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<document id="FADAC9F760BEFE153732CD988A460766" ID-CLB-Dataset="28064" ID-DOI="10.11646/zootaxa.4504.4.6" ID-GBIF-Dataset="e597b10d-8e4e-4ffa-8d36-87454b4b35b9" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606666" ID-ZooBank="FA17234E-046E-4165-BEB9-305195E7FD0C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553585146620" checkinUser="plazi" docAuthor="Ito, Tomiko &amp; Nozaki, Takao" docDate="2018" docId="551587C17F77FFD028FF3736E695D39A" docLanguage="en" docName="zootaxa.4504.4.6.pdf" docOrigin="Zootaxa 4504 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrostemum formosicolum" docType="treatment" docVersion="9" lastPageNumber="561" masterDocId="A92CFFB97F78FFC02868302BE400D16A" masterDocTitle="The family Hydropsychidae Curtis (Trichoptera) in the Ryukyu Archipelago, southwestern Japan" masterLastPageNumber="565" masterPageNumber="545" pageNumber="560" updateTime="1698712834044" updateUser="ExternalLinkService">
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<mods:title id="5981FC41FE96B369B9822C8DC7447DB5">The family Hydropsychidae Curtis (Trichoptera) in the Ryukyu Archipelago, southwestern Japan</mods:title>
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<mods:namePart id="94350E17073C035235817CAF4A1A0F9F">Ito, Tomiko</mods:namePart>
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<mods:namePart id="2355BF1FCDA5847D165BD1925A61DC51">Nozaki, Takao</mods:namePart>
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<treatment id="551587C17F77FFD028FF3736E695D39A" ID-DOI="http://doi.org/10.5281/zenodo.5975808" ID-GBIF-Taxon="154495228" ID-Zenodo-Dep="5975808" LSID="urn:lsid:plazi:treatment:551587C17F77FFD028FF3736E695D39A" httpUri="http://treatment.plazi.org/id/551587C17F77FFD028FF3736E695D39A" lastPageId="16" lastPageNumber="561" pageId="15" pageNumber="560">
<subSubSection id="95A6655C7F77FFCF28FF3736E6E5D65D" box="[151,741,1821,1847]" pageId="15" pageNumber="560" type="nomenclature">
<paragraph id="DD0336D77F77FFCF28FF3736E6E5D65D" blockId="15.[151,741,1821,1880]" box="[151,741,1821,1847]" pageId="15" pageNumber="560">
<heading id="864B81BB7F77FFCF28FF3736E6E5D65D" bold="true" box="[151,741,1821,1847]" fontSize="11" level="1" pageId="15" pageNumber="560" reason="1">
<taxonomicName id="1ABC4D547F77FFCF28FF3736E6E5D65D" ID-CoL="6QS78" authority="(Matsumura 1931)" baseAuthorityName="Matsumura" baseAuthorityYear="1931" box="[151,741,1821,1847]" class="Insecta" family="Hydropsychidae" genus="Macrostemum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="560" phylum="Arthropoda" rank="species" species="formosicolum">
<emphasis id="EFC8EAC57F77FFCF28FF3736E6E5D65D" bold="true" box="[151,741,1821,1847]" pageId="15" pageNumber="560">
<emphasis id="EFC8EAC57F77FFCF28FF3736E5F4D65D" bold="true" box="[151,500,1821,1847]" italics="true" pageId="15" pageNumber="560">Macrostemum formosicolum</emphasis>
(
<bibRefCitation id="B92D4B267F77FFCF2A6B3736E6DCD65D" author="Matsumura, S." box="[515,732,1821,1847]" pageId="15" pageNumber="560" refId="ref11129" refString="Matsumura, S. (1931) 6000 illustrated Insects of Japan Empire. Tokoshoin, Tokyo, 1497 pp. [in Japanese]" type="book" year="1931">Matsumura 1931</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95A6655C7F77FFCF28FF3714E533D632" box="[151,307,1855,1880]" pageId="15" pageNumber="560" type="description">
<paragraph id="DD0336D77F77FFCF28FF3714E533D632" blockId="15.[151,741,1821,1880]" box="[151,307,1855,1880]" pageId="15" pageNumber="560">
(
<figureCitation id="45872A527F77FFCF28F73714E4E4D632" box="[159,228,1855,1880]" captionStart="FIGURE 9" captionStartId="17.[151,250,1711,1733]" captionTargetBox="[234,1386,197,1613]" captionTargetId="figure@17.[199,1389,195,1729]" captionTargetPageId="17" captionText="FIGURE 9. Macrostemum formosicolum (Matsumura) and M. okinawanum (Matsumura). Macrostemum formosicolum (9A 9K): Holotype male from Taiwan (9A9F): 9A, habitus, dorsal; 9B, left forewing, dorsal, venation and dark markings; 9C, genitalia, left lateral; 9D, same, dorsal; 9E, right inferior appendage, ventral; 9F, phallic apparatus, left lateral; specimens from Ishigaki-jima (9G9K): 9G1, 9G2, right forewings, venation and dark markings (9G1, female; 9G2, male); 9H, male genitalia, left lateral; 9I, same, dorsal; 9J, phallic apparatus, left lateral; 9K, same, ventral. Macrostemum okinawanum (9L9Y): Holotype male from Okinawa-jima (9L9O): 9L, habitus, dorsal; 9M, right forewing, dorsal; 9N, genitalia, dorsal; 9O, right inferior appendage, ventral; specimens from Okinawa-jima (9P9T): 9P1, 9P2, male right forewings, venation and dark markings; 9Q, male genitalia, left lateral; 9R1, 9R2, male genitalia, dorsal; 9S, phallic apparatus, left lateral; 9T, same, ventral; specimens from Amami-o-shima (9U9Y): 9U1,9U2, male right wings, dorsal; 9V, male genitalia, lateral; 9W, same, dorsal; 9X, phallic apparatus, lateral; 9Y, same, ventral." httpUri="https://zenodo.org/record/2606684/files/figure.png" pageId="15" pageNumber="560">Figs 9</figureCitation>
A9K)
</paragraph>
</subSubSection>
<subSubSection id="95A6655C7F77FFD028FF37A2E5C7D076" lastPageId="16" lastPageNumber="561" pageId="15" pageNumber="560" type="reference_group">
<paragraph id="DD0336D77F77FFCF28FF37A2E76DD6F4" blockId="15.[151,1199,1928,1981]" box="[151,877,1928,1950]" pageId="15" pageNumber="560">
<taxonomicName id="1ABC4D547F77FFCF28FF37A2E694D6F4" authority="Matsumura 1931" authorityName="Matsumura" authorityYear="1931" box="[151,660,1928,1950]" class="Insecta" family="Hydropsychidae" genus="Macronema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="560" phylum="Arthropoda" rank="species" species="formosicolum">
<emphasis id="EFC8EAC57F77FFCF28FF37A2E5A1D6F4" box="[151,417,1928,1950]" italics="true" pageId="15" pageNumber="560">Macronema formosicolum</emphasis>
<bibRefCitation id="B92D4B267F77FFCF29C037A2E657D6F4" author="Matsumura, S." box="[424,599,1928,1950]" pageId="15" pageNumber="560" refId="ref11129" refString="Matsumura, S. (1931) 6000 illustrated Insects of Japan Empire. Tokoshoin, Tokyo, 1497 pp. [in Japanese]" type="book" year="1931">Matsumura 1931</bibRefCitation>
, 1132
</taxonomicName>
1133, male,
<collectingCountry id="A5AB76477F77FFCF2B7337A3E768D6F4" box="[795,872,1928,1950]" name="Taiwan" pageId="15" pageNumber="560">Taiwan</collectingCountry>
.
</paragraph>
<paragraph id="DD0336D77F77FFCF28FF3783E0AFD6D6" blockId="15.[151,1199,1928,1981]" box="[151,1199,1958,1981]" pageId="15" pageNumber="560">
<taxonomicName id="1ABC4D547F77FFCF28FF3783E655D6D6" ID-CoL="6QS78" authority=": Malicky 1998" authorityName=": Malicky" authorityYear="1998" box="[151,597,1958,1981]" class="Insecta" family="Hydropsychidae" genus="Macrostemum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="560" phylum="Arthropoda" rank="species" species="formosicolum">
<emphasis id="EFC8EAC57F77FFCF28FF3783E5B8D6D6" box="[151,440,1958,1981]" italics="true" pageId="15" pageNumber="560">Macrostemum formosicolum</emphasis>
:
<bibRefCitation id="B92D4B267F77FFCF29AD378DE655D6D6" author="Malicky, H." box="[453,597,1958,1980]" pageId="15" pageNumber="560" pagination="767 - 793" refId="ref10834" refString="Malicky, H. (1998) Ein Beitrag zur Kenntnis asiatischer Macronematini (Trichoptera, Hydropsychidae) (Zugleich 24. Arbeit uber thailandische Kocherfliegen). Linzer biologische Beitrage, 30, 767 - 793. [in German with English abstract]" type="journal article" year="1998">Malicky 1998</bibRefCitation>
</taxonomicName>
, 771, pls. 1, 6, male,
<collectingCountry id="A5AB76477F77FFCF2B44378DE77AD6D6" box="[812,890,1958,1980]" name="Taiwan" pageId="15" pageNumber="560">Taiwan</collectingCountry>
;
<bibRefCitation id="B92D4B267F77FFCF2BEC378DE014D6D6" author="Malicky, H." box="[900,1044,1958,1980]" pageId="15" pageNumber="560" pagination="1607 - 1646" refId="ref10930" refString="Malicky, H. (2014) Kocherfliegen (Trichoptera) von Taiwan, mit Neubeschreibungen. Linzer biologische Beitrage, 46, 1607 - 1646. [in German with English abstract]" type="journal article" year="2014">Malicky 2014</bibRefCitation>
, 1628,
<collectingCountry id="A5AB76477F77FFCF2C35378DE0ABD6D6" box="[1117,1195,1958,1980]" name="Taiwan" pageId="15" pageNumber="560">Taiwan</collectingCountry>
.
</paragraph>
<paragraph id="DD0336D77F68FFD028FF30BCE5C7D076" blockId="16.[151,1437,151,752]" pageId="16" pageNumber="561">
<emphasis id="EFC8EAC57F68FFD028FF30BCE515D1DA" bold="true" box="[151,277,151,176]" pageId="16" pageNumber="561">Specimens</emphasis>
.
<materialsCitation id="6DD43C8A7F68FFD0294B30BCE5C7D076" ID-GBIF-Occurrence="2028138227" collectingDate="2009-05-21" collectingDateMax="2011-04-11" collectingDateMin="2009-05-21" collectionCode="TI, MT" collectorName="K. Tojo &amp; Takeda &amp; M. Tanaka" country="Taiwan" county="Nagura-gawa" elevation="70" location="Ishigaki-jima" municipality="Nagura-gawa" pageId="16" pageNumber="561" specimenCount="6" specimenCount-female="4" specimenCount-male="2" typeStatus="holotype">
<typeStatus id="020788757F68FFD0294B30BCE58BD1DA" box="[291,395,151,176]" pageId="16" pageNumber="561" type="holotype">Holotype</typeStatus>
male (pinned, cleared abdomen was dried up in a small vial), labeled “Formosa Matsumura, Type Matsumura,
<taxonomicName id="1ABC4D547F68FFD0290A3096E5E7D1BE" ID-CoL="5JH3" box="[354,487,189,212]" class="Insecta" family="Hydropsychidae" genus="Macronema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="563" phylum="Arthropoda" rank="genus">Macronema</taxonomicName>
formosicot”,
<collectingCountry id="A5AB76477F68FFD02AE93097E6D3D1BF" box="[641,723,188,213]" name="Taiwan" pageId="16" pageNumber="561">Taiwan</collectingCountry>
(HU).
<emphasis id="EFC8EAC57F68FFD02B493097E7C1D1BF" bold="true" box="[801,961,188,213]" pageId="16" pageNumber="561">
<location id="D863600C7F68FFD02B493097E7C1D1BF" LSID="urn:lsid:plazi:treatment:551587C17F77FFD028FF3736E695D39A:D863600C7F68FFD02B493097E7C1D1BF" box="[801,961,188,213]" country="Taiwan" county="Nagura-gawa" municipality="Nagura-gawa" name="Ishigaki-jima" pageId="16" pageNumber="561">Ishigaki-jima</location>
</emphasis>
:
<specimenCount id="CBBAFD5E7F68FFD02BB83097E01DD1BF" box="[976,1053,188,213]" pageId="16" pageNumber="561" type="male">1 male</specimenCount>
,
<location id="D863600C7F68FFD02C403097E079D1BE" LSID="urn:lsid:plazi:treatment:551587C17F77FFD028FF3736E695D39A:D863600C7F68FFD02C403097E079D1BE" box="[1064,1145,188,212]" country="Taiwan" county="Nagura-gawa" municipality="Nagura-gawa" name="Omoto" pageId="16" pageNumber="561">Omoto</location>
,
<collectingMunicipality id="3D67ACAD7F68FFD02CEB3096E11DD1BF" box="[1155,1309,189,213]" pageId="16" pageNumber="561">Nagura-gawa</collectingMunicipality>
,
<date id="A90210177F68FFD02D4F3097E198D1BE" box="[1319,1432,188,212]" pageId="16" pageNumber="561" value="2009-05-21">
<collectingDate id="B946E9FF7F68FFD02D4F3097E198D1BE" box="[1319,1432,188,212]" pageId="16" pageNumber="561" value="2009-05-21">21.v.2009</collectingDate>
</date>
,
<collectorName id="704953017F68FFD028FF30CBE4F6D192" box="[151,246,223,248]" pageId="16" pageNumber="561">K. Tojo</collectorName>
;
<specimenCount id="CBBAFD5E7F68FFD0296E30CBE570D192" box="[262,368,223,248]" pageId="16" pageNumber="561" type="female">1 female</specimenCount>
,
<collectingCounty id="34624E5B7F68FFD029E830CBE61AD192" box="[384,538,224,248]" pageId="16" pageNumber="561">Nagura-gawa</collectingCounty>
, Nagura-dam-ue,
<elevation id="5691D1E47F68FFD02A8630CBE76FD192" box="[750,879,223,248]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="16" pageNumber="561" unit="m" value="70.0">
<quantity id="1A449B327F68FFD02A8630CBE72DD19D" box="[750,813,224,248]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="16" pageNumber="561" unit="m" value="70.0">70 m</quantity>
a.s.l.
</elevation>
,
<date id="A90210177F68FFD02BE930F4E7F7D192" box="[897,1015,223,248]" pageId="16" pageNumber="561" value="2011-04-11">
<collectingDate id="B946E9FF7F68FFD02BE930F4E7F7D192" box="[897,1015,223,248]" pageId="16" pageNumber="561" value="2011-04-11">11.iv.2011</collectingDate>
</date>
,
<collectionCode id="BBADAE127F68FFD02C6130CBE026D19D" box="[1033,1062,224,247]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="16" pageNumber="561">TI</collectionCode>
, P;
<specimenCount id="CBBAFD5E7F68FFD02C3330CBE0AED192" box="[1115,1198,223,248]" pageId="16" pageNumber="561" type="male">1 male</specimenCount>
,
<specimenCount id="CBBAFD5E7F68FFD02CD730CBE134D192" box="[1215,1332,223,248]" pageId="16" pageNumber="561" type="female">3 females</specimenCount>
,
<collectorName id="704953017F68FFD02D2D30F4E198D192" box="[1349,1432,223,248]" pageId="16" pageNumber="561">Takeda</collectorName>
,
<date id="A90210177F68FFD028FF312FE4FAD076" box="[151,250,260,284]" pageId="16" pageNumber="561" value="2010-05-08">
<collectingDate id="B946E9FF7F68FFD028FF312FE4FAD076" box="[151,250,260,284]" pageId="16" pageNumber="561" value="2010-05-08">8.v.2010</collectingDate>
</date>
,
<collectorName id="704953017F68FFD0296D312EE57CD077" box="[261,380,260,285]" pageId="16" pageNumber="561">M. Tanaka</collectorName>
(
<collectionCode id="BBADAE127F68FFD029E4312FE5BBD076" box="[396,443,260,284]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="16" pageNumber="561">MT</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="95A6655C7F68FFD028AF310CE7CAD055" box="[199,970,295,320]" pageId="16" pageNumber="561" type="distribution">
<paragraph id="DD0336D77F68FFD028AF310CE7CAD055" blockId="16.[151,1437,151,752]" box="[199,970,295,320]" pageId="16" pageNumber="561">
<emphasis id="EFC8EAC57F68FFD028AF310CE559D02A" bold="true" box="[199,345,295,320]" pageId="16" pageNumber="561">Distribution</emphasis>
.
<collectingCountry id="A5AB76477F68FFD0290F3103E5ABD055" box="[359,427,296,319]" name="Japan" pageId="16" pageNumber="561">Japan</collectingCountry>
: Ryukyu (Ishigaki-jima).
<collectingCountry id="A5AB76477F68FFD02AA2310CE720D02A" box="[714,800,295,320]" name="Taiwan" pageId="16" pageNumber="561">Taiwan</collectingCountry>
. New to
<collectingCountry id="A5AB76477F68FFD02BEB3103E7C7D055" box="[899,967,296,319]" name="Japan" pageId="16" pageNumber="561">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="95A6655C7F68FFD028AF3167E695D39A" pageId="16" pageNumber="561" type="discussion">
<paragraph id="DD0336D77F68FFD028AF3167E0D0D357" blockId="16.[151,1437,151,752]" pageId="16" pageNumber="561">
<emphasis id="EFC8EAC57F68FFD028AF3167E534D00F" bold="true" box="[199,308,332,357]" pageId="16" pageNumber="561">Remarks</emphasis>
. This species was originally described from
<collectingCountry id="A5AB76477F68FFD02B513167E78BD00F" box="[825,907,332,357]" name="Taiwan" pageId="16" pageNumber="561">Taiwan</collectingCountry>
by
<bibRefCitation id="B92D4B267F68FFD02BD23166E091D00E" author="Matsumura, S." box="[954,1169,332,356]" pageId="16" pageNumber="561" refId="ref11129" refString="Matsumura, S. (1931) 6000 illustrated Insects of Japan Empire. Tokoshoin, Tokyo, 1497 pp. [in Japanese]" type="book" year="1931">Matsumura (1931)</bibRefCitation>
, and previously known only from
<collectingCountry id="A5AB76477F68FFD029793144E563D0E2" box="[273,355,367,392]" name="Taiwan" pageId="16" pageNumber="561">Taiwan</collectingCountry>
(
<bibRefCitation id="B92D4B267F68FFD0291C3144E617D0E2" author="Malicky, H." box="[372,535,367,392]" pageId="16" pageNumber="561" pagination="767 - 793" refId="ref10834" refString="Malicky, H. (1998) Ein Beitrag zur Kenntnis asiatischer Macronematini (Trichoptera, Hydropsychidae) (Zugleich 24. Arbeit uber thailandische Kocherfliegen). Linzer biologische Beitrage, 30, 767 - 793. [in German with English abstract]" type="journal article" year="1998">Malicky 1998</bibRefCitation>
,
<bibRefCitation id="B92D4B267F68FFD02A4C315BE661D0E2" author="Malicky, H." box="[548,609,368,392]" pageId="16" pageNumber="561" pagination="1607 - 1646" refId="ref10930" refString="Malicky, H. (2014) Kocherfliegen (Trichoptera) von Taiwan, mit Neubeschreibungen. Linzer biologische Beitrage, 46, 1607 - 1646. [in German with English abstract]" type="journal article" year="2014">2014</bibRefCitation>
). We examined the
<typeStatus id="020788757F68FFD02B2C3144E7A6D0E2" box="[836,934,367,392]" pageId="16" pageNumber="561" type="holotype">holotype</typeStatus>
male of this species deposited in Hokkaido University (
<figureCitation id="45872A527F68FFD0297431BFE576D0C6" box="[284,374,404,429]" captionStart="FIGURE 9" captionStartId="17.[151,250,1711,1733]" captionTargetBox="[234,1386,197,1613]" captionTargetId="figure@17.[199,1389,195,1729]" captionTargetPageId="17" captionText="FIGURE 9. Macrostemum formosicolum (Matsumura) and M. okinawanum (Matsumura). Macrostemum formosicolum (9A 9K): Holotype male from Taiwan (9A9F): 9A, habitus, dorsal; 9B, left forewing, dorsal, venation and dark markings; 9C, genitalia, left lateral; 9D, same, dorsal; 9E, right inferior appendage, ventral; 9F, phallic apparatus, left lateral; specimens from Ishigaki-jima (9G9K): 9G1, 9G2, right forewings, venation and dark markings (9G1, female; 9G2, male); 9H, male genitalia, left lateral; 9I, same, dorsal; 9J, phallic apparatus, left lateral; 9K, same, ventral. Macrostemum okinawanum (9L9Y): Holotype male from Okinawa-jima (9L9O): 9L, habitus, dorsal; 9M, right forewing, dorsal; 9N, genitalia, dorsal; 9O, right inferior appendage, ventral; specimens from Okinawa-jima (9P9T): 9P1, 9P2, male right forewings, venation and dark markings; 9Q, male genitalia, left lateral; 9R1, 9R2, male genitalia, dorsal; 9S, phallic apparatus, left lateral; 9T, same, ventral; specimens from Amami-o-shima (9U9Y): 9U1,9U2, male right wings, dorsal; 9V, male genitalia, lateral; 9W, same, dorsal; 9X, phallic apparatus, lateral; 9Y, same, ventral." httpUri="https://zenodo.org/record/2606684/files/figure.png" pageId="16" pageNumber="561">Fig. 9A</figureCitation>
). The
<typeStatus id="020788757F68FFD029D431BFE61ED0C7" box="[444,542,404,429]" pageId="16" pageNumber="561" type="holotype">holotype</typeStatus>
has yellow forewings, and each wing has a large dark marking apically and five dark spots on the central area (
<figureCitation id="45872A527F68FFD02999319CE64AD0BA" box="[497,586,439,464]" captionStart="FIGURE 9" captionStartId="17.[151,250,1711,1733]" captionTargetBox="[234,1386,197,1613]" captionTargetId="figure@17.[199,1389,195,1729]" captionTargetPageId="17" captionText="FIGURE 9. Macrostemum formosicolum (Matsumura) and M. okinawanum (Matsumura). Macrostemum formosicolum (9A 9K): Holotype male from Taiwan (9A9F): 9A, habitus, dorsal; 9B, left forewing, dorsal, venation and dark markings; 9C, genitalia, left lateral; 9D, same, dorsal; 9E, right inferior appendage, ventral; 9F, phallic apparatus, left lateral; specimens from Ishigaki-jima (9G9K): 9G1, 9G2, right forewings, venation and dark markings (9G1, female; 9G2, male); 9H, male genitalia, left lateral; 9I, same, dorsal; 9J, phallic apparatus, left lateral; 9K, same, ventral. Macrostemum okinawanum (9L9Y): Holotype male from Okinawa-jima (9L9O): 9L, habitus, dorsal; 9M, right forewing, dorsal; 9N, genitalia, dorsal; 9O, right inferior appendage, ventral; specimens from Okinawa-jima (9P9T): 9P1, 9P2, male right forewings, venation and dark markings; 9Q, male genitalia, left lateral; 9R1, 9R2, male genitalia, dorsal; 9S, phallic apparatus, left lateral; 9T, same, ventral; specimens from Amami-o-shima (9U9Y): 9U1,9U2, male right wings, dorsal; 9V, male genitalia, lateral; 9W, same, dorsal; 9X, phallic apparatus, lateral; 9Y, same, ventral." httpUri="https://zenodo.org/record/2606684/files/figure.png" pageId="16" pageNumber="561">Fig. 9B</figureCitation>
). Specimens examined in this study also have the same coloration although the posterior spot near the anal veins is occasionally indistinct (
<figureCitation id="45872A527F68FFD02B1131F7E7D6D09E" box="[889,982,476,501]" captionStart="FIGURE 9" captionStartId="17.[151,250,1711,1733]" captionTargetBox="[234,1386,197,1613]" captionTargetId="figure@17.[199,1389,195,1729]" captionTargetPageId="17" captionText="FIGURE 9. Macrostemum formosicolum (Matsumura) and M. okinawanum (Matsumura). Macrostemum formosicolum (9A 9K): Holotype male from Taiwan (9A9F): 9A, habitus, dorsal; 9B, left forewing, dorsal, venation and dark markings; 9C, genitalia, left lateral; 9D, same, dorsal; 9E, right inferior appendage, ventral; 9F, phallic apparatus, left lateral; specimens from Ishigaki-jima (9G9K): 9G1, 9G2, right forewings, venation and dark markings (9G1, female; 9G2, male); 9H, male genitalia, left lateral; 9I, same, dorsal; 9J, phallic apparatus, left lateral; 9K, same, ventral. Macrostemum okinawanum (9L9Y): Holotype male from Okinawa-jima (9L9O): 9L, habitus, dorsal; 9M, right forewing, dorsal; 9N, genitalia, dorsal; 9O, right inferior appendage, ventral; specimens from Okinawa-jima (9P9T): 9P1, 9P2, male right forewings, venation and dark markings; 9Q, male genitalia, left lateral; 9R1, 9R2, male genitalia, dorsal; 9S, phallic apparatus, left lateral; 9T, same, ventral; specimens from Amami-o-shima (9U9Y): 9U1,9U2, male right wings, dorsal; 9V, male genitalia, lateral; 9W, same, dorsal; 9X, phallic apparatus, lateral; 9Y, same, ventral." httpUri="https://zenodo.org/record/2606684/files/figure.png" pageId="16" pageNumber="561">Fig. 9G</figureCitation>
<figureCitation id="45872A527F68FFD02BBD31F7E7E4D09E" box="[981,996,476,500]" captionStart="FIGURE 2" captionStartId="3.[151,250,1737,1759]" captionTargetBox="[193,1386,196,1705]" captionTargetId="figure@3.[188,1403,188,1718]" captionTargetPageId="3" captionText="FIGURE 2. Diplectrona burha Schmid and D. albofaciata Ulmer. Diplectrona burha (2A2I): male (2A2F): 2A, abdominal segments VX, left lateral; 2B, genitalia, left lateral; 2C, same, dorsal; 2D, left inferior appendage, ventral; 2E, phallic apparatus, left lateral; 2F, apex of phallic apparatus, ventral; female (2G2I): 2G, abdominal segment VIII, left lateral; 2H, abdominal segments IXX, left lateral; 2I, abdominal segments VIIIX, ventral. Diplectrona albofaciata (2J2R): male (2J 2O): 2J, abdominal segments IVX, left lateral; 2K, genitalia, left lateral; 2L, same, dorsal; 2M, left inferior appendage, ventral; 2N, phallic apparatus, left lateral; 2O, apex of phallic apparatus, ventral; female (2P2R): 2P, abdominal segment VIII, left lateral; 2Q, abdominal segments IXX, left lateral; 2R, abdominal segments VIIIX, ventral. Abbreviations: VX = abdominal segments VX; l.l. = lateral lobe (paired); m.c.l. = mesocaudal lobe (paired); p.a. = preanal appendage (paired). Arrows, see text." httpUri="https://zenodo.org/record/2606670/files/figure.png" pageId="16" pageNumber="561">2</figureCitation>
). The genitalic structure of two males collected from Ishigaki-jima agrees well with that of the
<typeStatus id="020788757F68FFD02B5831D4E792D372" box="[816,914,511,536]" pageId="16" pageNumber="561" type="holotype">holotype</typeStatus>
(
<figureCitation id="45872A527F68FFD02BCC31D4E7EBD372" box="[932,1003,511,536]" captionStart="FIGURE 8" captionStartId="16.[151,250,1908,1930]" captionTargetBox="[188,1418,885,1825]" captionTargetId="figure@16.[151,1435,860,1839]" captionTargetPageId="16" captionText="FIGURE 8. Hydropsyche verrucosa (Ulmer). Male (8A8F): 8A, genitalia, left lateral; 8B, same, dorsal; 8C, left inferior appendage, ventral; 8D, phallic apparatus, left lateral; 8E, apical part of phallic apparatus, dorsal; 8F, same, ventral. Female (8G, 8H): 8G, genitalia, left lateral; 8H, same, ventral. Abbreviations: VIIIX = abdominal segments VIIIX; rec. = receptacle of inferior appendage (paired)." httpUri="https://zenodo.org/record/2606682/files/figure.png" pageId="16" pageNumber="561">Figs 8</figureCitation>
CF, HK) although the phallotremal sclerites of the
<typeStatus id="020788757F68FFD02929320FE5A3D357" box="[321,419,548,573]" pageId="16" pageNumber="561" type="holotype">holotype</typeStatus>
are protruded from the apex of phallotheca, probably because of drying.
</paragraph>
<paragraph id="DD0336D77F68FFD028AF3263E562D3A7" blockId="16.[151,1437,151,752]" pageId="16" pageNumber="561">
In addition, one
<taxonomicName id="1ABC4D547F68FFD029163262E61DD30A" authorityName="Kolenati" authorityYear="1859" box="[382,541,585,608]" class="Insecta" family="Hydropsychidae" genus="Macrostemum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="561" phylum="Arthropoda" rank="genus">
<emphasis id="EFC8EAC57F68FFD029163262E61DD30A" box="[382,541,585,608]" italics="true" pageId="16" pageNumber="561">Macrostemum</emphasis>
</taxonomicName>
female was collected from Iriomote-jima (Aira-gawa,
<quantity id="1A449B327F68FFD02CED326CE0B0D335" box="[1157,1200,583,608]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="561" unit="m" value="5.0">5 m</quantity>
a.s.l.,
<date id="A90210177F68FFD02C91326CE175D30A" box="[1273,1397,583,608]" pageId="16" pageNumber="561" value="1999-03-25">25.iii.1999</date>
, by TI &amp; AO). Forewings of this female bear similar markings to those of
<taxonomicName id="1ABC4D547F68FFD02BC13246E068D3EE" authorityName=": Malicky" authorityYear="1998" box="[937,1128,620,644]" class="Insecta" family="Hydropsychidae" genus="Macrostemum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="561" phylum="Arthropoda" rank="species" species="formosicolum">
<emphasis id="EFC8EAC57F68FFD02BC13246E068D3EE" box="[937,1128,620,644]" italics="true" pageId="16" pageNumber="561">M. formosicolum</emphasis>
</taxonomicName>
, but lack a dark spot on the anal area. Although this female probably belongs to this species, we hold the identification until more specimens become available.
</paragraph>
<paragraph id="DD0336D77F68FFD028AF32FCE695D39A" blockId="16.[151,1437,151,752]" box="[199,661,727,752]" pageId="16" pageNumber="561">
<emphasis id="EFC8EAC57F68FFD028AF32FCE57CD39A" bold="true" box="[199,380,727,752]" pageId="16" pageNumber="561">Japanese name</emphasis>
. Taiwan-shima-tobikera.
</paragraph>
</subSubSection>
</treatment>
</document>