589,1531]" captionTargetId-2="figure@54.[151,1435,193,611]" captionTargetPageId-0="52" captionTargetPageId-1="53" captionTargetPageId-2="54" captionText-0="FIGURE 27. Map of the distribution of Trimma maiandros (solid diamonds) and T. meristum (solid square with white center). A solid circle with a white star indicates localities at which both species occur." captionText-1="FIGURE 28. Trimma meristum, 19.0 mm SL male, AMS I. 20793 - 090, dorsal view of head, stained with cyanine blue. Photo and image enhancement by R. Winterbottom." captionText-2="FIGURE 29. Trimma meristum, 19.0 mm SL male (preserved), AMS I. 20793 - 090, Cape York, Queensland. Photo by R. Winterbottom." httpUri-0="https://zenodo.org/record/236094/files/figure.png" httpUri-1="https://zenodo.org/record/236095/files/figure.png" httpUri-2="https://zenodo.org/record/236096/files/figure.png" pageId="52" pageNumber="53">Figs. 27–29</figureCitation>
|
||
</paragraph>
|
||
<paragraph id="DD0F368CB837F371FF1FFADC6903EE47" blockId="52.[151,590,1321,1343]" box="[151,590,1321,1343]" pageId="52" pageNumber="53">No published names pertain to this species.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="95AA6507B837F371FF1FFA9D6801ED40" pageId="52" pageNumber="53" type="materials_examined">
|
||
<paragraph id="DD0F368CB837F371FF1FFA9D6BBCEED0" blockId="52.[151,1437,1387,2024]" pageId="52" pageNumber="53">
|
||
<emphasis id="EFC4EA9EB837F371FF1FFA9D6ACCEEFC" bold="true" box="[151,385,1387,1412]" pageId="52" pageNumber="53">
|
||
Material.
|
||
<typeStatus id="020B882EB837F371FE85FA9D6A31EEFC" box="[269,380,1387,1412]" pageId="52" pageNumber="53" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectionCode id="BBA1AE49B837F371FE00FA9A6A88EEFC" box="[392,453,1388,1412]" pageId="52" pageNumber="53">AMS</collectionCode>
|
||
I.18353–037,
|
||
<quantity id="1A489B69B837F371FDEFFA9A6986EEFB" box="[615,715,1388,1412]" metricMagnitude="-2" metricUnit="m" metricValue="1.83" pageId="52" pageNumber="53" unit="mm" value="18.3">18.3 mm</quantity>
|
||
SL male,
|
||
<collectingCountry id="A5A7761CB837F371FCB6FA9D6825EEFC" box="[830,872,1387,1412]" name="Fiji" pageId="52" pageNumber="53">Fiji</collectingCountry>
|
||
,
|
||
<collectingCountry id="A5A7761CB837F371FCFBFA9D68D0EEFC" box="[883,925,1387,1412]" name="Fiji" pageId="52" pageNumber="53">Viti</collectingCountry>
|
||
Levu, Suva Harbour, Bird Id,
|
||
<date id="A90E104CB837F371FB7AFA9A6E7CEEFC" box="[1266,1329,1387,1412]" pageId="52" pageNumber="53">3 Jul.</date>
|
||
, 1974. B. Russell.
|
||
</paragraph>
|
||
<paragraph id="DD0F368CB837F371FF4FFA456801ED40" blockId="52.[151,1437,1387,2024]" pageId="52" pageNumber="53">
|
||
<emphasis id="EFC4EA9EB837F371FF4FFA456A0BEEB4" bold="true" box="[199,326,1459,1484]" pageId="52" pageNumber="53">
|
||
<typeStatus id="020B882EB837F371FF4FFA456A0CEEB4" box="[199,321,1459,1484]" pageId="52" pageNumber="53" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectionCode id="BBA1AE49B837F371FEC5FA426AC7EEB4" box="[333,394,1460,1484]" pageId="52" pageNumber="53">AMS</collectionCode>
|
||
I.18354-104, (19.1), same locality as
|
||
<typeStatus id="020B882EB837F371FCB0FA4568D0EEB4" box="[824,925,1459,1484]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
|
||
,
|
||
<specimenCount id="CBB6FD05B837F371FC21FA456891EEB3" box="[937,988,1459,1484]" pageId="52" pageNumber="53" type="generic">
|
||
<quantity id="1A489B69B837F371FC21FA456895EEB3" box="[937,984,1459,1484]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="52" pageNumber="53" unit="m" value="6.0">6 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
B. Russell,
|
||
<date id="A90E104CB837F371FBEDFA426FE8EEB4" box="[1125,1189,1459,1484]" pageId="52" pageNumber="53">9 Jul.</date>
|
||
, 1974.
|
||
<collectionCode id="BBA1AE49B837F371FB70FA426E78EEB4" box="[1272,1333,1460,1484]" pageId="52" pageNumber="53">AMS</collectionCode>
|
||
I.20793- 0 90, 2(13.5–19.0),
|
||
<collectingCountry id="A5A7761CB837F371FEF8FA216A97EE88" box="[368,474,1495,1520]" name="Australia" pageId="52" pageNumber="53">Australia</collectingCountry>
|
||
, Queensland, Cape York, NE point of Clack Id (
|
||
<geoCoordinate id="B884504BB837F371FB8AFA2E6F11EE88" box="[1026,1116,1496,1520]" direction="south" orientation="latitude" pageId="52" pageNumber="53" precision="925" value="-14.05">14°03'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B884504BB837F371FBE1FA216F99EE88" box="[1129,1236,1495,1520]" direction="east" orientation="longitude" pageId="52" pageNumber="53" precision="925" value="144.26666">144°16'E</geoCoordinate>
|
||
),
|
||
<quantity id="1A489B69B837F371FB61FA2E6E7EEE97" box="[1257,1331,1496,1520]" metricMagnitude="0" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="52" pageNumber="53" unit="m" value="5.0" valueMax="7.0" valueMin="3.0">3–7 m</quantity>
|
||
,
|
||
<date id="A90E104CB837F371FAC8FA2E6ED5EE88" box="[1344,1432,1495,1520]" pageId="52" pageNumber="53" value="1924-02">24 Feb.</date>
|
||
, 1979,
|
||
<collectionCode id="BBA1AE49B837F371FF6AFA0A6A53ED6C" box="[226,286,1532,1556]" pageId="52" pageNumber="53">AMS</collectionCode>
|
||
/
|
||
<collectionCode id="BBA1AE49B837F371FEAEFA0A6A21ED6C" box="[294,364,1532,1556]" pageId="52" pageNumber="53">AIMS</collectionCode>
|
||
party.
|
||
<collectionCode id="BBA1AE49B837F371FE4AFA0A695EED6C" LSID="urn:lsid:biocol.org:col:34871" box="[450,531,1532,1556]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="52" pageNumber="53">USNM</collectionCode>
|
||
264526, (18.7),
|
||
<collectingCountry id="A5A7761CB837F371FD54FA0A688DED6C" box="[732,960,1531,1556]" name="Papua New Guinea" pageId="52" pageNumber="53">Papua New Guinea</collectingCountry>
|
||
, Ninigo Ids, just SE of Ami Id (
|
||
<geoCoordinate id="B884504BB837F371FAC4FA0A6EDAED6C" box="[1356,1431,1532,1556]" direction="south" orientation="latitude" pageId="52" pageNumber="53" precision="925" value="-1.2333333">1°14'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B884504BB837F371FF1FF9D66A4CED40" box="[151,257,1568,1592]" direction="east" orientation="longitude" pageId="52" pageNumber="53" precision="925" value="144.36667">144°22'E</geoCoordinate>
|
||
), patch reef,
|
||
<quantity id="1A489B69B837F371FE1BF9E96A91ED4F" box="[403,476,1567,1592]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="6.0" metricValueMin="0.0" pageId="52" pageNumber="53" unit="m" value="3.0" valueMax="6.0" valueMin="0.0">0–6 m</quantity>
|
||
,
|
||
<date id="A90E104CB837F371FE6FF9D66970ED40" box="[487,573,1568,1592]" pageId="52" pageNumber="53" value="1922-10">22 Oct.</date>
|
||
, 1978, V. Springer
|
||
<emphasis id="EFC4EA9EB837F371FC9CF9D66801ED40" box="[788,844,1567,1592]" italics="true" pageId="52" pageNumber="53">et al.</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="95AA6507B837F371FF4FF9B56890ED94" pageId="52" pageNumber="53" type="diagnosis">
|
||
<paragraph id="DD0F368CB837F371FF4FF9B56890ED94" blockId="52.[151,1437,1387,2024]" pageId="52" pageNumber="53">
|
||
<emphasis id="EFC4EA9EB837F371FF4FF9B56A0CED24" bold="true" box="[199,321,1603,1628]" pageId="52" pageNumber="53">Diagnosis.</emphasis>
|
||
A species of
|
||
<taxonomicName id="1AB04D0FB837F371FE6EF9B36971ED24" box="[486,572,1605,1628]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
|
||
<emphasis id="EFC4EA9EB837F371FE6EF9B36971ED24" box="[486,572,1605,1628]" italics="true" pageId="52" pageNumber="53">Trimma</emphasis>
|
||
</taxonomicName>
|
||
with a concave bony interorbital less than half pupil-diameter in width; a slight or no groove posterodorsal to the eye; 8–10 scales in the midline of the predorsal; 7 anal-fin rays; fifth pelvic ray branched twice dichotomously and subequal to the fourth in length, with a full basal membrane; epaxial musculature reaching anteromedially to a point in line with the posterior margin of the pupil in adults; and scale pockets on the head and body strongly outlined by brown chromatophores.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="95AA6507B837F370FF4FF9016F1CECFF" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="description">
|
||
<paragraph id="DD0F368CB837F370FF4FF9016F62E96F" blockId="52.[151,1437,1387,2024]" lastBlockId="53.[151,1436,151,536]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
|
||
<emphasis id="EFC4EA9EB837F371FF4FF9016A14EC68" bold="true" box="[199,345,1783,1808]" pageId="52" pageNumber="53">Description.</emphasis>
|
||
Dorsal fins VI +
|
||
<date id="A90E104CB837F371FD97F90E690DEC68" box="[543,576,1784,1808]" pageId="52" pageNumber="53">I 8</date>
|
||
, second spine longest, reaching to base of spine or first ray of second dorsal fin when adpressed, first ray and anterior element of last ray unbranched; anal fin
|
||
<date id="A90E104CB837F371FBA3F8EA6F01EC4B" box="[1067,1100,1820,1843]" pageId="52" pageNumber="53">I 7</date>
|
||
, first ray and anterior element of last ray unbranched; pectoral fin 17 (once 18), uppermost 3–
|
||
<emphasis id="EFC4EA9EB837F371FCD4F8C96827EC20" bold="true" box="[860,874,1855,1880]" pageId="52" pageNumber="53">4</emphasis>
|
||
rays unbranched followed by about 5–
|
||
<emphasis id="EFC4EA9EB837F371FAA8F8C96E63EC20" bold="true" box="[1312,1326,1855,1880]" pageId="52" pageNumber="53">8</emphasis>
|
||
branched rays, ventralmost rays unbranched, fin reaching posteriorly to a vertical with bases of anteriormost few elements of anal fin; pelvic fin
|
||
<date id="A90E104CB837F371FEF8F87E6ADCECD8" box="[368,401,1927,1952]" pageId="52" pageNumber="53">I 5</date>
|
||
, first four rays with a single sequential branch, fifth ray branched twice dichotomously and subequal in length to fourth ray, which reaches posteriorly to base of first anal-fin ray, no fraenum, full basal membrane. Lateral scales 23; transverse scales 8;
|
||
<emphasis id="EFC4EA9EB837F371FD52F83969A5EC90" bold="true" box="[730,744,1999,2024]" pageId="52" pageNumber="53">8</emphasis>
|
||
–10 scales in predorsal midline, reaching to above posterior margin of pupil; scales on pectoral base, breast and anterior belly cycloid, other scales ctenoid; four vertical rows of scales on pectoral base, ventralmost smallest and opposite bases of ventralmost 1–3 fin rays, other scales increase in size dorsally; about 4 scale rows anterior to pelvic fin; no scales seen on cheeks or opercles. Teeth in outer and inner rows of both jaws consist of spaced, enlarged, curved canines, those in inner rows about two-thirds height of outer row teeth, two rows separated by 2–3 irregular rows of small conical teeth. Tongue truncately rounded and pupil-diameter in width. Gill opening extending anteroventrally to below anterior half of pupil; outer gill rakers on first gill arch
|
||
<emphasis id="EFC4EA9EB836F370FE4FFE996A98EAF0" bold="true" box="[455,469,367,392]" pageId="53" pageNumber="54">3</emphasis>
|
||
(once 4) + 15 = 18–19. Anterior nares at the end of a short tube, posterior nares a pore with a raised rim, nasal sac somewhat elevated, nasal apparatus confined to anterior half of snout. Interorbital trough moderate, about as broad as deep with steep sides, grading to shallow or absent posterodorsal to eye; bony interorbital width about equal to half-pupil diameter or less; epaxialis reaching anteriorly to a point above posterior margin of the pupil (
|
||
<figureCitation id="458B2A09B836F370FEF5FE096A9CE960" box="[381,465,511,536]" captionStart="FIGURE 28" captionStartId="53.[151,250,1553,1575]" captionTargetBox="[162,1435,601,1516]" captionTargetId="figure@53.[151,1435,589,1531]" captionTargetPageId="53" captionText="FIGURE 28. Trimma meristum, 19.0 mm SL male, AMS I. 20793 - 090, dorsal view of head, stained with cyanine blue. Photo and image enhancement by R. Winterbottom." httpUri="https://zenodo.org/record/236095/files/figure.png" pageId="53" pageNumber="54">Fig. 28</figureCitation>
|
||
). Vertebral pattern not recorded, but probably
|
||
<typeStatus id="020B882EB836F370FC5FFDF66F42E960" box="[983,1039,512,536]" pageId="53" pageNumber="54">Type</typeStatus>
|
||
B.
|
||
</paragraph>
|
||
<caption id="89CF6604B836F370FF1FF9E7692EED3D" httpUri="https://zenodo.org/record/236095/files/figure.png" pageId="53" pageNumber="54" targetBox="[162,1435,601,1516]" targetPageId="53">
|
||
<paragraph id="DD0F368CB836F370FF1FF9E7692EED3D" blockId="53.[151,1436,1553,1605]" pageId="53" pageNumber="54">
|
||
<emphasis id="EFC4EA9EB836F370FF1FF9E76A6DED5F" bold="true" box="[151,288,1553,1575]" pageId="53" pageNumber="54">FIGURE 28.</emphasis>
|
||
<taxonomicName id="1AB04D0FB836F370FEAFF9E46A96ED5F" box="[295,475,1554,1575]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="meristum">
|
||
<emphasis id="EFC4EA9EB836F370FEAFF9E46A96ED5F" box="[295,475,1554,1575]" italics="true" pageId="53" pageNumber="54">Trimma meristum</emphasis>
|
||
</taxonomicName>
|
||
, 19.0 mm SL male, AMS I.20793-090, dorsal view of head, stained with cyanine blue. Photo and image enhancement by R. Winterbottom.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DD0F368CB836F370FF4FF98469A8EDF3" blockId="53.[151,1436,1650,2035]" box="[199,741,1650,1675]" pageId="53" pageNumber="54">
|
||
<emphasis id="EFC4EA9EB836F370FF4FF9846971EDF3" box="[199,572,1650,1675]" italics="true" pageId="53" pageNumber="54">Colour pattern. Freshly collected</emphasis>
|
||
. Not recorded.
|
||
</paragraph>
|
||
<paragraph id="DD0F368CB836F370FF4FF9606F1CECFF" blockId="53.[151,1436,1650,2035]" pageId="53" pageNumber="54">
|
||
<emphasis id="EFC4EA9EB836F370FF4FF9606A7AEDD7" box="[199,311,1686,1711]" italics="true" pageId="53" pageNumber="54">Preserved</emphasis>
|
||
. Brownish with scale pockets on head and anterior body strongly outlined with dark brown chromatophores, fading posteriorly except in
|
||
<collectingCountry id="A5A7761CB836F370FD10F94C69F2EDAB" box="[664,703,1722,1747]" name="Fiji" pageId="53" pageNumber="54">Fiji</collectingCountry>
|
||
specimens; snout, cheek and dorsal rim of opercle fairly densely and evenly sprinkled with brown chromatophores, ventral portion of opercle and the subopercle without pigmentation; five small (half pupil diameter) saddles across dorsum, first just anterior to base of sixth dorsal spine, second at base of dorsal ray 3, third at base of last ray, and two on the peduncle; pectoral-fin base with evenly distributed brown chromatophores except for a irregularly defined dark bar on the upper two-thirds over the bases of the fin rays, which is made up of densely packed brown chromatophores (
|
||
<figureCitation id="458B2A09B836F370FC78F8986F08ECFF" box="[1008,1093,1902,1927]" captionStart="FIGURE 29" captionStartId="54.[151,250,632,654]" captionTargetBox="[151,1435,193,610]" captionTargetId="figure@54.[151,1435,193,611]" captionTargetPageId="54" captionText="FIGURE 29. Trimma meristum, 19.0 mm SL male (preserved), AMS I. 20793 - 090, Cape York, Queensland. Photo by R. Winterbottom." httpUri="https://zenodo.org/record/236096/files/figure.png" pageId="53" pageNumber="54">Fig. 29</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="95AA6507B836F370FF4FF8646F63ECB7" pageId="53" pageNumber="54" type="etymology">
|
||
<paragraph id="DD0F368CB836F370FF4FF8646F63ECB7" blockId="53.[151,1436,1650,2035]" pageId="53" pageNumber="54">
|
||
<emphasis id="EFC4EA9EB836F370FF4FF8646A03ECD3" bold="true" box="[199,334,1938,1963]" pageId="53" pageNumber="54">Etymology.</emphasis>
|
||
From the Greek ‘meristos’, meaning divided, in allusion to the two dichotomous branches in the fifth pelvic-fin ray resulting in four terminal tips characteristic of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="95AA6507B836F373FF4FF82C698EEE29" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54" type="distribution">
|
||
<paragraph id="DD0F368CB836F370FF4FF82C6ED5EC8A" blockId="53.[151,1436,1650,2035]" box="[199,1432,2010,2035]" pageId="53" pageNumber="54">
|
||
<emphasis id="EFC4EA9EB836F370FF4FF82C6A2DEC8B" bold="true" box="[199,352,2010,2035]" pageId="53" pageNumber="54">Distribution.</emphasis>
|
||
Currently known only from the Great Barrier Reef, the Bismark Archipelago, and
|
||
<collectingCountry id="A5A7761CB836F370FA89F82C6E65EC8B" box="[1281,1320,2010,2035]" name="Fiji" pageId="53" pageNumber="54">Fiji</collectingCountry>
|
||
(
|
||
<figureCitation id="458B2A09B836F370FAB0F82C6EC1EC8A" box="[1336,1420,2010,2035]" captionStart="FIGURE 27" captionStartId="52.[151,250,1055,1077]" captionTargetBox="[163,1427,203,1022]" captionTargetId="figure@52.[151,1436,193,1034]" captionTargetPageId="52" captionText="FIGURE 27. Map of the distribution of Trimma maiandros (solid diamonds) and T. meristum (solid square with white center). A solid circle with a white star indicates localities at which both species occur." httpUri="https://zenodo.org/record/236094/files/figure.png" pageId="53" pageNumber="54">Fig. 27</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="89CF6604B835F373FF1FFD8E6A66E9D5" httpUri="https://zenodo.org/record/236096/files/figure.png" pageId="54" pageNumber="55" targetBox="[151,1435,193,610]" targetPageId="54">
|
||
<paragraph id="DD0F368CB835F373FF1FFD8E6A66E9D5" blockId="54.[151,1436,632,685]" pageId="54" pageNumber="55">
|
||
<emphasis id="EFC4EA9EB835F373FF1FFD8E6A6EE9F6" bold="true" box="[151,291,632,654]" pageId="54" pageNumber="55">FIGURE 29.</emphasis>
|
||
<taxonomicName id="1AB04D0FB835F373FEA5FD8F6AA9E9F6" box="[301,484,633,654]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="meristum">
|
||
<emphasis id="EFC4EA9EB835F373FEA5FD8F6AA9E9F6" box="[301,484,633,654]" italics="true" pageId="54" pageNumber="55">Trimma meristum</emphasis>
|
||
</taxonomicName>
|
||
, 19.0 mm SL male (preserved), AMS I.20793-090, Cape York, Queensland. Photo by R. Winterbottom.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DD0F368CB835F373FF4FFD22698EEE29" blockId="54.[151,1436,724,1505]" pageId="54" pageNumber="55">
|
||
<emphasis id="EFC4EA9EB835F373FF4FFD226A21E995" bold="true" box="[199,364,724,749]" pageId="54" pageNumber="55">Comparisons.</emphasis>
|
||
There are two other undescribed species (one described here as
|
||
<taxonomicName id="1AB04D0FB835F373FBC2FD236F90E995" box="[1098,1245,724,749]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="pentherum" status="sp. nov.">
|
||
<emphasis id="EFC4EA9EB835F373FBC2FD236F90E995" box="[1098,1245,724,749]" italics="true" pageId="54" pageNumber="55">T. pentherum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F4F757E5B835F373FB6EFD226E0CE995" box="[1254,1345,724,749]" pageId="54" pageNumber="55" rank="species">
|
||
<emphasis id="EFC4EA9EB835F373FB6EFD226E0CE995" bold="true" box="[1254,1345,724,749]" pageId="54" pageNumber="55">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and the other currently designated as
|
||
<emphasis id="EFC4EA9EB835F373FE56FD0F6AA6E868" box="[478,491,761,784]" italics="true" pageId="54" pageNumber="55">T</emphasis>
|
||
. RW sp. 69—see below), and
|
||
<taxonomicName id="1AB04D0FB835F373FCB6FD0F6F38E869" authority="Winterbottom 2011" authorityName="Winterbottom" authorityYear="2011" box="[830,1141,760,785]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="cheni">
|
||
<emphasis id="EFC4EA9EB835F373FCB6FD0F68D9E869" box="[830,916,760,785]" italics="true" pageId="54" pageNumber="55">T. cheni</emphasis>
|
||
Winterbottom 2011
|
||
</taxonomicName>
|
||
which have outlined scale pockets, a branched fifth pelvic-fin ray, and at least some branched pectoral-fin rays.
|
||
<taxonomicName id="1AB04D0FB835F373FBC2FCEB6E5DE84C" box="[1098,1296,797,820]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="meristum">
|
||
<emphasis id="EFC4EA9EB835F373FBC2FCEB6E5DE84C" box="[1098,1296,797,820]" italics="true" pageId="54" pageNumber="55">Trimma meristum</emphasis>
|
||
</taxonomicName>
|
||
differs from all three of these species in having only seven anal-fin rays (vs. 8–10) and in the fifth pelvic-fin ray being branched twice and subequal to fourth (vs. branched once and 50–60% length of fourth). It differs further from the currently undescribed
|
||
<taxonomicName id="1AB04D0FB835F373FEACFC7F6A36E8D8" box="[292,379,905,928]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="EFC4EA9EB835F373FEACFC7F6A36E8D8" box="[292,379,905,928]" italics="true" pageId="54" pageNumber="55">Trimma</emphasis>
|
||
</taxonomicName>
|
||
RW sp. 69 (
|
||
<collectingCountry id="A5A7761CB835F373FD8CFC7E6938E8D9" box="[516,629,904,929]" name="Indonesia" pageId="54" pageNumber="55">Indonesia</collectingCountry>
|
||
,
|
||
<collectingCountry id="A5A7761CB835F373FD08FC7E684CE8D9" box="[640,769,904,929]" name="Philippines" pageId="54" pageNumber="55">Philippines</collectingCountry>
|
||
,
|
||
<collectingCountry id="A5A7761CB835F373FC83FC7F68A8E8D9" box="[779,997,904,929]" name="Papua New Guinea" pageId="54" pageNumber="55">Papua New Guinea</collectingCountry>
|
||
, Solomons,
|
||
<collectingCountry id="A5A7761CB835F373FBE4FC7F6F85E8D8" box="[1132,1224,905,928]" name="Vanuatu" pageId="54" pageNumber="55">Vanuatu</collectingCountry>
|
||
and
|
||
<collectingCountry id="A5A7761CB835F373FB75FC7E6E6BE8D9" box="[1277,1318,904,929]" name="Fiji" pageId="54" pageNumber="55">Fiji</collectingCountry>
|
||
) in having one less dorsal-fin ray (8 vs. 9), second spine not elongated (vs. elongated), a scaleless opercle (vs. 2–3 cycloid scales on opercle), and a uniformly pigmented cheek (vs. cheek with two light bars).
|
||
<taxonomicName id="1AB04D0FB835F373FBF7FC276E04E890" box="[1151,1353,977,1000]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="meristum">
|
||
<emphasis id="EFC4EA9EB835F373FBF7FC276E04E890" box="[1151,1353,977,1000]" italics="true" pageId="54" pageNumber="55">Trimma meristum</emphasis>
|
||
</taxonomicName>
|
||
differs further from the other undescribed species and from
|
||
<taxonomicName id="1AB04D0FB835F373FD73FC036819EF75" box="[763,852,1012,1037]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="cheni">
|
||
<emphasis id="EFC4EA9EB835F373FD73FC036819EF75" box="[763,852,1012,1037]" italics="true" pageId="54" pageNumber="55">T. cheni</emphasis>
|
||
</taxonomicName>
|
||
in having 17 (vs. 18–19) pectoral-fin rays, and in having the epaxialis reaching anteriorly to a vertical above the posterior margin of the pupil (vs. posterior margin of the eye). An additional difference from
|
||
<taxonomicName id="1AB04D0FB835F373FDD5FBCB69B7EF2D" box="[605,762,1084,1109]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="cheni">
|
||
<emphasis id="EFC4EA9EB835F373FDD5FBCB69B7EF2D" box="[605,762,1084,1109]" italics="true" pageId="54" pageNumber="55">Trimma cheni</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="A5A7761CB835F373FC82FBCA6836EF2D" box="[778,891,1084,1109]" name="Indonesia" pageId="54" pageNumber="55">Indonesia</collectingCountry>
|
||
,
|
||
<collectingCountry id="A5A7761CB835F373FC00FBCA6F44EF2D" box="[904,1033,1084,1109]" name="Philippines" pageId="54" pageNumber="55">Philippines</collectingCountry>
|
||
,
|
||
<collectingCountry id="A5A7761CB835F373FB9EFBCB6FBEEF2D" box="[1046,1267,1084,1109]" name="Papua New Guinea" pageId="54" pageNumber="55">Papua New Guinea</collectingCountry>
|
||
) is the hyaline dorsal fin (vs. a dark basal stripe), and from the undescribed
|
||
<emphasis id="EFC4EA9EB835F373FCD0FB976828EF00" box="[856,869,1121,1144]" italics="true" pageId="54" pageNumber="55">T</emphasis>
|
||
. RW sp. 54 (Line Islands,
|
||
<collectingCountry id="A5A7761CB835F373FB16FB966E52EF01" box="[1182,1311,1120,1145]" name="Micronesia" pageId="54" pageNumber="55">Micronesia</collectingCountry>
|
||
, Marianas,
|
||
<collectingCountry id="A5A7761CB835F373FF1FFB736A3CEFE5" box="[151,369,1156,1181]" name="Papua New Guinea" pageId="54" pageNumber="55">Papua New Guinea</collectingCountry>
|
||
) in the short second dorsal spine (vs. elongated) and the absence of the trough posterodorsal to the eye (vs. present).
|
||
<taxonomicName id="1AB04D0FB835F373FE03FB5F691EEFB8" box="[395,595,1193,1216]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="meristum">
|
||
<emphasis id="EFC4EA9EB835F373FE03FB5F691EEFB8" box="[395,595,1193,1216]" italics="true" pageId="54" pageNumber="55">Trimma meristum</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="1AB04D0FB835F373FD63FB5F68CDEFB9" box="[747,896,1192,1217]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="pentherum" status="sp. nov.">
|
||
<emphasis id="EFC4EA9EB835F373FD63FB5F68CDEFB9" box="[747,896,1192,1217]" italics="true" pageId="54" pageNumber="55">T. pentherum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F4F757E5B835F373FC00FB5E68AFEFB9" box="[904,994,1192,1217]" pageId="54" pageNumber="55" rank="species">
|
||
<emphasis id="EFC4EA9EB835F373FC00FB5E68AFEFB9" bold="true" box="[904,994,1192,1217]" pageId="54" pageNumber="55">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in the double branched fifth pelvic-fin ray (vs. branched once), branched pectoral-fin rays (vs. unbranched) with four vertical rows of scales on its base (vs. three rows), one more outer gill raker on the lower limb of the first arch (15 vs. 12–14), anterior extent of epaxialis (to posterior pupil vs. posterior eye), and in the less well defined and incomplete vertical bar on the pectoral-fin base (vs. well defined and complete).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="95AA6507B835F373FF4FFAAA6F73EE99" pageId="54" pageNumber="55" type="discussion">
|
||
<paragraph id="DD0F368CB835F373FF4FFAAA6F0FEEC5" blockId="54.[151,1436,724,1505]" pageId="54" pageNumber="55">
|
||
<emphasis id="EFC4EA9EB835F373FF4FFAAA6A01EE0D" bold="true" box="[199,332,1372,1397]" pageId="54" pageNumber="55">Discussion.</emphasis>
|
||
This appears to be a rare species (five known specimens), because it seems to occur in the relatively well-collected shallow waters of corals reefs. The species is normally found on shallow turbid reefs in depths of less than
|
||
<specimenCount id="CBB6FD05B835F373FEE5FA536AE0EEC4" box="[365,429,1445,1469]" pageId="54" pageNumber="55" type="generic">
|
||
<quantity id="1A489B69B835F373FEE5FA536AE4EEC4" box="[365,425,1445,1469]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="54" pageNumber="55" unit="m" value="10.0">10 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
In
|
||
<collectingCountry id="A5A7761CB835F373FE5BFA526977EEC5" box="[467,570,1444,1469]" name="Australia" pageId="54" pageNumber="55">Australia</collectingCountry>
|
||
it is known only from reefs close to the shore.
|
||
</paragraph>
|
||
<paragraph id="DD0F368CB835F373FF4FFA3F6F73EE99" blockId="54.[151,1436,724,1505]" box="[199,1086,1480,1505]" pageId="54" pageNumber="55">
|
||
It has informally been referred to as
|
||
<taxonomicName id="1AB04D0FB835F373FDD7FA3F69FBEE98" box="[607,694,1481,1504]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="EFC4EA9EB835F373FDD7FA3F69FBEE98" box="[607,694,1481,1504]" italics="true" pageId="54" pageNumber="55">Trimma</emphasis>
|
||
</taxonomicName>
|
||
DFH sp. 48 or
|
||
<taxonomicName id="1AB04D0FB835F373FCECFA3F68F7EE98" box="[868,954,1481,1504]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="EFC4EA9EB835F373FCECFA3F68F7EE98" box="[868,954,1481,1504]" italics="true" pageId="54" pageNumber="55">Trimma</emphasis>
|
||
</taxonomicName>
|
||
RW sp. 72.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |