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<mods:title id="1B5B1C9156C533647F7D7DD46E741CE2">The family Hydropsychidae Curtis (Trichoptera) in the Ryukyu Archipelago, southwestern Japan</mods:title>
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<mods:namePart id="8F044D7A3D5D4D49ADE2309ED7F77E4F">Ito, Tomiko</mods:namePart>
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<treatment id="551587C17F7EFFC628FF31A2E7A0D685" ID-DOI="http://doi.org/10.5281/zenodo.5975790" ID-GBIF-Taxon="154495211" ID-Zenodo-Dep="5975790" LSID="urn:lsid:plazi:treatment:551587C17F7EFFC628FF31A2E7A0D685" httpUri="http://treatment.plazi.org/id/551587C17F7EFFC628FF31A2E7A0D685" lastPageNumber="551" pageId="6" pageNumber="551">
<subSubSection id="95A6655C7F7EFFC628FF31A2E6FBD0C9" box="[151,763,393,419]" pageId="6" pageNumber="551" type="nomenclature">
<paragraph id="DD0336D77F7EFFC628FF31A2E6FBD0C9" blockId="6.[151,763,393,453]" box="[151,763,393,419]" pageId="6" pageNumber="551">
<heading id="864B81BB7F7EFFC628FF31A2E6FBD0C9" bold="true" box="[151,763,393,419]" fontSize="11" level="1" pageId="6" pageNumber="551" reason="1">
<emphasis id="EFC8EAC57F7EFFC628FF31A2E6FBD0C9" bold="true" box="[151,763,393,419]" pageId="6" pageNumber="551">
<taxonomicName id="1ABC4D547F7EFFC628FF31A2E696D0C9" authority="Nozaki and Ito" authorityName="Ito &amp; Nozaki" authorityYear="2018" box="[151,662,393,419]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="551" phylum="Arthropoda" rank="species" species="crassicornea" status="sp. nov.">
<emphasis id="EFC8EAC57F7EFFC628FF31A2E5D5D0C9" bold="true" box="[151,469,393,419]" italics="true" pageId="6" pageNumber="551">Homoplectra crassicornea</emphasis>
Nozaki and Ito
</taxonomicName>
<taxonomicNameLabel id="F4FB57BE7F7EFFC62AF531A2E6FBD0C9" box="[669,763,393,419]" pageId="6" pageNumber="551" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="95A6655C7F7EFFC628FF3187E6D0D360" pageId="6" pageNumber="551" type="description">
<paragraph id="DD0336D77F7EFFC628FF3187E528D0AE" blockId="6.[151,763,393,453]" box="[151,296,428,453]" pageId="6" pageNumber="551">
(
<figureCitation id="45872A527F7EFFC628F73187E4E5D0AE" box="[159,229,428,453]" captionStart="FIGURE 4" captionStartId="7.[151,250,1788,1810]" captionTargetBox="[212,1382,191,1769]" captionTargetId="figure@7.[201,1390,187,1781]" captionTargetPageId="7" captionText="FIGURE 4. Homoplectra crassicornea Nozaki &amp; Ito sp. nov. and H. trifurcata Nozaki &amp; Ito sp. nov. Homoplectra crassicornea (4A4I): male (4A4F): 4A, right wings, dorsal; 4B, genitalia, left lateral; 4C, same, dorsal; 4D, same, ventral; 4E, phallic apparatus, left lateral; 4F, same, dorsal; female (4G4I): 4G, genitalia, left lateral; 4H, same, dorsal; 4I, same, ventral. Homoplectra trifurcata (4J4Q): male (4J4N): 4J, genitalia, left lateral; 4K, same, dorsal; 4L, same, ventral; 4M, phallic apparatus, left lateral; 4N, same, dorsal; female (4O4Q): 4O, genitalia, left lateral; 4P, same, dorsal; 4Q, same, ventral. Abbreviations: IV = apical forks IV; VIIIX = abdominal segments VIIIX; sIX = sternite IX; ae. = aedeagus; d.p.a. = dorsolateral process of aedeagus (paired); d.p.p. = dorsal process of phallotheca (paired); end. = endotheca; l.p.p. = lateral process of phallotheca (paired); pha, phallotheca; v.p.a. = ventral process of aedeagus (paired)." httpUri="https://zenodo.org/record/2606674/files/figure.png" pageId="6" pageNumber="551">Figs 4</figureCitation>
A
<date id="A90210177F7EFFC629623187E520D0AE" box="[266,288,428,452]" pageId="6" pageNumber="551">4I</date>
)
</paragraph>
<paragraph id="DD0336D77F7EFFC628FF31DFE6D0D360" blockId="6.[151,720,500,522]" box="[151,720,500,522]" pageId="6" pageNumber="551">
<taxonomicName id="1ABC4D547F7EFFC628FF31DFE51DD360" box="[151,285,500,522]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="EFC8EAC57F7EFFC628FF31DFE51DD360" box="[151,285,500,522]" italics="true" pageId="6" pageNumber="551">Homoplectra</emphasis>
</taxonomicName>
sp.:
<bibRefCitation id="B92D4B267F7EFFC6292431DFE61FD360" author="Kuhara, N. &amp; Ito, T." box="[332,543,500,522]" pageId="6" pageNumber="551" pagination="11 - 20" refId="ref10739" refString="Kuhara, N. &amp; Ito, T. (2017) Caddisflies (Trichoptera) of Yakushima, northern Ryukyu, southwestern Japan. Biology of Inland Waters, 31, 11 - 20. [in Japanese with English abstract]." type="journal article" year="2017">Kuhara and Ito 2017</bibRefCitation>
, 15, Yaku-shima.
</paragraph>
</subSubSection>
<subSubSection id="95A6655C7F7EFFC628FF321DE6B1D221" pageId="6" pageNumber="551" type="diagnosis">
<paragraph id="DD0336D77F7EFFC628FF321DE52AD3B4" blockId="6.[151,1437,566,2031]" pageId="6" pageNumber="551">
<emphasis id="EFC8EAC57F7EFFC628FF321DE50AD325" bold="true" box="[151,266,566,591]" pageId="6" pageNumber="551">Diagnosis</emphasis>
. The male of this species can be recognized readily by its aedeagus having a pair of ventral processes as compared to a single ventral process with a bifid apex in other known males (
<bibRefCitation id="B92D4B267F7EFFC62C6D3270E09ED319" author="Weaver, J. S. III" box="[1029,1182,602,627]" pageId="6" pageNumber="551" pagination="71 - 77" refId="ref11939" refString="Weaver, J. S. III (1985) A new species and new generic synonym of the Nearctic caddisfly genus Homoplectra (Trichoptera: Hydropsychidae). Entomological News, 96, 71 - 77." type="journal article" year="1985">Weaver 1985</bibRefCitation>
,
<bibRefCitation id="B92D4B267F7EFFC62CC23271E133D319" author="Ruiter, D. E." box="[1194,1331,602,627]" pageId="6" pageNumber="551" pagination="54 - 57" refId="ref11538" refString="Ruiter, D. E. (2003) Two new Trichoptera (Hydropsychidae and Uenoidae) from the Sierra Nevada, California. Pan-Pacific Entomologist, 79, 54 - 57." type="journal article" year="2003">Ruiter 2003</bibRefCitation>
,
<bibRefCitation id="B92D4B267F7EFFC62D573271E4D4D3FD" author="Malicky, H." pageId="6" pageNumber="551" pagination="667 - 686" refId="ref10964" refString="Malicky, H. (2015) Einige neue chinesische Kocherfligen (Trichoptera). Linzer biologische Beitrage, 47, 667 - 686. [in German with English abstract]" type="journal article" year="2015">Malicky 2015</bibRefCitation>
). The female of this species can be distinguished from that of
<taxonomicName id="1ABC4D547F7EFFC62BE03254E015D3FD" authority="Ito &amp; Nozaki, 2018" authorityName="Ito &amp; Nozaki" authorityYear="2018" box="[904,1045,639,663]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="551" phylum="Arthropoda" rank="species" species="trifurcata" status="sp. nov.">
<emphasis id="EFC8EAC57F7EFFC62BE03254E015D3FD" box="[904,1045,639,663]" italics="true" pageId="6" pageNumber="551">H. trifurcata</emphasis>
</taxonomicName>
<emphasis id="EFC8EAC57F7EFFC62C743255E075D3FD" bold="true" box="[1052,1141,638,663]" pageId="6" pageNumber="551">
<taxonomicNameLabel id="F4FB57BE7F7EFFC62C743255E075D3FD" box="[1052,1141,638,663]" pageId="6" pageNumber="551" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the shape of the lateral lobe of the sternite VIII in lateral aspect: The posterodorsal corner of each lobe is round in this species, but acute in
<taxonomicName id="1ABC4D547F7EFFC628FF32ECE523D3B5" authorityName="Ito &amp; Nozaki" authorityYear="2018" box="[151,291,711,735]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="551" phylum="Arthropoda" rank="species" species="trifurcata">
<emphasis id="EFC8EAC57F7EFFC628FF32ECE523D3B5" box="[151,291,711,735]" italics="true" pageId="6" pageNumber="551">H. trifurcata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="DD0336D77F7EFFC628AF32C1E6B1D221" blockId="6.[151,1437,566,2031]" pageId="6" pageNumber="551">
<emphasis id="EFC8EAC57F7EFFC628AF32C1E50CD269" bold="true" box="[199,268,746,771]" pageId="6" pageNumber="551">Adult</emphasis>
(
<figureCitation id="45872A527F7EFFC6297332C1E564D269" box="[283,356,746,771]" captionStart="FIGURE 4" captionStartId="7.[151,250,1788,1810]" captionTargetBox="[212,1382,191,1769]" captionTargetId="figure@7.[201,1390,187,1781]" captionTargetPageId="7" captionText="FIGURE 4. Homoplectra crassicornea Nozaki &amp; Ito sp. nov. and H. trifurcata Nozaki &amp; Ito sp. nov. Homoplectra crassicornea (4A4I): male (4A4F): 4A, right wings, dorsal; 4B, genitalia, left lateral; 4C, same, dorsal; 4D, same, ventral; 4E, phallic apparatus, left lateral; 4F, same, dorsal; female (4G4I): 4G, genitalia, left lateral; 4H, same, dorsal; 4I, same, ventral. Homoplectra trifurcata (4J4Q): male (4J4N): 4J, genitalia, left lateral; 4K, same, dorsal; 4L, same, ventral; 4M, phallic apparatus, left lateral; 4N, same, dorsal; female (4O4Q): 4O, genitalia, left lateral; 4P, same, dorsal; 4Q, same, ventral. Abbreviations: IV = apical forks IV; VIIIX = abdominal segments VIIIX; sIX = sternite IX; ae. = aedeagus; d.p.a. = dorsolateral process of aedeagus (paired); d.p.p. = dorsal process of phallotheca (paired); end. = endotheca; l.p.p. = lateral process of phallotheca (paired); pha, phallotheca; v.p.a. = ventral process of aedeagus (paired)." httpUri="https://zenodo.org/record/2606674/files/figure.png" pageId="6" pageNumber="551">Figs 4</figureCitation>
A
<date id="A90210177F7EFFC629E132C0E5A1D269" box="[393,417,747,771]" pageId="6" pageNumber="551">4I</date>
). Body and forewings mostly brown in alcohol. Forewings each 8.19.0 mm long in male (n = 5),
<quantity id="1A449B327F7EFFC628913325E551D24C" box="[249,337,782,807]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="6" pageNumber="551" unit="mm" value="8.5">8.5 mm</quantity>
long in female (n = 1). Venation typical for the genus. Abdominal sternite V with pair of long finger-like processes near anterolateral margins.
</paragraph>
</subSubSection>
<subSubSection id="95A6655C7F7EFFC628AF337DE7DED49C" pageId="6" pageNumber="551" type="description">
<paragraph id="DD0336D77F7EFFC628AF337DE65CD40D" blockId="6.[151,1437,566,2031]" pageId="6" pageNumber="551">
<emphasis id="EFC8EAC57F7EFFC628AF337DE573D205" bold="true" box="[199,371,854,879]" pageId="6" pageNumber="551">Male genitalia</emphasis>
(
<figureCitation id="45872A527F7EFFC629EB337DE5CCD205" box="[387,460,854,879]" captionStart="FIGURE 4" captionStartId="7.[151,250,1788,1810]" captionTargetBox="[212,1382,191,1769]" captionTargetId="figure@7.[201,1390,187,1781]" captionTargetPageId="7" captionText="FIGURE 4. Homoplectra crassicornea Nozaki &amp; Ito sp. nov. and H. trifurcata Nozaki &amp; Ito sp. nov. Homoplectra crassicornea (4A4I): male (4A4F): 4A, right wings, dorsal; 4B, genitalia, left lateral; 4C, same, dorsal; 4D, same, ventral; 4E, phallic apparatus, left lateral; 4F, same, dorsal; female (4G4I): 4G, genitalia, left lateral; 4H, same, dorsal; 4I, same, ventral. Homoplectra trifurcata (4J4Q): male (4J4N): 4J, genitalia, left lateral; 4K, same, dorsal; 4L, same, ventral; 4M, phallic apparatus, left lateral; 4N, same, dorsal; female (4O4Q): 4O, genitalia, left lateral; 4P, same, dorsal; 4Q, same, ventral. Abbreviations: IV = apical forks IV; VIIIX = abdominal segments VIIIX; sIX = sternite IX; ae. = aedeagus; d.p.a. = dorsolateral process of aedeagus (paired); d.p.p. = dorsal process of phallotheca (paired); end. = endotheca; l.p.p. = lateral process of phallotheca (paired); pha, phallotheca; v.p.a. = ventral process of aedeagus (paired)." httpUri="https://zenodo.org/record/2606674/files/figure.png" pageId="6" pageNumber="551">Figs 4</figureCitation>
B4F). Anterior margin of sternite IX smoothly triangular in lateral aspect; tergite IX semi-sclerotized, mostly fused with segment X. Segment X large, semi-sclerotized dorsally, sclerotized laterally, membranous posteroventrally; spine-like process arising from posterolateral margins, directed dorsad. Inferior appendages long, 1-segmented, extending far beyond apex of segment X, apex with short spine-like setae mesally; in lateral aspect club-shaped, with subapex curved slightly upwards; in dorsal and ventral aspects slightly arched with apices pointed to each other. Phallic apparatus large, complex: Phallotheca (pha.) somewhat conical with pairs of dorsal processes (d.p.p.) and lateral processes (l.p.p); dorsal processes about 2/3 length of lateral ones, in lateral aspect arched, each apex with spine surrounded by hair, in dorsal aspect each constricted laterally at midway, forming square shoulder; lateral processes straight, each with subapex slightly bulged and then narrowed into point, with hair apicodorsally. Aedeagus (ae.) and two pairs of processes arising from endotheca (end.); aedeagus long, slightly shorter than lateral processes of phallotheca, with subapex narrower; dorsolateral processes of aedeagus (d.p.a.) in lateral aspect each thick and horn-like, with apex directed dorsad, in dorsal aspect each arched, with apices pointed to each other, each apex with single spine surrounded by hair; pair of ventral process of aedeagus (v.p.a.) slender, arising from aedeagus basolaterally, directed posteroventrad, shorter than aedeagus, with single spine surrounded by hair apically.
</paragraph>
<paragraph id="DD0336D77F7EFFC628AF3559E7DED49C" blockId="6.[151,1437,566,2031]" pageId="6" pageNumber="551">
<emphasis id="EFC8EAC57F7EFFC628AF3559E58CD4E1" bold="true" box="[199,396,1394,1419]" pageId="6" pageNumber="551">Female genitalia</emphasis>
(
<figureCitation id="45872A527F7EFFC629F43559E5E4D4E1" box="[412,484,1394,1419]" captionStart="FIGURE 4" captionStartId="7.[151,250,1788,1810]" captionTargetBox="[212,1382,191,1769]" captionTargetId="figure@7.[201,1390,187,1781]" captionTargetPageId="7" captionText="FIGURE 4. Homoplectra crassicornea Nozaki &amp; Ito sp. nov. and H. trifurcata Nozaki &amp; Ito sp. nov. Homoplectra crassicornea (4A4I): male (4A4F): 4A, right wings, dorsal; 4B, genitalia, left lateral; 4C, same, dorsal; 4D, same, ventral; 4E, phallic apparatus, left lateral; 4F, same, dorsal; female (4G4I): 4G, genitalia, left lateral; 4H, same, dorsal; 4I, same, ventral. Homoplectra trifurcata (4J4Q): male (4J4N): 4J, genitalia, left lateral; 4K, same, dorsal; 4L, same, ventral; 4M, phallic apparatus, left lateral; 4N, same, dorsal; female (4O4Q): 4O, genitalia, left lateral; 4P, same, dorsal; 4Q, same, ventral. Abbreviations: IV = apical forks IV; VIIIX = abdominal segments VIIIX; sIX = sternite IX; ae. = aedeagus; d.p.a. = dorsolateral process of aedeagus (paired); d.p.p. = dorsal process of phallotheca (paired); end. = endotheca; l.p.p. = lateral process of phallotheca (paired); pha, phallotheca; v.p.a. = ventral process of aedeagus (paired)." httpUri="https://zenodo.org/record/2606674/files/figure.png" pageId="6" pageNumber="551">Figs 4</figureCitation>
G
<date id="A90210177F7EFFC62A613558E621D4E1" box="[521,545,1395,1419]" pageId="6" pageNumber="551">4I</date>
). Sternite VIII cleft about half distance from base, forming pair of lateral lobes in ventral aspect, posterodorsal corner of each lobe round in lateral aspect. Segment IX triangular in dorsal aspect, oblique S-shaped in lateral aspect, each ventral margin protruding anteriorly. Segment X narrow rectangular in lateral aspect. Vulvar scale large, with X-shaped marking in ventral aspect.
</paragraph>
</subSubSection>
<subSubSection id="95A6655C7F7EFFC628AF3629E651D7C1" pageId="6" pageNumber="551" type="materials_examined">
<paragraph id="DD0336D77F7EFFC628AF3629E663D755" blockId="6.[151,1437,566,2031]" pageId="6" pageNumber="551">
<materialsCitation id="6DD43C8A7F7EFFC628AF3629E663D755" ID-GBIF-Occurrence="2028138231" collectingDate="1992-07-12" collectionCode="CBM-ZI" collectorName="T. Ogata" elevation="620" latitude="30.3794" pageId="6" pageNumber="551" specimenCode="CBM-ZI 166070" specimenCount="1" typeStatus="holotype">
<emphasis id="EFC8EAC57F7EFFC628AF3629E534D771" bold="true" box="[199,308,1538,1563]" pageId="6" pageNumber="551">
<typeStatus id="020788757F7EFFC628AF3629E534D771" box="[199,308,1538,1563]" pageId="6" pageNumber="551" type="holotype">Holotype</typeStatus>
</emphasis>
. Male,
<emphasis id="EFC8EAC57F7EFFC629F43629E628D771" bold="true" box="[412,552,1538,1563]" pageId="6" pageNumber="551">Yaku-shima</emphasis>
: Shiratani-unsui-kyo, Yaku-cho, (
<geoCoordinate id="B88850107F7EFFC62BAE3628E044D771" box="[966,1092,1539,1563]" direction="north" orientation="latitude" pageId="6" pageNumber="551" precision="5" value="30.3794">30.3794°N</geoCoordinate>
, 130.574°3E,
<elevation id="5691D1E47F7EFFC62C9E3629E18CD771" box="[1270,1420,1538,1563]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="6" pageNumber="551" unit="m" value="620.0">
<quantity id="1A449B327F7EFFC62C9E3629E147D770" box="[1270,1351,1538,1563]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="6" pageNumber="551" unit="m" value="620.0">620 m</quantity>
a.s.l.
</elevation>
),
<date id="A90210177F7EFFC628FF360DE519D755" box="[151,281,1574,1599]" pageId="6" pageNumber="551" value="1992-07-12">
<collectingDate id="B946E9FF7F7EFFC628FF360DE519D755" box="[151,281,1574,1599]" pageId="6" pageNumber="551" value="1992-07-12">12.vii.1992</collectingDate>
</date>
,
<collectorName id="704953017F7EFFC6294D360CE584D755" box="[293,388,1575,1599]" pageId="6" pageNumber="551">T. Ogata</collectorName>
(
<specimenCode id="8D1A9EAC7F7EFFC629FB360CE656D755" box="[403,598,1574,1599]" collectionCode="CBM-ZI" pageId="6" pageNumber="551">CBM-ZI 166070</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="DD0336D77F7EFFC628AF3661E7DDD709" blockId="6.[151,1437,566,2031]" box="[199,989,1610,1635]" pageId="6" pageNumber="551">
<materialsCitation id="6DD43C8A7F7EFFC628AF3661E7D9D709" ID-GBIF-Occurrence="2028138236" box="[199,985,1610,1635]" collectingDate="1992-07-12" collectionCode="CBM-ZI" collectorName="T. Ogata" elevation="620" latitude="30.3794" pageId="6" pageNumber="551" specimenCode="CBM-ZI 166071-166073" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<emphasis id="EFC8EAC57F7EFFC628AF3661E534D709" bold="true" box="[199,308,1610,1635]" pageId="6" pageNumber="551">
<typeStatus id="020788757F7EFFC628AF3661E534D709" box="[199,308,1610,1635]" pageId="6" pageNumber="551" type="paratype">Paratype</typeStatus>
</emphasis>
.
<specimenCount id="CBBAFD5E7F7EFFC6292A3660E59BD709" box="[322,411,1610,1635]" pageId="6" pageNumber="551" type="male">3 males</specimenCount>
, same data as holotype (
<specimenCode id="8D1A9EAC7F7EFFC62AC33660E7D1D709" box="[683,977,1610,1635]" collectionCode="CBM-ZI" pageId="6" pageNumber="551">CBM-ZI 166071166073</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="DD0336D77F7EFFC628AF3645E651D7C1" blockId="6.[151,1437,566,2031]" pageId="6" pageNumber="551">
<emphasis id="EFC8EAC57F7EFFC628AF3645E608D7ED" bold="true" box="[199,520,1646,1671]" pageId="6" pageNumber="551">Other specimens examined</emphasis>
.
<emphasis id="EFC8EAC57F7EFFC62A7D3645E6A1D7ED" bold="true" box="[533,673,1646,1671]" pageId="6" pageNumber="551">Yaku-shima</emphasis>
:
<materialsCitation id="6DD43C8A7F7EFFC62ADB3644E01AD7ED" ID-GBIF-Occurrence="2028138220" box="[691,1050,1646,1671]" collectingDate="1992-07-12" collectorName="T. Ogata" elevation="620" latitude="30.3794" pageId="6" pageNumber="551" specimenCount="1" specimenCount-female="1">
<specimenCount id="CBBAFD5E7F7EFFC62ADB3644E717D7ED" box="[691,791,1646,1671]" pageId="6" pageNumber="551" type="female">1 female</specimenCount>
, same data as holotype
</materialsCitation>
;
<materialsCitation id="6DD43C8A7F7EFFC62C763644E64DD7C1" ID-GBIF-Occurrence="2028138234" collectingDate="2006-05-12" collectionCode="TI" latitude="30.304" longLatPrecision="72" longitude="130.509" pageId="6" pageNumber="551" specimenCount="1" specimenCount-male="1">
<specimenCount id="CBBAFD5E7F7EFFC62C763644E06CD7ED" box="[1054,1132,1646,1671]" pageId="6" pageNumber="551" type="male">1 male</specimenCount>
, Ara-kawa, Yakusugi-land,
<geoCoordinate id="B88850107F7EFFC628FF36B8E506D7C1" box="[151,262,1683,1707]" direction="north" orientation="latitude" pageId="6" pageNumber="551" precision="55" value="30.304">30.304°N</geoCoordinate>
,
<geoCoordinate id="B88850107F7EFFC6297936B9E58BD7C1" box="[273,395,1682,1707]" direction="east" orientation="longitude" pageId="6" pageNumber="551" precision="55" value="130.509">130.509°E</geoCoordinate>
,
<date id="A90210177F7EFFC629FE36B9E607D7C1" box="[406,519,1682,1707]" pageId="6" pageNumber="551" value="2006-05-12">
<collectingDate id="B946E9FF7F7EFFC629FE36B9E607D7C1" box="[406,519,1682,1707]" pageId="6" pageNumber="551" value="2006-05-12">12.v.2006</collectingDate>
</date>
,
<collectionCode id="BBADAE127F7EFFC62A7B36B8E630D7C0" box="[531,560,1683,1706]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="6" pageNumber="551">TI</collectionCode>
, S
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95A6655C7F7EFFC628AF369DE19CD7A4" box="[199,1436,1718,1743]" pageId="6" pageNumber="551" type="etymology">
<paragraph id="DD0336D77F7EFFC628AF369DE19CD7A4" blockId="6.[151,1437,566,2031]" box="[199,1436,1718,1743]" pageId="6" pageNumber="551">
<emphasis id="EFC8EAC57F7EFFC628AF369DE546D7A5" bold="true" box="[199,326,1718,1743]" pageId="6" pageNumber="551">Etymology</emphasis>
. The species name (Latin adjective) refers to the thick horn-like processes of the phallic apparatus.
</paragraph>
</subSubSection>
<subSubSection id="95A6655C7F7EFFC628AF36F1E718D67D" pageId="6" pageNumber="551" type="distribution">
<paragraph id="DD0336D77F7EFFC628AF36F1E007D799" blockId="6.[151,1437,566,2031]" box="[199,1031,1754,1779]" pageId="6" pageNumber="551">
<emphasis id="EFC8EAC57F7EFFC628AF36F1E559D799" bold="true" box="[199,345,1754,1779]" pageId="6" pageNumber="551">Distribution</emphasis>
.
<collectingCountry id="A5AB76477F7EFFC6290F36F0E5ABD798" box="[359,427,1755,1778]" name="Japan" pageId="6" pageNumber="551">Japan</collectingCountry>
: Ryukyu (Yaku-shima). Endemic to northern Ryukyu.
</paragraph>
<paragraph id="DD0336D77F7EFFC628AF36D5E718D67D" blockId="6.[151,1437,566,2031]" box="[199,792,1790,1815]" pageId="6" pageNumber="551">
<emphasis id="EFC8EAC57F7EFFC628AF36D5E57CD67D" bold="true" box="[199,380,1790,1815]" pageId="6" pageNumber="551">Japanese name</emphasis>
. Tsuno-nisemiyama-shima-tobikera.
</paragraph>
</subSubSection>
<subSubSection id="95A6655C7F7EFFC628AF3709E7A0D685" pageId="6" pageNumber="551" type="discussion">
<paragraph id="DD0336D77F7EFFC628AF3709E7A0D685" blockId="6.[151,1437,566,2031]" pageId="6" pageNumber="551">
<emphasis id="EFC8EAC57F7EFFC628AF3709E534D651" bold="true" box="[199,308,1826,1851]" pageId="6" pageNumber="551">Remarks</emphasis>
. Several additional unnamed
<taxonomicName id="1ABC4D547F7EFFC62AF63708E732D651" box="[670,818,1827,1851]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="EFC8EAC57F7EFFC62AF63708E732D651" box="[670,818,1827,1851]" italics="true" pageId="6" pageNumber="551">Homoplectra</emphasis>
</taxonomicName>
species have been recorded from Japanese Islands including this species (
<bibRefCitation id="B92D4B267F7EFFC629C6376DE64CD635" author="Nozaki, T." box="[430,588,1862,1887]" pageId="6" pageNumber="551" refId="ref11283" refString="Nozaki, T. (2018) A catalogue of Japanese Trichoptera, Family Hydropsychidae Curtis. Available from: http: // tobikera. eco. coocan. jp / catalog / Hydropsychidae. html (accessed on 19 January 2018)" type="url" year="2018">Nozaki 2018</bibRefCitation>
). Homology of the phallic structure in Japanese and North American
<taxonomicName id="1ABC4D547F7EFFC628FF3740E52BD6E9" box="[151,299,1899,1923]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="EFC8EAC57F7EFFC628FF3740E52BD6E9" box="[151,299,1899,1923]" italics="true" pageId="6" pageNumber="551">Homoplectra</emphasis>
</taxonomicName>
spp. is not yet clear, but processes numbered (1) to (3) and (5) by
<bibRefCitation id="B92D4B267F7EFFC62C633740E092D6E9" author="Ross, H. H." box="[1035,1170,1899,1923]" pageId="6" pageNumber="551" pagination="117 - 124" refId="ref11508" refString="Ross, H. H. (1938) Descriptions of new North American Trichoptera. Proceedings of the Entomological Society of Washington, 40, 117 - 124." type="journal article" year="1938">Ross (1938)</bibRefCitation>
probably correspond to the pairs of dorsal processes and lateral processes of the phallotheca, and the pairs of dorsolateral processes and ventral processes of the aedeagus as they are named in this study, respectively.
<bibRefCitation id="B92D4B267F7EFFC62C333798E0E9D6A1" author="Ross, H. H." box="[1115,1257,1971,1995]" pageId="6" pageNumber="551" pagination="117 - 124" refId="ref11508" refString="Ross, H. H. (1938) Descriptions of new North American Trichoptera. Proceedings of the Entomological Society of Washington, 40, 117 - 124." type="journal article" year="1938">Ross (1938)</bibRefCitation>
numbered and described the aedeagus as “(4) rod-like apex of the aedeagus proper”.
</paragraph>
</subSubSection>
</treatment>
</document>