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<document id="54F426B1C79CDA97F2A6ABC5DEAFF116" ID-CLB-Dataset="28064" ID-DOI="10.11646/zootaxa.4504.4.6" ID-GBIF-Dataset="e597b10d-8e4e-4ffa-8d36-87454b4b35b9" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606666" ID-ZooBank="FA17234E-046E-4165-BEB9-305195E7FD0C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553585146620" checkinUser="plazi" docAuthor="Ito, Tomiko &amp; Nozaki, Takao" docDate="2018" docId="551587C17F73FFCD28FF32FCE6C9D077" docLanguage="en" docName="zootaxa.4504.4.6.pdf" docOrigin="Zootaxa 4504 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cheumatopsyche aira Ito &amp; Nozaki 2018, sp. nov." docType="treatment" docVersion="8" lastPageNumber="558" masterDocId="A92CFFB97F78FFC02868302BE400D16A" masterDocTitle="The family Hydropsychidae Curtis (Trichoptera) in the Ryukyu Archipelago, southwestern Japan" masterLastPageNumber="565" masterPageNumber="545" pageNumber="556" updateTime="1698712834044" updateUser="ExternalLinkService">
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<mods:titleInfo id="84C1DAEBC8B7FCF1BA46B924A92DB8CB">
<mods:title id="EAA9A8DA8EBC9F693BC09B63678F7AC8">The family Hydropsychidae Curtis (Trichoptera) in the Ryukyu Archipelago, southwestern Japan</mods:title>
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<mods:namePart id="8199D025CCF1A1212AC9924DF0DF6BE5">Ito, Tomiko</mods:namePart>
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<mods:namePart id="6421C1FDC8A06ABEE3D0EB47CFACA6D4">Nozaki, Takao</mods:namePart>
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<mods:title id="5EAAA1C43A6CD8454009E3E9C0E830B9">Zootaxa</mods:title>
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<mods:date id="B13591BBC9F480325AF15E49348D50FE">2018</mods:date>
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<mods:number id="B78AEFDE77A58B70AB7141630A46D745">2018-10-26</mods:number>
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<mods:number id="E2DDE9BA003F49A35AADF8F02442B4F0">4504</mods:number>
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<mods:classification id="802AFCFF786848395264A16C410934AF">journal article</mods:classification>
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<mods:identifier id="5584B4BCCD0E17CF0BACB511248D7F85" type="DOI">10.11646/zootaxa.4504.4.6</mods:identifier>
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<mods:identifier id="451B0252F684A882E06CDDBDA4BC1282" type="ISSN">1175-5326</mods:identifier>
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<treatment id="551587C17F73FFCD28FF32FCE6C9D077" ID-DOI="http://doi.org/10.5281/zenodo.5975800" ID-GBIF-Taxon="154495221" ID-Zenodo-Dep="5975800" LSID="urn:lsid:plazi:treatment:551587C17F73FFCD28FF32FCE6C9D077" httpUri="http://treatment.plazi.org/id/551587C17F73FFCD28FF32FCE6C9D077" lastPageId="13" lastPageNumber="558" pageId="11" pageNumber="556">
<subSubSection id="95A6655C7F73FFCB28FF32FCE6C3D398" box="[151,707,727,754]" pageId="11" pageNumber="556" type="nomenclature">
<paragraph id="DD0336D77F73FFCB28FF32FCE6C3D398" blockId="11.[151,707,727,787]" box="[151,707,727,754]" pageId="11" pageNumber="556">
<heading id="864B81BB7F73FFCB28FF32FCE6C3D398" bold="true" box="[151,707,727,754]" fontSize="11" level="1" pageId="11" pageNumber="556" reason="1">
<emphasis id="EFC8EAC57F73FFCB28FF32FCE6C3D398" bold="true" box="[151,707,727,754]" pageId="11" pageNumber="556">
<taxonomicName id="1ABC4D547F73FFCB28FF32FCE65FD398" authority="Ito and Nozaki" authorityName="Ito &amp; Nozaki" authorityYear="2018" box="[151,607,727,754]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="556" phylum="Arthropoda" rank="species" species="aira" status="sp. nov.">
<emphasis id="EFC8EAC57F73FFCB28FF32FCE59ED39B" bold="true" box="[151,414,727,753]" italics="true" pageId="11" pageNumber="556">Cheumatopsyche aira</emphasis>
Ito and Nozaki
</taxonomicName>
<taxonomicNameLabel id="F4FB57BE7F73FFCB2A0E32F3E6C3D398" box="[614,707,728,754]" pageId="11" pageNumber="556" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95A6655C7F73FFCB28FF32D1E528D279" box="[151,296,762,787]" pageId="11" pageNumber="556" type="description">
<paragraph id="DD0336D77F73FFCB28FF32D1E528D279" blockId="11.[151,707,727,787]" box="[151,296,762,787]" pageId="11" pageNumber="556">
(
<figureCitation id="45872A527F73FFCB28F732D1E4E4D279" box="[159,228,762,787]" captionStart="FIGURE 6" captionStartId="12.[151,250,1878,1900]" captionTargetBox="[181,1425,487,1837]" captionTargetId="figure@12.[161,1427,482,1850]" captionTargetPageId="12" captionText="FIGURE 6. Cheumatopsyche aira Ito &amp; Nozaki sp. nov. Male (6A–6F): 6A, right wings, dorsal, light color marks of forewing, venations of forewing and hind wing; 6B, genitalia, left lateral; 6C, same, dorsal; 6D, left inferior appendage, ventral; 6E, phallic apparatus, left lateral; 6F, apical part of phallus, ventral. Female (6G–6I): 6G, right wings, dorsal; 6H, genitalia, left lateral; 6I, same, ventral. Abbreviations: I–V = apical forks I–V; VIII–X = abdominal segments VIII–X; e.p. = endothecal process (paired); l.l. = lateral lobe (paired); m.c.l. = mesocaudal lobe; p.s. = phallotremal sclerite." httpUri="https://zenodo.org/record/2606678/files/figure.png" pageId="11" pageNumber="556">Figs 6</figureCitation>
A–
<date id="A90210177F73FFCB296232D1E520D279" box="[266,288,762,787]" pageId="11" pageNumber="556">6I</date>
)
</paragraph>
</subSubSection>
<subSubSection id="95A6655C7F73FFCB28FF3369E160D461" pageId="11" pageNumber="556" type="diagnosis">
<paragraph id="DD0336D77F73FFCB28FF3369E7F0D558" blockId="11.[151,1437,834,2011]" pageId="11" pageNumber="556">
<emphasis id="EFC8EAC57F73FFCB28FF3369E511D231" bold="true" box="[151,273,834,859]" pageId="11" pageNumber="556">Diagnosis.</emphasis>
The male of this species is similar to that of
<taxonomicName id="1ABC4D547F73FFCB2B543368E11FD231" authority="Martynov 1930" authorityName="Martynov" authorityYear="1930" box="[828,1311,835,859]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="556" phylum="Arthropoda" rank="species" species="clavalis">
<emphasis id="EFC8EAC57F73FFCB2B543368E05ED231" box="[828,1118,835,859]" italics="true" pageId="11" pageNumber="556">Cheumatopsyche clavalis</emphasis>
<bibRefCitation id="B92D4B267F73FFCB2C023368E11FD231" author="Martynov, A. V." box="[1130,1311,835,859]" pageId="11" pageNumber="556" pagination="65 - 112" refId="ref11048" refString="Martynov, A. V. (1930) On the trichopterous fauna of China and eastern Tibet. Proceedings of the Zoological Society of London, 5, 65 - 112, pl. 1." type="journal article" year="1930">Martynov 1930</bibRefCitation>
</taxonomicName>
, originally described from
<collectingCountry id="A5AB76477F73FFCB293A334DE5A7D215" box="[338,423,870,895]" name="Taiwan" pageId="11" pageNumber="556">Taiwan</collectingCountry>
, in having an almost straight posterior margin on each side of segment IX and low mesocaudal lobes of segment X (m.c.l.) (
<bibRefCitation id="B92D4B267F73FFCB2AE733A1E752D2C9" author="Kobayashi, M." box="[655,850,906,931]" pageId="11" pageNumber="556" pagination="37 - 48" refId="ref10696" refString="Kobayashi, M. (1987) Systematic study of the caddisflies from Taiwan, with descriptions of eleven new species (Trichoptera: Insecta). Bulletin of the Kanagawa Prefectural Museum, Natural Science, 17, 37 - 48." type="journal article" year="1987">Kobayashi 1987</bibRefCitation>
; Oláh
<emphasis id="EFC8EAC57F73FFCB2BC233A0E7E1D2C9" box="[938,993,907,931]" italics="true" pageId="11" pageNumber="556">et al</emphasis>
. 2008). However,
<taxonomicName id="1ABC4D547F73FFCB2CA133A0E121D2C8" authorityName="Ito &amp; Nozaki" authorityYear="2018" box="[1225,1313,907,930]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="556" phylum="Arthropoda" rank="species" species="aira">
<emphasis id="EFC8EAC57F73FFCB2CA133A0E121D2C8" box="[1225,1313,907,930]" italics="true" pageId="11" pageNumber="556">C. aira</emphasis>
</taxonomicName>
is clearly discriminated from
<taxonomicName id="1ABC4D547F73FFCB29103384E5F1D2AC" box="[376,497,942,966]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="556" phylum="Arthropoda" rank="species" species="clavalis">
<emphasis id="EFC8EAC57F73FFCB29103384E5F1D2AC" box="[376,497,942,966]" italics="true" pageId="11" pageNumber="556">C. clavalis</emphasis>
</taxonomicName>
as follows: Each lateral lobe of segment X (l.l.) is short, round, widely separated with each other in dorsal aspect in
<taxonomicName id="1ABC4D547F73FFCB2A4333F8E67DD280" authorityName="Ito &amp; Nozaki" authorityYear="2018" box="[555,637,979,1002]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="556" phylum="Arthropoda" rank="species" species="aira">
<emphasis id="EFC8EAC57F73FFCB2A4333F8E67DD280" box="[555,637,979,1002]" italics="true" pageId="11" pageNumber="556">C. aira</emphasis>
</taxonomicName>
, but moderately long with apicomesal acute corner, closely contacted each other in dorsal aspect in
<taxonomicName id="1ABC4D547F73FFCB299A33DCE66CD564" box="[498,620,1014,1038]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="556" phylum="Arthropoda" rank="species" species="clavalis">
<emphasis id="EFC8EAC57F73FFCB299A33DCE66CD564" box="[498,620,1014,1038]" italics="true" pageId="11" pageNumber="556">C. clavalis</emphasis>
</taxonomicName>
; distal segment of each inferior appendage is nearly parallel sided and subquadrate apically in
<taxonomicName id="1ABC4D547F73FFCB29C93430E5F1D558" authorityName="Ito &amp; Nozaki" authorityYear="2018" box="[417,497,1051,1074]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="556" phylum="Arthropoda" rank="species" species="aira">
<emphasis id="EFC8EAC57F73FFCB29C93430E5F1D558" box="[417,497,1051,1074]" italics="true" pageId="11" pageNumber="556">C. aira</emphasis>
</taxonomicName>
, but tapered and acute apically in
<taxonomicName id="1ABC4D547F73FFCB2B193430E7E9D559" box="[881,1001,1051,1075]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="556" phylum="Arthropoda" rank="species" species="clavalis">
<emphasis id="EFC8EAC57F73FFCB2B193430E7E9D559" box="[881,1001,1051,1075]" italics="true" pageId="11" pageNumber="556">C. clavalis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="DD0336D77F73FFCB28AF3415E160D461" blockId="11.[151,1437,834,2011]" pageId="11" pageNumber="556">
<emphasis id="EFC8EAC57F73FFCB28AF3415E50CD53D" bold="true" box="[199,268,1086,1111]" pageId="11" pageNumber="556">Adult</emphasis>
(
<figureCitation id="45872A527F73FFCB29773415E56BD53D" box="[287,363,1086,1111]" captionStart="FIGURE 6" captionStartId="12.[151,250,1878,1900]" captionTargetBox="[181,1425,487,1837]" captionTargetId="figure@12.[161,1427,482,1850]" captionTargetPageId="12" captionText="FIGURE 6. Cheumatopsyche aira Ito &amp; Nozaki sp. nov. Male (6A–6F): 6A, right wings, dorsal, light color marks of forewing, venations of forewing and hind wing; 6B, genitalia, left lateral; 6C, same, dorsal; 6D, left inferior appendage, ventral; 6E, phallic apparatus, left lateral; 6F, apical part of phallus, ventral. Female (6G–6I): 6G, right wings, dorsal; 6H, genitalia, left lateral; 6I, same, ventral. Abbreviations: I–V = apical forks I–V; VIII–X = abdominal segments VIII–X; e.p. = endothecal process (paired); l.l. = lateral lobe (paired); m.c.l. = mesocaudal lobe; p.s. = phallotremal sclerite." httpUri="https://zenodo.org/record/2606678/files/figure.png" pageId="11" pageNumber="556">Figs 6</figureCitation>
A–
<date id="A90210177F73FFCB29F93415E5A9D53D" box="[401,425,1086,1111]" pageId="11" pageNumber="556">6I</date>
). Wings brown, many light brown dots and patterns present mainly along margins in forewings, but often indistinct in alcohol specimens; light brown dots absent in hind wings. In males, forewings
<quantity id="1A449B327F73FFCB28FF34ADE51FD5F4" box="[151,287,1158,1183]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" metricValueMax="6.3" metricValueMin="5.5" pageId="11" pageNumber="556" unit="mm" value="5.9" valueMax="6.3" valueMin="5.5">5.5–6.3 mm</quantity>
long and hind wings
<quantity id="1A449B327F73FFCB2A7034ACE6A1D5F4" box="[536,673,1159,1183]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="4.8" metricValueMin="4.2" pageId="11" pageNumber="556" unit="mm" value="4.5" valueMax="4.8" valueMin="4.2">4.2–4.8 mm</quantity>
long (n = 5); apical forks I–V present, discoidal cell small, medial cell open in forewings; apical forks II, III and V present, discoidal cell short in hind wings. In females, forewings
<quantity id="1A449B327F73FFCB28FF34E5E51FD58C" box="[151,287,1230,1255]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.2" metricValueMin="5.8" pageId="11" pageNumber="556" unit="mm" value="6.0" valueMax="6.2" valueMin="5.8">5.8–6.2 mm</quantity>
long and hind wings
<quantity id="1A449B327F73FFCB2A7034E4E6A1D58C" box="[536,673,1231,1255]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="4.7" metricValueMin="4.3" pageId="11" pageNumber="556" unit="mm" value="4.5" valueMax="4.7" valueMin="4.3">4.3–4.7 mm</quantity>
long (n = 5); apical forks I–V present, discoidal cell small, medial cell longer than discoidal cell in forewing; apical forks II, III and V present, discoidal cell short in hind wing.
</paragraph>
</subSubSection>
<subSubSection id="95A6655C7F73FFCB28AF353DE17DD7B5" pageId="11" pageNumber="556" type="description">
<paragraph id="DD0336D77F73FFCB28AF353DE0A0D741" blockId="11.[151,1437,834,2011]" pageId="11" pageNumber="556">
<emphasis id="EFC8EAC57F73FFCB28AF353DE574D445" bold="true" box="[199,372,1302,1327]" pageId="11" pageNumber="556">Male genitalia</emphasis>
(
<figureCitation id="45872A527F73FFCB29ED353DE5CED445" box="[389,462,1302,1327]" captionStart="FIGURE 6" captionStartId="12.[151,250,1878,1900]" captionTargetBox="[181,1425,487,1837]" captionTargetId="figure@12.[161,1427,482,1850]" captionTargetPageId="12" captionText="FIGURE 6. Cheumatopsyche aira Ito &amp; Nozaki sp. nov. Male (6A–6F): 6A, right wings, dorsal, light color marks of forewing, venations of forewing and hind wing; 6B, genitalia, left lateral; 6C, same, dorsal; 6D, left inferior appendage, ventral; 6E, phallic apparatus, left lateral; 6F, apical part of phallus, ventral. Female (6G–6I): 6G, right wings, dorsal; 6H, genitalia, left lateral; 6I, same, ventral. Abbreviations: I–V = apical forks I–V; VIII–X = abdominal segments VIII–X; e.p. = endothecal process (paired); l.l. = lateral lobe (paired); m.c.l. = mesocaudal lobe; p.s. = phallotremal sclerite." httpUri="https://zenodo.org/record/2606678/files/figure.png" pageId="11" pageNumber="556">Figs 6</figureCitation>
B–6F). Segment IX short with gently protruding anterior margins and nearly straight caudal margins in lateral aspect. Segment X with very short mesocaudal lobe (m.c.l.) in lateral and dorsal aspects; lateral lobes (l.l.) short, with length almost as long as width and oval and widely separated each other in dorsal aspect, bar-like in lateral aspect. Inferior appendages slender, directed postero-dorsad in lateral aspect and gently curved posteromesad in ventral aspect; basal segment long, 3.3 times of distal segment; distal segment more slender than basal segment, almost parallel sided in ventral aspect, slightly curved in lateral aspect, each with apex subquadrate. Phallic apparatus with short endothecal process (e.p.), phallotremal sclerite (p.s.) C- and reverse Cshaped in ventral aspect; numerous very small spines on apical part of phallotheca ventrally.
</paragraph>
<paragraph id="DD0336D77F73FFCB28AF361DE17DD7B5" blockId="11.[151,1437,834,2011]" pageId="11" pageNumber="556">
<emphasis id="EFC8EAC57F73FFCB28AF361DE58CD725" bold="true" box="[199,396,1590,1615]" pageId="11" pageNumber="556">Female genitalia</emphasis>
(
<figureCitation id="45872A527F73FFCB29F3361DE5E3D725" box="[411,483,1590,1615]" captionStart="FIGURE 6" captionStartId="12.[151,250,1878,1900]" captionTargetBox="[181,1425,487,1837]" captionTargetId="figure@12.[161,1427,482,1850]" captionTargetPageId="12" captionText="FIGURE 6. Cheumatopsyche aira Ito &amp; Nozaki sp. nov. Male (6A–6F): 6A, right wings, dorsal, light color marks of forewing, venations of forewing and hind wing; 6B, genitalia, left lateral; 6C, same, dorsal; 6D, left inferior appendage, ventral; 6E, phallic apparatus, left lateral; 6F, apical part of phallus, ventral. Female (6G–6I): 6G, right wings, dorsal; 6H, genitalia, left lateral; 6I, same, ventral. Abbreviations: I–V = apical forks I–V; VIII–X = abdominal segments VIII–X; e.p. = endothecal process (paired); l.l. = lateral lobe (paired); m.c.l. = mesocaudal lobe; p.s. = phallotremal sclerite." httpUri="https://zenodo.org/record/2606678/files/figure.png" pageId="11" pageNumber="556">Figs 6</figureCitation>
H–
<date id="A90210177F73FFCB2A61361DE621D725" box="[521,545,1590,1615]" pageId="11" pageNumber="556">6I</date>
). Tergite VIII with almost straight anterior margins and convex caudal margins in lateral aspect. Sternite VIII subquadrate, larger than tergite VIII, with oblique straight dorsal margin and large subquadrate lateral lobes in lateral aspect, triangular lateral lobes in ventral aspect, each lobe wide at anterior margin, mesal margin gently curved laterad. Segment IX obliquely S-shaped in lateral aspects. Receptacle of inferior appendage (rec.) distinct, nearly question mark shaped in lateral aspect. Segment X oblique rectangular.
</paragraph>
</subSubSection>
<subSubSection id="95A6655C7F73FFCC28AF36C1E19DD0ED" lastPageId="12" lastPageNumber="557" pageId="11" pageNumber="556" type="materials_examined">
<paragraph id="DD0336D77F73FFCB28AF36C1E5ADD64D" blockId="11.[151,1437,834,2011]" pageId="11" pageNumber="556">
<materialsCitation id="6DD43C8A7F73FFCB28AF36C1E5ADD64D" ID-GBIF-Occurrence="2028138161" collectingDate="2012-10-28" collectingDateMax="2012-10-30" collectingDateMin="2012-10-28" collectionCode="TI" elevation="5" latitude="24.3332" longLatPrecision="7" longitude="123.9134" pageId="11" pageNumber="556" specimenCode="CBM-ZI 166013" specimenCount="1" typeStatus="holotype">
<emphasis id="EFC8EAC57F73FFCB28AF36C1E53DD669" bold="true" box="[199,317,1770,1795]" pageId="11" pageNumber="556">
<typeStatus id="020788757F73FFCB28AF36C1E538D669" box="[199,312,1770,1795]" pageId="11" pageNumber="556" type="holotype">Holotype</typeStatus>
:
</emphasis>
Male,
<emphasis id="EFC8EAC57F73FFCB29E536C1E635D669" bold="true" box="[397,565,1770,1795]" pageId="11" pageNumber="556">Iriomote-jima</emphasis>
, Taketomi-cho, Aira-gawa (
<geoCoordinate id="B88850107F73FFCB2B1B36C0E7E7D669" box="[883,999,1771,1795]" direction="north" orientation="latitude" pageId="11" pageNumber="556" precision="5" value="24.3332">24.3332N</geoCoordinate>
,
<geoCoordinate id="B88850107F73FFCB2B9B36C0E06FD669" box="[1011,1135,1771,1795]" direction="east" orientation="longitude" pageId="11" pageNumber="556" precision="5" value="123.9134">123.9134E</geoCoordinate>
,
<elevation id="5691D1E47F73FFCB2C1436C1E0E4D669" box="[1148,1252,1770,1795]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="556" unit="m" value="5.0">
<quantity id="1A449B327F73FFCB2C1436C1E0A7D668" box="[1148,1191,1770,1795]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="556" unit="m" value="5.0">5 m</quantity>
a.s.l.
</elevation>
),
<date id="A90210177F73FFCB2C9336C0E198D669" box="[1275,1432,1771,1795]" pageId="11" pageNumber="556" value="2012-10-28" valueMax="2012-10-30" valueMin="2012-10-28">
<collectingDate id="B946E9FF7F73FFCB2C9336C0E198D669" box="[1275,1432,1771,1795]" pageId="11" pageNumber="556" value="2012-10-28" valueMax="2012-10-30" valueMin="2012-10-28">28–30.x.2012</collectingDate>
</date>
,
<collectionCode id="BBADAE127F73FFCB28FF3724E4B4D64C" box="[151,180,1807,1830]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="11" pageNumber="556">TI</collectionCode>
, P (
<specimenCode id="8D1A9EAC7F73FFCB28B53724E5A1D64D" box="[221,417,1806,1831]" collectionCode="CBM-ZI" pageId="11" pageNumber="556">CBM-ZI 166013</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="DD0336D77F73FFCB28AF3719E062D621" blockId="11.[151,1437,834,2011]" box="[199,1122,1842,1867]" pageId="11" pageNumber="556">
<materialsCitation id="6DD43C8A7F73FFCB28AF3719E05ED621" ID-GBIF-Occurrence="2028138157" box="[199,1118,1842,1867]" collectingDate="2012-10-28" collectingDateMax="2012-10-30" collectingDateMin="2012-10-28" collectionCode="TI" elevation="5" latitude="24.3332" longLatPrecision="7" longitude="123.9134" pageId="11" pageNumber="556" specimenCode="CBM-ZI 166014-166023" specimenCount="10" specimenCount-female="5" specimenCount-male="5" typeStatus="paratype">
<emphasis id="EFC8EAC57F73FFCB28AF3719E53FD621" bold="true" box="[199,319,1842,1867]" pageId="11" pageNumber="556">
<typeStatus id="020788757F73FFCB28AF3719E53FD621" box="[199,319,1842,1867]" pageId="11" pageNumber="556" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<specimenCount id="CBBAFD5E7F73FFCB29243719E5A6D621" box="[332,422,1842,1867]" pageId="11" pageNumber="556" type="male">5 males</specimenCount>
,
<specimenCount id="CBBAFD5E7F73FFCB29D93719E620D621" box="[433,544,1842,1867]" pageId="11" pageNumber="556" type="female">5 females</specimenCount>
, same data as holotype (
<specimenCode id="8D1A9EAC7F73FFCB2B583718E056D621" box="[816,1110,1842,1867]" collectionCode="CBM-ZI" pageId="11" pageNumber="556">CBM-ZI 166014–166023</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="DD0336D77F73FFCC28AF377DE19DD0ED" blockId="11.[151,1437,834,2011]" lastBlockId="12.[151,1437,151,429]" lastPageId="12" lastPageNumber="557" pageId="11" pageNumber="556">
<emphasis id="EFC8EAC57F73FFCB28AF377DE614D605" bold="true" box="[199,532,1878,1903]" pageId="11" pageNumber="556">Other specimens examined</emphasis>
.
<emphasis id="EFC8EAC57F73FFCB2A4F377DE6CFD605" bold="true" box="[551,719,1878,1903]" pageId="11" pageNumber="556">Iriomote-jima</emphasis>
:
<materialsCitation id="6DD43C8A7F73FFCB2A8C377CE10DD605" ID-GBIF-Occurrence="2028138154" box="[740,1293,1878,1903]" collectingDate="2012-10-28" collectingDateMax="2012-10-30" collectingDateMin="2012-10-28" collectionCode="TI" elevation="5" latitude="24.3332" longLatPrecision="7" longitude="123.9134" pageId="11" pageNumber="556" specimenCount="288" specimenCount-female="271" specimenCount-male="17">
<specimenCount id="CBBAFD5E7F73FFCB2A8C377CE750D605" box="[740,848,1878,1903]" pageId="11" pageNumber="556" type="male">17 males</specimenCount>
,
<specimenCount id="CBBAFD5E7F73FFCB2B09377CE7F2D605" box="[865,1010,1878,1903]" pageId="11" pageNumber="556" type="female">271 females</specimenCount>
, same data as holotype
</materialsCitation>
;
<materialsCitation id="6DD43C8A7F73FFCB2D77377CE638D6F8" ID-GBIF-Occurrence="2028138174" collectingDate="1999-03-25" collectionCode="TI" pageId="11" pageNumber="556" specimenCount="16" specimenCount-female="7" specimenCount-male="9">
<specimenCount id="CBBAFD5E7F73FFCB2D77377CE17DD605" box="[1311,1405,1878,1903]" pageId="11" pageNumber="556" type="male">9 males</specimenCount>
,
<specimenCount id="CBBAFD5E7F73FFCB2DE6377CE4F1D6F9" pageId="11" pageNumber="556" type="female">7 females</specimenCount>
, type locality,
<date id="A90210177F73FFCB29FD3751E611D6F9" box="[405,529,1914,1939]" pageId="11" pageNumber="556" value="1999-03-25">
<collectingDate id="B946E9FF7F73FFCB29FD3751E611D6F9" box="[405,529,1914,1939]" pageId="11" pageNumber="556" value="1999-03-25">25.iii.1999</collectingDate>
</date>
,
<collectionCode id="BBADAE127F73FFCB2A753750E638D6F8" box="[541,568,1915,1938]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="11" pageNumber="556">TI</collectionCode>
</materialsCitation>
&amp; AO, L;
<materialsCitation id="6DD43C8A7F73FFCB2ADD3750E110D6F8" ID-GBIF-Occurrence="2028138169" box="[693,1296,1914,1939]" collectingDate="2013-11-28" collectingDateMax="2013-12-02" collectingDateMin="2013-11-28" collectionCode="TI" pageId="11" pageNumber="556" specimenCount="4" specimenCount-female="2" specimenCount-male="2">
<specimenCount id="CBBAFD5E7F73FFCB2ADD3750E70ED6F9" box="[693,782,1914,1939]" pageId="11" pageNumber="556" type="male">2 males</specimenCount>
,
<specimenCount id="CBBAFD5E7F73FFCB2B723750E78AD6F9" box="[794,906,1914,1939]" pageId="11" pageNumber="556" type="female">2 females</specimenCount>
, type locality,
<date id="A90210177F73FFCB2C463751E0EAD6F9" box="[1070,1258,1914,1939]" pageId="11" pageNumber="556" value="2013-11-28" valueMax="2013-12-02" valueMin="2013-11-28">
<collectingDate id="B946E9FF7F73FFCB2C463751E0EAD6F9" box="[1070,1258,1914,1939]" pageId="11" pageNumber="556" value="2013-11-28" valueMax="2013-12-02" valueMin="2013-11-28">28.xi–2.xii.2013</collectingDate>
</date>
,
<collectionCode id="BBADAE127F73FFCB2C9D3750E110D6F8" box="[1269,1296,1915,1938]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="11" pageNumber="556">TI</collectionCode>
</materialsCitation>
&amp; RS, M;
<materialsCitation id="6DD43C8A7F73FFCB2DE63750E6DAD6DC" ID-GBIF-Occurrence="2028138167" collectingDate="2016-03-21" collectingDateMax="2016-03-22" collectingDateMin="2016-03-21" collectionCode="TI" pageId="11" pageNumber="556" specimenCount="21" specimenCount-female="18" specimenCount-male="3">
<specimenCount id="CBBAFD5E7F73FFCB2DE63750E4DBD6DD" pageId="11" pageNumber="556" type="male">3 males</specimenCount>
,
<specimenCount id="CBBAFD5E7F73FFCB288F37B4E565D6DD" box="[231,357,1950,1975]" pageId="11" pageNumber="556" type="female">18 females</specimenCount>
, type locality,
<date id="A90210177F73FFCB2A6237B5E6B0D6DD" box="[522,688,1950,1975]" pageId="11" pageNumber="556" value="2016-03-21" valueMax="2016-03-22" valueMin="2016-03-21">
<collectingDate id="B946E9FF7F73FFCB2A6237B5E6B0D6DD" box="[522,688,1950,1975]" pageId="11" pageNumber="556" value="2016-03-21" valueMax="2016-03-22" valueMin="2016-03-21">21–22.iii.2016</collectingDate>
</date>
,
<collectionCode id="BBADAE127F73FFCB2AD537B4E6DAD6DC" box="[701,730,1951,1974]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="11" pageNumber="556">TI</collectionCode>
</materialsCitation>
, P;
<materialsCitation id="6DD43C8A7F73FFCB2B6D37B4E163D6DC" ID-GBIF-Occurrence="2028138160" box="[773,1379,1950,1975]" collectingDate="2016-10-20" collectingDateMax="2016-10-24" collectingDateMin="2016-10-20" collectionCode="TI" pageId="11" pageNumber="556" specimenCount="25" specimenCount-female="13" specimenCount-male="12">
<specimenCount id="CBBAFD5E7F73FFCB2B6D37B4E76DD6DD" box="[773,877,1950,1975]" pageId="11" pageNumber="556" type="male">12 males</specimenCount>
,
<specimenCount id="CBBAFD5E7F73FFCB2B1237B4E7F7D6DD" box="[890,1015,1950,1975]" pageId="11" pageNumber="556" type="female">13 females</specimenCount>
, type locality,
<date id="A90210177F73FFCB2CF537B5E13AD6DD" box="[1181,1338,1950,1975]" pageId="11" pageNumber="556" value="2016-10-20" valueMax="2016-10-24" valueMin="2016-10-20">
<collectingDate id="B946E9FF7F73FFCB2CF537B5E13AD6DD" box="[1181,1338,1950,1975]" pageId="11" pageNumber="556" value="2016-10-20" valueMax="2016-10-24" valueMin="2016-10-20">20–24.x.2016</collectingDate>
</date>
,
<collectionCode id="BBADAE127F73FFCB2D2E37B4E163D6DC" box="[1350,1379,1951,1974]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="11" pageNumber="556">TI</collectionCode>
</materialsCitation>
, P;
<materialsCitation id="6DD43C8A7F73FFCB2DE637B4E764D6B0" ID-GBIF-Occurrence="2028138155" collectingDate="2013-12-01" collectionCode="TI" elevation="35" pageId="11" pageNumber="556" specimenCount="17" specimenCount-female="15" specimenCount-male="2">
<specimenCount id="CBBAFD5E7F73FFCB2DE637B4E4DBD6B1" pageId="11" pageNumber="556" type="male">2 males</specimenCount>
,
<specimenCount id="CBBAFD5E7F73FFCB288237E9E56CD6B1" box="[234,364,1986,2011]" pageId="11" pageNumber="556" type="female">15 females</specimenCount>
, Aira-gawa,
<quantity id="1A449B327F73FFCB2A6B37E9E66DD6B0" box="[515,621,1986,2011]" metricMagnitude="1" metricUnit="m" metricValue="3.5" metricValueMax="4.5" metricValueMin="2.5" pageId="11" pageNumber="556" unit="m" value="35.0" valueMax="45.0" valueMin="25.0">
<elevation id="5691D1E47F73FFCB2A6B37E9E66DD6B0" box="[515,621,1986,2011]" metricMagnitude="1" metricUnit="m" metricValue="3.5" metricValueMax="4.5" metricValueMin="2.5" pageId="11" pageNumber="556" unit="m" value="35.0" valueMax="45.0" valueMin="25.0">25–45 m</elevation>
</quantity>
, a.s.l.,
<date id="A90210177F73FFCB2AAB37E9E737D6B1" box="[707,823,1986,2011]" pageId="11" pageNumber="556" value="2013-12-01">
<collectingDate id="B946E9FF7F73FFCB2AAB37E9E737D6B1" box="[707,823,1986,2011]" pageId="11" pageNumber="556" value="2013-12-01">1.xii.2013</collectingDate>
</date>
,
<collectionCode id="BBADAE127F73FFCB2B2F37E8E764D6B0" box="[839,868,1987,2010]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="11" pageNumber="556">TI</collectionCode>
</materialsCitation>
, L, P &amp; S;
<materialsCitation id="6DD43C8A7F73FFCC2B9D37E8E532D1C5" ID-GBIF-Occurrence="2028138164" collectingDate="2015-10-21" collectionCode="TI" elevation="35" lastPageId="12" lastPageNumber="557" pageId="11" pageNumber="556" specimenCount="22" specimenCount-female="11" specimenCount-male="11">
<specimenCount id="CBBAFD5E7F73FFCB2B9D37E8E05FD6B1" box="[1013,1119,1986,2011]" pageId="11" pageNumber="556" type="male">11 males</specimenCount>
,
<specimenCount id="CBBAFD5E7F73FFCB2C0737E8E0EFD6B1" box="[1135,1263,1986,2011]" pageId="11" pageNumber="556" type="female">11 females</specimenCount>
, same locality,
<date id="A90210177F74FFCC28FF30BCE509D1DA" box="[151,265,151,176]" pageId="12" pageNumber="557" value="2015-10-21">
<collectingDate id="B946E9FF7F74FFCC28FF30BCE509D1DA" box="[151,265,151,176]" pageId="12" pageNumber="557" value="2015-10-21">21.x.2015</collectingDate>
</date>
,
<collectionCode id="BBADAE127F74FFCC297F30B3E532D1C5" box="[279,306,152,175]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="12" pageNumber="557">TI</collectionCode>
</materialsCitation>
&amp; RS, L;
<materialsCitation id="6DD43C8A7F74FFCC29D830B3E002D1C5" ID-GBIF-Occurrence="2028138151" box="[432,1026,151,176]" collectingDate="2016-03-21" collectionCode="TI" elevation="35" pageId="12" pageNumber="557" specimenCount="43" specimenCount-female="31" specimenCount-male="12">
<specimenCount id="CBBAFD5E7F74FFCC29D830B3E61AD1DA" box="[432,538,151,176]" pageId="12" pageNumber="557" type="male">12 males</specimenCount>
,
<specimenCount id="CBBAFD5E7F74FFCC2A4030B3E6A8D1DA" box="[552,680,151,176]" pageId="12" pageNumber="557" type="female">31 females</specimenCount>
, same locality,
<date id="A90210177F74FFCC2B3330BCE7D7D1DA" box="[859,983,151,176]" pageId="12" pageNumber="557" value="2016-03-21">
<collectingDate id="B946E9FF7F74FFCC2B3330BCE7D7D1DA" box="[859,983,151,176]" pageId="12" pageNumber="557" value="2016-03-21">21.iii.2016</collectingDate>
</date>
,
<collectionCode id="BBADAE127F74FFCC2B8D30B3E002D1C5" box="[997,1026,152,175]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="12" pageNumber="557">TI</collectionCode>
</materialsCitation>
, L;
<specimenCount id="CBBAFD5E7F74FFCC2C5B30B3E083D1DA" box="[1075,1155,151,176]" pageId="12" pageNumber="557" type="male">1 male</specimenCount>
,
<specimenCount id="CBBAFD5E7F74FFCC2CFA30BCE111D1DA" box="[1170,1297,151,176]" pageId="12" pageNumber="557" type="female">16 females</specimenCount>
, Aira-gawa,
<date id="A90210177F74FFCC28FF3097E513D1BF" box="[151,275,188,213]" pageId="12" pageNumber="557" value="2002-03-15">15.iii.2002</date>
, I. Oshima
<emphasis id="EFC8EAC57F74FFCC29F53096E5D2D1BE" box="[413,466,188,212]" italics="true" pageId="12" pageNumber="557">et al</emphasis>
., L;
<materialsCitation id="6DD43C8A7F74FFCC2A653097E0ACD1BE" ID-GBIF-Occurrence="2028138177" box="[525,1196,188,213]" collectingDate="2009-05-27" collectionCode="TI" collectorName="K. Tojo" elevation="35" pageId="12" pageNumber="557" specimenCount="18" specimenCount-female="8" specimenCount-male="10">
<specimenCount id="CBBAFD5E7F74FFCC2A653097E678D1BF" box="[525,632,188,213]" pageId="12" pageNumber="557" type="male">10 males</specimenCount>
,
<specimenCount id="CBBAFD5E7F74FFCC2AEE3097E6F8D1BF" box="[646,760,188,213]" pageId="12" pageNumber="557" type="female">8 females</specimenCount>
, same locality,
<date id="A90210177F74FFCC2BC63097E01FD1BE" box="[942,1055,188,212]" pageId="12" pageNumber="557" value="2009-05-27">
<collectingDate id="B946E9FF7F74FFCC2BC63097E01FD1BE" box="[942,1055,188,212]" pageId="12" pageNumber="557" value="2009-05-27">27.v.2009</collectingDate>
</date>
,
<collectorName id="704953017F74FFCC2C463096E089D1BF" box="[1070,1161,188,213]" pageId="12" pageNumber="557">K. Tojo</collectorName>
, L
</materialsCitation>
;
<materialsCitation id="6DD43C8A7F74FFCC2CD33097E73BD19D" ID-GBIF-Occurrence="2028138183" collectingDate="1999-03-23" collectionCode="TI" pageId="12" pageNumber="557" specimenCount="5" specimenCount-female="3" specimenCount-male="2">
<specimenCount id="CBBAFD5E7F74FFCC2CD33097E117D1BF" box="[1211,1303,188,213]" pageId="12" pageNumber="557" type="male">2 males</specimenCount>
,
<specimenCount id="CBBAFD5E7F74FFCC2D4E3097E198D1BF" box="[1318,1432,188,213]" pageId="12" pageNumber="557" type="female">3 females</specimenCount>
, Nishi-funatsuki-gawa, Nishi-funatsuki-bashi,
<date id="A90210177F74FFCC2AF130F4E715D192" box="[665,789,223,248]" pageId="12" pageNumber="557" value="1999-03-23">
<collectingDate id="B946E9FF7F74FFCC2AF130F4E715D192" box="[665,789,223,248]" pageId="12" pageNumber="557" value="1999-03-23">23.iii.1999</collectingDate>
</date>
,
<collectionCode id="BBADAE127F74FFCC2B4830CBE73BD19D" box="[800,827,224,247]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="12" pageNumber="557">TI</collectionCode>
</materialsCitation>
&amp; AO, L;
<materialsCitation id="6DD43C8A7F74FFCC2BDF30CBE178D19D" ID-GBIF-Occurrence="2028138199" box="[951,1400,223,248]" collectingDate="2016-03-23" collectionCode="TI" pageId="12" pageNumber="557" specimenCount="4" specimenCount-female="4">
<specimenCount id="CBBAFD5E7F74FFCC2BDF30CBE026D192" box="[951,1062,223,248]" pageId="12" pageNumber="557" type="female">4 females</specimenCount>
, same locality,
<date id="A90210177F74FFCC2CBB30F4E14ED192" box="[1235,1358,223,248]" pageId="12" pageNumber="557" value="2016-03-23">
<collectingDate id="B946E9FF7F74FFCC2CBB30F4E14ED192" box="[1235,1358,223,248]" pageId="12" pageNumber="557" value="2016-03-23">23.iii.2016</collectingDate>
</date>
,
<collectionCode id="BBADAE127F74FFCC2D3330CBE178D19D" box="[1371,1400,224,247]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="12" pageNumber="557">TI</collectionCode>
</materialsCitation>
, L;
<materialsCitation id="6DD43C8A7F74FFCC28FF312FE6AFD076" ID-GBIF-Occurrence="2028138158" box="[151,687,260,285]" collectingDate="2012-10-28" collectionCode="TI" elevation="9" pageId="12" pageNumber="557" specimenCount="1" specimenCount-female="1">
<specimenCount id="CBBAFD5E7F74FFCC28FF312FE4FBD077" box="[151,251,260,285]" pageId="12" pageNumber="557" type="female">1 female</specimenCount>
, Maera-gawa,
<elevation id="5691D1E47F74FFCC29C8312FE608D077" box="[416,520,260,285]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="557" unit="m" value="9.0">
<quantity id="1A449B327F74FFCC29C8312FE5CBD076" box="[416,459,260,284]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="557" unit="m" value="9.0">9 m</quantity>
a.s.l.
</elevation>
,
<date id="A90210177F74FFCC2A7C312FE687D076" box="[532,647,260,284]" pageId="12" pageNumber="557" value="2012-10-28">
<collectingDate id="B946E9FF7F74FFCC2A7C312FE687D076" box="[532,647,260,284]" pageId="12" pageNumber="557" value="2012-10-28">28.x.2012</collectingDate>
</date>
,
<collectionCode id="BBADAE127F74FFCC2AFA312EE6AFD076" box="[658,687,261,284]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="12" pageNumber="557">TI</collectionCode>
</materialsCitation>
, L;
<materialsCitation id="6DD43C8A7F74FFCC2AB2312FE178D076" ID-GBIF-Occurrence="2028138184" box="[730,1400,260,285]" collectingDate="2015-10-23" collectionCode="TI" elevation="10" pageId="12" pageNumber="557" specimenCount="6" specimenCount-female="2" specimenCount-male="4">
<specimenCount id="CBBAFD5E7F74FFCC2AB2312FE734D077" box="[730,820,260,285]" pageId="12" pageNumber="557" type="male">4 males</specimenCount>
,
<specimenCount id="CBBAFD5E7F74FFCC2B57312FE7AED077" box="[831,942,260,285]" pageId="12" pageNumber="557" type="female">2 females</specimenCount>
, Yuchin-gawa,
<elevation id="5691D1E47F74FFCC2C33312FE0D0D077" box="[1115,1232,260,285]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="557" unit="m" value="10.0">
<quantity id="1A449B327F74FFCC2C33312FE094D076" box="[1115,1172,260,284]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="557" unit="m" value="10.0">10 m</quantity>
a.s.l.
</elevation>
,
<date id="A90210177F74FFCC2CB5312FE150D077" box="[1245,1360,260,285]" pageId="12" pageNumber="557" value="2015-10-23">
<collectingDate id="B946E9FF7F74FFCC2CB5312FE150D077" box="[1245,1360,260,285]" pageId="12" pageNumber="557" value="2015-10-23">23.x.2015</collectingDate>
</date>
,
<collectionCode id="BBADAE127F74FFCC2D33312EE178D076" box="[1371,1400,261,284]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="12" pageNumber="557">TI</collectionCode>
</materialsCitation>
, L;
<materialsCitation id="6DD43C8A7F74FFCC28FF310CE6AED055" ID-GBIF-Occurrence="2028138153" box="[151,686,295,320]" collectingDate="2016-10-22" collectionCode="TI" elevation="10" pageId="12" pageNumber="557" specimenCount="9" specimenCount-female="4" specimenCount-male="5">
<specimenCount id="CBBAFD5E7F74FFCC28FF310CE4EFD02A" box="[151,239,295,320]" pageId="12" pageNumber="557" type="male">5 males</specimenCount>
,
<specimenCount id="CBBAFD5E7F74FFCC28923103E568D02A" box="[250,360,295,320]" pageId="12" pageNumber="557" type="female">4 females</specimenCount>
, same locality,
<date id="A90210177F74FFCC2A7B310CE685D02A" box="[531,645,295,320]" pageId="12" pageNumber="557" value="2016-10-22">
<collectingDate id="B946E9FF7F74FFCC2A7B310CE685D02A" box="[531,645,295,320]" pageId="12" pageNumber="557" value="2016-10-22">22.x.2016</collectingDate>
</date>
,
<collectionCode id="BBADAE127F74FFCC2AF93103E6AED055" box="[657,686,296,319]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="12" pageNumber="557">TI</collectionCode>
</materialsCitation>
, L;
<materialsCitation id="6DD43C8A7F74FFCC2AB03103E4B2D00E" ID-GBIF-Occurrence="2028138192" collectingDate="1999-03-24" collectionCode="TI" pageId="12" pageNumber="557" specimenCount="44" specimenCount-female="25" specimenCount-male="19">
<specimenCount id="CBBAFD5E7F74FFCC2AB03103E73ED02A" box="[728,830,295,320]" pageId="12" pageNumber="557" type="male">19 males</specimenCount>
,
<specimenCount id="CBBAFD5E7F74FFCC2B21310CE7C5D02A" box="[841,965,295,320]" pageId="12" pageNumber="557" type="female">25 females</specimenCount>
, Omijya-gawa, Omijya-bashi,
<date id="A90210177F74FFCC2D74310CE198D02A" box="[1308,1432,295,320]" pageId="12" pageNumber="557" value="1999-03-24">
<collectingDate id="B946E9FF7F74FFCC2D74310CE198D02A" box="[1308,1432,295,320]" pageId="12" pageNumber="557" value="1999-03-24">24.iii.1999</collectingDate>
</date>
,
<collectionCode id="BBADAE127F74FFCC28FF3166E4B2D00E" box="[151,178,333,356]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="12" pageNumber="557">TI</collectionCode>
</materialsCitation>
&amp; AO, L;
<materialsCitation id="6DD43C8A7F74FFCC29463167E742D00E" ID-GBIF-Occurrence="2028138163" box="[302,834,332,357]" collectingDate="2009-05-26" collectionCode="TI" collectorName="K. Tojo" pageId="12" pageNumber="557" specimenCount="3" specimenCount-female="3">
<specimenCount id="CBBAFD5E7F74FFCC29463167E59ED00F" box="[302,414,332,357]" pageId="12" pageNumber="557" type="female">3 females</specimenCount>
, same locality,
<date id="A90210177F74FFCC2A243167E6BDD00F" box="[588,701,332,357]" pageId="12" pageNumber="557" value="2009-05-26">
<collectingDate id="B946E9FF7F74FFCC2A243167E6BDD00F" box="[588,701,332,357]" pageId="12" pageNumber="557" value="2009-05-26">26.v.2009</collectingDate>
</date>
,
<collectorName id="704953017F74FFCC2AA13166E722D00F" box="[713,802,332,357]" pageId="12" pageNumber="557">K. Tojo</collectorName>
, L
</materialsCitation>
;
<materialsCitation id="6DD43C8A7F74FFCC2B263167E108D00E" ID-GBIF-Occurrence="2028138188" box="[846,1288,332,357]" collectingDate="2012-10-29" collectionCode="TI" collectorName="K. Tojo" pageId="12" pageNumber="557" specimenCount="8" specimenCount-female="8">
<specimenCount id="CBBAFD5E7F74FFCC2B263167E7BED00F" box="[846,958,332,357]" pageId="12" pageNumber="557" type="female">8 females</specimenCount>
, same locality,
<date id="A90210177F74FFCC2C043167E0DFD00E" box="[1132,1247,332,356]" pageId="12" pageNumber="557" value="2012-10-29">
<collectingDate id="B946E9FF7F74FFCC2C043167E0DFD00E" box="[1132,1247,332,356]" pageId="12" pageNumber="557" value="2012-10-29">29.x.2012</collectingDate>
</date>
,
<collectionCode id="BBADAE127F74FFCC2C833166E108D00E" box="[1259,1288,333,356]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="12" pageNumber="557">TI</collectionCode>
</materialsCitation>
, P;
<materialsCitation id="6DD43C8A7F74FFCC2D5B3167E5D9D0ED" ID-GBIF-Occurrence="2028138190" collectingDate="2013-11-30" collectionCode="TI" collectorName="K. Tojo" pageId="12" pageNumber="557" specimenCount="1" specimenCount-female="1">
<specimenCount id="CBBAFD5E7F74FFCC2D5B3167E198D00F" box="[1331,1432,332,357]" pageId="12" pageNumber="557" type="female">1 female</specimenCount>
, same locality,
<date id="A90210177F74FFCC295E3144E5B1D0E2" box="[310,433,367,392]" pageId="12" pageNumber="557" value="2013-11-30">
<collectingDate id="B946E9FF7F74FFCC295E3144E5B1D0E2" box="[310,433,367,392]" pageId="12" pageNumber="557" value="2013-11-30">30.xi.2013</collectingDate>
</date>
,
<collectionCode id="BBADAE127F74FFCC29D4315BE5D9D0ED" box="[444,473,368,391]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="12" pageNumber="557">TI</collectionCode>
</materialsCitation>
, P;
<materialsCitation id="6DD43C8A7F74FFCC2A6A315BE7B6D0ED" ID-GBIF-Occurrence="2028138178" box="[514,950,367,392]" collectingDate="2015-10-21" collectionCode="TI" collectorName="K. Tojo" pageId="12" pageNumber="557" specimenCount="2" specimenCount-female="2">
<specimenCount id="CBBAFD5E7F74FFCC2A6A315BE670D0E2" box="[514,624,367,392]" pageId="12" pageNumber="557" type="female">2 females</specimenCount>
, same locality,
<date id="A90210177F74FFCC2B733144E78DD0E2" box="[795,909,367,392]" pageId="12" pageNumber="557" value="2015-10-21">
<collectingDate id="B946E9FF7F74FFCC2B733144E78DD0E2" box="[795,909,367,392]" pageId="12" pageNumber="557" value="2015-10-21">21.x.2015</collectingDate>
</date>
,
<collectionCode id="BBADAE127F74FFCC2BF0315BE7B6D0ED" box="[920,950,368,391]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="12" pageNumber="557">TI</collectionCode>
</materialsCitation>
;
<materialsCitation id="6DD43C8A7F74FFCC2BA93144E17ED0ED" ID-GBIF-Occurrence="2028138185" box="[961,1406,367,392]" collectingDate="2016-03-23" collectionCode="TI" collectorName="K. Tojo" pageId="12" pageNumber="557" specimenCount="5" specimenCount-female="5">
<specimenCount id="CBBAFD5E7F74FFCC2BA93144E02FD0E2" box="[961,1071,367,392]" pageId="12" pageNumber="557" type="female">5 females</specimenCount>
, same locality,
<date id="A90210177F74FFCC2CB23144E156D0E2" box="[1242,1366,367,392]" pageId="12" pageNumber="557" value="2016-03-23">
<collectingDate id="B946E9FF7F74FFCC2CB23144E156D0E2" box="[1242,1366,367,392]" pageId="12" pageNumber="557" value="2016-03-23">23.iii.2016</collectingDate>
</date>
,
<collectionCode id="BBADAE127F74FFCC2D09315BE17ED0ED" box="[1377,1406,368,391]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="12" pageNumber="557">TI</collectionCode>
</materialsCitation>
, P.
</paragraph>
</subSubSection>
<subSubSection id="95A6655C7F74FFCC28AF31BFE06ED0C7" box="[199,1134,404,429]" pageId="12" pageNumber="557" type="etymology">
<paragraph id="DD0336D77F74FFCC28AF31BFE06ED0C7" blockId="12.[151,1437,151,429]" box="[199,1134,404,429]" pageId="12" pageNumber="557">
<emphasis id="EFC8EAC57F74FFCC28AF31BFE547D0C7" bold="true" box="[199,327,404,429]" pageId="12" pageNumber="557">Etymology</emphasis>
. The name “
<taxonomicName id="1ABC4D547F74FFCC29BF31BEE605D0C6" authorityName="Ito &amp; Nozaki" authorityYear="2018" box="[471,517,405,428]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="557" phylum="Arthropoda" rank="species" species="aira">
<emphasis id="EFC8EAC57F74FFCC29BF31BEE605D0C6" box="[471,517,405,428]" italics="true" pageId="12" pageNumber="557">aira</emphasis>
</taxonomicName>
” is a noun in apposition, coined from the
<typeStatus id="020788757F74FFCC2BB531BEE00DD0C7" box="[989,1037,405,429]" pageId="12" pageNumber="557">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<caption id="89C3665F7F74FFCC28FF377DE070D68D" httpUri="https://zenodo.org/record/2606678/files/figure.png" pageId="12" pageNumber="557" startId="12.[151,250,1878,1900]" targetBox="[181,1425,487,1837]" targetPageId="12">
<paragraph id="DD0336D77F74FFCC28FF377DE070D68D" blockId="12.[151,1436,1878,2023]" pageId="12" pageNumber="557">
<emphasis id="EFC8EAC57F74FFCC28FF377DE512D601" bold="true" box="[151,274,1878,1900]" pageId="12" pageNumber="557">FIGURE 6.</emphasis>
<taxonomicName id="1ABC4D547F74FFCC2970377DE67FD606" authority="Ito &amp; Nozaki" authorityName="Ito &amp; Nozaki" authorityYear="2018" box="[280,639,1878,1900]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="557" phylum="Arthropoda" rank="species" species="aira" status="sp. nov.">
<emphasis id="EFC8EAC57F74FFCC2970377DE5F5D606" box="[280,501,1878,1900]" italics="true" pageId="12" pageNumber="557">Cheumatopsyche aira</emphasis>
Ito &amp; Nozaki
</taxonomicName>
<emphasis id="EFC8EAC57F74FFCC2AED377CE6D5D606" bold="true" box="[645,725,1879,1900]" pageId="12" pageNumber="557">
<taxonomicNameLabel id="F4FB57BE7F74FFCC2AED377CE6D5D606" box="[645,725,1879,1900]" pageId="12" pageNumber="557" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Male (6A–6F): 6A, right wings, dorsal, light color marks of forewing, venations of forewing and hind wing; 6B, genitalia, left lateral; 6C, same, dorsal; 6D, left inferior appendage, ventral; 6E, phallic apparatus, left lateral; 6F, apical part of phallus, ventral. Female (6G–6I): 6G, right wings, dorsal; 6H, genitalia, left lateral; 6I, same, ventral. Abbreviations: I–V = apical forks I–V; VIII–X = abdominal segments VIII–X; e.p. = endothecal process (paired); l.l. = lateral lobe (paired); m.c.l. = mesocaudal lobe; p.s. = phallotremal sclerite.
</paragraph>
</caption>
<subSubSection id="95A6655C7F75FFCD28AF30BCE01ED1DA" box="[199,1054,151,176]" pageId="13" pageNumber="558" type="distribution">
<paragraph id="DD0336D77F75FFCD28AF30BCE01ED1DA" blockId="13.[151,1436,151,285]" box="[199,1054,151,176]" pageId="13" pageNumber="558">
<emphasis id="EFC8EAC57F75FFCD28AF30BCE559D1DA" bold="true" box="[199,345,151,176]" pageId="13" pageNumber="558">Distribution</emphasis>
.
<collectingCountry id="A5AB76477F75FFCD290F30B3E5ABD1C5" box="[359,427,152,175]" name="Japan" pageId="13" pageNumber="558">Japan</collectingCountry>
: Ryukyu (Iriomote-jima). Endemic to southern Ryukyu.
</paragraph>
</subSubSection>
<subSubSection id="95A6655C7F75FFCD28AF3097E6C9D077" pageId="13" pageNumber="558" type="discussion">
<paragraph id="DD0336D77F75FFCD28AF3097E5B1D19D" blockId="13.[151,1436,151,285]" pageId="13" pageNumber="558">
<emphasis id="EFC8EAC57F75FFCD28AF3097E534D1BF" bold="true" box="[199,308,188,213]" pageId="13" pageNumber="558">Remarks</emphasis>
. This is the most common hydropsychid caddisflies in Iriomote-jima and sometimes is collected together with
<taxonomicName id="1ABC4D547F75FFCD295A30CBE5AAD192" box="[306,426,224,248]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="558" phylum="Arthropoda" rank="species" species="clavalis">
<emphasis id="EFC8EAC57F75FFCD295A30CBE5AAD192" box="[306,426,224,248]" italics="true" pageId="13" pageNumber="558">C. clavalis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="DD0336D77F75FFCD28AF312FE6C9D077" blockId="13.[151,1436,151,285]" box="[199,713,260,285]" pageId="13" pageNumber="558">
<emphasis id="EFC8EAC57F75FFCD28AF312FE57CD077" bold="true" box="[199,380,260,285]" pageId="13" pageNumber="558">Japanese name</emphasis>
. Aira-kogata-shima-tobikera.
</paragraph>
</subSubSection>
</treatment>
</document>
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