487 lines
54 KiB
XML
487 lines
54 KiB
XML
<document id="AF8B1C41202A235729F1D684B5E538C9" ID-DOI="10.11646/zootaxa.3934.1.1" ID-GBIF-Dataset="0abffe3e-a2c6-4623-b7c6-c8b26cee4a2b" ID-ISSN="1175-5326" ID-Zenodo-Dep="236066" ID-ZooBank="11C2A2CB-30B3-4694-B379-AE9D47332F0C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461012229557" checkinUser="plazi" docAuthor="Winterbottom, Richard & Hoese, Douglass F." docDate="2015" docId="5519879AB804F34FFF1FFC7068FEE927" docLanguage="en" docName="zt03934p102.pdf" docOrigin="Zootaxa 3934 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Trimma anaima Winterbottom 2000" docType="treatment" docVersion="11" lastPageNumber="11" masterDocId="A920FFE2B803F345FF88FFF66B4DEB78" masterDocTitle="A revision of the Australian species of Trimma (Actinopterygii, Gobiidae), with descriptions of six new species and redescriptions of twenty-three valid species" masterLastPageNumber="102" masterPageNumber="1" pageNumber="8" updateTime="1698567702668" updateUser="plazi">
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<mods:title id="DB0EAFD1DBDA7FE8CE201C412A4AE56C">A revision of the Australian species of Trimma (Actinopterygii, Gobiidae), with descriptions of six new species and redescriptions of twenty-three valid species</mods:title>
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<mods:namePart id="1A2D494245830E1B02E47A415B9DE703">Winterbottom, Richard</mods:namePart>
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<mods:namePart id="EFBA0E154B108DB98D93434DA9631BF1">Hoese, Douglass F.</mods:namePart>
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<mods:title id="28B7E7D6A49C5D93CDE8BBCFB283C8A5">Zootaxa</mods:title>
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<mods:part id="E72FAE9DCAE74BE2F96CC170A082924D">
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<mods:date id="8A0346727EC50893740BE87031E0D8F9">2015</mods:date>
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<paragraph id="DD0F368CB804F342FF1FFC706910E8D8" blockId="7.[151,605,902,997]" box="[151,605,902,928]" pageId="7" pageNumber="8">
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<heading id="864781E0B804F342FF1FFC706910E8D8" bold="true" box="[151,605,902,928]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
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<taxonomicName id="1AB04D0FB804F342FF1FFC706910E8D8" ID-CoL="58V4J" authority="Winterbottom, 2000" authorityName="Winterbottom" authorityYear="2000" box="[151,605,902,928]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="anaima">
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<emphasis id="EFC4EA9EB804F342FF1FFC706910E8D8" bold="true" box="[151,605,902,928]" pageId="7" pageNumber="8">
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<emphasis id="EFC4EA9EB804F342FF1FFC706A17E8D8" bold="true" box="[151,346,902,928]" italics="true" pageId="7" pageNumber="8">Trimma anaima</emphasis>
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<bibRefCitation id="B9214B7DB804F342FEE9FC706910E8D8" author="Winterbottom" box="[353,605,902,928]" pageId="7" pageNumber="8" refString="Winterbottom, R. (2000) Four new species of Trimma (Gobiidae), from the Indian and western Pacific oceans. aqua, Journal of Ichthyology and Aquatic Biology, 4 (2), 57 - 66." type="journal article" year="2000">Winterbottom, 2000</bibRefCitation>
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<paragraph id="DD0F368CB804F342FF1FFC5E6AC0E8B9" blockId="7.[151,605,902,997]" box="[151,397,936,961]" pageId="7" pageNumber="8">Sharpeye Pygmygoby</paragraph>
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<paragraph id="DD0F368CB804F342FF1FFC3A6A11E89D" blockId="7.[151,605,902,997]" box="[151,348,972,997]" pageId="7" pageNumber="8">
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<figureCitation id="458B2A09B804F342FF1FFC3A6BA8E89D" box="[151,229,972,997]" captionStart="FIGURE 2" captionStartId="8.[151,250,1868,1890]" captionTargetBox="[171,1372,1003,1809]" captionTargetId="figure@8.[152,1435,985,1847]" captionTargetPageId="8" captionText="FIGURE 2. Trimma anaima, A: 13.7 mm SL male, ROM 83022; B: 14.0 mm SL male, ROM 80303 (note parasitic copepod attached to belly), both from Palau. Photos by R. Winterbottom." httpUri="https://zenodo.org/record/236068/files/figure.png" pageId="7" pageNumber="8">Figs. 2</figureCitation>
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,
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<figureCitation id="458B2A09B804F342FF79FC3B6A4CE89D" box="[241,257,973,997]" captionStart="FIGURE 3" captionStartId="9.[151,250,1948,1970]" captionTargetBox="[162,1425,1246,1915]" captionTargetId="figure@9.[151,1436,1237,1927]" captionTargetPageId="9" captionText="FIGURE 3. Map of the distributions of Trimma anaima (solid circles) and T. annosum (solid triangles). A black star indicates localities at which both species occur." httpUri="https://zenodo.org/record/236069/files/figure.png" pageId="7" pageNumber="8">3</figureCitation>
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; Pl. 1 A
|
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</paragraph>
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<subSubSection id="95AA6507B804F342FF1FFBE369F8EE49" pageId="7" pageNumber="8" type="reference_group">
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<taxonomicName id="1AB04D0FB804F342FF1FFBE36966EF52" ID-CoL="58V4J" authority="Winterbottom, 2000:58" authorityName="Winterbottom" authorityPageNumber="58" authorityYear="2000" box="[151,555,1044,1066]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="anaima">
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<emphasis id="EFC4EA9EB804F342FF1FFBE36A75EF52" box="[151,312,1045,1066]" italics="true" pageId="7" pageNumber="8">Trimma anaima</emphasis>
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<treatmentCitation id="5C11109DB804F342FEB7FBE26966EF52" author="Winterbottom" box="[319,555,1044,1066]" page="58" pageId="7" pageNumber="8" year="2000">
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<bibRefCitation id="B9214B7DB804F342FEB7FBE26942EF52" author="Winterbottom" box="[319,527,1044,1066]" pageId="7" pageNumber="8" refString="Winterbottom, R. (2000) Four new species of Trimma (Gobiidae), from the Indian and western Pacific oceans. aqua, Journal of Ichthyology and Aquatic Biology, 4 (2), 57 - 66." type="journal article" year="2000">Winterbottom, 2000</bibRefCitation>
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:58
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||
</treatmentCitation>
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||
</taxonomicName>
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||
(Anjouan Id,
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<collectingCountry id="A5A7761CB804F342FD32FBE26857EF52" box="[698,794,1044,1066]" name="Comoros" pageId="7" pageNumber="8">Comores</collectingCountry>
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);
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<treatmentCitation id="5C11109DB804F342FCA4FBE26F15EF52" author="Hayashi" box="[812,1112,1044,1066]" page="41" pageId="7" pageNumber="8" year="2003">
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<bibRefCitation id="B9214B7DB804F342FCA4FBE26F76EF52" author="Hayashi" box="[812,1083,1044,1066]" pageId="7" pageNumber="8" refString="Hayashi, M. & Shiratori, T. (2003) Gobies of Japanese Waters. Hankyu Books, Osaka, 224 pp. [in Japanese]" type="book" year="2003">Hayashi & Shiratori, 2003</bibRefCitation>
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||
:41
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</treatmentCitation>
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(
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||
<collectingCountry id="A5A7761CB804F342FBEEFBE26FE8EF52" box="[1126,1189,1044,1066]" name="Japan" pageId="7" pageNumber="8">Japan</collectingCountry>
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);
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<treatmentCitation id="5C11109DB804F342FB3FFBE26ED5EF52" author="Allen" box="[1207,1432,1044,1066]" page="331" pageId="7" pageNumber="8" year="2004">
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<bibRefCitation id="B9214B7DB804F342FB3FFBE26E20EF52" author="Allen" box="[1207,1389,1044,1066]" pageId="7" pageNumber="8" refString="Allen, G., Steene, R., Humann, P. & DeLoach, N. (2004) Reef Fish Identification. Tropical Pacific. New World Publications Ltd., Florida, 457 pp." type="book" year="2004">
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||
Allen
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||
<emphasis id="EFC4EA9EB804F342FB7DFBE36E63EF52" box="[1269,1326,1044,1066]" italics="true" pageId="7" pageNumber="8">et al.,</emphasis>
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||
2004
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||
</bibRefCitation>
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||
:331
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||
</treatmentCitation>
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;
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||
<treatmentCitation id="5C11109DB804F342FF4FFBC56ADBEF31" author="Anderson" box="[199,406,1075,1097]" page="104" pageId="7" pageNumber="8" year="2005">
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<bibRefCitation id="B9214B7DB804F342FF4FFBC56A21EF31" author="Anderson" box="[199,364,1075,1097]" pageId="7" pageNumber="8" refString="Anderson, R. C. (2005) Reef Fishes of the Maldives. Manta Marine Pvt Ltd., Maldives, vi + 130 pp." type="book" year="2005">Anderson, 2005</bibRefCitation>
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||
:104
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||
</treatmentCitation>
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(
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||
<collectingCountry id="A5A7761CB804F342FE2BFBC5694AEF31" box="[419,519,1075,1097]" name="Maldives" pageId="7" pageNumber="8">Maldives</collectingCountry>
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);
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||
<treatmentCitation id="5C11109DB804F342FD90FBC56874EF31" author="Allen" box="[536,825,1075,1097]" page="934" pageId="7" pageNumber="8" year="2012">
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<bibRefCitation id="B9214B7DB804F342FD90FBC56842EF31" author="Allen" box="[536,783,1075,1097]" pageId="7" pageNumber="8" refString="Allen, G. R. & Erdmann, M. V. (2012) Reef fishes of the East Indies. Vol. III. Tropical Reef Research, Perth, pp. 857 - 1292." type="book chapter" year="2012">Allen & Erdmann, 2012</bibRefCitation>
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||
:934
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||
</treatmentCitation>
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||
(Indo-west Pacific).
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="DD0F368CB804F342FF1FFBA56820EF10" blockId="7.[151,1436,1044,1189]" box="[151,877,1106,1128]" pageId="7" pageNumber="8">
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<treatmentCitationGroup id="FDA011A2B804F342FF1FFBA56820EF10" box="[151,877,1106,1128]" pageId="7" pageNumber="8">
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<taxonomicName id="1AB04D0FB804F342FF1FFBA56A75EF10" box="[151,312,1107,1128]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="amaina">
|
||
<emphasis id="EFC4EA9EB804F342FF1FFBA56A75EF10" box="[151,312,1107,1128]" italics="true" pageId="7" pageNumber="8">Trimma amaina</emphasis>
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||
</taxonomicName>
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||
(lapsus):
|
||
<treatmentCitation id="5C11109DB804F342FE14FBA469F2EF10" author="Kuiter" box="[412,703,1106,1128]" page="73" pageId="7" pageNumber="8" year="2004">
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<bibRefCitation id="B9214B7DB804F342FE14FBA469EEEF10" author="Kuiter" box="[412,675,1106,1128]" pageId="7" pageNumber="8" refString="Kuiter, R. H. & Tonozuka, T. (2004) Pictorial Guide to Indonesian Reef Fishes. Part 3. PT Dive & Dive's, Bali, Indonesia, iv + 623 - 893 pp." type="book chapter" year="2004">Kuiter & Tonozuka, 2004</bibRefCitation>
|
||
:73
|
||
</treatmentCitation>
|
||
(
|
||
<date id="A90E104CB804F342FD46FBA469BDEF10" box="[718,752,1106,1128]" pageId="7" pageNumber="8">x 2</date>
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||
,
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||
<collectingCountry id="A5A7761CB804F342FD73FBA4682FEF10" box="[763,866,1106,1128]" name="Indonesia" pageId="7" pageNumber="8">Indonesia</collectingCountry>
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).
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="DD0F368CB804F342FF1FFB87691FEFFE" blockId="7.[151,1436,1044,1189]" box="[151,594,1136,1158]" pageId="7" pageNumber="8">
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<treatmentCitationGroup id="FDA011A2B804F342FF1FFB87691FEFFE" box="[151,594,1136,1158]" pageId="7" pageNumber="8">
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<taxonomicName id="1AB04D0FB804F342FF1FFB876BABEFFE" ID-CoL="827C" box="[151,230,1137,1158]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
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||
<emphasis id="EFC4EA9EB804F342FF1FFB876BABEFFE" box="[151,230,1137,1158]" italics="true" pageId="7" pageNumber="8">Trimma</emphasis>
|
||
</taxonomicName>
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||
sp. 5:
|
||
<treatmentCitation id="5C11109DB804F342FEA1FB866A9BEFFE" author="Kuiter" box="[297,470,1136,1158]" page="209" pageId="7" pageNumber="8" year="1998">
|
||
<bibRefCitation id="B9214B7DB804F342FEA1FB866AE4EFFE" author="Kuiter" box="[297,425,1136,1158]" pageId="7" pageNumber="8" refString="Kuiter, R. H. (1998) Photo Guide to the Fishes of the Maldives. Atoll Editions, Victoria, 257 pp." type="book" year="1998">Kuiter, 1998</bibRefCitation>
|
||
:209
|
||
</treatmentCitation>
|
||
(
|
||
<collectingCountry id="A5A7761CB804F342FE6AFB86690BEFFE" box="[482,582,1136,1158]" name="Maldives" pageId="7" pageNumber="8">Maldives</collectingCountry>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="DD0F368CB804F342FF1FFB6668D3EFDD" blockId="7.[151,1436,1044,1189]" box="[151,926,1167,1189]" pageId="7" pageNumber="8">
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||
<treatmentCitationGroup id="FDA011A2B804F342FF1FFB6668D3EFDD" box="[151,926,1167,1189]" pageId="7" pageNumber="8">
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||
<taxonomicName id="1AB04D0FB804F342FF1FFB666BABEFDD" ID-CoL="827C" box="[151,230,1168,1189]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
|
||
<emphasis id="EFC4EA9EB804F342FF1FFB666BABEFDD" box="[151,230,1168,1189]" italics="true" pageId="7" pageNumber="8">Trimma</emphasis>
|
||
</taxonomicName>
|
||
RW sp. 28:
|
||
<treatmentCitation id="5C11109DB804F342FEEBFB796910EFDD" author="Winterbottom" box="[355,605,1167,1189]" page="255" pageId="7" pageNumber="8" year="1990">
|
||
<bibRefCitation id="B9214B7DB804F342FEEBFB79697FEFDD" author="Winterbottom" box="[355,562,1167,1189]" pageId="7" pageNumber="8" refString="Winterbottom, R. (1990) The Trimmatom nanus species complex (Actinopterygii, Gobiidae): phylogeny and progenetic heterochrony. Systematic Zoology, 39, 253 - 265. http: // dx. doi. org / 10.2307 / 2992185" type="journal article" year="1990">Winterbottom, 1990</bibRefCitation>
|
||
:255
|
||
</treatmentCitation>
|
||
(comparison with
|
||
<taxonomicName id="1AB04D0FB804F342FC93FB6668DDEFDD" ID-CoL="827D" box="[795,912,1168,1189]" class="Actinopterygii" family="Gobiidae" genus="Trimmatom" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
|
||
<emphasis id="EFC4EA9EB804F342FC93FB6668DDEFDD" box="[795,912,1168,1189]" italics="true" pageId="7" pageNumber="8">Trimmatom</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="DD0F368CB804F342FF1FFB2769F8EE49" blockId="7.[151,1437,1233,2015]" pageId="7" pageNumber="8">
|
||
<emphasis id="EFC4EA9EB804F342FF1FFB276969EF92" bold="true" box="[151,548,1233,1258]" pageId="7" pageNumber="8">Australian Material.Queensland:</emphasis>
|
||
<collectingCountry id="A5A7761CB804F342FDA7FB2769DBEF92" box="[559,662,1233,1258]" name="Ashmore and Cartier Islands" pageId="7" pageNumber="8">Ashmore</collectingCountry>
|
||
Reef: AMS I.33728-043, 6(11–12),
|
||
<quantity id="1A489B69B804F342FBC0FB276FFFEF91" box="[1096,1202,1233,1258]" metricMagnitude="1" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="7" pageNumber="8" unit="m" value="13.0" valueMax="15.0" valueMin="11.0">11–15 m</quantity>
|
||
; AMS I.33736-050, (13),
|
||
<quantity id="1A489B69B804F342FF5CFB006A77EE76" box="[212,314,1270,1294]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="7" pageNumber="8" unit="m" value="25.0" valueMax="30.0" valueMin="20.0">
|
||
20–
|
||
<specimenCount id="CBB6FD05B804F342FF76FB006A77EE76" box="[254,314,1270,1294]" pageId="7" pageNumber="8" type="generic">30 m</specimenCount>
|
||
</quantity>
|
||
. Near Lizard Id: WAM P.28531-010, (15),
|
||
<quantity id="1A489B69B804F342FC94FB0368CFEE76" box="[796,898,1269,1294]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="7" pageNumber="8" unit="m" value="12.5" valueMax="15.0" valueMin="10.0">
|
||
10–
|
||
<specimenCount id="CBB6FD05B804F342FCC0FB0368CFEE76" box="[840,898,1269,1294]" pageId="7" pageNumber="8" type="generic">15 m</specimenCount>
|
||
</quantity>
|
||
. Osprey Reef: AMS I.25112-036, (14),
|
||
<quantity id="1A489B69B804F342FAC8FB036ED5EE76" box="[1344,1432,1269,1294]" metricMagnitude="0" metricUnit="m" metricValue="8.0" metricValueMax="15.0" metricValueMin="1.0" pageId="7" pageNumber="8" unit="m" value="8.0" valueMax="15.0" valueMin="1.0">
|
||
1–
|
||
<specimenCount id="CBB6FD05B804F342FAD3FB036ED5EE76" box="[1371,1432,1269,1294]" pageId="7" pageNumber="8" type="generic">15 m</specimenCount>
|
||
</quantity>
|
||
. Portlock Reef: AMS I.33751-071, (12),
|
||
<quantity id="1A489B69B804F342FDD0FAEF69FDEE49" box="[600,688,1305,1330]" metricMagnitude="1" metricUnit="m" metricValue="1.8" metricValueMax="3.1" metricValueMin="0.5" pageId="7" pageNumber="8" unit="m" value="18.0" valueMax="31.0" valueMin="5.0">
|
||
5–
|
||
<specimenCount id="CBB6FD05B804F342FDFDFAEF69FDEE49" box="[629,688,1305,1330]" pageId="7" pageNumber="8" type="generic">31 m</specimenCount>
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="95AA6507B804F342FF4FFACB6A42ED0E" pageId="7" pageNumber="8" type="materials_examined">
|
||
<paragraph id="DD0F368CB804F342FF4FFACB6A42ED0E" blockId="7.[151,1437,1233,2015]" pageId="7" pageNumber="8">
|
||
<emphasis id="EFC4EA9EB804F342FF4FFACB6AC8EE2E" bold="true" box="[199,389,1341,1366]" pageId="7" pageNumber="8">Other material.</emphasis>
|
||
See
|
||
<bibRefCitation id="B9214B7DB804F342FE4AFACB69FEEE2E" author="Winterbottom" box="[450,691,1341,1366]" pageId="7" pageNumber="8" refString="Winterbottom, R. (2000) Four new species of Trimma (Gobiidae), from the Indian and western Pacific oceans. aqua, Journal of Ichthyology and Aquatic Biology, 4 (2), 57 - 66." type="journal article" year="2000">Winterbottom (2000)</bibRefCitation>
|
||
for a list of material examined in the original description. New material includes:
|
||
<emphasis id="EFC4EA9EB804F342FEEEFA976AA9EE02" bold="true" box="[358,484,1377,1402]" pageId="7" pageNumber="8">
|
||
<collectingCountry id="A5A7761CB804F342FEEEFA976A92EE02" box="[358,479,1377,1402]" name="Indonesia" pageId="7" pageNumber="8">Indonesia</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
Raja Ampat,
|
||
<collectionCode id="BBA1AE49B804F342FD08FA94698DEE02" LSID="urn:lsid:biocol.org:col:34778" box="[640,704,1378,1402]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="7" pageNumber="8">ROM</collectionCode>
|
||
85248, 2(11–13),
|
||
<quantity id="1A489B69B804F342FC07FA9768BBEE01" box="[911,1014,1377,1402]" metricMagnitude="1" metricUnit="m" metricValue="1.9" metricValueMax="2.6" metricValueMin="1.2" pageId="7" pageNumber="8" unit="m" value="19.0" valueMax="26.0" valueMin="12.0">
|
||
12–
|
||
<specimenCount id="CBB6FD05B804F342FC31FA9768BBEE01" box="[953,1014,1377,1402]" pageId="7" pageNumber="8" type="generic">26 m</specimenCount>
|
||
</quantity>
|
||
.
|
||
<emphasis id="EFC4EA9EB804F342FB88FA976FF2EE02" bold="true" box="[1024,1215,1377,1402]" pageId="7" pageNumber="8">
|
||
<collectingCountry id="A5A7761CB804F342FB88FA976FF6EE02" box="[1024,1211,1377,1402]" name="New Caledonia" pageId="7" pageNumber="8">New Caledonia</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
Loyalty Id:
|
||
<collectionCode id="BBA1AE49B804F342FAC3FA946ED1EE02" LSID="urn:lsid:biocol.org:col:34871" box="[1355,1436,1378,1402]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="8">USNM</collectionCode>
|
||
321802, 2,
|
||
<quantity id="1A489B69B804F342FE91FA706ACFEEE6" box="[281,386,1414,1438]" metricMagnitude="1" metricUnit="m" metricValue="2.6" metricValueMax="2.9" metricValueMin="2.3" pageId="7" pageNumber="8" unit="m" value="26.0" valueMax="29.0" valueMin="23.0">
|
||
23–
|
||
<specimenCount id="CBB6FD05B804F342FECBFA706ACFEEE6" box="[323,386,1414,1438]" pageId="7" pageNumber="8" type="generic">29 m</specimenCount>
|
||
</quantity>
|
||
.
|
||
<emphasis id="EFC4EA9EB804F342FE06FA736A91EEE6" bold="true" box="[398,476,1413,1438]" pageId="7" pageNumber="8">
|
||
<collectingCountry id="A5A7761CB804F342FE06FA736A9AEEE6" box="[398,471,1413,1438]" name="Palau" pageId="7" pageNumber="8">Palau</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
Main Ids:
|
||
<collectionCode id="BBA1AE49B804F342FDD5FA7069D0EEE6" LSID="urn:lsid:biocol.org:col:34778" box="[605,669,1414,1438]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="7" pageNumber="8">ROM</collectionCode>
|
||
80303, (14),
|
||
<quantity id="1A489B69B804F342FCB4FA7068E9EEE6" box="[828,932,1414,1438]" metricMagnitude="1" metricUnit="m" metricValue="3.45" metricValueMax="4.2" metricValueMin="2.7" pageId="7" pageNumber="8" unit="m" value="34.5" valueMax="42.0" valueMin="27.0">27–42 m</quantity>
|
||
;
|
||
<collectionCode id="BBA1AE49B804F342FC39FA7068BCEEE6" LSID="urn:lsid:biocol.org:col:34778" box="[945,1009,1414,1438]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="7" pageNumber="8">ROM</collectionCode>
|
||
80028, (12),
|
||
<quantity id="1A489B69B804F342FB07FA736FB5EEE6" box="[1167,1272,1413,1438]" metricMagnitude="1" metricUnit="m" metricValue="3.95" metricValueMax="4.5" metricValueMin="3.4" pageId="7" pageNumber="8" unit="m" value="39.5" valueMax="45.0" valueMin="34.0">34–45 m</quantity>
|
||
;
|
||
<collectionCode id="BBA1AE49B804F342FA8EFA706E0BEEE6" LSID="urn:lsid:biocol.org:col:34778" box="[1286,1350,1414,1438]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="7" pageNumber="8">ROM</collectionCode>
|
||
80351, 2(14–15),
|
||
<quantity id="1A489B69B804F342FE85FA5C6A0FEEBA" box="[269,322,1450,1474]" metricMagnitude="1" metricUnit="m" metricValue="7.3" pageId="7" pageNumber="8" unit="m" value="73.0">73m</quantity>
|
||
;
|
||
<collectionCode id="BBA1AE49B804F342FEC7FA5C6AC2EEBA" LSID="urn:lsid:biocol.org:col:34778" box="[335,399,1450,1474]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="7" pageNumber="8">ROM</collectionCode>
|
||
80395, 2(11–11),
|
||
<quantity id="1A489B69B804F342FDE9FA5C6985EEB9" box="[609,712,1450,1474]" metricMagnitude="1" metricUnit="m" metricValue="2.35" metricValueMax="2.7" metricValueMin="2.0" pageId="7" pageNumber="8" unit="m" value="23.5" valueMax="27.0" valueMin="20.0">20–27 m</quantity>
|
||
;
|
||
<collectionCode id="BBA1AE49B804F342FD5DFA5C6858EEBA" LSID="urn:lsid:biocol.org:col:34778" box="[725,789,1450,1474]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="7" pageNumber="8">ROM</collectionCode>
|
||
80404, 2(13–14),
|
||
<quantity id="1A489B69B804F342FC61FA5C6F05EEBA" box="[1001,1096,1450,1474]" metricMagnitude="1" metricUnit="m" metricValue="3.9" metricValueMax="4.4" metricValueMin="3.4" pageId="7" pageNumber="8" unit="m" value="39.0" valueMax="44.0" valueMin="34.0">34–44m</quantity>
|
||
;
|
||
<collectionCode id="BBA1AE49B804F342FBDDFA5C6FD8EEBA" LSID="urn:lsid:biocol.org:col:34778" box="[1109,1173,1450,1474]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="7" pageNumber="8">ROM</collectionCode>
|
||
80469, (12),
|
||
<quantity id="1A489B69B804F342FAB9FA5C6ED5EEB9" box="[1329,1432,1450,1474]" metricMagnitude="1" metricUnit="m" metricValue="4.45" metricValueMax="4.9" metricValueMin="4.0" pageId="7" pageNumber="8" unit="m" value="44.5" valueMax="49.0" valueMin="40.0">
|
||
40–
|
||
<specimenCount id="CBB6FD05B804F342FAD3FA5C6ED5EEB9" box="[1371,1432,1450,1474]" pageId="7" pageNumber="8" type="generic">49 m</specimenCount>
|
||
</quantity>
|
||
. Southwest Ids:
|
||
<collectionCode id="BBA1AE49B804F342FECBFA386ACEEE9E" LSID="urn:lsid:biocol.org:col:34778" box="[323,387,1486,1510]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="7" pageNumber="8">ROM</collectionCode>
|
||
82989, (12.0),
|
||
<quantity id="1A489B69B804F342FDA6FA3B69DEEE9E" box="[558,659,1485,1510]" metricMagnitude="1" metricUnit="m" metricValue="2.8" metricValueMax="3.6" metricValueMin="2.0" pageId="7" pageNumber="8" unit="m" value="28.0" valueMax="36.0" valueMin="20.0">20–36 m</quantity>
|
||
;
|
||
<collectionCode id="BBA1AE49B804F342FD16FA386993EE9E" LSID="urn:lsid:biocol.org:col:34778" box="[670,734,1486,1510]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="7" pageNumber="8">ROM</collectionCode>
|
||
83022, (13.7),
|
||
<quantity id="1A489B69B804F342FC01FA3868A2EE9E" box="[905,1007,1486,1510]" metricMagnitude="1" metricUnit="m" metricValue="2.55" metricValueMax="3.1" metricValueMin="2.0" pageId="7" pageNumber="8" unit="m" value="25.5" valueMax="31.0" valueMin="20.0">20–31 m</quantity>
|
||
;
|
||
<collectionCode id="BBA1AE49B804F342FC72FA386F77EE9E" LSID="urn:lsid:biocol.org:col:34778" box="[1018,1082,1486,1510]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="7" pageNumber="8">ROM</collectionCode>
|
||
83048, 2(13.6–15.5),
|
||
<quantity id="1A489B69B804F342FABAFA386ED5EE9E" box="[1330,1432,1486,1510]" metricMagnitude="1" metricUnit="m" metricValue="2.1" metricValueMax="3.0" metricValueMin="1.2" pageId="7" pageNumber="8" unit="m" value="21.0" valueMax="30.0" valueMin="12.0">12–30 m</quantity>
|
||
;
|
||
<collectionCode id="BBA1AE49B804F342FF1FFA046B9AED72" LSID="urn:lsid:biocol.org:col:34778" box="[151,215,1522,1546]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="7" pageNumber="8">ROM</collectionCode>
|
||
83071, (12.9),
|
||
<quantity id="1A489B69B804F342FE02FA076ABFED71" box="[394,498,1521,1546]" metricMagnitude="1" metricUnit="m" metricValue="2.2" metricValueMax="2.8" metricValueMin="1.6" pageId="7" pageNumber="8" unit="m" value="22.0" valueMax="28.0" valueMin="16.0">16–28 m</quantity>
|
||
;
|
||
<collectionCode id="BBA1AE49B804F342FD88FA04690DED72" LSID="urn:lsid:biocol.org:col:34778" box="[512,576,1522,1546]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="7" pageNumber="8">ROM</collectionCode>
|
||
83262, 8(9.8–11.7),
|
||
<quantity id="1A489B69B804F342FCBAFA0768D7ED71" box="[818,922,1521,1546]" metricMagnitude="1" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="7" pageNumber="8" unit="m" value="27.5" valueMax="35.0" valueMin="20.0">20–35 m</quantity>
|
||
;
|
||
<collectionCode id="BBA1AE49B804F342FC20FA0468A5ED72" LSID="urn:lsid:biocol.org:col:34778" box="[936,1000,1522,1546]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="7" pageNumber="8">ROM</collectionCode>
|
||
83269, 2(11.5–11.9),
|
||
<quantity id="1A489B69B804F342FB6EFA046E03ED71" box="[1254,1358,1522,1546]" metricMagnitude="1" metricUnit="m" metricValue="2.8" metricValueMax="3.4" metricValueMin="2.2" pageId="7" pageNumber="8" unit="m" value="28.0" valueMax="34.0" valueMin="22.0">22–34 m</quantity>
|
||
;
|
||
<collectionCode id="BBA1AE49B804F342FAD4FA046ED1ED72" LSID="urn:lsid:biocol.org:col:34778" box="[1372,1436,1522,1546]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="7" pageNumber="8">ROM</collectionCode>
|
||
83331, (11.8),
|
||
<quantity id="1A489B69B804F342FECEF9E06AFCED56" box="[326,433,1558,1582]" metricMagnitude="1" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="7" pageNumber="8" unit="m" value="20.0" valueMax="30.0" valueMin="10.0">10–30 m</quantity>
|
||
;
|
||
<collectionCode id="BBA1AE49B804F342FE4AF9E0694FED56" LSID="urn:lsid:biocol.org:col:34778" box="[450,514,1558,1582]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="7" pageNumber="8">ROM</collectionCode>
|
||
83373, 5(9.5–12.5),
|
||
<quantity id="1A489B69B804F342FD75F9E36825ED56" box="[765,872,1557,1582]" metricMagnitude="1" metricUnit="m" metricValue="2.95" metricValueMax="3.6" metricValueMin="2.3" pageId="7" pageNumber="8" unit="m" value="29.5" valueMax="36.0" valueMin="23.0">
|
||
23–
|
||
<specimenCount id="CBB6FD05B804F342FCAFF9E36825ED56" box="[807,872,1557,1582]" pageId="7" pageNumber="8" type="generic">36 m</specimenCount>
|
||
</quantity>
|
||
.
|
||
<emphasis id="EFC4EA9EB804F342FCF0F9E36F4AED56" bold="true" box="[888,1031,1557,1582]" pageId="7" pageNumber="8">
|
||
<collectingCountry id="A5A7761CB804F342FCF0F9E36F4FED56" box="[888,1026,1557,1582]" name="Philippines" pageId="7" pageNumber="8">Philippines</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
Palawan, Dilumacad Id:
|
||
<collectionCode id="BBA1AE49B804F342FAB0F9E06E30ED56" LSID="urn:lsid:biocol.org:col:34981" box="[1336,1405,1558,1582]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="7" pageNumber="8">WAM</collectionCode>
|
||
P. 32877.001, 2(19–20),
|
||
<quantity id="1A489B69B804F342FE1BF9CC6AB7ED29" box="[403,506,1594,1618]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="7" pageNumber="8" unit="m" value="25.0" valueMax="30.0" valueMin="20.0">
|
||
20–
|
||
<specimenCount id="CBB6FD05B804F342FE35F9CC6AB7ED29" box="[445,506,1594,1618]" pageId="7" pageNumber="8" type="generic">30 m</specimenCount>
|
||
</quantity>
|
||
. Pamilacan Id:
|
||
<collectionCode id="BBA1AE49B804F342FD21F9CC69B7ED2A" LSID="urn:lsid:biocol.org:col:34871" box="[681,762,1594,1618]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="8">USNM</collectionCode>
|
||
244051,
|
||
<quantity id="1A489B69B804F342FCEEF9CF689EED29" box="[870,979,1593,1618]" metricMagnitude="1" metricUnit="m" metricValue="1.675" metricValueMax="3.35" metricValueMin="0.0" pageId="7" pageNumber="8" unit="m" value="16.75" valueMax="33.5" valueMin="0.0">0–33.5 m</quantity>
|
||
.
|
||
<emphasis id="EFC4EA9EB804F342FC57F9CF6F11ED2A" bold="true" box="[991,1116,1593,1618]" pageId="7" pageNumber="8">Solomons:</emphasis>
|
||
Stewart Id:
|
||
<collectionCode id="BBA1AE49B804F342FB60F9CC6E74ED2A" LSID="urn:lsid:biocol.org:col:34871" box="[1256,1337,1594,1618]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="8">USNM</collectionCode>
|
||
365584, 2,
|
||
<quantity id="1A489B69B804F342FF3BF9A86A46ED0E" box="[179,267,1630,1654]" metricMagnitude="1" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="7" pageNumber="8" unit="m" value="10.0" valueMax="20.0" valueMin="0.0">
|
||
0–
|
||
<specimenCount id="CBB6FD05B804F342FF47F9A86A46ED0E" box="[207,267,1630,1654]" pageId="7" pageNumber="8" type="generic">20 m</specimenCount>
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="95AA6507B804F342FF4FF9776950ECEE" pageId="7" pageNumber="8" type="diagnosis">
|
||
<paragraph id="DD0F368CB804F342FF4FF9776950ECEE" blockId="7.[151,1437,1233,2015]" pageId="7" pageNumber="8">
|
||
<emphasis id="EFC4EA9EB804F342FF4FF9776A0CEDE2" bold="true" box="[199,321,1665,1690]" pageId="7" pageNumber="8">Diagnosis.</emphasis>
|
||
A species of
|
||
<taxonomicName id="1AB04D0FB804F342FE6BF9756977EDE2" box="[483,570,1667,1690]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
|
||
<emphasis id="EFC4EA9EB804F342FE6BF9756977EDE2" box="[483,570,1667,1690]" italics="true" pageId="7" pageNumber="8">Trimma</emphasis>
|
||
</taxonomicName>
|
||
with a bony interorbital about two-thirds pupil diameter in width and only slightly concave; no troughs or trenches in the interorbital or posterodorsal orbital regions; no scales in the predorsal midline; fifth pelvic-fin ray unbranched and 50–60% fourth ray; pectoral-fin rays unbranched; epaxialis extending anteriorly to a vertical in line with the posterior margin of the orbit; no spines of first dorsal fin elongated; body straw-yellow (translucent above midlateral septum, reddish-orange below in life) with most specimens possessing a small dark spot posterodorsally just behind the eye, four small (one-quarter pupil diameter) dark saddles in the dorsal midline at the origins of the first and second dorsal fins, the last dorsal ray and about the mid-length of the caudal peduncle.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="95AA6507B804F34CFF4FF8576F32E8DD" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8" type="description">
|
||
<paragraph id="DD0F368CB804F34DFF4FF857699FE8DC" blockId="7.[151,1437,1233,2015]" lastBlockId="8.[151,1437,151,933]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
|
||
<emphasis id="EFC4EA9EB804F342FF4FF8576A14ECC2" bold="true" box="[199,345,1953,1978]" pageId="7" pageNumber="8">Description.</emphasis>
|
||
The description is based primarily on the
|
||
<typeStatus id="020B882EB804F342FCC8F854683DECC2" box="[832,880,1954,1978]" pageId="7" pageNumber="8">type</typeStatus>
|
||
material from the
|
||
<collectingCountry id="A5A7761CB804F342FBC1F8546FE3ECC2" box="[1097,1198,1954,1978]" name="Comoros" pageId="7" pageNumber="8">Comores</collectingCountry>
|
||
Archipelago. Dorsal fins VI +
|
||
<date id="A90E104CB804F342FE80F8316A64ECA6" box="[264,297,1990,2014]" pageId="7" pageNumber="8">I 8</date>
|
||
, second spine longest (but not elongated), reaching posteriorly to bases of spine or first two rays of second dorsal fin when adpressed, first ray and anterior element of last ray branched or unbranched, last ray reaching posteriorly to midpeduncle in females and juveniles, to or almost to procurrent rays of caudal fin in males; anal fin
|
||
<date id="A90E104CB80BF34DFF7BFF166A5EEB80" box="[243,275,224,248]" pageId="8" pageNumber="9">I 8</date>
|
||
, condition of first and last rays as for dorsal fin; pectoral fin 15–16 (once 17, mean = 15.8), all fin rays unbranched, fin reaching posteriorly to above anterior elements of anal fin; pelvic fin I, 5, first four rays with a single sequential branch, fifth ray unbranched and 50–60% length of fourth, which reaches posteriorly to bases of first few elements of anal fin; fraenum absent, basal membrane vestigial. Lateral scales 23; transverse scales 7; two specimens (not largest) in very good condition with one or two deciduous scales on upper part of opercle (no trace of scales or scale pockets discerned in other specimens); two large vertically aligned scales on pectoral-fin base with a small third scale ventrally; 4–5 irregular rows of scales across midline anterior to pelvic fin; scales on nape, opercle (when present), pectoral base, breast and anterior midline of belly cycloid; others ctenoid; body scales extending anteriorly on either side of midline almost to eye but very deciduous and often missing (with little or no trace of scale pockets). Teeth in inner and outer rows of lower jaw and outer row of upper jaw of curved, spaced, enlarged canines separated by irregular rows of small conical teeth. Tongue narrow and pointed, about half pupil diameter in width. Gill opening to below mid-pupil; outer gill rakers on first arch 2–3 + 11–12 (mean = 2.6 + 11.7). Anterior nares in a short tube, posterior pore-like with a raised rim, nasal sac somewhat elevated, nasal apparatus confined to anterior half of snout. Bony interorbital about two-thirds pupil diameter in width, slightly concave with no interorbital or posterodorsal orbital troughs; epaxialis musculature reaching to above posterior margin of orbit. Abdominal/caudal vertebral transition
|
||
<typeStatus id="020B882EB80BF34DFDCEFD0B693BE86D" box="[582,630,765,789]" pageId="8" pageNumber="9">type</typeStatus>
|
||
intermediate, modified
|
||
<typeStatus id="020B882EB80BF34DFC0AFD0B68F7E86D" box="[898,954,765,789]" pageId="8" pageNumber="9">Type</typeStatus>
|
||
B, with ninth abdominal vertebra lacking a 'haemal arch' (based on two cleared and stained specimens, ROM 1149CS, although there is a medial projection of left haemal arch that does not reach midline on one side of one specimen), tenth vertebra with such an arch, and first abdominal vertebra with two foraminae.
|
||
<collectingCountry id="A5A7761CB80BF34DFD37FC9E6993E8F8" box="[703,734,872,896]" name="American Samoa" pageId="8" pageNumber="9">As</collectingCountry>
|
||
far as is known, this combination, if confirmed on more specimens, is unique within the species of
|
||
<taxonomicName id="1AB04D0FB80BF34DFDFCFC7B6987E8DC" box="[628,714,909,932]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
|
||
<emphasis id="EFC4EA9EB80BF34DFDFCFC7B6987E8DC" box="[628,714,909,932]" italics="true" pageId="8" pageNumber="9">Trimma</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="89CF6604B80BF34DFF1FF8BA686DECF9" httpUri="https://zenodo.org/record/236068/files/figure.png" pageId="8" pageNumber="9" targetBox="[171,1372,1003,1809]" targetPageId="8">
|
||
<paragraph id="DD0F368CB80BF34DFF1FF8BA686DECF9" blockId="8.[151,1436,1868,1921]" pageId="8" pageNumber="9">
|
||
<emphasis id="EFC4EA9EB80BF34DFF1FF8BA6A5EEC1A" bold="true" box="[151,275,1868,1890]" pageId="8" pageNumber="9">FIGURE 2.</emphasis>
|
||
<taxonomicName id="1AB04D0FB80BF34DFE93F8BB6AF0EC1A" box="[283,445,1869,1890]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="anaima">
|
||
<emphasis id="EFC4EA9EB80BF34DFE93F8BB6AF0EC1A" box="[283,445,1869,1890]" italics="true" pageId="8" pageNumber="9">Trimma anaima</emphasis>
|
||
</taxonomicName>
|
||
, A: 13.7 mm SL male, ROM 83022; B: 14.0 mm SL male, ROM 80303 (note parasitic copepod attached to belly), both from Palau. Photos by R. Winterbottom.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DD0F368CB80AF34CFF4FFF616955E9B5" blockId="9.[151,1437,151,1184]" pageId="9" pageNumber="10">
|
||
<emphasis id="EFC4EA9EB80AF34CFF4FFF6169E3EBC8" box="[199,686,151,176]" italics="true" pageId="9" pageNumber="10">Colour pattern. Live and freshly collected</emphasis>
|
||
. Based on colour slides of living specimens from the
|
||
<collectingCountry id="A5A7761CB80AF34CFAA5FF616ED5EBC8" box="[1325,1432,151,176]" name="Maldives" pageId="9" pageNumber="10">Maldives</collectingCountry>
|
||
,
|
||
<collectingCountry id="A5A7761CB80AF34CFF1FFF4A6A49EBAD" box="[151,260,188,213]" name="Indonesia" pageId="9" pageNumber="10">Indonesia</collectingCountry>
|
||
(Pl. 1 A),
|
||
<collectingCountry id="A5A7761CB80AF34CFEF7FF4B6917EBAD" box="[383,602,188,213]" name="Papua New Guinea" pageId="9" pageNumber="10">Papua New Guinea</collectingCountry>
|
||
and
|
||
<collectingCountry id="A5A7761CB80AF34CFD1BFF4A6997EBAC" box="[659,730,188,212]" name="Guam" pageId="9" pageNumber="10">Guam</collectingCountry>
|
||
, and freshly collected specimens from the
|
||
<collectingCountry id="A5A7761CB80AF34CFB49FF4A6E6BEBAC" box="[1217,1318,188,212]" name="Comoros" pageId="9" pageNumber="10">Comores</collectingCountry>
|
||
(13.0 mm SL male
|
||
<typeStatus id="020B882EB80AF34CFE88FF296A28EB80" box="[256,357,223,248]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
|
||
, and
|
||
<quantity id="1A489B69B80AF34CFE2CFF166947EB8F" box="[420,522,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="9" pageNumber="10" unit="mm" value="13.8">13.8 mm</quantity>
|
||
SL female),
|
||
<collectingCountry id="A5A7761CB80AF34CFD2AFF2969ADEB80" box="[674,736,223,248]" name="Palau" pageId="9" pageNumber="10">Palau</collectingCountry>
|
||
(
|
||
<figureCitation id="458B2A09B80AF34CFD7AFF296874EB80" box="[754,825,223,248]" captionStart="FIGURE 2" captionStartId="8.[151,250,1868,1890]" captionTargetBox="[171,1372,1003,1809]" captionTargetId="figure@8.[152,1435,985,1847]" captionTargetPageId="8" captionText="FIGURE 2. Trimma anaima, A: 13.7 mm SL male, ROM 83022; B: 14.0 mm SL male, ROM 80303 (note parasitic copepod attached to belly), both from Palau. Photos by R. Winterbottom." httpUri="https://zenodo.org/record/236068/files/figure.png" pageId="9" pageNumber="10">Fig. 2</figureCitation>
|
||
, 13.7 and
|
||
<quantity id="1A489B69B80AF34CFC3BFF166F54EB8F" box="[947,1049,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="9" pageNumber="10" unit="mm" value="14.2">14.2 mm</quantity>
|
||
SL) and
|
||
<collectingCountry id="A5A7761CB80AF34CFB0EFF296FE0EB80" box="[1158,1197,223,248]" name="Fiji" pageId="9" pageNumber="10">Fiji</collectingCountry>
|
||
(
|
||
<quantity id="1A489B69B80AF34CFB36FF296E6BEB8F" box="[1214,1318,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="9" pageNumber="10" unit="mm" value="13.6">13.6 mm</quantity>
|
||
SL). Live
|
||
<collectingCountry id="A5A7761CB80AF34CFF1FFEF26BB2EA65" box="[151,255,260,285]" name="Maldives" pageId="9" pageNumber="10">Maldives</collectingCountry>
|
||
specimens translucent with reddish wash from below midlateral line to just above ventral midline on body (to just below orbit on head), interorbital region red which may continue posteriorly as an increasingly poorly defined mid-dorsal stripe as it grades to orange (no longer discernible posterior to origin of second dorsal fin), snout orange-red except for a white spot in anterior midline, iris ringed with red with a dorsomedial stripe of electric blue, caudal fin straw-coloured, other fins not discernible. Indonesian specimen similar, with translucent fins and white spot on snout not apparent in available image.
|
||
<collectingCountry id="A5A7761CB80AF34CFCFAFE4E6F1FEAA8" box="[882,1106,439,464]" name="Papua New Guinea" pageId="9" pageNumber="10">Papua New Guinea</collectingCountry>
|
||
specimen with two rows of indistinct reddish orange spots in dorsal fins, a bluish-white bar through top of iris, and iridocytes on cheek just beneath a horizontal line touching ventral margin of orbit that marks ventral extent of reddish body wash.
|
||
<collectingCountry id="A5A7761CB80AF34CFAD0FDF66ED0E960" box="[1368,1437,512,536]" name="Guam" pageId="9" pageNumber="10">Guam</collectingCountry>
|
||
specimen with only a hint of reddish wash on lower half of body, whole fish translucent greyish, with a strong iridescent stripe just below eye. Freshly collected specimens similar (although colouration of dorsal midline cannot be seen) but distal portions of dorsal, anal and pelvic fins may be light orange or yellow, and proximal areas usually hyaline (although colour may persist to distal part of anal fin). Fijian specimen with scattering of iridocytes proximally in dorsal and anal fins.
|
||
</paragraph>
|
||
<paragraph id="DD0F368CB80AF34CFF4FFD216F32E8DD" blockId="9.[151,1437,151,1184]" pageId="9" pageNumber="10">
|
||
<emphasis id="EFC4EA9EB80AF34CFF4FFD216A7AE988" box="[199,311,727,752]" italics="true" pageId="9" pageNumber="10">Preserved</emphasis>
|
||
. Plain straw coloured with a small (one-quarter pupil diameter) black spot made up of melanophores just posterodorsal to orbit (two o'clock position), and four similar-sized dark saddles across dorsum just anterior to origin of first and second dorsal fins, in gap between two dorsal fins, at base of last dorsal-fin ray, and at middle of length of caudal peduncle (all spots may be faint or absent); fins hyaline except in some specimens, where first dorsal fin may have a scattering of melanophores, especially distally. A dark line of melanophores adjacent to base of the anal-fin, continuing in midventral septum to the base of the first procurrent-fin ray.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="95AA6507B80AF34CFF4FFC5968A5E895" pageId="9" pageNumber="10" type="etymology">
|
||
<paragraph id="DD0F368CB80AF34CFF4FFC5968A5E895" blockId="9.[151,1437,151,1184]" pageId="9" pageNumber="10">
|
||
<emphasis id="EFC4EA9EB80AF34CFF4FFC596A03E8B0" bold="true" box="[199,334,943,968]" pageId="9" pageNumber="10">Etymology.</emphasis>
|
||
From the Greek 'anaimos', meaning bloodless, pallid, pale or anaemic, in allusion to the pale, almost washed-out colouration of the species. Used as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="95AA6507B80AF34FFF4FFC016A7DEAA8" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="distribution">
|
||
<paragraph id="DD0F368CB80AF34CFF4FFC016A1EEFD8" blockId="9.[151,1437,151,1184]" pageId="9" pageNumber="10">
|
||
<emphasis id="EFC4EA9EB80AF34CFF4FFC016A2DEF68" bold="true" box="[199,352,1015,1040]" pageId="9" pageNumber="10">Distribution.</emphasis>
|
||
<taxonomicName id="1AB04D0FB80AF34CFEE0FC0F6951EF68" box="[360,540,1017,1040]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="anaima">
|
||
<emphasis id="EFC4EA9EB80AF34CFEE0FC0F6951EF68" box="[360,540,1017,1040]" italics="true" pageId="9" pageNumber="10">Trimma anaima</emphasis>
|
||
</taxonomicName>
|
||
is a relatively rare species in collections, and has usually been taken in lots of one or two specimens. An exception to this is
|
||
<collectingCountry id="A5A7761CB80AF34CFD24FBEA699BEF4D" box="[684,726,1052,1077]" name="Fiji" pageId="9" pageNumber="10">Fiji</collectingCountry>
|
||
, where the ROM has lots of 6, 19 and
|
||
<specimenCount id="CBB6FD05B80AF34CFB10FBEA6E7FEF4D" box="[1176,1330,1052,1077]" pageId="9" pageNumber="10" type="generic">22 specimens</specimenCount>
|
||
from the Great Astrolabe Reef. It has been found from the
|
||
<collectingCountry id="A5A7761CB80AF34CFD45FBB6687FEF20" box="[717,818,1088,1112]" name="Comoros" pageId="9" pageNumber="10">Comores</collectingCountry>
|
||
and
|
||
<collectingCountry id="A5A7761CB80AF34CFCE3FBC9689EEF20" box="[875,979,1087,1112]" name="Maldives" pageId="9" pageNumber="10">Maldives</collectingCountry>
|
||
(photographic record) eastward through the central
|
||
<collectingCountry id="A5A7761CB80AF34CFE9AFB926ADDEF05" box="[274,400,1124,1149]" name="Philippines" pageId="9" pageNumber="10">Philippines</collectingCountry>
|
||
and
|
||
<collectingCountry id="A5A7761CB80AF34CFE4DFB92694EEF05" box="[453,515,1124,1149]" name="Palau" pageId="9" pageNumber="10">Palau</collectingCountry>
|
||
(new record) to
|
||
<collectingCountry id="A5A7761CB80AF34CFD33FB9269AFEF05" box="[699,738,1124,1149]" name="Fiji" pageId="9" pageNumber="10">Fiji</collectingCountry>
|
||
(
|
||
<figureCitation id="458B2A09B80AF34CFD79FB926879EF04" box="[753,820,1124,1149]" captionStart="FIGURE 3" captionStartId="9.[151,250,1948,1970]" captionTargetBox="[162,1425,1246,1915]" captionTargetId="figure@9.[151,1436,1237,1927]" captionTargetPageId="9" captionText="FIGURE 3. Map of the distributions of Trimma anaima (solid circles) and T. annosum (solid triangles). A black star indicates localities at which both species occur." httpUri="https://zenodo.org/record/236069/files/figure.png" pageId="9" pageNumber="10">Fig. 3</figureCitation>
|
||
). There are currently no records from mainland Asia or the Pacific plate.
|
||
</paragraph>
|
||
<caption id="89CF6604B80AF34CFF1FF86A695BECA9" httpUri="https://zenodo.org/record/236069/files/figure.png" pageId="9" pageNumber="10" targetBox="[162,1425,1246,1915]" targetPageId="9">
|
||
<paragraph id="DD0F368CB80AF34CFF1FF86A695BECA9" blockId="9.[151,1436,1948,2001]" pageId="9" pageNumber="10">
|
||
<emphasis id="EFC4EA9EB80AF34CFF1FF86A6A5EECCA" bold="true" box="[151,275,1948,1970]" pageId="9" pageNumber="10">FIGURE 3.</emphasis>
|
||
Map of the distributions of
|
||
<taxonomicName id="1AB04D0FB80AF34CFDBAF868699EECCB" box="[562,723,1950,1971]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="anaima">
|
||
<emphasis id="EFC4EA9EB80AF34CFDBAF868699EECCB" box="[562,723,1950,1971]" italics="true" pageId="9" pageNumber="10">Trimma anaima</emphasis>
|
||
</taxonomicName>
|
||
(solid circles) and
|
||
<taxonomicName id="1AB04D0FB80AF34CFC1FF8686F41ECCB" box="[919,1036,1950,1971]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="annosum">
|
||
<emphasis id="EFC4EA9EB80AF34CFC1FF8686F41ECCB" box="[919,1036,1950,1971]" italics="true" pageId="9" pageNumber="10">T. annosum</emphasis>
|
||
</taxonomicName>
|
||
(solid triangles). A black star indicates localities at which both species occur.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DD0F368CB809F34FFF4FFF616A7DEAA8" blockId="10.[151,1437,151,608]" pageId="10" pageNumber="11">
|
||
<emphasis id="EFC4EA9EB809F34FFF4FFF616A21EBC8" bold="true" box="[199,364,151,176]" pageId="10" pageNumber="11">Comparisons.</emphasis>
|
||
<taxonomicName id="1AB04D0FB809F34FFEF2FF6F697EEBC8" box="[378,563,153,176]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="anaima">
|
||
<emphasis id="EFC4EA9EB809F34FFEF2FF6F697EEBC8" box="[378,563,153,176]" italics="true" pageId="10" pageNumber="11">Trimma anaima</emphasis>
|
||
</taxonomicName>
|
||
is a small (maximum recorded SL =
|
||
<quantity id="1A489B69B809F34FFB80FF616F39EBC8" box="[1032,1140,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.53" pageId="10" pageNumber="11" unit="mm" value="15.3">15.3 mm</quantity>
|
||
) distinctive species. The relatively broad interorbital is somewhat reminiscent of the
|
||
<taxonomicName id="1AB04D0FB809F34FFCB9FF4B68EFEBAC" box="[817,930,189,212]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="tevegae">
|
||
<emphasis id="EFC4EA9EB809F34FFCB9FF4B6873EBAC" box="[817,830,189,212]" italics="true" pageId="10" pageNumber="11">T</emphasis>
|
||
.
|
||
<emphasis id="EFC4EA9EB809F34FFCC4FF4B68EFEBAC" box="[844,930,189,212]" italics="true" pageId="10" pageNumber="11">tevegae</emphasis>
|
||
</taxonomicName>
|
||
species group, all of which have scales in the predorsal midline and a different configuration of the abdominal/caudal vertebrae (see Introduction). At least one other species (
|
||
<taxonomicName id="1AB04D0FB809F34FFEB6FEF369C9EA64" authority="Winterbottom, 2004" authorityName="Winterbottom" authorityYear="2004" box="[318,644,260,285]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="cana">
|
||
<emphasis id="EFC4EA9EB809F34FFEB6FEF36ADEEA64" box="[318,403,261,284]" italics="true" pageId="10" pageNumber="11">T. cana</emphasis>
|
||
Winterbottom, 2004
|
||
</taxonomicName>
|
||
, from the
|
||
<collectingCountry id="A5A7761CB809F34FFD76FEF26831EA65" box="[766,892,260,285]" name="Philippines" pageId="10" pageNumber="11">Philippines</collectingCountry>
|
||
and
|
||
<collectingCountry id="A5A7761CB809F34FFC30FEF268BBEA65" box="[952,1014,260,285]" name="Palau" pageId="10" pageNumber="11">Palau</collectingCountry>
|
||
to the Marshalls Islands and
|
||
<collectingCountry id="A5A7761CB809F34FFAC7FEF26E35EA65" box="[1359,1400,260,285]" name="Fiji" pageId="10" pageNumber="11">Fiji</collectingCountry>
|
||
) is known to have a bright white spot in the midline at the tip of the snout when alive (RW, pers.obs.); this species has red bands on the body (which disappear after preservation), an elongate second dorsal-fin spine, 9–10 dorsal- and anal-fin rays, and 16–18 pectoral-fin rays. Currently, only two specimens (from Raja Ampat,
|
||
<collectingCountry id="A5A7761CB809F34FFB26FE996E50EAF0" box="[1198,1309,367,392]" name="Indonesia" pageId="10" pageNumber="11">Indonesia</collectingCountry>
|
||
) have been analysed for the CO1 gene. They are well separated (ca. 25%) from all other species of the genus based on the distance data.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="95AA6507B809F34FFF4FFE2A68FEE927" pageId="10" pageNumber="11" type="discussion">
|
||
<paragraph id="DD0F368CB809F34FFF4FFE2A68FEE927" blockId="10.[151,1437,151,608]" pageId="10" pageNumber="11">
|
||
<emphasis id="EFC4EA9EB809F34FFF4FFE2A6A01EA8D" bold="true" box="[199,332,476,501]" pageId="10" pageNumber="11">Discussion.</emphasis>
|
||
This species, referred to as
|
||
<taxonomicName id="1AB04D0FB809F34FFD01FE2B69ADEA8C" box="[649,736,477,500]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
|
||
<emphasis id="EFC4EA9EB809F34FFD01FE2B69ADEA8C" box="[649,736,477,500]" italics="true" pageId="10" pageNumber="11">Trimma</emphasis>
|
||
</taxonomicName>
|
||
RW sp. 28, has been suggested as the possible sister group to the genus
|
||
<taxonomicName id="1AB04D0FB809F34FFE80FDF76AC4E960" box="[264,393,513,536]" class="Actinopterygii" family="Gobiidae" genus="Trimmatom" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
|
||
<emphasis id="EFC4EA9EB809F34FFE80FDF76AC4E960" box="[264,393,513,536]" italics="true" pageId="10" pageNumber="11">Trimmatom</emphasis>
|
||
</taxonomicName>
|
||
on the basis of its small adult size, unbranched pectoral-fin rays, unbranched fifth pelvic-fin, and the small dark saddles over the dorsal midline (
|
||
<bibRefCitation id="B9214B7DB809F34FFD60FDD2689FE944" author="Winterbottom" box="[744,978,548,573]" pageId="10" pageNumber="11" refString="Winterbottom, R. (1990) The Trimmatom nanus species complex (Actinopterygii, Gobiidae): phylogeny and progenetic heterochrony. Systematic Zoology, 39, 253 - 265. http: // dx. doi. org / 10.2307 / 2992185" type="journal article" year="1990">Winterbottom, 1990</bibRefCitation>
|
||
). Testing of this hypothesis will have to await a detailed phylogenetic analysis of both
|
||
<taxonomicName id="1AB04D0FB809F34FFD15FDBF69B9E918" box="[669,756,585,608]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
|
||
<emphasis id="EFC4EA9EB809F34FFD15FDBF69B9E918" box="[669,756,585,608]" italics="true" pageId="10" pageNumber="11">Trimma</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="1AB04D0FB809F34FFCA3FDBF68E1E918" box="[811,940,585,608]" class="Actinopterygii" family="Gobiidae" genus="Trimmatom" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
|
||
<emphasis id="EFC4EA9EB809F34FFCA3FDBF68E1E918" box="[811,940,585,608]" italics="true" pageId="10" pageNumber="11">Trimmatom</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |