258 lines
27 KiB
XML
258 lines
27 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.277.3594" ID-GBIF-Dataset="e4370703-19bb-4809-9e86-c4fcb410964c" ID-PMC="PMC3677373" ID-Pensoft-Pub="1313-2970-277-69" ID-PubMed="23794824" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-277-69" ModsDocOrigin="ZooKeys 277" ModsDocTitle="Two sympatric new species of woodlizards (Hoplocercinae, Enyalioides) from Cordillera Azul National Park in northeastern Peru" checkinTime="1451247556685" checkinUser="pensoft" docAuthor="Venegas, Pablo J., Torres-Carvajal, Omar, Duran, Vilma & Queiroz, Kevin de" docDate="2013" docId="55048C72369F5C621DAC379D23A59E0E" docLanguage="en" docName="ZooKeys 277: 69-90" docOrigin="ZooKeys 277" docSource="http://dx.doi.org/10.3897/zookeys.277.3594" docTitle="Enyalioides azulae Venegas, Torres-Carvajal, Duran & Queiroz, 2013, sp. n." docType="treatment" docVersion="4" lastPageNumber="77" masterDocId="FFB68F4D5E1E0521FFC7E22BFFB77E0D" masterDocTitle="Two sympatric new species of woodlizards (Hoplocercinae, Enyalioides) from Cordillera Azul National Park in northeastern Peru" masterLastPageNumber="90" masterPageNumber="69" pageNumber="71" updateTime="1668155493910" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Two sympatric new species of woodlizards (Hoplocercinae, Enyalioides) from Cordillera Azul National Park in northeastern Peru</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Venegas, Pablo J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Torres-Carvajal, Omar</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Duran, Vilma</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Queiroz, Kevin de</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>277</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>69</mods:start>
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<mods:end>90</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.277.3594</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.277.3594</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-277-69</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152041610" LSID="urn:lsid:zoobank.org:act:FADE520D-B1C5-4C5A-A54D-6E8A682E1E29" httpUri="http://treatment.plazi.org/id/55048C72369F5C621DAC379D23A59E0E" lastPageId="8" lastPageNumber="77" pageId="2" pageNumber="71">
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<subSubSection pageId="2" pageNumber="71" type="nomenclature">
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<paragraph pageId="2" pageNumber="71">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:FADE520D-B1C5-4C5A-A54D-6E8A682E1E29" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides azulae" order="Squamata" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="azulae">Enyalioides azulae</taxonomicName>
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<taxonomicNameLabel pageId="2" pageNumber="71">sp. n.</taxonomicNameLabel>
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Figs 1-4
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="71" type="holotype">
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<paragraph pageId="2" pageNumber="71">Holotype.</paragraph>
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<paragraph pageId="2" pageNumber="71">
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CORBIDI 06772 (Fig. 1), an adult male from Chambirillo close to Checkpoint 16 of the CAZNP (
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-7.069139">07°04'8.9"S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-76.01422">76°00'51.2"W</geoCoordinate>
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, 1122 m), Provincia de Picota,
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<normalizedToken originalValue="Región">Region</normalizedToken>
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San
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<normalizedToken originalValue="Martín">Martin</normalizedToken>
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,
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<normalizedToken originalValue="Perú">Peru</normalizedToken>
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, collected on 1 May 2010 by P. J. Venegas.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="71" type="paratypes">
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<paragraph pageId="2" pageNumber="71">Paratypes.</paragraph>
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<paragraph pageId="2" pageNumber="71">CORBIDI 8825, 8826, adult females collected on 30 October 2010 by P. J. Venegas; CORBIDI 08786, 08790, 08791, adult male, juvenile female, and juvenile male, respectively, collected on 21 January 2011 by P. J. Venegas and V. Duran; CORBIDI 09213, 09214, juvenile male and female, respectively, collected on 8 May 2011 by P. J. Venegas and V. Duran. All paratypes are from the type locality.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="72" pageId="2" pageNumber="71" type="diagnosis">
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<paragraph pageId="2" pageNumber="71">Diagnosis.</paragraph>
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<paragraph lastPageId="3" lastPageNumber="72" pageId="2" pageNumber="71">
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<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides azulae" order="Squamata" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="azulae">Enyalioides azulae</taxonomicName>
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can be distinguished from other species of
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<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides" order="Squamata" pageId="2" pageNumber="71" phylum="Chordata" rank="genus">Enyalioides</taxonomicName>
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, except
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<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides microlepis" order="Squamata" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="microlepis">Enyalioides microlepis</taxonomicName>
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and
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<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides cofanorum" order="Squamata" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="cofanorum">Enyalioides cofanorum</taxonomicName>
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, by the combination of the following characters: (1) strongly keeled ventral scales; (2) more than 37 longitudinal rows of dorsals in a transverse line between the dorsolateral crests at midbody; and (3) absence of superciliary flaps projecting over each orbit (present only in
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<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides palpebralis" order="Squamata" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="palpebralis">Enyalioides palpebralis</taxonomicName>
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).
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<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides azulae" order="Squamata" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="azulae">Enyalioides azulae</taxonomicName>
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differs from
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<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides cofanorum" order="Squamata" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="cofanorum">Enyalioides cofanorum</taxonomicName>
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and
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<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides microlepis" order="Squamata" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="microlepis">Enyalioides microlepis</taxonomicName>
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in having more gulars (45-57, mean = 51.13
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<normalizedToken originalValue="±">+/-</normalizedToken>
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4.05, versus 34-41, mean = 36.13
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<normalizedToken originalValue="±">+/-</normalizedToken>
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2.00 in
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<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides cofanorum" order="Squamata" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="cofanorum">Enyalioides cofanorum</taxonomicName>
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and 34-49, mean = 37.88
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<normalizedToken originalValue="±">+/-</normalizedToken>
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3.44 in
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<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides microlepis" order="Squamata" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="microlepis">Enyalioides microlepis</taxonomicName>
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), a smaller body size (maximum SVL = 96 mm in both males and females, versus 107 mm in males and 109 mm in females of
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<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides cofanorum" order="Squamata" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="cofanorum">Enyalioides cofanorum</taxonomicName>
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, and 127 mm in males and 116 mm in females of
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<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides microlepis" order="Squamata" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="microlepis">Enyalioides microlepis</taxonomicName>
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), a lower vertebral crest on the neck, a narrower snout in dorsal view, and in lacking blue on the gular region in males. Additionally,
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<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides azulae" order="Squamata" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="azulae">Enyalioides azulae</taxonomicName>
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has a marked sexual dichromatism, with males having greenish and females brownish background coloration (Fig. 2), whereas the other two species have brownish background coloration in both sexes.
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<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides azulae" order="Squamata" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="azulae">Enyalioides azulae</taxonomicName>
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further
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<pageBreakToken pageId="3" pageNumber="72" start="start">differs</pageBreakToken>
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from
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<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides cofanorum" order="Squamata" pageId="3" pageNumber="72" phylum="Chordata" rank="species" species="cofanorum">Enyalioides cofanorum</taxonomicName>
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in lacking scattered enlarged scales on the dorsum, well-developed dorsolateral crests between the hind limbs, and a dark gular patch in females.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="75" pageId="3" pageNumber="72" type="description">
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<paragraph pageId="3" pageNumber="72">Description of holotype.</paragraph>
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<paragraph pageId="3" pageNumber="72">
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Male (Fig. 1); SVL = 96 mm; TL = 140 mm; maximum head width = 21.28 mm; head length = 26.35 mm; head height = 17.95 mm; dorsal head scales uni- or multicarinate, those on parietal region projected dorsally; parietal eye present; scales immediately posterior to superciliares conical and as dorsally projected as adjacent parietals and temporals; temporal scales small, granular and multicarinate; one enlarged pretympanic scale; 14 superciliares; six canthals; five postrostrals; 11 (left or right) supralabials counted to a point below middle of eye; rostral (2.57
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<normalizedToken originalValue="×">x</normalizedToken>
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1.16 mm) about twice as wide as adjacent supralabials; two longitudinal rows of lorilabials between suboculars and supralabials at level of middle of eye, 3-4 longitudinal rows of lorilabials anterior to this point; loreal region broken into small, multicarinate, and juxtaposed scales; nasal at level of supralabials
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<normalizedToken originalValue="III–IV">III-IV</normalizedToken>
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; 10 (left or right) infralabials counted to a point right below middle of eye; mental (2.51
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<normalizedToken originalValue="×">x</normalizedToken>
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1.53 mm) wider and longer than adjacent infralabials; two postmentals; gulars ventrally projected; gular fold complete midventrally, extending dorsally and posteriorly to form antehumeral fold; neck with several oblique folds and a dorsolateral row of enlarged scales.
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</paragraph>
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<paragraph lastPageId="4" lastPageNumber="73" pageId="3" pageNumber="72">
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Vertebral crest not strongly projected, with vertebrals on neck similar in size to those between hind limbs; crest bifurcates posteriorly and extends onto tail less than
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<normalizedToken originalValue="¼">1/4</normalizedToken>
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its length; body flanks between fore and hind limbs without folds; irregular dorsolateral row of 1-2 keeled, enlarged scales (i.e., approximately twice as large as adjacent scales); dorsal scales between dorsolateral scale rows and vertebral crest small, keeled and subimbricate towards vertebral crest, granular towards dorsolateral scale rows;
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<pageBreakToken pageId="4" pageNumber="73" start="start">scales</pageBreakToken>
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on flanks similar in size to lateralmost dorsal scales; ventral scales subimbricate, keeled, subrectangular, with a posterolateral mucron; ventrals more than twice the length of dorsals.
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</paragraph>
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<paragraph lastPageId="5" lastPageNumber="74" pageId="4" pageNumber="73">
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Limb scales keeled and imbricate dorsally and ventrally; scales on dorsal and posterior aspects of thighs keeled and imbricate, with most scales less than half the size of those on anterior and ventral aspects; 19 subdigitals on manual digit
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<pageBreakToken pageId="5" pageNumber="74" start="start">IV</pageBreakToken>
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; 26 subdigitals on pedal digit IV; one femoral pore on each side; tail laterally compressed and gradually decreasing in relative height towards tip; caudal scales strongly keeled and imbricate, moderately increasing in size posteriorly on lateral and dorsal aspects of each autotomic segment; ventral caudals larger than dorsal caudals, with individual vertebral segments three scales long ventrally and six scales long dorsally.
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</paragraph>
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<paragraph lastPageId="6" lastPageNumber="75" pageId="5" pageNumber="74">
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Color in life of holotype (Fig. 1). Dorsal surface of head dark brown with light green flecks; lateral surface of head green with lorilabial and pretympanic regions turquoise and a black narrow supratemporal stripe; a black oblique stripe extending from
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<pageBreakToken pageId="6" pageNumber="75" start="start">eye</pageBreakToken>
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to commisure of the mouth; an orange cream oblique stripe on suboculars, posterior labials and adjacent gulars; labials cream; rostral and mental light green; wide, cream longitudinal stripe extending from above tympanum to scapular region; gular region dirty cream with dark spots and flecks; dark brown patch on medial aspect of gular fold; dorsal background green with diffuse, transverse dark brown bars on body, limbs, and tail; flanks covered with dark brown reticulations; ventral surface of body, limbs and tail tan with diffuse darker brown spots on thighs; iris reddish copper with a fine golden ring around the pupil.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="8" lastPageNumber="77" pageId="6" pageNumber="75" type="intraspecific variation">
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<paragraph pageId="6" pageNumber="75">Intraspecific variation.</paragraph>
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<paragraph pageId="6" pageNumber="75">
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Meristic and morphometric characters of
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<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides azulae" order="Squamata" pageId="6" pageNumber="75" phylum="Chordata" rank="species" species="azulae">Enyalioides azulae</taxonomicName>
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are summarized in Table 1. Male paratypes (CORBIDI 08786, 09213) are very similar in coloration to the holotype (Fig. 3
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<normalizedToken originalValue="A–D">A-D</normalizedToken>
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). The dark patch on the gular region of adult males is also present in juvenile male specimens.
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</paragraph>
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<paragraph lastPageId="7" lastPageNumber="76" pageId="6" pageNumber="75">
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Adult (CORBIDI 08825-08826) and juvenile (CORBIDI 08791, 09214) females share similar color patterns (Figs. 3
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<normalizedToken originalValue="E–F">E-F</normalizedToken>
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, 4): head brown with a narrow dark brown supratemporal stripe; broad subocular dark stripe extending from eye to commisure of mouth, with a parallel, conspicuous white or cream stripe immediately anterior to it; pale, wide longitudinal stripe extending from tympanum to scapular region; gular region pale brown without dark markings, or white with faint reddish brown reticulation as in specimen CORBIDI 08826; dorsal background light brown, with a greenish tone in CORBIDI 08826 (Fig. 3
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<normalizedToken originalValue="E–F">E-F</normalizedToken>
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) and coppery tone in CORBIDI 09214
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<pageBreakToken pageId="7" pageNumber="76" start="start">(</pageBreakToken>
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Fig. 4E); transverse dark brown bars on dorsal aspect of body, limbs, and tail; ventral surface of body, limbs and tail light brown (CORBIDI 08825; Fig. 4B) or white (CORBIDI 08826; Fig. 4F); iris reddish brown.
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</paragraph>
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<paragraph pageId="8" pageNumber="77">
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<pageBreakToken pageId="8" pageNumber="77" start="start">Although</pageBreakToken>
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this species seems to have a marked sexual dichromatism in background colors (green in males, brown in females, see Fig. 2), one male specimen (the holotype) exhibited metachromatism consisting of dark brown tones being replaced by green tones.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="77" type="distribution">
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<paragraph pageId="8" pageNumber="77">Distribution and natural history.</paragraph>
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<paragraph pageId="8" pageNumber="77">
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<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides azulae" order="Squamata" pageId="8" pageNumber="77" phylum="Chordata" rank="species" species="azulae">Enyalioides azulae</taxonomicName>
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is known only from its type locality in the montane rainforest of the
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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Huallaga basin (Fig. 5) in northeastern Peru at an elevation of 1100 m. This locality lies within the CAZNP, on a mountain ridge between the
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<normalizedToken originalValue="Región">Region</normalizedToken>
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San
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<normalizedToken originalValue="Martín">Martin</normalizedToken>
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and
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<normalizedToken originalValue="Región">Region</normalizedToken>
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Loreto (Fig. 6). Seven of the eight individuals of
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<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides azulae" order="Squamata" pageId="8" pageNumber="77" phylum="Chordata" rank="species" species="azulae">Enyalioides azulae</taxonomicName>
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reported in this paper were collected at night sleeping on low vertical stems of bushes 15-80 cm above the ground. One adult male (the holotype) was collected during the day on a narrow trail after a rain; when approached, it fled and hid under a fallen log. This species is found in sympatry and possibly syntopy with
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<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides binzayedi" order="Squamata" pageId="8" pageNumber="77" phylum="Chordata" rank="species" species="binzayedi">Enyalioides binzayedi</taxonomicName>
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sp. n. (see below) and
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<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides laticeps" order="Squamata" pageId="8" pageNumber="77" phylum="Chordata" rank="species" species="laticeps">Enyalioides laticeps</taxonomicName>
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. The smallest individuals (CORBIDI 08790-08791, SVL = 61 and 62 mm, respectively) were collected in January. Other species of squamate reptiles collected at the same locality include
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<taxonomicName class="Reptilia" family="Gymnophthalmidae" genus="Alopoglossus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alopoglossus angulatus" order="Squamata" pageId="8" pageNumber="77" phylum="Chordata" rank="species" species="angulatus">Alopoglossus angulatus</taxonomicName>
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,
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<taxonomicName class="Reptilia" family="Dactyloidae" genus="Anolis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anolis fuscoauratus" order="Squamata" pageId="8" pageNumber="77" phylum="Chordata" rank="species" species="fuscoauratus">Anolis fuscoauratus</taxonomicName>
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,
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<taxonomicName class="Reptilia" family="Dactyloidae" genus="Anolis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anolis transversalis" order="Squamata" pageId="8" pageNumber="77" phylum="Chordata" rank="species" species="transversalis">Anolis transversalis</taxonomicName>
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,
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<taxonomicName class="Reptilia" family="Gymnophthalmidae" genus="Cercosaura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercosaura manicata" order="Squamata" pageId="8" pageNumber="77" phylum="Chordata" rank="species" species="manicata">Cercosaura manicata</taxonomicName>
|
||
,
|
||
<taxonomicName class="Reptilia" family="Gymnophthalmidae" genus="Potamites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamites ecpleopus" order="Squamata" pageId="8" pageNumber="77" phylum="Chordata" rank="species" species="ecpleopus">Potamites ecpleopus</taxonomicName>
|
||
,
|
||
<taxonomicName class="Reptilia" family="Gymnophthalmidae" genus="Potamites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamites strangulatus" order="Squamata" pageId="8" pageNumber="77" phylum="Chordata" rank="species" species="strangulatus">Potamites strangulatus</taxonomicName>
|
||
,
|
||
<taxonomicName class="Reptilia" family="Gymnophthalmidae" genus="Potamites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamites" order="Squamata" pageId="8" pageNumber="77" phylum="Chordata" rank="genus">Potamites</taxonomicName>
|
||
sp.,
|
||
<taxonomicName class="Reptilia" family="Colubridae" genus="Chironius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chironius fuscus" order="Squamata" pageId="8" pageNumber="77" phylum="Chordata" rank="species" species="fuscus">Chironius fuscus</taxonomicName>
|
||
,
|
||
<taxonomicName class="Reptilia" family="Dipsadidae" genus="Dipsas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dipsas indica" order="Squamata" pageId="8" pageNumber="77" phylum="Chordata" rank="species" species="indica">Dipsas indica</taxonomicName>
|
||
,
|
||
<taxonomicName class="Reptilia" family="Dipsadidae" genus="Imantodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imantodes cenchoa" order="Squamata" pageId="8" pageNumber="77" phylum="Chordata" rank="species" species="cenchoa">Imantodes cenchoa</taxonomicName>
|
||
,
|
||
<taxonomicName class="Reptilia" family="Dipsadidae" genus="Imantodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imantodes lentiferus" order="Squamata" pageId="8" pageNumber="77" phylum="Chordata" rank="species" species="lentiferus">Imantodes lentiferus</taxonomicName>
|
||
,
|
||
<taxonomicName class="Reptilia" family="Elapidae" genus="Micrurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Micrurus obscurus" order="Squamata" pageId="8" pageNumber="77" phylum="Chordata" rank="species" species="obscurus">Micrurus obscurus</taxonomicName>
|
||
,
|
||
<taxonomicName class="Reptilia" family="Dipsadidae" genus="Oxyrhopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxyrhopus petola" order="Squamata" pageId="8" pageNumber="77" phylum="Chordata" rank="species" species="petola">Oxyrhopus petola</taxonomicName>
|
||
, and
|
||
<taxonomicName class="Reptilia" family="Dipsadidae" genus="Xenopholis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenopholis scalaris" order="Squamata" pageId="8" pageNumber="77" phylum="Chordata" rank="species" species="scalaris">Xenopholis scalaris</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="77" type="etymology">
|
||
<paragraph pageId="8" pageNumber="77">Etymology.</paragraph>
|
||
<paragraph pageId="8" pageNumber="77">The specific epithet is a noun derived from the Spanish word "azul" (blue) in the genitive case; it refers to the Cordillera Azul, the mountain range after which the National Park where this species was discovered is named. Although the word "azul" in "Cordillera Azul" is an adjective, and the Spanish noun "azul" is masculine, we are here treating "azulae" as a feminine noun that is an abbreviation for "Cordillera Azul" and is therefore to be interpreted as meaning "of the [Cordillera] Azul."</paragraph>
|
||
<caption pageId="8" pageNumber="77">
|
||
<paragraph pageId="8" pageNumber="77">
|
||
Figure 1. Holotype of
|
||
<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides azulae" order="Squamata" pageId="8" pageNumber="77" phylum="Chordata" rank="species" species="azulae">Enyalioides azulae</taxonomicName>
|
||
sp. n. (CORBIDI 06772, adult male, SVL = 96 mm). Top: lateral view; middle: close-up of head; bottom: ventral view. Photographs by P.J. Venegas.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="8" pageNumber="77">
|
||
<paragraph pageId="8" pageNumber="77">
|
||
Figure 2. Male (top, CORBIDI 09213) and female (bottom, CORBIDI 09214) of
|
||
<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides azulae" order="Squamata" pageId="8" pageNumber="77" phylum="Chordata" rank="species" species="azulae">Enyalioides azulae</taxonomicName>
|
||
sp. n. Photograph by P.J. Venegas.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="8" pageNumber="77">
|
||
<paragraph pageId="8" pageNumber="77">
|
||
Figure 3. Adult male (A, B, C CORBIDI 08786), juvenile male (D CORBIDI 09213), and adult female (E, F CORBIDI 08826) of
|
||
<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides azulae" order="Squamata" pageId="8" pageNumber="77" phylum="Chordata" rank="species" species="azulae">Enyalioides azulae</taxonomicName>
|
||
. Photographs by P.J. Venegas.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="8" pageNumber="77">
|
||
<paragraph pageId="8" pageNumber="77">
|
||
Figure 4. Adult (A, B, C CORBIDI 08825) and juvenile (D CORBIDI 08791 E, F CORBIDI 09214) females of
|
||
<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Enyalioides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enyalioides azulae" order="Squamata" pageId="8" pageNumber="77" phylum="Chordata" rank="species" species="azulae">Enyalioides azulae</taxonomicName>
|
||
. Photographs by P.J. Venegas.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |