treatments-xml/data/55/0B/87/550B87C3C162FFBAFCEC72AFFE700368.xml
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<document id="A4C3A95E403F79B90683FF12F44D6AF4" ID-DOI="10.5281/zenodo.4651080" ID-ISSN="1638-9395" ID-Zenodo-Dep="4651080" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617195131926" checkinUser="felipe" docAuthor="Bianucci, Giovanni, Lambert, Olivier &amp; Post, Klaas" docDate="2007" docId="550B87C3C162FFBAFCEC72AFFE700368" docLanguage="en" docName="Geodiversitas.29.4.561-618.pdf" docOrigin="Geodiversitas 29 (4)" docStyle="DocumentStyle:94524207C6714CF92F879FE43E306202.3:Geodiversitas.2000-2008.journal_article.type2" docStyleId="94524207C6714CF92F879FE43E306202" docStyleName="Geodiversitas.2000-2008.journal_article.type2" docStyleVersion="3" docTitle="Nenga meganasalis Bianucci &amp; Lambert &amp; Post 2007, n. sp." docType="treatment" docUuid="1D401A65-E781-4E70-9AF8-04059BE605EE" docUuidSource="ZooBank" docVersion="4" lastPageNumber="592" masterDocId="A932FFBBC17EFFA5FFC47277FFBE056E" masterDocTitle="A high diversity in fossil beaked whales (Mammalia, Odontoceti, Ziphiidae) recovered by trawling from the sea floor off South Africa" masterLastPageNumber="618" masterPageNumber="561" pageNumber="589" updateTime="1698897450673" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="FB8529285C869F39278001BA0058D52C">A high diversity in fossil beaked whales (Mammalia, Odontoceti, Ziphiidae) recovered by trawling from the sea floor off South Africa</mods:title>
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<mods:namePart id="F92E53908B1734234D715B4FAB1C9358">Bianucci, Giovanni</mods:namePart>
<mods:affiliation id="D22464C304532F51DC41BE09ECB719D0">Dipartimento di Scienze della Terra, Università di Pisa, via S. Maria, 53, I- 56126 Pisa (Italy) bianucci @ dst. unipi. it</mods:affiliation>
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<mods:namePart id="4C01D4C1A3EFBAB0A026BEF4A72443BA">Lambert, Olivier</mods:namePart>
<mods:affiliation id="1EEEB0D82880BB2162AC7BF5DF0D441F">Institut royal des Sciences naturelles de Belgique, Département de Paléontologie, rue Vautier, 29, B- 1000 Brussels (Belgium) olivier. lambert @ naturalsciences. be</mods:affiliation>
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<mods:namePart id="6EB6157CF81C4D45DC7B2ADB3122BCC9">Post, Klaas</mods:namePart>
<mods:affiliation id="8EE08610F3A3293F8646D184DE1BECFD">Natuurhistorisch Museum Rotterdam, P. O. Box 23452, NL- 3001 Rotterdam (The Netherlands) klaas @ fiskano. nl</mods:affiliation>
<mods:nameIdentifier id="14462B97312EAC4EB21B70800D708DFB" type="email">klaas@fiskano.nl</mods:nameIdentifier>
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<mods:title id="8C50406F59CD332657C76981CC1C4CA7">Geodiversitas</mods:title>
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<mods:date id="7F027CEEC4F647DB845BB6BD407CAC38">2007</mods:date>
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<subSubSection id="95B8655EC162FFB9FCEC72AFFB88059C" box="[808,1078,215,242]" pageId="28" pageNumber="589" type="nomenclature">
<paragraph id="DD1D36D5C162FFB9FCEC72AFFB88059C" blockId="28.[808,1078,215,242]" box="[808,1078,215,242]" pageId="28" pageNumber="589">
<heading id="865581B9C162FFB9FCEC72AFFB88059C" box="[808,1078,215,242]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="589" reason="2">
<taxonomicName id="1AA24D56C162FFB9FCEC72AFFC4B059C" authority="Bianucci &amp; Lambert &amp; Post, 2007" authorityName="Bianucci &amp; Lambert &amp; Post" authorityYear="2007" box="[808,1013,215,242]" class="Liliopsida" family="Arecaceae" genus="Nenga" kingdom="Plantae" order="Arecales" pageId="28" pageNumber="589" phylum="Tracheophyta" rank="species" species="meganasalis" status="sp. nov.">
<emphasis id="EFD6EAC7C162FFB9FCEC72AFFC4B059C" bold="true" box="[808,1013,215,242]" italics="true" pageId="28" pageNumber="589">Nenga meganasalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4E557BCC162FFB9FC3872AFFB88059C" box="[1020,1078,216,242]" pageId="28" pageNumber="589" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95B8655EC162FFB9FD617366FC3F0742" pageId="28" pageNumber="589" type="materials_examined">
<paragraph id="DD1D36D5C162FFB9FD617366FD770433" blockId="28.[677,1209,273,349]" pageId="28" pageNumber="589">
<materialsCitation id="6DCA3C88C162FFB9FD617366FD770433" collectionCode="SAM" pageId="28" pageNumber="589" specimenCode="PQ 69675" specimenCount="1" typeStatus="holotype">
<typeStatus id="02198877C162FFB9FD617366FCB20449" box="[677,780,273,296]" pageId="28" pageNumber="589" type="holotype">HOLOTYPE</typeStatus>
. —
<collectionCode id="BBB3AE10C162FFB9FCF27366FCD70447" box="[822,873,273,297]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="28" pageNumber="589">SAM</collectionCode>
<specimenCode id="8D049EAEC162FFB9FCA97366FC6A0447" box="[877,980,273,297]" collectionCode="SAM" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="28" pageNumber="589">PQ 69675</specimenCode>
, a partial skull including the rostrum, the premaxillary sac fossae, and the vertex.
</materialsCitation>
</paragraph>
<paragraph id="DD1D36D5C162FFB9FD617301FC3F0742" blockId="28.[677,1210,373,556]" pageId="28" pageNumber="589">
REFERRED SPECIMENS. —
<collectionCode id="BBB3AE10C162FFB9FC707302FC5404E3" box="[948,1002,373,397]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="28" pageNumber="589">SAM</collectionCode>
PQ 2117, a partial rostrum, trawled off the South African coast;
<collectionCode id="BBB3AE10C162FFB9FB9A73E7FB2F04C6" box="[1118,1169,400,424]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="28" pageNumber="589">SAM</collectionCode>
PQ 2339, a partial skull including the rostrum base, the premaxillary sac fossae, and the vertex, trawled off
<collectingRegion id="1F66F837C162FFB9FB4373B2FB0404B3" box="[1159,1210,453,477]" country="South Africa" name="Western Cape" pageId="28" pageNumber="589">Cape</collectingRegion>
coast;
<collectionCode id="BBB3AE10C162FFB9FD2773A8FCA80499" box="[739,790,479,503]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="28" pageNumber="589">SAM</collectionCode>
PQ 69676, a partial rostrum and anterior of the cranium, trawled west of
<collectingRegion id="1F66F837C162FFB9FC2F738DFBA1077C" box="[1003,1055,506,530]" country="South Africa" name="Western Cape" pageId="28" pageNumber="589">Cape</collectingRegion>
Point, Atlantic Ocean (depth
<quantity id="1A5A9B30C162FFB9FCF67063FCCB0742" box="[818,885,532,556]" metricMagnitude="2" metricUnit="m" metricValue="3.47" pageId="28" pageNumber="589" unit="m" value="347.0">347 m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="95B8655EC162FFB9FD617033FBC90718" pageId="28" pageNumber="589" type="etymology">
<paragraph id="DD1D36D5C162FFB9FD617033FBC90718" blockId="28.[677,1210,580,630]" pageId="28" pageNumber="589">ETYMOLOGY. — From the Ancient Greek “megas”, large; “meganasalis” for the large nasals of this species.</paragraph>
</subSubSection>
<subSubSection id="95B8655EC162FFB9FD6170F9FC0207AE" pageId="28" pageNumber="589" type="materials_examined">
<paragraph id="DD1D36D5C162FFB9FD6170F9FC0207AE" blockId="28.[677,1210,654,704]" pageId="28" pageNumber="589">
<collectionCode id="BBB3AE10C162FFB9FD6170F9FC8407CA" box="[677,826,654,677]" pageId="28" pageNumber="589">TYPE LOCALITY</collectionCode>
. — No exact locality. Trawled west of
<collectingRegion id="1F66F837C162FFB9FD6170DFFD6607AE" box="[677,728,680,704]" country="South Africa" name="Western Cape" pageId="28" pageNumber="589">Cape</collectingRegion>
Town, Atlantic Ocean.
</paragraph>
</subSubSection>
<subSubSection id="95B8655EC162FFB9FD6170AFFBD00143" pageId="28" pageNumber="589" type="diagnosis">
<paragraph id="DD1D36D5C162FFB9FD6170AFFBD00143" blockId="28.[677,1211,728,1069]" pageId="28" pageNumber="589">
DIAGNOSIS. —
<taxonomicName id="1AA24D56C162FFB9FCF870AEFC52079E" authority="Bianucci &amp; Lambert &amp; Post, 2007" authorityName="Bianucci &amp; Lambert &amp; Post" authorityYear="2007" box="[828,1004,728,752]" class="Liliopsida" family="Arecaceae" genus="Nenga" kingdom="Plantae" order="Arecales" pageId="28" pageNumber="589" phylum="Tracheophyta" rank="species" species="meganasalis" status="n. gen., n. sp.">
<emphasis id="EFD6EAC7C162FFB9FCF870AEFC52079E" box="[828,1004,728,752]" italics="true" pageId="28" pageNumber="589">Nenga meganasalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4E557BCC162FFB9FC3770AFFBC60781" box="[1011,1144,728,752]" pageId="28" pageNumber="589" rank="species">n. gen., n. sp.</taxonomicNameLabel>
differs from all the other
<taxonomicName id="1AA24D56C162FFB9FC8D7085FC1A0664" authorityName="Gray" authorityYear="1850" box="[841,932,754,778]" class="Mammalia" family="Ziphiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="28" pageNumber="589" phylum="Chordata" rank="family">Ziphiidae</taxonomicName>
in the nearly rectangular wide nasals occupying most of the vertex dorsally. It differs from the
<taxonomicName id="1AA24D56C162FFB9FD397150FCE20651" authorityName="Gray" authorityYear="1850" box="[765,860,807,831]" class="Mammalia" family="Ziphiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="28" pageNumber="589" phylum="Chordata" rank="subFamily" subFamily="Ziphiinae">Ziphiinae</taxonomicName>
in the mesorostral ossification of the mesethmoid and it lacks the main synapomorphy of the subfamily, the anterolaterally directed premaxillary crest. It differs from the
<taxonomicName id="1AA24D56C162FFB9FC927100FBB806E1" authorityName="Gray" authorityYear="1866" box="[854,1030,887,911]" class="Mammalia" family="Ziphiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="28" pageNumber="589" phylum="Chordata" rank="subFamily" subFamily="Hyperoodontinae">Hyperoodontinae</taxonomicName>
,
<taxonomicName id="1AA24D56C162FFB9FBCB7100FBD506E0" authority="Bianucci &amp; Lambert &amp; Post, 2007" authorityName="Bianucci &amp; Lambert &amp; Post" authorityYear="2007" box="[1039,1131,887,910]" class="Mammalia" family="Ziphiidae" genus="Pterocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="28" pageNumber="589" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="EFD6EAC7C162FFB9FBCB7100FBD506E0" box="[1039,1131,887,910]" italics="true" pageId="28" pageNumber="589">Pterocetus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4E557BCC162FFB9FBB57100FB0806E1" box="[1137,1206,887,911]" pageId="28" pageNumber="589" rank="genus">n. gen.</taxonomicNameLabel>
,
<taxonomicName id="1AA24D56C162FFB9FD6171E5FCB306C7" authorityName="Oliver" authorityYear="1937" box="[677,781,914,937]" class="Mammalia" family="Hyperoodontidae" genus="Tasmacetus" kingdom="Animalia" order="Cetacea" pageId="28" pageNumber="589" phylum="Chordata" rank="genus">
<emphasis id="EFD6EAC7C162FFB9FD6171E5FCB306C7" box="[677,781,914,937]" italics="true" pageId="28" pageNumber="589">Tasmacetus</emphasis>
</taxonomicName>
, and
<taxonomicName id="1AA24D56C162FFB9FC8C71E6FC1506C7" authority="Bianucci &amp; Lambert &amp; Post, 2007" authorityName="Bianucci &amp; Lambert &amp; Post" authorityYear="2007" box="[840,939,913,937]" class="Mammalia" family="Ziphiidae" genus="Xhosacetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="28" pageNumber="589" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="EFD6EAC7C162FFB9FC8C71E6FC1506C7" box="[840,939,913,937]" italics="true" pageId="28" pageNumber="589">Xhosacetus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4E557BCC162FFB9FC7771E5FC4506C4" box="[947,1019,914,938]" pageId="28" pageNumber="589" rank="genus">n. gen.</taxonomicNameLabel>
, in the mesorostral ossification of the mesethmoid and lacks the constriction of the ascending process of the premaxilla and the intrusion of the nasal in the narrow premaxillary crest. It differs from
<taxonomicName id="1AA24D56C162FFB9FCE4718CFC2D017D" authorityName="de Muizon" authorityYear="1983" box="[800,915,1019,1043]" class="Mammalia" family="Ziphiidae" genus="Ninoziphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="28" pageNumber="589" phylum="Chordata" rank="genus">
<emphasis id="EFD6EAC7C162FFB9FCE4718CFC2D017D" box="[800,915,1019,1043]" italics="true" pageId="28" pageNumber="589">Ninoziphius</emphasis>
</taxonomicName>
in the rostrum elevated at its base and the reduction of the maxillary alveoli.
</paragraph>
</subSubSection>
<subSubSection id="95B8655EC162FFBAFD617638FE700368" lastPageId="31" lastPageNumber="592" pageId="28" pageNumber="589" type="description">
<paragraph id="DD1D36D5C162FFB9FD617638FB930104" blockId="28.[677,1210,1103,1699]" box="[677,1069,1103,1130]" pageId="28" pageNumber="589">
DESCRIPTION (
<figureCitation id="45992A50C162FFB9FC817627FC7E0104" box="[837,960,1104,1130]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="29.[113,124,1451,1468]" captionStartId-1="30.[113,124,1634,1651]" captionStartId-2="31.[113,124,592,609]" captionTargetBox-0="[150,1176,266,1382]" captionTargetBox-1="[125,1201,219,1516]" captionTargetBox-2="[212,1104,276,557]" captionTargetId-0="figure-18@29.[132,1219,262,1428]" captionTargetId-1="figure-22@30.[119,1208,214,1538]" captionTargetId-2="figure-18@31.[197,1128,208,593]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionTargetPageId-2="31" captionText-0="FIG. 21. — Skull of Nenga meganasalis n. gen., n. sp. (SAM PQ 69675, holotype): A, dorsal view; B, lateral view; C, ventral view. Scale bar: 10 cm." captionText-1="FIG. 22. — Nenga meganasalis n. gen., n. sp.: A-D, skull (SAM PQ 69675, holotype); A, detail of the vertex in dorsal view; B, corresponding line drawing; C, anterior view; D, corresponding line drawing; E, incomplete skull (SAM PQ 2339), dorsal view. Scale bars: A, B, 5 cm; C-E, 10 cm." captionText-2="FIG. 23. — Rostrum of Nenga meganasalis n. gen., n. sp. (SAM PQ 69676), in dorsal view showing the mesorostral groove partially filled by the mesethmoid. Scale bar: 10 cm." figureDoi-0="http://doi.org/10.5281/zenodo.4651122" figureDoi-1="http://doi.org/10.5281/zenodo.4651124" figureDoi-2="http://doi.org/10.5281/zenodo.4651128" httpUri-0="https://zenodo.org/record/4651122/files/figure.png" httpUri-1="https://zenodo.org/record/4651124/files/figure.png" httpUri-2="https://zenodo.org/record/4651128/files/figure.png" pageId="28" pageNumber="589">FIGS 21-23</figureCitation>
;
<tableCitation id="9020036EC162FFB9FC0F7638FB9A0104" box="[971,1060,1103,1130]" captionStart="TABLE" captionStartId="54.[113,122,220,237]" captionText="TABLE 5. — Measurements (in mm) on the skulls of Ziphiidae incertae sedis from the Neogene of South Africa. Abbreviations: e, estimate; +, nearly complete; -, no data." pageId="28" pageNumber="589">TABLE 5</tableCitation>
)
</paragraph>
<paragraph id="DD1D36D5C162FFB9FD617619FB0403CD" blockId="28.[677,1210,1103,1699]" pageId="28" pageNumber="589">
The almost complete rostrum of the
<typeStatus id="02198877C162FFB9FBE67619FB3E01E6" box="[1058,1152,1134,1160]" pageId="28" pageNumber="589" type="holotype">holotype</typeStatus>
is robust and long. The maxilla ends more than
<quantity id="1A5A9B30C162FFB9FBAE76F9FB0701C7" box="[1130,1209,1166,1193]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="28" pageNumber="589" unit="mm" value="45.0">45 mm</quantity>
from the apex. At mid-length, the rostrum is wider than high. The morphology of the rostrum base varies within the species: it is wider in SAM PQ 2339 and SAM PQ 69676 with a wide concave subhorizontal surface of the maxilla following anteriorly the large maxillary foramen. In the
<typeStatus id="02198877C162FFB9FBEE773BFB370008" box="[1066,1161,1356,1382]" pageId="28" pageNumber="589" type="holotype">holotype</typeStatus>
and SAM PQ 2117, the dorsal surface of the maxilla narrows more rapidly anteriorly, while the lateral slope is more pronounced. The lateral margin of the maxilla at the rostrum base is acute. The vomer is weakly thickened or not at all in the widely open mesorostral groove. In specimen SAM PQ 69676, the mesorostral groove is posteriorly filled by the ossified mesethmoid for a length of more than
<quantity id="1A5A9B30C162FFB9FD61741DFCBC03EA" box="[677,770,1642,1668]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="589" unit="mm" value="100.0">100 mm</quantity>
(specimen incomplete anteriorly); in this species the ossified mesethmoid might have occupied
</paragraph>
<paragraph id="DD1D36D5C163FFB8FF0272ACFF69059D" blockId="29.[198,215,219,243]" box="[198,215,219,243]" pageId="29" pageNumber="590">A</paragraph>
<paragraph id="DD1D36D5C163FFB8FEFE7706FE3E00EA" blockId="29.[113,1209,1395,1491]" box="[314,384,1393,1412]" pageId="29" pageNumber="590">choana</paragraph>
<caption id="89DD665DC163FFB8FFB577DCFF6C00BD" ID-DOI="http://doi.org/10.5281/zenodo.4651122" ID-Zenodo-Dep="4651122" httpUri="https://zenodo.org/record/4651122/files/figure.png" pageId="29" pageNumber="590" startId="29.[113,124,1451,1468]" targetBox="[150,1176,266,1382]" targetPageId="29">
<paragraph id="DD1D36D5C163FFB8FFB577DCFF6C00BD" blockId="29.[113,1209,1395,1491]" pageId="29" pageNumber="590">
FIG. 21. — Skull of
<taxonomicName id="1AA24D56C163FFB8FEC977DCFE1300D2" authority="Bianucci &amp; Lambert &amp; Post, 2007" authorityName="Bianucci &amp; Lambert &amp; Post" authorityYear="2007" box="[269,429,1451,1468]" class="Liliopsida" family="Arecaceae" genus="Nenga" kingdom="Plantae" order="Arecales" pageId="29" pageNumber="590" phylum="Tracheophyta" rank="species" species="meganasalis" status="n. gen., n. sp.">
<emphasis id="EFD6EAC7C163FFB8FEC977DCFE1300D2" box="[269,429,1451,1468]" italics="true" pageId="29" pageNumber="590">Nenga meganasalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4E557BCC163FFB8FE7677DCFD9A00D2" box="[434,548,1451,1468]" pageId="29" pageNumber="590" rank="species">n. gen., n. sp.</taxonomicNameLabel>
(SAM PQ 69675, holotype):
<emphasis id="EFD6EAC7C163FFB8FCC977DCFCA400D2" bold="true" box="[781,794,1451,1468]" pageId="29" pageNumber="590">A</emphasis>
, dorsal view;
<emphasis id="EFD6EAC7C163FFB8FC4F77DCFC2600D2" bold="true" box="[907,920,1451,1468]" pageId="29" pageNumber="590">B</emphasis>
, lateral view;
<emphasis id="EFD6EAC7C163FFB8FBCC77DCFBA800D2" bold="true" box="[1032,1046,1451,1468]" pageId="29" pageNumber="590">C</emphasis>
, ventral view. Scale bar: 10 cm.
</paragraph>
</caption>
<paragraph id="DD1D36D5C163FFB8FFB5747CFE7B030A" blockId="29.[113,647,1546,1700]" pageId="29" pageNumber="590">
most of the length of the mesorostral groove. Shallow alveoli marks are still visible on the maxillary alveolar groove of the
<typeStatus id="02198877C163FFB8FE9B743DFE7F030A" box="[351,449,1610,1636]" pageId="29" pageNumber="590" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="DD1D36D5C163FFB8FF4C741EFB0403CA" blockId="29.[113,647,1546,1700]" lastBlockId="29.[677,1210,1546,1700]" pageId="29" pageNumber="590">The premaxillary sac fossa is large and slightly concave; it rises slowly posteriorly until the ascending process. The ascent to the vertex from this point is more abrupt, but barely reaching vertical. In anterior view, the lateral margins of the premaxillae are roughly parallel until the transversely directed, weakly laterally developed, premaxillary crests.</paragraph>
<paragraph id="DD1D36D5C160FFBBFE6A779CFC96036B" blockId="30.[318,808,1482,1556]" box="[430,808,1515,1541]" pageId="30" pageNumber="591">premaxillary premaxillary foramen</paragraph>
<paragraph id="DD1D36D5C160FFBBFE6A7476FDB7037A" blockId="30.[318,808,1482,1556]" box="[430,521,1537,1556]" pageId="30" pageNumber="591">sac fossa</paragraph>
<caption id="89DD665DC160FFBBFFB57415FE8603CE" ID-DOI="http://doi.org/10.5281/zenodo.4651124" ID-Zenodo-Dep="4651124" httpUri="https://zenodo.org/record/4651124/files/figure.png" pageId="30" pageNumber="591" startId="30.[113,124,1634,1651]" targetBox="[125,1201,219,1516]" targetPageId="30">
<paragraph id="DD1D36D5C160FFBBFFB57415FE8603CE" blockId="30.[113,1211,1634,1696]" pageId="30" pageNumber="591">
FIG. 22. —
<taxonomicName id="1AA24D56C160FFBBFF087415FED0031D" authority="Bianucci &amp; Lambert &amp; Post, 2007" authorityName="Bianucci &amp; Lambert &amp; Post" authorityYear="2007" box="[204,366,1634,1651]" class="Liliopsida" family="Arecaceae" genus="Nenga" kingdom="Plantae" order="Arecales" pageId="30" pageNumber="591" phylum="Tracheophyta" rank="species" species="meganasalis" status="n. gen., n. sp.">
<emphasis id="EFD6EAC7C160FFBBFF087415FED0031D" box="[204,366,1634,1651]" italics="true" pageId="30" pageNumber="591">Nenga meganasalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4E557BCC160FFBBFEB07415FE54031D" box="[372,490,1634,1651]" pageId="30" pageNumber="591" rank="species">n. gen., n. sp.</taxonomicNameLabel>
:
<emphasis id="EFD6EAC7C160FFBBFE357415FDAD031D" bold="true" box="[497,531,1634,1651]" pageId="30" pageNumber="591">A -D</emphasis>
, skull (SAM PQ 69675, holotype);
<emphasis id="EFD6EAC7C160FFBBFCF57415FC80031D" bold="true" box="[817,830,1634,1651]" pageId="30" pageNumber="591">A</emphasis>
, detail of the vertex in dorsal view;
<emphasis id="EFD6EAC7C160FFBBFBA27415FBCD031D" bold="true" box="[1126,1139,1634,1651]" pageId="30" pageNumber="591">B</emphasis>
, corresponding line drawing;
<emphasis id="EFD6EAC7C160FFBBFEE8740EFE8403E4" bold="true" box="[300,314,1657,1674]" pageId="30" pageNumber="591">C</emphasis>
, anterior view;
<emphasis id="EFD6EAC7C160FFBBFE7E740EFE7603E4" bold="true" box="[442,456,1657,1674]" pageId="30" pageNumber="591">D</emphasis>
, corresponding line drawing;
<emphasis id="EFD6EAC7C160FFBBFD06740EFD7003E4" bold="true" box="[706,718,1657,1674]" pageId="30" pageNumber="591">E</emphasis>
, incomplete skull (SAM PQ 2339), dorsal view. Scale bars: A, B, 5 cm; C-E, 10 cm.
</paragraph>
</caption>
<paragraph id="DD1D36D5C161FFBAFE227293FD200599" blockId="31.[486,1005,228,321]" box="[486,670,228,247]" pageId="31" pageNumber="592">mesorostral groove</paragraph>
<paragraph id="DD1D36D5C161FFBAFE22728AFC7B0463" blockId="31.[486,1005,228,321]" box="[486,965,250,272]" pageId="31" pageNumber="592">filled by the mesethmoid concave dorsal surface</paragraph>
<caption id="89DD665DC161FFBAFFB57027FE620716" ID-DOI="http://doi.org/10.5281/zenodo.4651128" ID-Zenodo-Dep="4651128" httpUri="https://zenodo.org/record/4651128/files/figure.png" pageId="31" pageNumber="592" startId="31.[113,124,592,609]" targetBox="[212,1104,276,557]" targetPageId="31">
<paragraph id="DD1D36D5C161FFBAFFB57027FE620716" blockId="31.[113,1209,538,632]" pageId="31" pageNumber="592">
FIG. 23. — Rostrum of
<taxonomicName id="1AA24D56C161FFBAFEF57027FE6D070F" authority="Bianucci &amp; Lambert &amp; Post, 2007" authorityName="Bianucci &amp; Lambert &amp; Post" authorityYear="2007" box="[305,467,592,609]" class="Liliopsida" family="Arecaceae" genus="Nenga" kingdom="Plantae" order="Arecales" pageId="31" pageNumber="592" phylum="Tracheophyta" rank="species" species="meganasalis" status="n. gen., n. sp.">
<emphasis id="EFD6EAC7C161FFBAFEF57027FE6D070F" box="[305,467,592,609]" italics="true" pageId="31" pageNumber="592">Nenga meganasalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4E557BCC161FFBAFE1D7027FDF1070F" box="[473,591,592,609]" pageId="31" pageNumber="592" rank="species">n. gen., n. sp.</taxonomicNameLabel>
(SAM PQ 69676), in dorsal view showing the mesorostral groove partially filled by the mesethmoid. Scale bar: 10 cm.
</paragraph>
</caption>
<paragraph id="DD1D36D5C161FFBAFFB570C4FD9E0642" blockId="31.[113,647,691,1542]" pageId="31" pageNumber="592">The posterior projection of the premaxilla along the nasal contacts the frontal. The outline of the bony nares varies within the species being more V-shaped and longer in SAM PQ 2339.</paragraph>
<paragraph id="DD1D36D5C161FFBAFF4C7145FEAC0684" blockId="31.[113,647,691,1542]" pageId="31" pageNumber="592">
In the
<typeStatus id="02198877C161FFBAFF0A7145FE8C0622" box="[206,306,818,844]" pageId="31" pageNumber="592" type="holotype">holotype</typeStatus>
, several smaller foramina pierce the maxilla just behind the large maxillary foramen at the rostrum base. The relative position of maxillary and premaxillary foramina varies within the species, around the level of the antorbital notch.
</paragraph>
<paragraph id="DD1D36D5C161FFBAFF4C7187FE700368" blockId="31.[113,647,691,1542]" pageId="31" pageNumber="592">
The nasals occupy a large nearly rectangular surface on the wide and low vertex, much longer than the short strip of frontals (complete on SAM PQ2339), and wider than long. The lateral margin of the nasal is moderately convex and its anterolateral corner forms only a small part of the premaxillary crest. A vertical groove excavates the anterior margin of each nasal and the rounded anterior point of the nasals is only slightly more anterior than the premaxillary crests. The naso-frontal suture is roughly rectilinear. The morphology of the posterior portion of the vertex is not as well preserved as in
<taxonomicName id="1AA24D56C161FFBAFF63771BFEFA00E8" authority="Bianucci &amp; Lambert &amp; Post, 2007" authorityName="Bianucci &amp; Lambert &amp; Post" authorityYear="2007" box="[167,324,1388,1414]" class="Mammalia" family="Ziphiidae" genus="Microberardius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="31" pageNumber="592" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="EFD6EAC7C161FFBAFF63771BFEFA00E8" box="[167,324,1388,1414]" italics="true" pageId="31" pageNumber="592">Microberardius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4E557BCC161FFBAFE8F771AFE2A00E9" box="[331,404,1389,1415]" pageId="31" pageNumber="592" rank="genus">n. gen.</taxonomicNameLabel>
and in the
<taxonomicName id="1AA24D56C161FFBAFDCF771BFD3800E8" box="[523,646,1388,1414]" class="Mammalia" family="Ziphiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="31" pageNumber="592" phylum="Chordata" rank="subTribe" subTribe="Berardiinae">Berardiinae</taxonomicName>
indet. described above. We therefore are unable to detect the presence or absence of the interparietal, preventing firm attribution to the subfamily
<taxonomicName id="1AA24D56C161FFBAFDA577BBFF6E036B" class="Mammalia" family="Ziphiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="31" pageNumber="592" phylum="Chordata" rank="subTribe" subTribe="Berardiinae">Berardiinae</taxonomicName>
(see phylogeny below).
</paragraph>
</subSubSection>
</treatment>
</document>