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<document id="6C455B3668BEC5822228ACD996066D5F" ID-DOI="10.11646/zootaxa.3934.1.1" ID-GBIF-Dataset="0abffe3e-a2c6-4623-b7c6-c8b26cee4a2b" ID-ISSN="1175-5326" ID-Zenodo-Dep="236066" ID-ZooBank="11C2A2CB-30B3-4694-B379-AE9D47332F0C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461012229557" checkinUser="plazi" docAuthor="Winterbottom, Richard &amp; Hoese, Douglass F." docDate="2015" docId="5519879AB81DF364FF1FFF6E69BCEA8C" docLanguage="en" docName="zt03934p102.pdf" docOrigin="Zootaxa 3934 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Trimma flavatrum Hagiwara &amp; Winterbottom 2007" docType="treatment" docVersion="11" lastPageNumber="34" masterDocId="A920FFE2B803F345FF88FFF66B4DEB78" masterDocTitle="A revision of the Australian species of Trimma (Actinopterygii, Gobiidae), with descriptions of six new species and redescriptions of twenty-three valid species" masterLastPageNumber="102" masterPageNumber="1" pageNumber="31" updateTime="1698567702668" updateUser="plazi">
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<mods:title id="4EBE5F83EC16B0BE0404F49559228355">A revision of the Australian species of Trimma (Actinopterygii, Gobiidae), with descriptions of six new species and redescriptions of twenty-three valid species</mods:title>
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<mods:namePart id="7156D89E33DECD7EB74982E80B2C0B50">Winterbottom, Richard</mods:namePart>
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<mods:namePart id="19E5E554A8718B7FA1FB6D8FD641D77E">Hoese, Douglass F.</mods:namePart>
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<treatment id="5519879AB81DF364FF1FFF6E69BCEA8C" ID-DOI="http://doi.org/10.5281/zenodo.5621510" ID-GBIF-Taxon="119600481" ID-Zenodo-Dep="5621510" LSID="urn:lsid:plazi:treatment:5519879AB81DF364FF1FFF6E69BCEA8C" httpUri="http://treatment.plazi.org/id/5519879AB81DF364FF1FFF6E69BCEA8C" lastPageId="33" lastPageNumber="34" pageId="30" pageNumber="31">
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<taxonomicName id="1AB04D0FB81DF35BFF1FFF6E6851EBCA" ID-CoL="58V5M" authority="Hagiwara &amp; Winterbottom, 2007" authorityName="Hagiwara &amp; Winterbottom" authorityYear="2007" box="[151,796,152,178]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="flavatrum">
<emphasis id="EFC4EA9EB81DF35BFF1FFF6E6851EBCA" bold="true" box="[151,796,152,178]" pageId="30" pageNumber="31">
<emphasis id="EFC4EA9EB81DF35BFF1FFF6E6A3BEBCA" bold="true" box="[151,374,152,178]" italics="true" pageId="30" pageNumber="31">Trimma flavatrum</emphasis>
<bibRefCitation id="B9214B7DB81DF35BFEF5FF6E6851EBCA" author="Hagiwara" box="[381,796,152,178]" pageId="30" pageNumber="31" refString="Hagiwara, K. &amp; Winterbottom, R. (2007) Two new species of Trimma from the western Pacific. Bulletin of the National Museum of Natural Science (Tokyo), Series A, 1 (Supplement), 163 - 174." type="journal article" year="2007">Hagiwara &amp; Winterbottom, 2007</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="DD0F368CB81DF35BFF1FFF4D6A2FEBAB" blockId="30.[151,796,152,247]" box="[151,354,186,211]" pageId="30" pageNumber="31">Wasp Pygmygoby</paragraph>
<paragraph id="DD0F368CB81DF35BFF1FFF286A21EB8F" blockId="30.[151,796,152,247]" box="[151,364,222,247]" pageId="30" pageNumber="31">
<figureCitation id="458B2A09B81DF35BFF1FFF286A5FEB8F" box="[151,274,222,247]" captionStart-0="FIGURE 14" captionStart-1="FIGURE 15" captionStartId-0="31.[151,250,1452,1474]" captionStartId-1="32.[151,250,1940,1962]" captionTargetBox-0="[151,1435,645,1430]" captionTargetBox-1="[157,1427,1235,1908]" captionTargetId-0="figure@31.[151,1435,627,1431]" captionTargetId-1="figure@32.[151,1436,1226,1918]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionText-0="FIGURE 14. Trimma flavatrum. A: 18 mm SL, AMS I. 20775 - 056, Raine Id., Queensland. B: 15.5 mm SL female, ROM 83348, Helen Reef, Palau. Photos by D. F. Hoese (A) and R. Winterbottom (B)." captionText-1="FIGURE 15. Map of the distributions of Trimma flavatrum (solid inverted triangles) and T. hoesei (solid pentagons). A solid square with a white center indicates localities at which both species occur." httpUri-0="https://zenodo.org/record/236080/files/figure.png" httpUri-1="https://zenodo.org/record/236081/files/figure.png" pageId="30" pageNumber="31">Fig. 1415</figureCitation>
, Pl. 1 G
</paragraph>
</subSubSection>
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<taxonomicName id="1AB04D0FB81DF35BFF1FFEDE69AEEA44" ID-CoL="58V5M" authority="Hagiwara &amp; Winterbottom, 2007:164" authorityName="Hagiwara &amp; Winterbottom" authorityPageNumber="164" authorityYear="2007" box="[151,739,294,317]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="flavatrum">
<emphasis id="EFC4EA9EB81DF35BFF1FFEDE6A1FEA44" box="[151,338,294,317]" italics="true" pageId="30" pageNumber="31">Trimma flavatrum</emphasis>
<treatmentCitation id="5C11109DB81DF35BFED3FED069AEEA44" author="Hagiwara" box="[347,739,294,316]" page="164" pageId="30" pageNumber="31" year="2007">
<bibRefCitation id="B9214B7DB81DF35BFED3FED069F5EA44" author="Hagiwara" box="[347,696,294,316]" pageId="30" pageNumber="31" refString="Hagiwara, K. &amp; Winterbottom, R. (2007) Two new species of Trimma from the western Pacific. Bulletin of the National Museum of Natural Science (Tokyo), Series A, 1 (Supplement), 163 - 174." type="journal article" year="2007">Hagiwara &amp; Winterbottom, 2007</bibRefCitation>
:164
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</taxonomicName>
, figs 16 (Nishikomi Cave, Amamioshima Id, Nansei Ids,
<collectingCountry id="A5A7761CB81DF35BFADBFED16EDCEA44" box="[1363,1425,295,316]" name="Japan" pageId="30" pageNumber="31">Japan</collectingCountry>
);
<treatmentCitation id="5C11109DB81DF35BFF4FFEB36AA5EA23" author="Allen" box="[199,488,325,347]" page="937" pageId="30" pageNumber="31" year="2012">
<bibRefCitation id="B9214B7DB81DF35BFF4FFEB36AF3EA23" author="Allen" box="[199,446,325,347]" pageId="30" pageNumber="31" refString="Allen, G. R. &amp; Erdmann, M. V. (2012) Reef fishes of the East Indies. Vol. III. Tropical Reef Research, Perth, pp. 857 - 1292." type="book chapter" year="2012">Allen &amp; Erdmann, 2012</bibRefCitation>
:937
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(western Pacific)
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</paragraph>
<paragraph id="DD0F368CB81DF35BFF1FFE71695BE974" blockId="30.[151,1437,391,2037]" pageId="30" pageNumber="31">
<emphasis id="EFC4EA9EB81DF35BFF1FFE716965EAD8" bold="true" box="[151,552,391,416]" pageId="30" pageNumber="31">Australian Material. Queensland</emphasis>
: Escape Reef: AMS I.22580-025, (17),
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; Raine Id: AMS I.20757-067, (20),
<quantity id="1A489B69B81DF35BFF5DFE5A6A63EABC" box="[213,302,428,452]" metricMagnitude="1" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="30" pageNumber="31" unit="m" value="10.0" valueMax="20.0" valueMin="0.0">020 m</quantity>
; AMS I.20775-056, (18),
<quantity id="1A489B69B81DF35BFDD0FE5A69FCEABC" box="[600,689,428,452]" metricMagnitude="1" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="30" pageNumber="31" unit="m" value="10.0" valueMax="20.0" valueMin="0.0">020 m</quantity>
; Tijou Reef: AMS I.20779-142, (18),
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; Yonge Reef: AMS I.19472-089, (16),
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7
<specimenCount id="CBB6FD05B81DF35BFE02FE396A88EA9F" box="[394,453,463,488]" pageId="30" pageNumber="31" type="generic">15 m</specimenCount>
</quantity>
.
<emphasis id="EFC4EA9EB81DF35BFE59FE3969F8EA90" bold="true" box="[465,693,463,488]" pageId="30" pageNumber="31">
Western
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:
</emphasis>
<collectingCountry id="A5A7761CB81DF35BFD35FE396846EA90" box="[701,779,463,488]" name="Ashmore and Cartier Islands" pageId="30" pageNumber="31">Cartier</collectingCountry>
Reef: NTM S.12883-010, (13),
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13
<specimenCount id="CBB6FD05B81DF35BFB2BFE266F92EA9F" box="[1187,1247,464,488]" pageId="30" pageNumber="31" type="generic">14 m</specimenCount>
</quantity>
. Rowley Shoals: WAM P.28024-33, (18),
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35
<specimenCount id="CBB6FD05B81DF35BFE5DFE02695FE974" box="[469,530,500,525]" pageId="30" pageNumber="31" type="generic">40 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
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<emphasis id="EFC4EA9EB81DF35BFF4FFDE16ACFE948" bold="true" box="[199,386,535,560]" pageId="30" pageNumber="31">Other material.</emphasis>
We give details of the collections because of the relative rarity of this species in collections.
<emphasis id="EFC4EA9EB81DF35BFF1FFDCA6A3FE92D" bold="true" box="[151,370,572,597]" pageId="30" pageNumber="31">
<collectingCountry id="A5A7761CB81DF35BFF1FFDCA6A20E92D" box="[151,365,572,597]" name="American Samoa" pageId="30" pageNumber="31">American Samoa</collectingCountry>
:
</emphasis>
<collectionCode id="BBA1AE49B81DF35BFEF5FDCA6AF7E92C" box="[381,442,572,596]" pageId="30" pageNumber="31">AMS</collectionCode>
I.21998-002, 2(1415),
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.
</specimenCount>
<emphasis id="EFC4EA9EB81DF35BFCA1FDCA688CE92D" bold="true" box="[809,961,572,597]" pageId="30" pageNumber="31">Caroline Ids</emphasis>
:
<collectionCode id="BBA1AE49B81DF35BFC5CFDCB6F6FE92C" LSID="urn:lsid:biocol.org:col:34697" box="[980,1058,573,596]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="30" pageNumber="31">BPBM</collectionCode>
uncat., 3(1718),
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;
<collectionCode id="BBA1AE49B81DF35BFAC3FDCA6ED1E92C" LSID="urn:lsid:biocol.org:col:34871" box="[1355,1436,572,596]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="30" pageNumber="31">USNM</collectionCode>
313299, (14),
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;
<collectionCode id="BBA1AE49B81DF35BFE2DFD966ABBE900" LSID="urn:lsid:biocol.org:col:34871" box="[421,502,608,632]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="30" pageNumber="31">USNM</collectionCode>
298765, 3(1217),
<quantity id="1A489B69B81DF35BFD55FDA9687AE90F" box="[733,823,607,632]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="2.5" metricValueMin="0.0" pageId="30" pageNumber="31" unit="m" value="12.5" valueMax="25.0" valueMin="0.0">025 m</quantity>
;
<collectionCode id="BBA1AE49B81DF35BFCCEFD9668DAE900" LSID="urn:lsid:biocol.org:col:34871" box="[838,919,608,632]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="30" pageNumber="31">USNM</collectionCode>
223193, 3(1113),
<quantity id="1A489B69B81DF35BFBF5FD966F95E90F" box="[1149,1240,608,632]" metricMagnitude="0" metricUnit="m" metricValue="8.5" metricValueMax="17.0" metricValueMin="0.0" pageId="30" pageNumber="31" unit="m" value="8.5" valueMax="17.0" valueMin="0.0">017 m</quantity>
;
<collectionCode id="BBA1AE49B81DF35BFB6EFD966E7AE900" LSID="urn:lsid:biocol.org:col:34871" box="[1254,1335,608,632]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="30" pageNumber="31">USNM</collectionCode>
223195, 3(1114),
<quantity id="1A489B69B81DF35BFE82FD726A2CE9E4" box="[266,353,644,668]" metricMagnitude="0" metricUnit="m" metricValue="8.5" metricValueMax="17.0" metricValueMin="0.0" pageId="30" pageNumber="31" unit="m" value="8.5" valueMax="17.0" valueMin="0.0">017 m</quantity>
;
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298765, 3(1217),
<quantity id="1A489B69B81DF35BFD10FD7269BDE9E4" box="[664,752,644,668]" metricMagnitude="1" metricUnit="m" metricValue="1.2" metricValueMax="2.4" metricValueMin="0.0" pageId="30" pageNumber="31" unit="m" value="12.0" valueMax="24.0" valueMin="0.0">
0
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.
<emphasis id="EFC4EA9EB81DF35BFD72FD726860E9E5" bold="true" box="[762,813,644,669]" pageId="30" pageNumber="31">
<collectingCountry id="A5A7761CB81DF35BFD72FD726865E9E5" box="[762,808,644,669]" name="Fiji" pageId="30" pageNumber="31">Fiji</collectingCountry>
:
</emphasis>
<collectionCode id="BBA1AE49B81DF35BFCBBFD72683EE9E4" LSID="urn:lsid:biocol.org:col:34778" box="[819,883,644,668]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="30" pageNumber="31">ROM</collectionCode>
46001, 2(910),
<quantity id="1A489B69B81DF35BFBBBFD726FD5E9E4" box="[1075,1176,644,668]" metricMagnitude="1" metricUnit="m" metricValue="2.65" metricValueMax="3.0" metricValueMin="2.3" pageId="30" pageNumber="31" unit="m" value="26.5" valueMax="30.0" valueMin="23.0">2330 m</quantity>
;
<collectionCode id="BBA1AE49B81DF35BFB2BFD726FAEE9E4" LSID="urn:lsid:biocol.org:col:34778" box="[1187,1251,644,668]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="30" pageNumber="31">ROM</collectionCode>
46002, 2(718),
<quantity id="1A489B69B81DF35BFF1FFD516BB0E9C7" box="[151,253,679,704]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="30" pageNumber="31" unit="m" value="12.5" valueMax="15.0" valueMin="10.0">1015 m</quantity>
;
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236756, (18),
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:
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85385, 1(18),
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, 3(1616),
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244186, 8(1618),
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Ids
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:
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12181, (18),
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:
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31421, 4(1617),
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<collectionCode id="BBA1AE49B81DF35BFE85FD066A00E870" LSID="urn:lsid:biocol.org:col:34778" box="[269,333,752,776]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="30" pageNumber="31">ROM</collectionCode>
74801, 3(1415),
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74802, 4(716),
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74926, 3(1417),
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<collectionCode id="BBA1AE49B81DF35BFF1FFCE26B9AE854" LSID="urn:lsid:biocol.org:col:34778" box="[151,215,788,812]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="30" pageNumber="31">ROM</collectionCode>
80391, (14),
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80410, (18),
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80489, (16),
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:
</emphasis>
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244098, (18),
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</collectingCountry>
:
<collectionCode id="BBA1AE49B81DF35BFD0BFCCE698EE828" LSID="urn:lsid:biocol.org:col:34778" box="[643,707,824,848]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="30" pageNumber="31">ROM</collectionCode>
52976, 2(1718),
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<collectionCode id="BBA1AE49B81DF35BFB84FCCE6F01E828" LSID="urn:lsid:biocol.org:col:34778" box="[1036,1100,824,848]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="30" pageNumber="31">ROM</collectionCode>
52977, (17),
<quantity id="1A489B69B81DF35BFB78FCCE6E01E837" box="[1264,1356,824,848]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="2.1" metricValueMin="0.9" pageId="30" pageNumber="31" unit="m" value="15.0" valueMax="21.0" valueMin="9.0">921 m</quantity>
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<collectionCode id="BBA1AE49B81DF35BFAD4FCCE6ED1E828" LSID="urn:lsid:biocol.org:col:34778" box="[1372,1436,824,848]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="30" pageNumber="31">ROM</collectionCode>
52978, (15),
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<collectionCode id="BBA1AE49B81DF35BFE29FCAA6ABFE80C" LSID="urn:lsid:biocol.org:col:34871" box="[417,498,860,884]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="30" pageNumber="31">USNM</collectionCode>
243918, 28(1118),
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<collectionCode id="BBA1AE49B81DF35BFCC3FCAA68D1E80C" LSID="urn:lsid:biocol.org:col:34871" box="[843,924,860,884]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="30" pageNumber="31">USNM</collectionCode>
243919, 8(1621),
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;
<collectionCode id="BBA1AE49B81DF35BFB6FFCAA6E75E80C" LSID="urn:lsid:biocol.org:col:34871" box="[1255,1336,860,884]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="30" pageNumber="31">USNM</collectionCode>
264524, (19),
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295300, (14),
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6
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.
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B81DF35BFF4FFC526E05EE30" pageId="30" pageNumber="31" type="diagnosis">
<paragraph id="DD0F368CB81DF35BFF4FFC526E05EE30" blockId="30.[151,1437,391,2037]" pageId="30" pageNumber="31">
<emphasis id="EFC4EA9EB81DF35BFF4FFC526A0CE8C5" bold="true" box="[199,321,932,957]" pageId="30" pageNumber="31">Diagnosis.</emphasis>
A species of
<taxonomicName id="1AB04D0FB81DF35BFE6EFC536971E8C4" box="[486,572,933,956]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="30" pageNumber="31" phylum="Chordata" rank="genus">
<emphasis id="EFC4EA9EB81DF35BFE6EFC536971E8C4" box="[486,572,933,956]" italics="true" pageId="30" pageNumber="31">Trimma</emphasis>
</taxonomicName>
with a broad interorbital, with a shallow pit above the middle of the eye, followed by a very short median fleshy ridge (often collapsed) posteriorly between two large papillae, no groove behind the eye; predorsal completely scaled forward to above the middle of eye, anterior-most scale enlarged, posterior scales ctenoid, anterior 23 rows of predorsal scales cycloid; cheek with a single row of cycloid scales; operculum with 57 cycloid scales; pectoral-fin base covered with 5 or 6 cycloid scales, uppermost 12 scales largest (distinctly enlarged); prepelvic area covered with 5 or 6 rows of cycloid scale, segmented caudal rays usually 15, rarely 16; pectoral rays unbranched; pelvic fins largely separate, connected only at the base, pelvic fins widely separate, distance between the bases of the fins about half to three-quarters of the base of each pelvic fin; fifth pelvic ray usually unbranched, others with a single branch (2 terminal tips); 4060% the length of the fourth; second and third dorsal spines subequal in length, none filamentous; dorsal and anal rays usually
<date id="A90E104CB81DF35BFB7CFB1E6E5BEE78" box="[1268,1302,1256,1280]" pageId="30" pageNumber="31">I 8</date>
; body from second dorsal fin origin posteriorly darker than the front, becoming progressively darker posteriorly; a black stripe along the bases of each dorsal fin; pelvic spine usually with a dark brown margin; caudal fin clear to white.
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B81DF365FF4FFAA26AECE8E2" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31" type="description">
<paragraph id="DD0F368CB81DF35BFF4FFAA26ED1EC8C" blockId="30.[151,1437,391,2037]" pageId="30" pageNumber="31">
<emphasis id="EFC4EA9EB81DF35BFF4FFAA26A14EE15" bold="true" box="[199,345,1364,1389]" pageId="30" pageNumber="31">Description.</emphasis>
The description is primarily based on specimens from the Great Barrier Reef,
<collectingCountry id="A5A7761CB81DF35BFA92FAA26E2BEE14" box="[1306,1382,1364,1388]" name="Samoa" pageId="30" pageNumber="31">Samoa</collectingCountry>
and
<collectingCountry id="A5A7761CB81DF35BFF1FFA816A45EEE8" box="[151,264,1399,1424]" name="Indonesia" pageId="30" pageNumber="31">Indonesia</collectingCountry>
. Dorsal fin usually VI +
<date id="A90E104CB81DF35BFDA0FA8E6905EEE8" box="[552,584,1400,1424]" pageId="30" pageNumber="31">I 8</date>
(
<date id="A90E104CB81DF35BFDD2FA8E6936EEF7" box="[602,635,1400,1424]" pageId="30" pageNumber="31">
I
<quantity id="1A489B69B81DF35BFDE5FA8E6936EEF7" box="[621,635,1400,1423]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="30" pageNumber="31" unit="in" value="7.0">7</quantity>
</date>
in 22% of individuals), second to fourth rays longest and subequal in length, reaching to interspace between two dorsal fins when adpressed, first ray of dorsal fin usually branched, anterior element of last element branched; anal fin
<date id="A90E104CB81DF35BFD66FA36685DEEA0" box="[750,784,1472,1496]" pageId="30" pageNumber="31">I 8</date>
(rarely
<date id="A90E104CB81DF35BFCE7FA3668D9EEA0" box="[879,916,1472,1496]" pageId="30" pageNumber="31">I 9</date>
, in 6% of individuals), first ray unbranched, anterior element of last ray branched; pectoral fin 1215 (see Variation, discussion below), reaching just posteriorly to a vertical in line from just beyond anal spine to about second segmented ray, rays unbranched; pelvic fin
<date id="A90E104CB81DF35BFACEF9FE6E2BED58" box="[1350,1382,1543,1568]" pageId="30" pageNumber="31">I 5</date>
, fins widely separate with very low connecting membrane, distance between bases of fins; subequal to base of each pelvic fin; first four rays with one sequential branch, fifth ray with one dichotomous branch (two terminal tips) or rarely unbranched, 40%65% length of fourth ray, which reaches posteriorly to below anal spine to about second segmented ray. Lateral scales 2226; transverse scales 69; scales on preopercle, opercle and pectoral-fin base cycloid, breast scales cycloid on midline, ctenoid or cycloid to sides, but scale between bases of pelvic fins cycloid, and sometimes anteriormost 12 scales on midline of belly cycloid, remaining scales ctenoid; scales extending to above middle of eye; scales below dorsal fin with small accessory scales; pectoral-fin base with two vertical rows of cycloid scales, central scale vertically elongate in anterior row and dorsal-most scale vertically elongate in second row; prepelvic area with 57 rows of cycloid and ctenoid scales. Teeth in lower jaw consist of an enlarged outer row of curved, slightly enlarged canines and an inner row of similar, but smaller canines with irregular rows of small conical teeth in between; outer row of teeth in upper jaw similar to those of lower jaw, with small irregular inner rows of teeth. Tongue tip rounded. Gill opening extending anteroventrally to below posterior margin of pupil; outer gill rakers on first arch 3 + 1214 = 1517 (
<bibRefCitation id="B9214B7DB81DF35BFD28F82A6EC1EC8D" author="Hagiwara" box="[672,1420,2012,2037]" pageId="30" pageNumber="31" refString="Hagiwara, K. &amp; Winterbottom, R. (2007) Two new species of Trimma from the western Pacific. Bulletin of the National Museum of Natural Science (Tokyo), Series A, 1 (Supplement), 163 - 174." type="journal article" year="2007">Hagiwara &amp; Winterbottom, 2007 reported 34 + 1113 = 1517</bibRefCitation>
).
</paragraph>
<paragraph id="DD0F368CB81CF35AFF1FFF61690EEB8F" blockId="31.[151,1437,151,608]" pageId="31" pageNumber="32">
Anterior nares a narrow tube, posterior nares a large pore with a raised rim; nasal sac slightly elevated as a raised oval sac located just above upper lip, openings separated by about 12 nares diameters. Interorbital as described in diagnosis. Vertebral pattern is
<typeStatus id="020B882EB81CF35AFE63FF16696EEB80" box="[491,547,224,248]" pageId="31" pageNumber="32">Type</typeStatus>
B.
</paragraph>
<paragraph id="DD0F368CB81CF35AFF4FFEF26834E960" blockId="31.[151,1437,151,608]" pageId="31" pageNumber="32">
<emphasis id="EFC4EA9EB81CF35AFF4FFEF2690EEA65" box="[199,579,260,285]" italics="true" pageId="31" pageNumber="32">Colour pattern. Freshly collected</emphasis>
. From slides of freshly collected specimen from Raine
<collectingCountry id="A5A7761CB81CF35AFB4EFEF26E47EA65" box="[1222,1290,260,285]" name="Iceland" pageId="31" pageNumber="32">Island</collectingCountry>
(
<figureCitation id="458B2A09B81CF35AFA94FEF26E23EA64" box="[1308,1390,260,285]" captionStart="FIGURE 14" captionStartId="31.[151,250,1452,1474]" captionTargetBox="[151,1435,645,1430]" captionTargetId="figure@31.[151,1435,627,1431]" captionTargetPageId="31" captionText="FIGURE 14. Trimma flavatrum. A: 18 mm SL, AMS I. 20775 - 056, Raine Id., Queensland. B: 15.5 mm SL female, ROM 83348, Helen Reef, Palau. Photos by D. F. Hoese (A) and R. Winterbottom (B)." httpUri="https://zenodo.org/record/236080/files/figure.png" pageId="31" pageNumber="32">Fig. 14</figureCitation>
A), Yonge Reef and Escape Reef, Queensland. Head and anterior half of body largely yellow with scattered grey spots due to dense concentrations of melanophores; cheek yellowish-grey; posterior half of body (from below posterior end of first dorsal fin) grading anteriorly from yellowish grey to black on caudal peduncle; first dorsal fin yellowish to largely grey, with yellowish tinge; second dorsal and anal fins with thin basal black stripe, followed distally by broad yellowish-orange stripe and then a thin submarginal black line; distal margin of fins white; caudal fin white; pelvic fins yellowish-grey, with white outer margin; pectoral fin a translucent orange. See under Variation for differences in colour pattern from other localities, and live colour.
</paragraph>
<paragraph id="DD0F368CB81CF35AFF4FFDD26B9EE918" blockId="31.[151,1437,151,608]" pageId="31" pageNumber="32">
<emphasis id="EFC4EA9EB81CF35AFF4FFDD26A7AE945" box="[199,311,548,573]" italics="true" pageId="31" pageNumber="32">Preserved</emphasis>
. Head, body and fins brown, except for caudal fin; caudal peduncle grey to black; caudal fin white to clear.
</paragraph>
<caption id="89CF6604B81CF35AFF1FFA5A68F1EE99" httpUri="https://zenodo.org/record/236080/files/figure.png" pageId="31" pageNumber="32" targetBox="[151,1435,645,1430]" targetPageId="31">
<paragraph id="DD0F368CB81CF35AFF1FFA5A68F1EE99" blockId="31.[151,1436,1452,1505]" pageId="31" pageNumber="32">
<emphasis id="EFC4EA9EB81CF35AFF1FFA5A6A6FEEBA" bold="true" box="[151,290,1452,1474]" pageId="31" pageNumber="32">FIGURE 14.</emphasis>
<taxonomicName id="1AB04D0FB81CF35AFEA3FA5B6AABEEBA" box="[299,486,1452,1474]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="flavatrum">
<emphasis id="EFC4EA9EB81CF35AFEA3FA5B6AABEEBA" box="[299,486,1452,1474]" italics="true" pageId="31" pageNumber="32">Trimma flavatrum</emphasis>
</taxonomicName>
. A: 18 mm SL, AMS I. 20775-056, Raine Id., Queensland. B: 15.5 mm SL female, ROM 83348, Helen Reef, Palau. Photos by D. F. Hoese (A) and R. Winterbottom (B).
</paragraph>
</caption>
<paragraph id="DD0F368CB81CF35AFF4FF9FE6A63EDAD" blockId="31.[151,1436,1544,2037]" pageId="31" pageNumber="32">
<emphasis id="EFC4EA9EB81CF35AFF4FF9FE6A76ED59" bold="true" box="[199,315,1544,1569]" pageId="31" pageNumber="32">Variation.</emphasis>
Insufficient material was available for any statistical analysis of the geographical variation of meristic features. Also, many specimens were in poor conditions and obtaining accurate counts was not always possible. However pectoral ray counts may vary geographically.
<bibRefCitation id="B9214B7DB81CF35AFC1CF9A66E7EED10" author="Hagiwara" box="[916,1331,1616,1641]" pageId="31" pageNumber="32" refString="Hagiwara, K. &amp; Winterbottom, R. (2007) Two new species of Trimma from the western Pacific. Bulletin of the National Museum of Natural Science (Tokyo), Series A, 1 (Supplement), 163 - 174." type="journal article" year="2007">Hagiwara and Winterbottom (2007)</bibRefCitation>
reported 1315 pectoral rays with an average of 14 rays. Many of the specimens studied for this paper were in poor condition with a damaged pectoral fin. However, most from
<collectingCountry id="A5A7761CB81CF35AFCDCF96E688CEDC9" box="[852,961,1688,1713]" name="Indonesia" pageId="31" pageNumber="32">Indonesia</collectingCountry>
and
<collectingCountry id="A5A7761CB81CF35AFC75F96E6F29EDC9" box="[1021,1124,1688,1713]" name="Australia" pageId="31" pageNumber="32">Australia</collectingCountry>
appeared to have 12 or 13 pectoral rays.
</paragraph>
<paragraph id="DD0F368CB81CF35AFF4FF91668A4EC8D" blockId="31.[151,1436,1544,2037]" pageId="31" pageNumber="32">
Specimens from
<collectingCountry id="A5A7761CB81CF35AFE01F9166ABDED81" box="[393,496,1760,1785]" name="Australia" pageId="31" pageNumber="32">Australia</collectingCountry>
and
<collectingCountry id="A5A7761CB81CF35AFDA2F916693BED80" box="[554,630,1760,1784]" name="Samoa" pageId="31" pageNumber="32">Samoa</collectingCountry>
show considerable differences from material described from
<collectingCountry id="A5A7761CB81CF35AFABFF9176E3AED80" box="[1335,1399,1761,1784]" name="Japan" pageId="31" pageNumber="32">Japan</collectingCountry>
by
<bibRefCitation id="B9214B7DB81CF35AFF1FF8F26979EC64" author="Hagiwara" box="[151,564,1796,1821]" pageId="31" pageNumber="32" refString="Hagiwara, K. &amp; Winterbottom, R. (2007) Two new species of Trimma from the western Pacific. Bulletin of the National Museum of Natural Science (Tokyo), Series A, 1 (Supplement), 163 - 174." type="journal article" year="2007">Hagiwara and Winterbottom (2007)</bibRefCitation>
. In particular the black pigment on the caudal peduncle extends forward to under the second dorsal fin and the dark colouration decreases forward, but there are dense concentrations of melanophores up to the head. Specimens from
<collectingCountry id="A5A7761CB81CF35AFD24F8BB69A1EC1C" box="[684,748,1869,1892]" name="Japan" pageId="31" pageNumber="32">Japan</collectingCountry>
and the
<collectingCountry id="A5A7761CB81CF35AFCC6F8BA6881EC1D" box="[846,972,1868,1893]" name="Philippines" pageId="31" pageNumber="32">Philippines</collectingCountry>
generally have the black bar confined to the caudal peduncle. In addition, fresh specimens from
<collectingCountry id="A5A7761CB81CF35AFC9EF8866831ECF1" box="[790,892,1904,1929]" name="Australia" pageId="31" pageNumber="32">Australia</collectingCountry>
and
<collectingCountry id="A5A7761CB81CF35AFC30F8866F49ECF0" box="[952,1028,1904,1928]" name="Samoa" pageId="31" pageNumber="32">Samoa</collectingCountry>
typically have a black submarginal band on the second dorsal and anal fin, followed by white at the tips of the fins.
<bibRefCitation id="B9214B7DB81CF35AFB85F8626ED1ECD4" author="Hagiwara" box="[1037,1436,1940,1965]" pageId="31" pageNumber="32" refString="Hagiwara, K. &amp; Winterbottom, R. (2007) Two new species of Trimma from the western Pacific. Bulletin of the National Museum of Natural Science (Tokyo), Series A, 1 (Supplement), 163 - 174." type="journal article" year="2007">Hagiwara and Winterbottom (2007)</bibRefCitation>
reported a similar colouration in a specimen from
<collectingCountry id="A5A7761CB81CF35AFD5FF84E6863ECA9" box="[727,814,1976,2001]" name="Indonesia" pageId="31" pageNumber="32">Ambon</collectingCountry>
,
<collectingCountry id="A5A7761CB81CF35AFCB3F84E68E1ECA9" box="[827,940,1976,2001]" name="Indonesia" pageId="31" pageNumber="32">Indonesia</collectingCountry>
. Specimens from
<collectingCountry id="A5A7761CB81CF35AFBF1F84F6FF4ECA8" box="[1145,1209,1977,2000]" name="Japan" pageId="31" pageNumber="32">Japan</collectingCountry>
and the
<collectingCountry id="A5A7761CB81CF35AFA96F84E6ED1ECA9" box="[1310,1436,1976,2001]" name="Philippines" pageId="31" pageNumber="32">Philippines</collectingCountry>
generally have a broad light area distally on the second dorsal and anal fins.
</paragraph>
<paragraph id="DD0F368CB823F365FF4FFF6E6A65EAB4" blockId="32.[151,1437,151,1174]" pageId="32" pageNumber="33">
A photo of a freshly collected specimen from
<collectingCountry id="A5A7761CB823F365FD6BFF6E6862EBC8" box="[739,815,152,176]" name="Samoa" pageId="32" pageNumber="33">Samoa</collectingCountry>
shows a similar colouration to Australian specimens, except that the anterior body is darker, with only a trace of yellowish-orange pigment. Colouration of fresh material from the
<collectingCountry id="A5A7761CB823F365FF77FF286A30EB8F" box="[255,381,222,247]" name="Philippines" pageId="32" pageNumber="33">Philippines</collectingCountry>
and
<collectingCountry id="A5A7761CB823F365FE3CFF286ABFEB8F" box="[436,498,222,247]" name="Palau" pageId="32" pageNumber="33">Palau</collectingCountry>
(
<figureCitation id="458B2A09B823F365FD8AFF28691EEB8F" box="[514,595,222,247]" captionStart="FIGURE 14" captionStartId="31.[151,250,1452,1474]" captionTargetBox="[151,1435,645,1430]" captionTargetId="figure@31.[151,1435,627,1431]" captionTargetPageId="31" captionText="FIGURE 14. Trimma flavatrum. A: 18 mm SL, AMS I. 20775 - 056, Raine Id., Queensland. B: 15.5 mm SL female, ROM 83348, Helen Reef, Palau. Photos by D. F. Hoese (A) and R. Winterbottom (B)." httpUri="https://zenodo.org/record/236080/files/figure.png" pageId="32" pageNumber="33">Fig. 14</figureCitation>
B) was described by
<bibRefCitation id="B9214B7DB823F365FCC0FF286F93EB8F" author="Hagiwara" box="[840,1246,222,247]" pageId="32" pageNumber="33" refString="Hagiwara, K. &amp; Winterbottom, R. (2007) Two new species of Trimma from the western Pacific. Bulletin of the National Museum of Natural Science (Tokyo), Series A, 1 (Supplement), 163 - 174." type="journal article" year="2007">Hagiwara and Winterbottom (2007)</bibRefCitation>
. The colouration in these specimens and one from the
<collectingCountry id="A5A7761CB823F365FDB7FEF469B4EA63" box="[575,761,258,283]" name="Marshall Islands" pageId="32" pageNumber="33">Marshall Islands</collectingCountry>
is generally lighter than Australian and Samoan specimens and there is often yellow on the caudal fin. A live specimen from Raja Ampat (Pl. 1 G) has a mostly yellow body and head, with a dark red-brown posterior part of the caudal peduncle, more intense and extending further anteriorly along the ventral surface. There is a dark basal band in the dorsal and anal fins, followed by a slightly wider yellow stripe; the distal half to one-third of the fins is hyaline; and the pelvic fin is yellow grading to light pink distally.
</paragraph>
<paragraph id="DD0F368CB823F365FF4FFE21688DE9D9" blockId="32.[151,1437,151,1174]" pageId="32" pageNumber="33">
<bibRefCitation id="B9214B7DB823F365FF4FFE216923EA97" author="Hagiwara" box="[199,622,471,496]" pageId="32" pageNumber="33" refString="Hagiwara, K. &amp; Winterbottom, R. (2007) Two new species of Trimma from the western Pacific. Bulletin of the National Museum of Natural Science (Tokyo), Series A, 1 (Supplement), 163 - 174." type="journal article" year="2007">Hagiwara and Winterbottom (2007)</bibRefCitation>
reported completely separate pelvic fins, but material studied here (including the
<typeStatus id="020B882EB823F365FECCFE0C6AE5E96B" box="[324,424,506,531]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
) shows a low membrane connecting the bases of the two fins. The
<typeStatus id="020B882EB823F365FB50FE0C6E77E96B" box="[1240,1338,506,531]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
and the specimen from
<collectingCountry id="A5A7761CB823F365FEC1FDEB6ACAE94E" box="[329,391,541,566]" name="Palau" pageId="32" pageNumber="33">Palau</collectingCountry>
figured by
<bibRefCitation id="B9214B7DB823F365FD86FDEB68E8E94E" author="Hagiwara" box="[526,933,541,566]" pageId="32" pageNumber="33" refString="Hagiwara, K. &amp; Winterbottom, R. (2007) Two new species of Trimma from the western Pacific. Bulletin of the National Museum of Natural Science (Tokyo), Series A, 1 (Supplement), 163 - 174." type="journal article" year="2007">Hagiwara and Winterbottom (2007)</bibRefCitation>
have a snout pointed in lateral view, with a convex dorsal margin and very narrow suborbital. All specimens examined in this study have the snout with a notch or concave dorsal margin and a broader suborbital. It is not clear if these differences are due to different methods of preservation, abnormalities or indicative of separate species.
</paragraph>
<paragraph id="DD0F368CB823F365FF4FFD5A6A43E82A" blockId="32.[151,1437,151,1174]" pageId="32" pageNumber="33">
The interorbital fleshy ridge is often collapsed. The
<typeStatus id="020B882EB823F365FC84FD5B6871E9BD" box="[780,828,685,709]" pageId="32" pageNumber="33">type</typeStatus>
material largely lacks the ridge, but it is discernible in some of that material, although only extending forward to between the interorbital papillae above the middle of the eye. In Australian material the ridge normally extends well forward of these interorbital papillae. However, because it is collapsed in most specimens there is not adequate material to determine if the differences are consistent.
</paragraph>
<paragraph id="DD0F368CB823F365FF4FFCAB6AECE8E2" blockId="32.[151,1437,151,1174]" pageId="32" pageNumber="33">Given the colour and head shape variation and possible pectoral-fin count variation, it is likely that more than one species is involved.</paragraph>
</subSubSection>
<subSubSection id="95AA6507B823F365FF4FFC53697AE89A" pageId="32" pageNumber="33" type="etymology">
<paragraph id="DD0F368CB823F365FF4FFC53697AE89A" blockId="32.[151,1437,151,1174]" pageId="32" pageNumber="33">
<emphasis id="EFC4EA9EB823F365FF4FFC536A03E8C6" bold="true" box="[199,334,933,958]" pageId="32" pageNumber="33">Etymology.</emphasis>
From the Latin flavus, meaning yellow, and atrum, black, in allusion to the unusual yellow and black colouration of the new species.
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B823F364FF4FFC1B6ECAEAD4" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="distribution">
<paragraph id="DD0F368CB823F365FF4FFC1B6A1BEFEE" blockId="32.[151,1437,151,1174]" pageId="32" pageNumber="33">
<emphasis id="EFC4EA9EB823F365FF4FFC1B6A2DEF7E" bold="true" box="[199,352,1005,1030]" pageId="32" pageNumber="33">Distribution.</emphasis>
<taxonomicName id="1AB04D0FB823F365FEEEFC18697CEF7E" box="[358,561,1005,1030]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="flavatrum">
<emphasis id="EFC4EA9EB823F365FEEEFC18697CEF7E" box="[358,561,1005,1030]" italics="true" pageId="32" pageNumber="33">Trimma flavatrum</emphasis>
</taxonomicName>
is known from outer edge of the Great Barrier Reef from Tijou Reef to Escape and from
<collectingCountry id="A5A7761CB823F365FE8BFBE76A1CEF52" box="[259,337,1041,1066]" name="Ashmore and Cartier Islands" pageId="32" pageNumber="33">Cartier</collectingCountry>
Reef in the Timor Sea off Western
<collectingCountry id="A5A7761CB823F365FD52FBE76809EF52" box="[730,836,1041,1066]" name="Australia" pageId="32" pageNumber="33">Australia</collectingCountry>
. The species is not known from mid- or in-shore reefs on the Great Barrier Reef. In
<collectingCountry id="A5A7761CB823F365FE6AFBC36905EF36" box="[482,584,1077,1102]" name="Australia" pageId="32" pageNumber="33">Australia</collectingCountry>
the species is known from depths of
<quantity id="1A489B69B823F365FC63FBC06F0EEF35" box="[1003,1091,1078,1102]" metricMagnitude="1" metricUnit="m" metricValue="2.2" metricValueMax="3.7" metricValueMin="0.7" pageId="32" pageNumber="33" unit="m" value="22.0" valueMax="37.0" valueMin="7.0">737 m</quantity>
, and to depths of
<quantity id="1A489B69B823F365FA83FBC36E09EF35" box="[1291,1348,1077,1102]" metricMagnitude="1" metricUnit="m" metricValue="4.6" pageId="32" pageNumber="33" unit="m" value="46.0">46 m</quantity>
outside of
<collectingCountry id="A5A7761CB823F365FF3EFBAF6A52EF0A" box="[182,287,1113,1138]" name="Australia" pageId="32" pageNumber="33">Australia</collectingCountry>
, where it occurs eastward to
<collectingCountry id="A5A7761CB823F365FDE4FBAC69F6EF0A" box="[620,699,1114,1138]" name="Samoa" pageId="32" pageNumber="33">Samoa</collectingCountry>
, and north through
<collectingCountry id="A5A7761CB823F365FC12FBAF6895EF0A" box="[922,984,1113,1138]" name="Palau" pageId="32" pageNumber="33">Palau</collectingCountry>
and the
<collectingCountry id="A5A7761CB823F365FBB4FBAF6FB9EF0A" box="[1084,1268,1113,1138]" name="Micronesia" pageId="32" pageNumber="33">Caroline Islands</collectingCountry>
to the Ryukyu Islands (
<figureCitation id="458B2A09B823F365FF7DFB8B6A04EFEE" box="[245,329,1149,1174]" captionStart="FIGURE 15" captionStartId="32.[151,250,1940,1962]" captionTargetBox="[157,1427,1235,1908]" captionTargetId="figure@32.[151,1436,1226,1918]" captionTargetPageId="32" captionText="FIGURE 15. Map of the distributions of Trimma flavatrum (solid inverted triangles) and T. hoesei (solid pentagons). A solid square with a white center indicates localities at which both species occur." httpUri="https://zenodo.org/record/236081/files/figure.png" pageId="32" pageNumber="33">Fig. 15</figureCitation>
).
</paragraph>
<caption id="89CF6604B823F365FF1FF86268CBECB0" httpUri="https://zenodo.org/record/236081/files/figure.png" pageId="32" pageNumber="33" targetBox="[157,1427,1235,1908]" targetPageId="32">
<paragraph id="DD0F368CB823F365FF1FF86268CBECB0" blockId="32.[151,1436,1940,1992]" pageId="32" pageNumber="33">
<emphasis id="EFC4EA9EB823F365FF1FF8626A6DECD2" bold="true" box="[151,288,1940,1962]" pageId="32" pageNumber="33">FIGURE 15.</emphasis>
Map of the distributions of
<taxonomicName id="1AB04D0FB823F365FDCBF86369B6ECD2" box="[579,763,1940,1962]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="flavatrum">
<emphasis id="EFC4EA9EB823F365FDCBF86369B6ECD2" box="[579,763,1940,1962]" italics="true" pageId="32" pageNumber="33">Trimma flavatrum</emphasis>
</taxonomicName>
(solid inverted triangles) and
<taxonomicName id="1AB04D0FB823F365FBB9F8636FC7ECD2" box="[1073,1162,1940,1962]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="hoesei">
<emphasis id="EFC4EA9EB823F365FBB9F8636FC7ECD2" box="[1073,1162,1940,1962]" italics="true" pageId="32" pageNumber="33">T. hoesei</emphasis>
</taxonomicName>
(solid pentagons). A solid square with a white center indicates localities at which both species occur.
</paragraph>
</caption>
<paragraph id="DD0F368CB822F364FF4FFF616ECAEAD4" blockId="33.[151,1437,151,501]" pageId="33" pageNumber="34">
<emphasis id="EFC4EA9EB822F364FF4FFF616A21EBC8" bold="true" box="[199,364,151,176]" pageId="33" pageNumber="34">Comparisons.</emphasis>
<bibRefCitation id="B9214B7DB822F364FEFCFF616848EBC8" author="Hagiwara" box="[372,773,151,176]" pageId="33" pageNumber="34" refString="Hagiwara, K. &amp; Winterbottom, R. (2007) Two new species of Trimma from the western Pacific. Bulletin of the National Museum of Natural Science (Tokyo), Series A, 1 (Supplement), 163 - 174." type="journal article" year="2007">Hagiwara and Winterbottom (2007)</bibRefCitation>
compared
<taxonomicName id="1AB04D0FB822F364FC09FF6F6F01EBC8" box="[897,1100,151,176]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="flavatrum">
<emphasis id="EFC4EA9EB822F364FC09FF6F6F01EBC8" box="[897,1100,151,176]" italics="true" pageId="33" pageNumber="34">Trimma flavatrum</emphasis>
</taxonomicName>
with
<taxonomicName id="1AB04D0FB822F364FB04FF6F6FB0EBC8" box="[1164,1277,153,176]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="tevegae">
<emphasis id="EFC4EA9EB822F364FB04FF6F6FB0EBC8" box="[1164,1277,153,176]" italics="true" pageId="33" pageNumber="34">T. tevegae</emphasis>
</taxonomicName>
stating that
<taxonomicName id="1AB04D0FB822F364FA0FFF6F6BA0EBAC" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="tevegae">
<emphasis id="EFC4EA9EB822F364FA0FFF6F6BA0EBAC" italics="true" pageId="33" pageNumber="34">T. tevegae</emphasis>
</taxonomicName>
possessed a median fleshy ridge in the interorbital ridge, which is lacking in
<taxonomicName id="1AB04D0FB822F364FB18FF4B6E56EBAD" box="[1168,1307,188,213]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="flavatrum">
<emphasis id="EFC4EA9EB822F364FB18FF4B6E56EBAD" box="[1168,1307,188,213]" italics="true" pageId="33" pageNumber="34">T. flavatrum</emphasis>
</taxonomicName>
. However, elsewhere in the paper they record a median fleshy ridge in the interorbital in
<taxonomicName id="1AB04D0FB822F364FC72FF176F89EB80" box="[1018,1220,223,248]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="flavatrum">
<emphasis id="EFC4EA9EB822F364FC72FF176F89EB80" box="[1018,1220,223,248]" italics="true" pageId="33" pageNumber="34">Trimma flavatrum</emphasis>
</taxonomicName>
. Both species have the ridge, but it is better developed in
<taxonomicName id="1AB04D0FB822F364FDC6FEF3698DEA64" box="[590,704,261,284]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="tevegae">
<emphasis id="EFC4EA9EB822F364FDC6FEF3698DEA64" box="[590,704,261,284]" italics="true" pageId="33" pageNumber="34">T. tevegae</emphasis>
</taxonomicName>
as a rod-shaped ridge reaching to between the papillae near the anterodorsal margin of the eye, while the ridge is triangular in
<taxonomicName id="1AB04D0FB822F364FCD4FEDF68A9EA38" box="[860,996,295,320]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="flavatrum">
<emphasis id="EFC4EA9EB822F364FCD4FEDF68A9EA38" box="[860,996,295,320]" italics="true" pageId="33" pageNumber="34">T. flavatrum</emphasis>
</taxonomicName>
, only reaching to between the papillae above the middle of the eye with a pit anterior to the ridge.
<taxonomicName id="1AB04D0FB822F364FCBFFEBB68A6EA1C" box="[823,1003,333,356]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="tevegae">
<emphasis id="EFC4EA9EB822F364FCBFFEBB68A6EA1C" box="[823,1003,333,356]" italics="true" pageId="33" pageNumber="34">Trimma tevegae</emphasis>
</taxonomicName>
lacks the pit in the interorbital region. The combination of the colouration, interorbital and 15 segmented caudal rays readily separates this species from all others. Only two specimens, both from
<collectingCountry id="A5A7761CB822F364FDFDFE6269FBEAD5" box="[629,694,404,429]" name="Palau" pageId="33" pageNumber="34">Palau</collectingCountry>
, were available for genetic analysis (
<bibRefCitation id="B9214B7DB822F364FBDBFE626E36EAD4" author="Winterbottom" box="[1107,1403,404,429]" pageId="33" pageNumber="34" refString="Winterbottom, R., Hanner, R. H., Burridge, M. &amp; Zur, M. (2014) A cornucopia of cryptic species-a DNA barcode analysis of the gobiid fish genus Trimma (Percomorpha, Gobiiformes). ZooKeys, 381, 79 - 111. http: // dx. doi. org / 10.3897 / zookeys. 381.6445" type="journal article" year="2014">
Winterbottom
<emphasis id="EFC4EA9EB822F364FB70FE636E7DEAD5" box="[1272,1328,404,429]" italics="true" pageId="33" pageNumber="34">et al.</emphasis>
, 2014
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="95AA6507B822F364FF4FFE4169BCEA8C" pageId="33" pageNumber="34" type="discussion">
<paragraph id="DD0F368CB822F364FF4FFE4169BCEA8C" blockId="33.[151,1437,151,501]" pageId="33" pageNumber="34">
<emphasis id="EFC4EA9EB822F364FF4FFE416A01EAA8" bold="true" box="[199,332,439,464]" pageId="33" pageNumber="34">Discussion.</emphasis>
This species has been referred to informally as
<taxonomicName id="1AB04D0FB822F364FCE6FE4F6888EAA8" box="[878,965,441,464]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="genus">
<emphasis id="EFC4EA9EB822F364FCE6FE4F6888EAA8" box="[878,965,441,464]" italics="true" pageId="33" pageNumber="34">Trimma</emphasis>
</taxonomicName>
DFH sp. 14 and
<taxonomicName id="1AB04D0FB822F364FB02FE4F6FACEAA8" box="[1162,1249,441,464]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="genus">
<emphasis id="EFC4EA9EB822F364FB02FE4F6FACEAA8" box="[1162,1249,441,464]" italics="true" pageId="33" pageNumber="34">Trimma</emphasis>
</taxonomicName>
RW sp. 25. The
<typeStatus id="020B882EB822F364FF1FFE2A6BB4EA8D" box="[151,249,476,501]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
is the largest specimen known at
<quantity id="1A489B69B822F364FDFBFE2A698FEA8C" box="[627,706,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="33" pageNumber="34" unit="mm" value="23.0">23 mm</quantity>
SL.
</paragraph>
</subSubSection>
</treatment>
</document>