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<document id="54FC78F4BB6F9950F819E93DF7FB4FE7" ID-DOI="10.11646/zootaxa.3934.1.1" ID-GBIF-Dataset="0abffe3e-a2c6-4623-b7c6-c8b26cee4a2b" ID-ISSN="1175-5326" ID-Zenodo-Dep="236066" ID-ZooBank="11C2A2CB-30B3-4694-B379-AE9D47332F0C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461012229557" checkinUser="plazi" docAuthor="Winterbottom, Richard &amp; Hoese, Douglass F." docDate="2015" docId="5519879AB835F37CFF1FF9C269CAE8F8" docLanguage="en" docName="zt03934p102.pdf" docOrigin="Zootaxa 3934 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Trimma milta Winterbottom 2002" docType="treatment" docVersion="11" lastPageNumber="58" masterDocId="A920FFE2B803F345FF88FFF66B4DEB78" masterDocTitle="A revision of the Australian species of Trimma (Actinopterygii, Gobiidae), with descriptions of six new species and redescriptions of twenty-three valid species" masterLastPageNumber="102" masterPageNumber="1" pageNumber="55" updateTime="1698567702668" updateUser="plazi">
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<mods:title id="49F888D017C5948F10AC83F09A366C69">A revision of the Australian species of Trimma (Actinopterygii, Gobiidae), with descriptions of six new species and redescriptions of twenty-three valid species</mods:title>
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<mods:namePart id="C5AAC33AC4B746C0901C8EC66BC5E364">Winterbottom, Richard</mods:namePart>
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<mods:namePart id="791D5702860AB148430D3079DCD0F38F">Hoese, Douglass F.</mods:namePart>
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<mods:date id="9786C655F08828774CBB0FA2CCFE711D">2015</mods:date>
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<paragraph id="DD0F368CB835F373FF1FF9C2690DED36" blockId="54.[151,660,1588,1684]" box="[151,576,1588,1614]" pageId="54" pageNumber="55">
<heading id="864781E0B835F373FF1FF9C2690DED36" bold="true" box="[151,576,1588,1614]" fontSize="11" level="1" pageId="54" pageNumber="55" reason="1">
<taxonomicName id="1AB04D0FB835F373FF1FF9C2690DED36" ID-CoL="7D5HT" authority="Winterbottom, 2002" authorityName="Winterbottom" authorityYear="2002" box="[151,576,1588,1614]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="milta">
<emphasis id="EFC4EA9EB835F373FF1FF9C2690DED36" bold="true" box="[151,576,1588,1614]" pageId="54" pageNumber="55">
<emphasis id="EFC4EA9EB835F373FF1FF9C26A71ED36" bold="true" box="[151,316,1588,1614]" italics="true" pageId="54" pageNumber="55">Trimma milta</emphasis>
<bibRefCitation id="B9214B7DB835F373FECCF9C2690DED36" author="Winterbottom" box="[324,576,1588,1614]" pageId="54" pageNumber="55" refString="Winterbottom, R. (2002) Two new species of Trimma from the central, western and south Pacific. aqua, Journal of Ichthyology and Aquatic Biology, 5 (1), 45 - 52." type="journal article" year="2002">Winterbottom, 2002</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="DD0F368CB835F373FF1FF9A16A39EDEB" blockId="54.[151,660,1588,1684]" pageId="54" pageNumber="55">
Red-earth Pygmygoby (Moorea Pygmygoby)
<figureCitation id="458B2A09B835F373FF1FF98D6A53EDEB" box="[151,286,1659,1684]" captionStart-0="FIGURE 30" captionStart-1="FIGURE 31" captionStartId-0="56.[151,250,702,724]" captionStartId-1="56.[151,250,1748,1770]" captionTargetBox-0="[151,1435,193,680]" captionTargetBox-1="[158,1425,992,1720]" captionTargetId-0="figure@56.[151,1435,193,681]" captionTargetId-1="figure@56.[151,1436,984,1726]" captionTargetPageId-0="56" captionTargetPageId-1="56" captionText-0="FIGURE 30. Trimma milta, 17.6 mm SL female, ROM 85066, Quoy Id., Raja Ampat Ids. Photo by R. Winterbottom." captionText-1="FIGURE 31. Map of the distribution of Trimma milta (solid inverted triangles) and T. nasa (solid circles with white star)." httpUri-0="https://zenodo.org/record/236097/files/figure.png" httpUri-1="https://zenodo.org/record/236098/files/figure.png" pageId="54" pageNumber="55">Figs. 3031</figureCitation>
, Pl. 2 E
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B835F373FF1FF9326AB0EC0A" pageId="54" pageNumber="55" type="reference_group">
<paragraph id="DD0F368CB835F373FF1FF9326A08EC4D" blockId="54.[151,1437,1731,1906]" pageId="54" pageNumber="55">
<treatmentCitationGroup id="FDA011A2B835F373FF1FF9326A08EC4D" pageId="54" pageNumber="55">
<taxonomicName id="1AB04D0FB835F373FF1FF9326952EDA1" ID-CoL="7D5HT" authority="Winterbottom, 2002:46" authorityName="Winterbottom" authorityPageNumber="46" authorityYear="2002" box="[151,543,1731,1753]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="milta">
<emphasis id="EFC4EA9EB835F373FF1FF9326A69EDA1" box="[151,292,1731,1753]" italics="true" pageId="54" pageNumber="55">Trimma milta</emphasis>
<treatmentCitation id="5C11109DB835F373FEA6F9356952EDA1" author="Winterbottom" box="[302,543,1731,1753]" page="46" pageId="54" pageNumber="55" year="2002">
<bibRefCitation id="B9214B7DB835F373FEA6F935694FEDA1" author="Winterbottom" box="[302,514,1731,1753]" pageId="54" pageNumber="55" refString="Winterbottom, R. (2002) Two new species of Trimma from the central, western and south Pacific. aqua, Journal of Ichthyology and Aquatic Biology, 5 (1), 45 - 52." type="journal article" year="2002">Winterbottom, 2002</bibRefCitation>
:46
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</taxonomicName>
(Moorea, Society Ids, widespread in western Pacific, 4 figs.);
<treatmentCitation id="5C11109DB835F373FB48F9356A6AED80" author="Kimura" page="192" pageId="54" pageNumber="55" year="2003">
<bibRefCitation id="B9214B7DB835F373FB48F9356BB0ED80" author="Kimura" pageId="54" pageNumber="55" refString="Kimura, S. &amp; Matsuura, K. (2003) Fishes of Bitung, Northern Tip of Sulawesi, Indonesia. Ocean Research Institute, University of Tokyo, vi + 244 pp." type="book" year="2003">Kimura &amp; Matsuura, 2003</bibRefCitation>
:192
</treatmentCitation>
(Bitung, N. Sulawesi);
<treatmentCitation id="5C11109DB835F373FD9CF9146982ED80" author="Randall" box="[532,719,1762,1784]" page="553" pageId="54" pageNumber="55" year="2005">
<bibRefCitation id="B9214B7DB835F373FD9CF91469E9ED80" author="Randall" box="[532,676,1762,1784]" pageId="54" pageNumber="55" refString="Randall, J. E. (2005) Reef and Shore Fishes of the South Pacific. Univ. of Hawai'i Press, Honolulu, xii + 707 pp." type="book" year="2005">Randall, 2005</bibRefCitation>
:553
</treatmentCitation>
(photo from Hawaii);
<treatmentCitation id="5C11109DB835F373FC3AF9146FE4ED80" author="Bacchet" box="[946,1193,1762,1784]" page="456" pageId="54" pageNumber="55" year="2006">
<bibRefCitation id="B9214B7DB835F373FC3AF9146F33ED80" author="Bacchet" box="[946,1150,1762,1784]" pageId="54" pageNumber="55" refString="Bacchet, P., Zysman, T. &amp; Lefevre, Y. (2006) Guide des Poissons de Tahiti et ses iles. Au Vent Des Iles, Tahiti, 607 pp." type="book" year="2006">
Bacchet
<emphasis id="EFC4EA9EB835F373FB81F9156F71ED80" box="[1033,1084,1762,1784]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
, 2006
</bibRefCitation>
:456
</treatmentCitation>
;
<treatmentCitation id="5C11109DB835F373FB3BF9146ED1ED80" author="Suzuki" box="[1203,1436,1762,1784]" page="77" pageId="54" pageNumber="55" year="2007">
<bibRefCitation id="B9214B7DB835F373FB3BF9146E33ED80" author="Suzuki" box="[1203,1406,1762,1784]" pageId="54" pageNumber="55" refString="Suzuki, T., Senou, H., Shibukawa, K. &amp; Yano, K. (2007 a). First record of a gobiid fish Trimma milta (Pisces, Gobiidae) from Japan. Bulletin of the Biogeographic Society of Japan, 62, 77 - 82. [in Japanese]" type="journal article" year="2007" yearSuffix="a">
Suzuki
<emphasis id="EFC4EA9EB835F373FB77F9156E7FED80" box="[1279,1330,1762,1784]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
, 2007a
</bibRefCitation>
:77
</treatmentCitation>
(first record from
<collectingCountry id="A5A7761CB835F373FE0AF8F76A8DEC6E" box="[386,448,1793,1814]" name="Japan" pageId="54" pageNumber="55">Japan</collectingCountry>
);
<treatmentCitation id="5C11109DB835F373FE5CF8F669B7EC6E" author="Allen" box="[468,762,1792,1814]" page="942" pageId="54" pageNumber="55" year="2012">
<bibRefCitation id="B9214B7DB835F373FE5CF8F6699DEC6E" author="Allen" box="[468,720,1792,1814]" pageId="54" pageNumber="55" refString="Allen, G. R. &amp; Erdmann, M. V. (2012) Reef fishes of the East Indies. Vol. III. Tropical Reef Research, Perth, pp. 857 - 1292." type="book chapter" year="2012">Allen &amp; Erdmann, 2012</bibRefCitation>
:942
</treatmentCitation>
(
<collectingCountry id="A5A7761CB835F373FC82F8F6681BEC6E" box="[778,854,1792,1814]" name="Taiwan" pageId="54" pageNumber="55">Taiwan</collectingCountry>
to Solomon Ids, and
<collectingCountry id="A5A7761CB835F373FBBFF8F66FD5EC6E" box="[1079,1176,1792,1814]" name="Australia" pageId="54" pageNumber="55">Australia</collectingCountry>
);
<treatmentCitation id="5C11109DB835F373FB24F8F76BA0EC4D" author="Motomura" page="338" pageId="54" pageNumber="55" year="2013">
<bibRefCitation id="B9214B7DB835F373FB24F8F76ED4EC6E" author="Motomura" box="[1196,1433,1792,1814]" pageId="54" pageNumber="55" refString="Motomura, H., Dewa, S., Furuta, K. &amp; Matsuura, K. (Eds.) (2013) Fishes of Iou-jima and Take-shima Islands, Mishima, Kagoshima. The Kagoshima University Museum, Kagoshima, 390 pp." type="book" year="2013">
Motomura
<emphasis id="EFC4EA9EB835F373FA97F8F76E19EC6E" box="[1311,1364,1792,1814]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
, 2013
</bibRefCitation>
: 338
</treatmentCitation>
(
<collectingCountry id="A5A7761CB835F373FF72F8E96A74EC4D" box="[250,313,1823,1845]" name="Japan" pageId="54" pageNumber="55">Japan</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="DD0F368CB835F373FF1FF8C96AB0EC0A" blockId="54.[151,1437,1731,1906]" pageId="54" pageNumber="55">
<treatmentCitationGroup id="FDA011A2B835F373FF1FF8C96AB0EC0A" pageId="54" pageNumber="55">
<taxonomicName id="1AB04D0FB835F373FF1FF8C96A7DEC2B" ID-CoL="7CTHS" box="[151,304,1853,1876]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="winchi">
<emphasis id="EFC4EA9EB835F373FF1FF8C96A7DEC2B" box="[151,304,1853,1876]" italics="true" pageId="54" pageNumber="55">Trimma winchi</emphasis>
</taxonomicName>
(
<emphasis id="EFC4EA9EB835F373FEB6F8C96A29EC2C" box="[318,356,1855,1876]" italics="true" pageId="54" pageNumber="55">non</emphasis>
<bibRefCitation id="B9214B7DB835F373FEE3F8CB6976EC2B" author="Winterbottom" box="[363,571,1853,1875]" pageId="54" pageNumber="55" refString="Winterbottom, R. (1984) A review of the gobiid fish genus Trimma from the Chagos Archipelago, central Indian Ocean, with the description of seven new species. Canadian Journal of Zoology, 62, 695 - 715." type="journal article" year="1984">Winterbottom,1984</bibRefCitation>
):
<treatmentCitation id="5C11109DB835F373FDC4F8CB6835EC2B" author="Hayashi" box="[588,888,1853,1875]" page="39" pageId="54" pageNumber="55" year="2003">
<bibRefCitation id="B9214B7DB835F373FDC4F8CB6816EC2B" author="Hayashi" box="[588,859,1853,1875]" pageId="54" pageNumber="55" refString="Hayashi, M. &amp; Shiratori, T. (2003) Gobies of Japanese Waters. Hankyu Books, Osaka, 224 pp. [in Japanese]" type="book" year="2003">Hayashi &amp; Shiratori, 2003</bibRefCitation>
:39
</treatmentCitation>
(identity uncertain).
<taxonomicName id="1AB04D0FB835F373FF1FF8AB6BABEC0A" ID-CoL="827C" box="[151,230,1885,1906]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="genus">
<emphasis id="EFC4EA9EB835F373FF1FF8AB6BABEC0A" box="[151,230,1885,1906]" italics="true" pageId="54" pageNumber="55">Trimma</emphasis>
</taxonomicName>
sp. A:
<bibRefCitation id="B9214B7DB835F373FEA7F8AB6AF9EC0A" author="Myers" box="[303,436,1884,1906]" pageId="54" pageNumber="55" refString="Myers, R. F. (1999) Micronesian Reef Fishes. 3 rd Edition. Coral Graphics, Guam, 330 pp." type="book" year="1999">Myers, 1999</bibRefCitation>
:pl.163.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B835F372FF1FF8696800E96F" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55" type="materials_examined">
<paragraph id="DD0F368CB835F373FF1FF8696A35ECA4" blockId="54.[151,1437,1951,2012]" pageId="54" pageNumber="55">
<emphasis id="EFC4EA9EB835F373FF1FF8696A4BECC0" bold="true" box="[151,262,1951,1976]" pageId="54" pageNumber="55">Material.</emphasis>
In addition to the
<typeStatus id="020B882EB835F373FE79F856696CECC0" box="[497,545,1952,1976]" pageId="54" pageNumber="55">type</typeStatus>
material from the Society Islands, we have examined specimens from the following localities:
</paragraph>
<paragraph id="DD0F368CB834F372FF4FFF616EDEEA1C" blockId="55.[151,1437,151,2030]" pageId="55" pageNumber="56">
<emphasis id="EFC4EA9EB834F372FF4FFF6169E1EBC8" bold="true" box="[199,684,151,176]" pageId="55" pageNumber="56">Australian Material. Great Barrier Reef</emphasis>
: Escape Reef:
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I.22580-021, 6(1720),
<quantity id="1A489B69B834F372FB25FF6E6FA7EBD7" box="[1197,1258,152,176]" metricMagnitude="1" metricUnit="m" metricValue="3.7" pageId="55" pageNumber="56" unit="m" value="37.0">37 m</quantity>
;
<collectionCode id="BBA1AE49B834F372FB7FFF6E6E79EBC8" box="[1271,1332,152,176]" pageId="55" pageNumber="56">AMS</collectionCode>
I.22587- 0 21, 8(1621),
<quantity id="1A489B69B834F372FEC0FF4A6ACBEBAC" box="[328,390,188,212]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="55" pageNumber="56" unit="m" value="40.0">40 m</quantity>
;
<collectionCode id="BBA1AE49B834F372FE1CFF4A6A9CEBAC" box="[404,465,188,212]" pageId="55" pageNumber="56">AMS</collectionCode>
I.22613-043, (1720),
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;
<collectionCode id="BBA1AE49B834F372FCA1FF4A682BEBAC" box="[809,870,188,212]" pageId="55" pageNumber="56">AMS</collectionCode>
I.22627-016, 6(1923),
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;
<collectionCode id="BBA1AE49B834F372FB7FFF4A6E79EBAC" box="[1271,1332,188,212]" pageId="55" pageNumber="56">AMS</collectionCode>
I.22629- 0 0 8, 13(1521),
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;
<collectionCode id="BBA1AE49B834F372FE5BFF16695DEB80" box="[467,528,224,248]" pageId="55" pageNumber="56">AMS</collectionCode>
I.22637-047, (18),
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3
<specimenCount id="CBB6FD05B834F372FC9CFF29681BEB8F" box="[788,854,223,248]" pageId="55" pageNumber="56" type="generic">15 m</specimenCount>
</quantity>
.;
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. Herald Cays:
<collectionCode id="BBA1AE49B834F372FBE7FF166FF9EB80" LSID="urn:lsid:biocol.org:col:34981" box="[1135,1204,224,248]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="55" pageNumber="56">WAM</collectionCode>
P. 28537.029, (18),
<quantity id="1A489B69B834F372FF1FFEF26BB2EA64" box="[151,255,260,285]" metricMagnitude="1" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="55" pageNumber="56" unit="m" value="20.0" valueMax="25.0" valueMin="15.0">
15
<specimenCount id="CBB6FD05B834F372FF48FEF26BB2EA64" box="[192,255,260,285]" pageId="55" pageNumber="56" type="generic">25 m</specimenCount>
</quantity>
. Osprey Reef:
<collectionCode id="BBA1AE49B834F372FE24FEF26AA4EA64" box="[428,489,260,284]" pageId="55" pageNumber="56">AMS</collectionCode>
I.25107-051 3(1721),
<quantity id="1A489B69B834F372FD73FEF2682EEA64" box="[763,867,260,285]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="55" pageNumber="56" unit="m" value="17.5" valueMax="25.0" valueMin="10.0">
10
<specimenCount id="CBB6FD05B834F372FCADFEF2682EEA64" box="[805,867,260,285]" pageId="55" pageNumber="56" type="generic">25 m</specimenCount>
</quantity>
. Raine Id:
<collectionCode id="BBA1AE49B834F372FC6AFEF26F6FEA64" box="[994,1058,260,284]" pageId="55" pageNumber="56">AMS</collectionCode>
. Yonge Reef:
<collectionCode id="BBA1AE49B834F372FB4DFEF26E4FEA64" box="[1221,1282,260,284]" pageId="55" pageNumber="56">AMS</collectionCode>
I.19454-062, 20(1022),
<quantity id="1A489B69B834F372FE92FEDE6A3FEA47" box="[282,370,296,320]" metricMagnitude="0" metricUnit="m" metricValue="9.5" metricValueMax="18.0" metricValueMin="1.0" pageId="55" pageNumber="56" unit="m" value="9.5" valueMax="18.0" valueMin="1.0">
1
<specimenCount id="CBB6FD05B834F372FEBDFEDE6A3FEA47" box="[309,370,296,320]" pageId="55" pageNumber="56" type="generic">18 m</specimenCount>
</quantity>
.
<emphasis id="EFC4EA9EB834F372FEF5FED16912EA38" bold="true" box="[381,607,295,320]" pageId="55" pageNumber="56">
Western
<collectingCountry id="A5A7761CB834F372FE6FFED16917EA38" box="[487,602,295,320]" name="Australia" pageId="55" pageNumber="56">Australia</collectingCountry>
:
</emphasis>
<collectingCountry id="A5A7761CB834F372FDE0FED16982EA38" box="[616,719,295,320]" name="Ashmore and Cartier Islands" pageId="55" pageNumber="56">Ashmore</collectingCountry>
Reef:
<collectionCode id="BBA1AE49B834F372FC91FEDE6813EA38" LSID="urn:lsid:biocol.org:col:34981" box="[793,862,296,320]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="55" pageNumber="56">WAM</collectionCode>
P. 29047.036, 2(1919),
<quantity id="1A489B69B834F372FBF2FED16FADEA47" box="[1146,1248,295,320]" metricMagnitude="1" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="55" pageNumber="56" unit="m" value="22.5" valueMax="25.0" valueMin="20.0">
20
<specimenCount id="CBB6FD05B834F372FB2DFED16FADEA47" box="[1189,1248,295,320]" pageId="55" pageNumber="56" type="generic">25 m</specimenCount>
</quantity>
. Rowley Shoals:
<collectionCode id="BBA1AE49B834F372FF1FFEBA6B91EA1C" LSID="urn:lsid:biocol.org:col:34981" box="[151,220,332,356]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="55" pageNumber="56">WAM</collectionCode>
P. 28024.022, 7(1418),
<quantity id="1A489B69B834F372FE7EFEBA6911EA1C" box="[502,604,332,357]" metricMagnitude="1" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="55" pageNumber="56" unit="m" value="37.5" valueMax="40.0" valueMin="35.0">3540 m</quantity>
;
<collectionCode id="BBA1AE49B834F372FDE0FEBA69E0EA1C" LSID="urn:lsid:biocol.org:col:34981" box="[616,685,332,356]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="55" pageNumber="56">WAM</collectionCode>
P. 28026.026, (22),
<quantity id="1A489B69B834F372FC18FEBA68B8EA1C" box="[912,1013,332,356]" metricMagnitude="1" metricUnit="m" metricValue="3.65" metricValueMax="4.0" metricValueMin="3.3" pageId="55" pageNumber="56" unit="m" value="36.5" valueMax="40.0" valueMin="33.0">3340 m</quantity>
;
<collectionCode id="BBA1AE49B834F372FB89FEBA6F0BEA1C" LSID="urn:lsid:biocol.org:col:34981" box="[1025,1094,332,356]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="55" pageNumber="56">WAM</collectionCode>
P. 28034.016, (17),
<quantity id="1A489B69B834F372FAA1FEBA6EC2EA1C" box="[1321,1423,332,356]" metricMagnitude="1" metricUnit="m" metricValue="3.65" metricValueMax="4.0" metricValueMin="3.3" pageId="55" pageNumber="56" unit="m" value="36.5" valueMax="40.0" valueMin="33.0">
33
<specimenCount id="CBB6FD05B834F372FADAFEBA6EC2EA1C" box="[1362,1423,332,356]" pageId="55" pageNumber="56" type="generic">40 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="DD0F368CB834F372FF4FFE996800E96F" blockId="55.[151,1437,151,2030]" pageId="55" pageNumber="56">
<emphasis id="EFC4EA9EB834F372FF4FFE996A86EAF0" bold="true" box="[199,459,367,392]" pageId="55" pageNumber="56">
Other Material.
<collectingCountry id="A5A7761CB834F372FE10FE996A8BEAF0" box="[408,454,367,392]" name="Fiji" pageId="55" pageNumber="56">Fiji</collectingCountry>
:
</emphasis>
<collectingCountry id="A5A7761CB834F372FE50FE99694FEAF0" box="[472,514,367,392]" name="Fiji" pageId="55" pageNumber="56">Viti</collectingCountry>
Levu:
<collectionCode id="BBA1AE49B834F372FDD5FE8669D0EAF0" box="[605,669,368,392]" pageId="55" pageNumber="56">AMS</collectionCode>
,
<collectionCode id="BBA1AE49B834F372FD26FE8669BCEAF0" LSID="urn:lsid:biocol.org:col:34778" box="[686,753,368,392]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="55" pageNumber="56">ROM</collectionCode>
.
<emphasis id="EFC4EA9EB834F372FC8AFE99681AEAF0" bold="true" box="[770,855,367,392]" pageId="55" pageNumber="56">Hawaii</emphasis>
:
<collectionCode id="BBA1AE49B834F372FCE4FE8668F0EAFF" LSID="urn:lsid:biocol.org:col:34697" box="[876,957,368,391]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="55" pageNumber="56">BPBM</collectionCode>
,
<collectionCode id="BBA1AE49B834F372FC46FE866F5CEAF0" LSID="urn:lsid:biocol.org:col:34778" box="[974,1041,368,392]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="55" pageNumber="56">ROM</collectionCode>
.
<emphasis id="EFC4EA9EB834F372FBAAFE996FECEAF0" bold="true" box="[1058,1185,367,392]" pageId="55" pageNumber="56">
<collectingCountry id="A5A7761CB834F372FBAAFE996FD6EAF0" box="[1058,1179,367,392]" name="Indonesia" pageId="55" pageNumber="56">Indonesia</collectingCountry>
:
</emphasis>
Lucipara Ids:
<collectionCode id="BBA1AE49B834F372FADDFE866ED5EAF0" LSID="urn:lsid:biocol.org:col:34778" box="[1365,1432,368,392]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="55" pageNumber="56">ROM</collectionCode>
.
<collectingCountry id="A5A7761CB834F372FF1FFE626A45EAD5" box="[151,264,404,429]" name="Indonesia" pageId="55" pageNumber="56">Moluccas</collectingCountry>
:
<collectionCode id="BBA1AE49B834F372FE9DFE626A27EAD4" LSID="urn:lsid:biocol.org:col:34871" box="[277,362,404,428]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="55" pageNumber="56">USNM</collectionCode>
. Raja Ampat Ids:
<collectionCode id="BBA1AE49B834F372FDB5FE6269CDEAD4" LSID="urn:lsid:biocol.org:col:34778" box="[573,640,404,428]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="55" pageNumber="56">ROM</collectionCode>
.
<emphasis id="EFC4EA9EB834F372FD05FE626843EAD5" bold="true" box="[653,782,404,429]" pageId="55" pageNumber="56">Marshalls:</emphasis>
Enewetok:
<collectionCode id="BBA1AE49B834F372FC11FE6368A7EAD4" LSID="urn:lsid:biocol.org:col:34697" box="[921,1002,405,428]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="55" pageNumber="56">BPBM</collectionCode>
. Kwajalein:
<collectionCode id="BBA1AE49B834F372FBF4FE636F80EAD4" LSID="urn:lsid:biocol.org:col:34697" box="[1148,1229,405,428]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="55" pageNumber="56">BPBM</collectionCode>
.
<emphasis id="EFC4EA9EB834F372FB52FE626E25EAD5" bold="true" box="[1242,1384,404,429]" pageId="55" pageNumber="56">
<collectingCountry id="A5A7761CB834F372FB52FE626E29EAD5" box="[1242,1380,404,429]" name="Micronesia" pageId="55" pageNumber="56">Micronesia</collectingCountry>
:
</emphasis>
Ant Atoll:
<collectionCode id="BBA1AE49B834F372FF6AFE4E6A7AEAA8" LSID="urn:lsid:biocol.org:col:34871" box="[226,311,440,464]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="55" pageNumber="56">USNM</collectionCode>
.
<collectingCountry id="A5A7761CB834F372FECBFE4E6AD7EAB7" box="[323,410,440,463]" name="Micronesia" pageId="55" pageNumber="56">Ponape</collectingCountry>
:
<collectionCode id="BBA1AE49B834F372FE2FFE4E6AB6EAA8" LSID="urn:lsid:biocol.org:col:34871" box="[423,507,440,464]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="55" pageNumber="56">USNM</collectionCode>
. Ulithi Atoll:
<collectionCode id="BBA1AE49B834F372FD16FE4E69A2EAB7" LSID="urn:lsid:biocol.org:col:34697" box="[670,751,440,463]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="55" pageNumber="56">BPBM</collectionCode>
.
<emphasis id="EFC4EA9EB834F372FD74FE416807EAA8" bold="true" box="[764,842,439,464]" pageId="55" pageNumber="56">
<collectingCountry id="A5A7761CB834F372FD74FE416808EAA8" box="[764,837,439,464]" name="Palau" pageId="55" pageNumber="56">Palau</collectingCountry>
:
</emphasis>
<collectionCode id="BBA1AE49B834F372FCDBFE4E68DEEAA8" box="[851,915,440,464]" pageId="55" pageNumber="56">AMS</collectionCode>
,
<collectionCode id="BBA1AE49B834F372FC28FE4E68BCEAB7" LSID="urn:lsid:biocol.org:col:34697" box="[928,1009,440,463]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="55" pageNumber="56">BPBM</collectionCode>
,
<collectionCode id="BBA1AE49B834F372FC76FE4E6F0CEAA8" LSID="urn:lsid:biocol.org:col:34778" box="[1022,1089,440,464]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="55" pageNumber="56">ROM</collectionCode>
.
<emphasis id="EFC4EA9EB834F372FBC7FE416E0EEAA8" bold="true" box="[1103,1347,439,464]" pageId="55" pageNumber="56">
<collectingCountry id="A5A7761CB834F372FBC7FE416E72EAA8" box="[1103,1343,439,464]" name="Papua New Guinea" pageId="55" pageNumber="56">Papua New Guinea</collectingCountry>
:
</emphasis>
Hermit Ids:
<collectionCode id="BBA1AE49B834F372FF41FE2A6A50EA8C" LSID="urn:lsid:biocol.org:col:34871" box="[201,285,476,500]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="55" pageNumber="56">USNM</collectionCode>
. Manus Id:
<collectionCode id="BBA1AE49B834F372FE2CFE2A6AA1EA8C" LSID="urn:lsid:biocol.org:col:34981" box="[420,492,476,500]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="55" pageNumber="56">WAM</collectionCode>
. Rabaul:
<collectionCode id="BBA1AE49B834F372FDDFFE2A69D6EA8C" LSID="urn:lsid:biocol.org:col:34778" box="[599,667,476,500]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="55" pageNumber="56">ROM</collectionCode>
.
<emphasis id="EFC4EA9EB834F372FD2EFE2A6878EA8D" bold="true" box="[678,821,476,501]" pageId="55" pageNumber="56">
<collectingCountry id="A5A7761CB834F372FD2EFE2A687DEA8D" box="[678,816,476,501]" name="Philippines" pageId="55" pageNumber="56">Philippines</collectingCountry>
:
</emphasis>
Balicasag Id:
<collectionCode id="BBA1AE49B834F372FC51FE2A6F63EA8C" LSID="urn:lsid:biocol.org:col:34871" box="[985,1070,476,500]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="55" pageNumber="56">USNM</collectionCode>
. Cocoro Id:
<collectionCode id="BBA1AE49B834F372FB33FE2A6E5DEA8C" LSID="urn:lsid:biocol.org:col:34871" box="[1211,1296,476,500]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="55" pageNumber="56">USNM</collectionCode>
. Siquijor Id:
<collectionCode id="BBA1AE49B834F372FF1FFDF66B97E960" LSID="urn:lsid:biocol.org:col:34778" box="[151,218,512,536]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="55" pageNumber="56">ROM</collectionCode>
,
<collectionCode id="BBA1AE49B834F372FF6CFDF66A74E960" LSID="urn:lsid:biocol.org:col:34871" box="[228,313,512,536]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="55" pageNumber="56">USNM</collectionCode>
.
<emphasis id="EFC4EA9EB834F372FECCFE096A8DE960" bold="true" box="[324,448,511,536]" pageId="55" pageNumber="56">Solomons:</emphasis>
Florida Id:
<collectionCode id="BBA1AE49B834F372FDCCFDF669C9E960" box="[580,644,512,536]" pageId="55" pageNumber="56">AMS</collectionCode>
.
<emphasis id="EFC4EA9EB834F372FD07FE0969A3E960" bold="true" box="[655,750,511,536]" pageId="55" pageNumber="56">
<collectingCountry id="A5A7761CB834F372FD07FE0969A9E960" box="[655,740,511,536]" name="Taiwan" pageId="55" pageNumber="56">Taiwan</collectingCountry>
:
</emphasis>
<collectionCode id="BBA1AE49B834F372FD7FFDF66804E96F" LSID="urn:lsid:biocol.org:col:34697" box="[759,841,512,535]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="55" pageNumber="56">BPBM</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B834F372FF4FFDD26F22E988" pageId="55" pageNumber="56" type="diagnosis">
<paragraph id="DD0F368CB834F372FF4FFDD26F22E988" blockId="55.[151,1437,151,2030]" pageId="55" pageNumber="56">
<emphasis id="EFC4EA9EB834F372FF4FFDD26A0CE945" bold="true" box="[199,321,548,573]" pageId="55" pageNumber="56">Diagnosis.</emphasis>
A species of
<taxonomicName id="1AB04D0FB834F372FE5FFDD36960E944" box="[471,557,549,572]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="55" pageNumber="56" phylum="Chordata" rank="genus">
<emphasis id="EFC4EA9EB834F372FE5FFDD36960E944" box="[471,557,549,572]" italics="true" pageId="55" pageNumber="56">Trimma</emphasis>
</taxonomicName>
with a concave bony interorbital less than half the pupil-diameter in width, the scale pockets strongly outlined with chromatophores, 68 scales in the midline of the predorsal region and 23 cycloid scales on the upper margin of the opercle, fifth pelvic-fin ray unbranched and about half the length of the fourth, no trough or groove posterodorsal to the eye, epaxial musculature reaching anteromedially to a point in line with the posterior margins of the pupils in adults, and no greatly elongated spines in the first dorsal fin (longest spine reaching to, or slightly beyond, origin of second dorsal fin when fin is adpressed).
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B834F372FF4FFD0A6956EC95" pageId="55" pageNumber="56" type="description">
<paragraph id="DD0F368CB834F372FF4FFD0A6BA2EE17" blockId="55.[151,1437,151,2030]" pageId="55" pageNumber="56">
<emphasis id="EFC4EA9EB834F372FF4FFD0A6A14E86D" bold="true" box="[199,345,764,789]" pageId="55" pageNumber="56">Description.</emphasis>
The description is based primarily on the
<typeStatus id="020B882EB834F372FCB7FD0A68E9E86D" box="[831,932,764,789]" pageId="55" pageNumber="56" type="holotype">holotype</typeStatus>
, and 19
<typeStatus id="020B882EB834F372FB8FFD0B6F3FE86D" box="[1031,1138,765,789]" pageId="55" pageNumber="56" type="paratype">paratypes</typeStatus>
(
<specimenCount id="CBB6FD05B834F372FB0CFD0A6F98E86D" box="[1156,1237,764,789]" pageId="55" pageNumber="56" type="male">9 male</specimenCount>
,
<specimenCount id="CBB6FD05B834F372FB6AFD0A6E18E86D" box="[1250,1365,764,789]" pageId="55" pageNumber="56" type="female">10 female</specimenCount>
) from ROM 59751, and taken from
<bibRefCitation id="B9214B7DB834F372FE63FCE96990E840" author="Winterbottom" box="[491,733,799,824]" pageId="55" pageNumber="56" refString="Winterbottom, R. (2002) Two new species of Trimma from the central, western and south Pacific. aqua, Journal of Ichthyology and Aquatic Biology, 5 (1), 45 - 52." type="journal article" year="2002">Winterbottom (2002)</bibRefCitation>
. Dorsal fins VI +
<date id="A90E104CB834F372FC27FCD6689DE840" box="[943,976,800,824]" pageId="55" pageNumber="56">I 9</date>
, second and third spines longest but not elongated, reaching about to origin of second dorsal fin when adpressed, first ray of second dorsal usually branched; anal fin
<date id="A90E104CB834F372FEE1FC916AC4E807" box="[361,393,871,895]" pageId="55" pageNumber="56">I 8</date>
, first ray usually unbranched; pectoral fin 1718 (mean = 17.3), at least upper 3 and lower 4 unbranched, although all rays may be unbranched in small specimens (about
<quantity id="1A489B69B834F372FB8CFC7C6F1EE8D9" box="[1028,1107,906,930]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="55" pageNumber="56" unit="mm" value="11.0">11 mm</quantity>
SL); pelvic fin
<date id="A90E104CB834F372FA81FC7C6E66E8DA" box="[1289,1323,905,930]" pageId="55" pageNumber="56">I 5</date>
, first four rays with single sequential branching pattern, fifth ray unbranched, and about half length of longer of two branches of fourth ray, no fraenum, basal membrane restricted to distal one-tenth of inter-pelvic gap. Lateral scales 22 (1), 23, 24 (mean = 23.1); predorsal scales 6, 7, 8 (mean = 7.1); transverse scales 6.5; nape, opercle, pectoral-fin base, breast and midline of belly with cycloid scales, ctenoid scales posterior to line between upper opercular margin and origin of first dorsal fin; opercle with a single horizontal row of 23 scales across upper margin; pectoral-fin base margined by four scales. Teeth of outer row of upper jaw, and outer and inner rows of lower jaw consist of enlarged, curved, evenly spaced canines, with several irregular rows of small conical teeth behind outer rows. Tongue broadly rounded, about half pupil-diameter in width. Gill opening to below midpoint of pupil; outer gill rakers of first gill arch 34 + 1215 (
<typeStatus id="020B882EB834F372FDB3FB3F69D2EF9A" box="[571,671,1225,1250]" pageId="55" pageNumber="56" type="holotype">holotype</typeStatus>
14, mean = 13.7). Anterior nares a short tube, posterior nares porelike with raised rim, nasal sac slightly raised, with nasal apparatus confined to anterior half of snout. A shallow interorbital trough; none posterodorsal to eye; bony interorbital width equal to less than half-pupil diameter; epaxialis musculature reaching anteriorly to above posterior margin of pupil. Abdominal/caudal vertebral transition
<typeStatus id="020B882EB834F372FF1FFAAE6B82EE08" box="[151,207,1368,1392]" pageId="55" pageNumber="56">Type</typeStatus>
B.
</paragraph>
<paragraph id="DD0F368CB834F372FF4FFA8C68E7EC18" blockId="55.[151,1437,151,2030]" pageId="55" pageNumber="56">
<emphasis id="EFC4EA9EB834F372FF4FFA8C6970EEEB" box="[199,573,1402,1427]" italics="true" pageId="55" pageNumber="56">Colour pattern. Freshly collected</emphasis>
. Based on
<quantity id="1A489B69B834F372FD32FA8C6844EEEA" box="[698,777,1402,1427]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="55" pageNumber="56" unit="mm" value="35.0">35 mm</quantity>
colour slides of
<specimenCount id="CBB6FD05B834F372FC4BFA8D6F51EEEB" box="[963,1052,1402,1427]" pageId="55" pageNumber="56" type="male">4 males</specimenCount>
,
<quantity id="1A489B69B834F372FBA0FA8D6F86EEEA" box="[1064,1227,1403,1427]" metricMagnitude="-2" metricUnit="m" metricValue="2.065" metricValueMax="2.21" metricValueMin="1.92" pageId="55" pageNumber="56" unit="mm" value="20.65" valueMax="22.1" valueMin="19.2">19.222.1 mm</quantity>
SL and
<specimenCount id="CBB6FD05B834F372FAA1FA8D6ED5EEEB" box="[1321,1432,1402,1427]" pageId="55" pageNumber="56" type="female">4 females</specimenCount>
,
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SL). Overall colour brownish red with centres of scales lighter orange-red, mingled with melanophores, especially on head and anterior body. First dorsal fin with row of yellow spots just distal to base of fin, one spot on each spine, or spots tending to form or forming a yellow stripe, rest of fin membrane sprinkled with melanophores. Second dorsal fin with similar row of yellow spots on each fin ray (not forming a stripe); some individuals with additional but more diffuse row of yellow spots centred on fin rays in region of first branch; rest of fin sprinkled with melanophores. Caudal-fin membrane with melanophores, and with 03 vertical rows of light yellow spots. Anal fin with melanophores, sometimes diffuse light yellow band or few yellow spots just distal to fin base. Pectoral and pelvic fins hyaline, iris red with dark patches. Two very diffuse lighter bars under eye, third along vertical limb of preopercle. One specimen (female,
<quantity id="1A489B69B834F372FC93F94C6833EDA9" box="[795,894,1722,1746]" metricMagnitude="-2" metricUnit="m" metricValue="1.92" pageId="55" pageNumber="56" unit="mm" value="19.2">19.2 mm</quantity>
SL) more yellowish than others, with three light cheek bars better defined (see also
<figureCitation id="458B2A09B834F372FD97F92B693FED8D" box="[543,626,1757,1782]" captionStart="FIGURE 30" captionStartId="56.[151,250,702,724]" captionTargetBox="[151,1435,193,680]" captionTargetId="figure@56.[151,1435,193,681]" captionTargetPageId="56" captionText="FIGURE 30. Trimma milta, 17.6 mm SL female, ROM 85066, Quoy Id., Raja Ampat Ids. Photo by R. Winterbottom." httpUri="https://zenodo.org/record/236097/files/figure.png" pageId="55" pageNumber="56">Fig. 30</figureCitation>
). Raja Ampat female (
<quantity id="1A489B69B834F372FCE7F92B6899ED8D" box="[879,980,1757,1782]" metricMagnitude="-2" metricUnit="m" metricValue="1.76" pageId="55" pageNumber="56" unit="mm" value="17.6">17.6 mm</quantity>
SL) with translucent pale pink body and margins of scales (especially anteriorly) outlined with magenta, lighter bars on cheek and along vertical limb of preopercle are yellow (
<figureCitation id="458B2A09B834F372FE14F8D26ABFEC44" box="[412,498,1828,1853]" captionStart="FIGURE 30" captionStartId="56.[151,250,702,724]" captionTargetBox="[151,1435,193,680]" captionTargetId="figure@56.[151,1435,193,681]" captionTargetPageId="56" captionText="FIGURE 30. Trimma milta, 17.6 mm SL female, ROM 85066, Quoy Id., Raja Ampat Ids. Photo by R. Winterbottom." httpUri="https://zenodo.org/record/236097/files/figure.png" pageId="55" pageNumber="56">Fig. 30</figureCitation>
). This is similar to live specimen from Wetar,
<collectingCountry id="A5A7761CB834F372FB89F8D26F23EC45" box="[1025,1134,1828,1853]" name="Indonesia" pageId="55" pageNumber="56">Indonesia</collectingCountry>
(Pl. 2 D), which differs in having scale margins brown and in lacking yellow bars below the eye.
</paragraph>
<paragraph id="DD0F368CB834F372FF4FF89D6956EC95" blockId="55.[151,1437,151,2030]" pageId="55" pageNumber="56">
<emphasis id="EFC4EA9EB834F372FF4FF89D6A7AECFC" box="[199,311,1899,1924]" italics="true" pageId="55" pageNumber="56">Preserved</emphasis>
. Scale margins, pockets, and head with dark brown chromatophores and melanophores, centres of scales and yellow spots/stripe pale. Degree of chromatophore/melanophore intensity decreases posteriorly along body, where some specimens are pale straw-coloured. Two to four narrow transverse bands of melanophores across dorsal rim of orbit usually present.
</paragraph>
</subSubSection>
<caption id="89CF6604B83BF37DFF1FFD486E0FE9AC" box="[151,1346,702,724]" httpUri="https://zenodo.org/record/236097/files/figure.png" pageId="56" pageNumber="57" targetBox="[151,1435,193,680]" targetPageId="56">
<paragraph id="DD0F368CB83BF37DFF1FFD486E0FE9AC" blockId="56.[151,1346,702,724]" box="[151,1346,702,724]" pageId="56" pageNumber="57">
<emphasis id="EFC4EA9EB83BF37DFF1FFD486A52E9AC" bold="true" box="[151,287,702,724]" pageId="56" pageNumber="57">FIGURE 30.</emphasis>
<taxonomicName id="1AB04D0FB83BF37DFEADFD496AE2E9AC" box="[293,431,702,724]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="milta">
<emphasis id="EFC4EA9EB83BF37DFEADFD496AE2E9AC" box="[293,431,702,724]" italics="true" pageId="56" pageNumber="57">Trimma milta</emphasis>
</taxonomicName>
, 17.6 mm SL female, ROM 85066, Quoy Id., Raja Ampat Ids. Photo by R. Winterbottom.
</paragraph>
</caption>
<subSubSection id="95AA6507B83BF37DFF4FFD0D6801E840" pageId="56" pageNumber="57" type="etymology">
<paragraph id="DD0F368CB83BF37DFF4FFD0D6801E840" blockId="56.[151,1436,763,932]" pageId="56" pageNumber="57">
<emphasis id="EFC4EA9EB83BF37DFF4FFD0D6A03E86C" bold="true" box="[199,334,763,788]" pageId="56" pageNumber="57">Etymology.</emphasis>
From the Greek 'miltos' for red earth, in allusion to the predominant ground colour of the species and the colour of the topsoil of the islands in the
<typeStatus id="020B882EB83BF37DFD33FCD669A6E840" box="[699,747,800,824]" pageId="56" pageNumber="57">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B83BF37CFF4FFCB5698BEAA8" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57" type="distribution">
<paragraph id="DD0F368CB83BF37DFF4FFCB56A21E8DC" blockId="56.[151,1436,763,932]" pageId="56" pageNumber="57">
<emphasis id="EFC4EA9EB83BF37DFF4FFCB56A2DE824" bold="true" box="[199,352,835,860]" pageId="56" pageNumber="57">Distribution.</emphasis>
Recorded from numerous Pacific localities, from Rowley Shoals north to southern
<collectingCountry id="A5A7761CB83BF37DFACBFCB56ED5E824" box="[1347,1432,835,860]" name="Taiwan" pageId="56" pageNumber="57">Taiwan</collectingCountry>
, Hawaii, Society Islands (where it is numerically the dominate species of the genus), west to northern
<collectingCountry id="A5A7761CB83BF37DFA8EFC916E20E8F8" box="[1286,1389,871,896]" name="Australia" pageId="56" pageNumber="57">Australia</collectingCountry>
and Sulawesi (
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).
</paragraph>
<caption id="89CF6604B83BF37DFF1FF9226E25ED92" box="[151,1384,1748,1770]" httpUri="https://zenodo.org/record/236098/files/figure.png" pageId="56" pageNumber="57" targetBox="[158,1425,992,1720]" targetPageId="56">
<paragraph id="DD0F368CB83BF37DFF1FF9226E25ED92" blockId="56.[151,1384,1748,1770]" box="[151,1384,1748,1770]" pageId="56" pageNumber="57">
<emphasis id="EFC4EA9EB83BF37DFF1FF9226A52ED92" bold="true" box="[151,287,1748,1770]" pageId="56" pageNumber="57">FIGURE 31.</emphasis>
Map of the distribution of
<taxonomicName id="1AB04D0FB83BF37DFDB9F92369F7ED92" box="[561,698,1748,1770]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="milta">
<emphasis id="EFC4EA9EB83BF37DFDB9F92369F7ED92" box="[561,698,1748,1770]" italics="true" pageId="56" pageNumber="57">Trimma milta</emphasis>
</taxonomicName>
(solid inverted triangles) and
<taxonomicName id="1AB04D0FB83BF37DFC63F9236F7FED92" box="[1003,1074,1749,1770]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="nasa">
<emphasis id="EFC4EA9EB83BF37DFC63F9236F7FED92" box="[1003,1074,1749,1770]" italics="true" pageId="56" pageNumber="57">T. nasa</emphasis>
</taxonomicName>
(solid circles with white star).
</paragraph>
</caption>
<paragraph id="DD0F368CB83BF37CFF4FF8E7688EEBAD" blockId="56.[151,1436,1809,2014]" lastBlockId="57.[151,1437,151,896]" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
<emphasis id="EFC4EA9EB83BF37DFF4FF8E76A21EC52" bold="true" box="[199,364,1809,1834]" pageId="56" pageNumber="57">Comparisons.</emphasis>
This species belongs to a complex of 10 species possessing a scaled predorsal region and scale pockets on the body outlined with chromatophores and/or melanophores. Three species in the complex have been named,
<taxonomicName id="1AB04D0FB83BF37DFF7AF8AC6934EC0A" authority="Winterbottom, 2006" authorityName="Winterbottom" authorityYear="2006" box="[242,633,1881,1906]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="anthrenum">
<emphasis id="EFC4EA9EB83BF37DFF7AF8AC6AC5EC0A" box="[242,392,1881,1906]" italics="true" pageId="56" pageNumber="57">T. anthrenum</emphasis>
<bibRefCitation id="B9214B7DB83BF37DFE19F8AF6934EC0A" author="Winterbottom" box="[401,633,1881,1906]" pageId="56" pageNumber="57" refString="Winterbottom, R. (2006) Two new species of the gobiid fish Trimma from the coral reefs of the western Pacific Ocean (Pisces; Perciformes; Gobioidei). Zootaxa, 1331, 55 - 68." type="journal article" year="2006">Winterbottom, 2006</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="1AB04D0FB83BF37DFD0EF8AC69BDEC09" box="[646,752,1882,1905]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="emeryi">
<emphasis id="EFC4EA9EB83BF37DFD0EF8AC69BDEC09" box="[646,752,1882,1905]" italics="true" pageId="56" pageNumber="57">T. emeryi</emphasis>
</taxonomicName>
and
<taxonomicName id="1AB04D0FB83BF37DFCA4F8AC68F6EC0A" box="[812,955,1881,1906]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="preclarum">
<emphasis id="EFC4EA9EB83BF37DFCA4F8AC68F6EC0A" box="[812,955,1881,1906]" italics="true" pageId="56" pageNumber="57">T. preclarum</emphasis>
</taxonomicName>
(see
<bibRefCitation id="B9214B7DB83BF37DFC75F8AF6FACEC0A" author="Winterbottom" box="[1021,1249,1881,1906]" pageId="56" pageNumber="57" refString="Winterbottom, R. (2006) Two new species of the gobiid fish Trimma from the coral reefs of the western Pacific Ocean (Pisces; Perciformes; Gobioidei). Zootaxa, 1331, 55 - 68." type="journal article" year="2006">Winterbottom, 2006</bibRefCitation>
for descriptions and comparisons).
<taxonomicName id="1AB04D0FB83BF37DFEFDF888695FECEE" box="[373,530,1917,1942]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="milta">
<emphasis id="EFC4EA9EB83BF37DFEFDF888695FECEE" box="[373,530,1917,1942]" italics="true" pageId="56" pageNumber="57">Trimma milta</emphasis>
</taxonomicName>
is the only species in the complex which possesses opercular scales and an unbranched fifth pelvic fin-ray (occasional specimens of
<taxonomicName id="1AB04D0FB83BF37DFC9CF85468EEECC2" box="[788,931,1953,1978]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="preclarum">
<emphasis id="EFC4EA9EB83BF37DFC9CF85468EEECC2" box="[788,931,1953,1978]" italics="true" pageId="56" pageNumber="57">T. preclarum</emphasis>
</taxonomicName>
may have an unbranched fifth pelvic ray in a few localities). It can be further distinguished from an undescribed species which has opercular scales and an unbranched fifth pelvic-fin ray by the lack of a elongated second spine of the first dorsal fin and in having fewer lower gill rakers on the outer surface of the first gill arch (1314 vs. 16).
</paragraph>
<paragraph id="DD0F368CB83AF37CFF4FFF29698BEAA8" blockId="57.[151,1437,151,896]" pageId="57" pageNumber="58">
Analysis of the CO1 genome of
<specimenCount id="CBB6FD05B83AF37CFDB8FF166985EB80" box="[560,712,223,248]" pageId="57" pageNumber="58" type="generic">21 specimens</specimenCount>
resulted in the identification of four haplogroups, three of which are currently restricted to a single area (
<bibRefCitation id="B9214B7DB83AF37CFDDCFEF2683BEA64" author="Winterbottom" box="[596,886,260,285]" pageId="57" pageNumber="58" refString="Winterbottom, R., Hanner, R. H., Burridge, M. &amp; Zur, M. (2014) A cornucopia of cryptic species-a DNA barcode analysis of the gobiid fish genus Trimma (Percomorpha, Gobiiformes). ZooKeys, 381, 79 - 111. http: // dx. doi. org / 10.3897 / zookeys. 381.6445" type="journal article" year="2014">
Winterbottom
<emphasis id="EFC4EA9EB83AF37CFD72FEF3687FEA65" box="[762,818,260,285]" italics="true" pageId="57" pageNumber="58">et al.</emphasis>
2014
</bibRefCitation>
). The first three are Moorea, Society Islands (the
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locality, n = 4),
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(n = 2), and
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(n = 10). The fourth group contains specimens from Raja Ampat,
<collectingCountry id="A5A7761CB83AF37CFF1FFEBA6A49EA1D" box="[151,260,332,357]" name="Indonesia" pageId="57" pageNumber="58">Indonesia</collectingCountry>
(n = 2),
<collectingCountry id="A5A7761CB83AF37CFEEEFEBA6AE9EA1D" box="[358,420,332,357]" name="Palau" pageId="57" pageNumber="58">Palau</collectingCountry>
(n = 1) and
<collectingCountry id="A5A7761CB83AF37CFDA6FEBA69BDEA1D" box="[558,752,332,357]" name="Indonesia" pageId="57" pageNumber="58">Flores, Indonesia</collectingCountry>
(n = 1). Further studies of both morphology and genetics are clearly needed to resolve, if possible, the variation described above. For the present, in view of the absence of positive evidence and the lack of tissue samples, we retain the Australian population in
<taxonomicName id="1AB04D0FB83AF37CFBEBFE636FF5EAD5" box="[1123,1208,404,429]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="milta">
<emphasis id="EFC4EA9EB83AF37CFBEBFE636FF5EAD5" box="[1123,1208,404,429]" italics="true" pageId="57" pageNumber="58">T. milta</emphasis>
</taxonomicName>
. The species has the Australian standard name of Moorea Pygmygoby.
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B83AF37CFF4FFE2A69CAE8F8" pageId="57" pageNumber="58" type="discussion">
<paragraph id="DD0F368CB83AF37CFF4FFE2A69CAE8F8" blockId="57.[151,1437,151,896]" pageId="57" pageNumber="58">
<emphasis id="EFC4EA9EB83AF37CFF4FFE2A6A01EA8D" bold="true" box="[199,332,476,501]" pageId="57" pageNumber="58">Discussion.</emphasis>
Specimens from the Great Barrier Reef have a slightly higher average pectoral-fin ray counts (mean = 17.8, n = 9), lateral-scale counts (mean = 23.9, n = 8) and outer gill rakers (mean = 3.8 + 14.4, n = 8), whereas those from Western
<collectingCountry id="A5A7761CB83AF37CFE55FDD26909E945" box="[477,580,548,573]" name="Australia" pageId="57" pageNumber="58">Australia</collectingCountry>
were more similar in these values to the
<typeStatus id="020B882EB83AF37CFB9EFDD36F0BE945" box="[1046,1094,549,573]" pageId="57" pageNumber="58">type</typeStatus>
specimens but differed in that two (of seven) specimens had eight (instead of nine) rays in the second dorsal fin. The fresh colouration of Australian specimens has yet to be recorded. Several freshly collected photographed specimens from
<collectingCountry id="A5A7761CB83AF37CFA97FD9A6E10E9FD" box="[1311,1373,620,645]" name="Palau" pageId="57" pageNumber="58">Palau</collectingCountry>
were yellow with a yellow stripe in each of the dorsal fins just above a basal dark stripe. In addition, specimens from
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and the Hermit Islands do not seem to reach as large a size as those from elsewhere (maximum recorded SL was
<quantity id="1A489B69B83AF37CFED4FD2E6A8CE997" box="[348,449,728,752]" metricMagnitude="-2" metricUnit="m" metricValue="1.8199999999999998" pageId="57" pageNumber="58" unit="mm" value="18.2">18.2 mm</quantity>
out of
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vs. a maximum of nearly
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SL elsewhere). Finally, none of the Micronesian/New
<collectingCountry id="A5A7761CB83AF37CFE1AFD0A6AAEE86D" box="[402,483,764,789]" name="Guinea-Bissau" pageId="57" pageNumber="58">Guinea</collectingCountry>
specimens examined had the short transverse bars across the top of the eye that are present in most (but not all) specimens from other areas. The species is found in depths of
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3
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</quantity>
. In
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it is known only from clear water outer reefs on the Great Barrier Reef and Timor Sea. It has been referred to informally as
<taxonomicName id="1AB04D0FB83AF37CFF3DFC9F6A41E8F8" box="[181,268,873,896]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="57" pageNumber="58" phylum="Chordata" rank="genus">
<emphasis id="EFC4EA9EB83AF37CFF3DFC9F6A41E8F8" box="[181,268,873,896]" italics="true" pageId="57" pageNumber="58">Trimma</emphasis>
</taxonomicName>
DFH sp. 7 or
<taxonomicName id="1AB04D0FB83AF37CFE24FC9F694EE8F8" box="[428,515,873,896]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="57" pageNumber="58" phylum="Chordata" rank="genus">
<emphasis id="EFC4EA9EB83AF37CFE24FC9F694EE8F8" box="[428,515,873,896]" italics="true" pageId="57" pageNumber="58">Trimma</emphasis>
</taxonomicName>
RW sp. 34.
</paragraph>
</subSubSection>
</treatment>
</document>