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<document id="6616AD129AF31EC890FDDFA35FE8D01E" ID-DOI="10.3897/zitteliana.98.e138605" ID-ZooBank="9FB6F185-9EA1-48EB-A5C9-632BD52A8B6F" ID-publisher-id="138605" URI-arpha="88AF8F00-FD88-5C61-809C-0C929B9B2F5A" URI-zoobank="http://zoobank.org/9FB6F185-9EA1-48EB-A5C9-632BD52A8B6F" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1733930160955" checkinUser="pensoft" docAuthor="Gründel, Joachim &amp; Nützel, Alexander" docDate="2024" docId="54F018CA13545B3CAC84EAD61F0C1E08" docLanguage="en" docName="Zitteliana 98: 19-72" docOrigin="Zitteliana 98" docSource="https://zitteliana.pensoft.net/article/138605/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Pseudorhytidopilus quadratus Gründel &amp; Nützel, 2024, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="72" masterDocId="88AF8F00FD885C61809C0C929B9B2F5A" masterDocTitle="Late Jurassic (Upper Kimmeridgian) gastropods from Saal near Kelheim (Germany, Bavaria)" masterLastPageNumber="72" masterPageNumber="19" pageNumber="19" updateTime="1733940029114" updateUser="admin">
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<mods:title id="98FE68DF30493EE90AFFDFE7EB978649">Late Jurassic (Upper Kimmeridgian) gastropods from Saal near Kelheim (Germany, Bavaria)</mods:title>
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<mods:name id="332A822036F06CC4F4BDCD819E90CFED" type="personal">
<mods:role id="2FA531E21CA25DD0F05FE98483B22558">
<mods:roleTerm id="742492DDD686FC9EBA021A39B7805510">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7F3C006396D00919A145B1666185B2BD">Gründel, Joachim</mods:namePart>
<mods:affiliation id="6570FE52F117AAC67D3B5C3834B3EC7C">Institut für Geowissenschaften, Fachrichtung Paläontologie, Freie Universität Berlin, Malteserstraße 74 100, 12249 Berlin, Germany</mods:affiliation>
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</mods:role>
<mods:namePart id="B903C0974601C7EB564FA02DDAAC701D">Nützel, Alexander</mods:namePart>
<mods:nameIdentifier id="43A96A191CA3B3D436E87B24ACA7AB66" type="ORCID">0000-0002-8852-7688</mods:nameIdentifier>
<mods:affiliation id="FC3F1869DB401C2DCC12D3BC82D7DED8">SNSB-Bayerische Staatssammlung für Paläontologie und Geologie, Richard-Wagner-Str. 10, 80333 München, Germany &amp; Department of Earth and Environmental Sciences, Paleontology and Geobiology, Ludwig-Maximilians-Universität München, Richard-Wagner-Str. 10, 80333 München, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource id="760CF567FDF3944CF66067672A6FD334">text</mods:typeOfResource>
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<mods:title id="6F24061703A9F4C6BF9635F640DBBDF5">Zitteliana</mods:title>
</mods:titleInfo>
<mods:part id="790BAE931C3EBE8247F8E2468CEA78E3">
<mods:date id="762CF93B7416078C8DE22B8926821C4D">2024</mods:date>
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<mods:number id="30EB154B93A6930449EF6BF38EB81A8D">2024-12-11</mods:number>
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<mods:number id="D1C2257ABD9C4049E9885ADD1F28C13D">98</mods:number>
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<mods:classification id="B87D76CEAE92738F1F23F2CA446B3F23">journal article</mods:classification>
<mods:identifier id="5B40B33DF11947E3DB683196E50F8B16" type="DOI">10.3897/zitteliana.98.e138605</mods:identifier>
<mods:identifier id="BC29E2D128551E543EEB46D890F95B7D" type="ZooBank">9FB6F185-9EA1-48EB-A5C9-632BD52A8B6F</mods:identifier>
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<treatment id="54F018CA13545B3CAC84EAD61F0C1E08" ID-arpha="54F018CA-1354-5B3C-AC84-EAD61F0C1E08" ID-zoobank="https://zoobank.org/B6959D20-CF2C-4189-A9AA-DCB15B3AAF2B" LSID="urn:lsid:plazi:treatment:54F018CA13545B3CAC84EAD61F0C1E08" LSID-ZBK="urn:lsid:zoobank.org:act:B6959D20-CF2C-4189-A9AA-DCB15B3AAF2B" httpUri="http://treatment.plazi.org/id/54F018CA13545B3CAC84EAD61F0C1E08" scope_family="Acmaeidae" scope_subClass="Patellogastropoda" scope_superFamily="Lottioidea">
<subSubSection id="77C7F3C56173DE53ACDB8FF85FF8C87D" type="nomenclature">
<paragraph id="FAA76A30F4FDD4C78D65610D2808ABB0">
<taxonomicName id="0CEFE9570678C2A0213BC7A578D6CA09" authorityName="Gründel &amp; Nützel" authorityYear="2024" class="Gastropoda" family="Acmaeidae" genus="Pseudorhytidopilus" kingdom="Animalia" phylum="Mollusca" rank="species" species="quadratus" status="sp. nov." subClass="Patellogastropoda">
<emphasis id="A3C75E9EEF23FCB73686749B228124DA" italics="true">Pseudorhytidopilus</emphasis>
?
<emphasis id="79317AFB59EF3952C9524539AD385227" italics="true">quadratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="749D4AAEBEFD4C590ED0CCA35102A2C6" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="21654D311EC6C0D32DA05AEDFB78BC07" type="description">
<paragraph id="9A484C0EEF602E904CA21BD4F6D4FA22">
<figureCitation id="2934F48E07C9D057514C30DD898DDC73" captionStart="Plate 1" captionStartId="F1" captionText="Plate 1. (1) Patella sp., SNSB BSPG 2016 XXI 1601, collection O. Neubauer, dorsal view, length 7 mm. (24) Scurriopsis cragolis sp. nov. holotype, SNSB BSPG 2016 XXI 1602, (23) lateral and dorsal views, length 16 mm, (4) detail of ornament, height ca. 5.8 mm. (58) Scurriopsis sp., SNSB BSPG 2016 XXI 1606, (5) dorsal view, length 8 mm, (6) detail in dorsal view, width 3.5 mm, (78) lateral view and detail of ornament, height 6 mm. (910) Pseudorhytidopilus? quadratus sp. nov., holotype, SNSB BSPG 2016 XXI 1607, dorsal and lateral views, largest diameter 21 mm. (11) Pseudorhytidopilus? quadratus sp. nov., paratype, SNSB BSPG 2016 XXI 1608, dorsal view, largest diameter 16 mm. (12) Leptomaria goldfussi (Sieberer, 1907), SNSB BSPG 2016 XXI 1616, lateral view, height 25 mm. (13, 14) Leptomaria goldfussi (Sieberer, 1907), SNSB BSPG 2016 XXI 1617, early whorls and detail of ornament, (13) height 16.5 mm, (14) height 14 mm. (15, 16) Leptomaria goldfussi (Sieberer, 1907), SNSB BSPG 2016 XXI 1618, aperture and base, width 24 mm." figureDoi="10.3897/zitteliana.98.e138605.plate1" httpUri="https://binary.pensoft.net/fig/1196357">Plate 1: figs 911</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECPAE" type="etymology">
<paragraph id="C4DD12743F847D46349B5576045B10C9">
<heading id="46B31A9F59CACBDB4EA03466C80AACA0" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="1FE65CFB18CB7AF610EA0BC405201714">
Latin
<tp_taxon-name id="E5158075E6B31AC1B8A481E5A968E208">
<emphasis id="374DC150EE1AC7F4539D0E8DF65A735B" italics="true">
<tp_taxon-name-part id="2787D35A941BD978A9B900290B82A9A5" reg="quadratus" taxon-name-part-type="species">quadratus</tp_taxon-name-part>
</emphasis>
</tp_taxon-name>
due to the quadratic outline of the shell.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERPAE" type="Holotype">
<paragraph id="7D9E1595E00C1C9028884C7984075D48">
<heading id="21C3A48F090ED06B5A9FCD2D957133AE" reason="title">
<typeStatus id="0682AD09F2AD4BF2E903F73BC7956812" type="holotype">Holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="02CC19A16FFD5253DA0E4C5420E0E360">
<materialsCitation id="43C874DA25EA36683D53495BA5609345" collectionCode="SNSB" specimenCode="SNSB - BSPG 2016 XXI 1607" specimenCount="1" typeStatus="holotype">
<specimenCode id="F73505020141BECB19C570D1A9082FF5">SNSBBSPG 2016 XXI 1607</specimenCode>
(Plate
<figureCitation id="3FCF29E26A5C64914E4FD85A097E3F8E" captionStart="Plate 1" captionStartId="F1" captionText="Plate 1. (1) Patella sp., SNSB BSPG 2016 XXI 1601, collection O. Neubauer, dorsal view, length 7 mm. (24) Scurriopsis cragolis sp. nov. holotype, SNSB BSPG 2016 XXI 1602, (23) lateral and dorsal views, length 16 mm, (4) detail of ornament, height ca. 5.8 mm. (58) Scurriopsis sp., SNSB BSPG 2016 XXI 1606, (5) dorsal view, length 8 mm, (6) detail in dorsal view, width 3.5 mm, (78) lateral view and detail of ornament, height 6 mm. (910) Pseudorhytidopilus? quadratus sp. nov., holotype, SNSB BSPG 2016 XXI 1607, dorsal and lateral views, largest diameter 21 mm. (11) Pseudorhytidopilus? quadratus sp. nov., paratype, SNSB BSPG 2016 XXI 1608, dorsal view, largest diameter 16 mm. (12) Leptomaria goldfussi (Sieberer, 1907), SNSB BSPG 2016 XXI 1616, lateral view, height 25 mm. (13, 14) Leptomaria goldfussi (Sieberer, 1907), SNSB BSPG 2016 XXI 1617, early whorls and detail of ornament, (13) height 16.5 mm, (14) height 14 mm. (15, 16) Leptomaria goldfussi (Sieberer, 1907), SNSB BSPG 2016 XXI 1618, aperture and base, width 24 mm." figureDoi="10.3897/zitteliana.98.e138605.plate1" httpUri="https://binary.pensoft.net/fig/1196357">1</figureCitation>
: figs 910).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2PAE" type="Paratypes">
<paragraph id="60EA13960FE0280F011173372EF13DF9">
<heading id="EEE4C1402D9416E4A37B7F59AC4E6AEE" reason="title">
<typeStatus id="730A997941D8C261C7A8B4A83A011397" type="paratype">Paratypes</typeStatus>
.
</heading>
</paragraph>
<paragraph id="4AED1B4178EABB52338ADD55DEC46CD2">
<materialsCitation id="A88BC0F2D3617806AD5B528290407E0A" collectionCode="SNSB" specimenCode="SNSB-BSPG 2016 XXI 1608" specimenCount="6" typeStatus="holotype">
<specimenCount id="F8195FFD0387D5135B8C621B6B87F8FF" count="6" type="generic">6 specimens</specimenCount>
<specimenCode id="331E98BCA52421608150E8538E3E7D09">SNSBBSPG 2016 XXI 1608</specimenCode>
1613.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EAQAE" type="diagnosis">
<paragraph id="81F960888AD6A46B48691E0F34311B22">
<heading id="BAA83EC3A0C892B78B209177FE7F5641" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="DB561678C9F3448FD1071C59BA42BB09">Shell thin, with trapezoidal to subquadratic outline with rounded edges in dorsal view; apex approximately in central position; shell ornamented with irregular concentric bulges.</paragraph>
</subSubSection>
<subSubSection id="SECID0EFQAE" type="type locality">
<paragraph id="EBE571951C6A9C9BE8BC384888E64162">
<heading id="2DEA0CF2BC183D07C0A9400036AD03C0" reason="title">Type locality and stratum.</heading>
</paragraph>
<paragraph id="318DC67E11AAD35AABD6AC4E21257585">
Upper Jurassic (Kimmeridgian) reefal limestones from the locality Saal near Kelheim, Lower
<collectingRegion id="C0C72DFFAA354BAA0F21E35F3803583D" country="Germany" name="Bayern">Bavaria</collectingRegion>
(
<bibRefCitation id="2B8E662F129B830D977D8B99EEB42C24" author="Grundel" etAl="et al." firstAuthor="Grundel" journalOrPublisher="Zitteliana A" pagination="77-106" refId="B84" refString="Gründel J, Keupp H, Lang F (2015) Die Arten der Unterklasse Neritimorpha Koken, 1896 (Gastropoda) aus der Korallenfazies des oberen Kimmeridgiums (oberer Jura) von Saal bei Kelheim und dem Gebiet Nattheim (Süddeutschland). Zitteliana A 55: 77106." title="Die Arten der Unterklasse Neritimorpha Koken, 1896 (Gastropoda) aus der Korallenfazies des oberen Kimmeridgiums (oberer Jura) von Saal bei Kelheim und dem Gebiet Nattheim (Süddeutschland)." volume="55" year="2015">Gründel et al. 2015</bibRefCitation>
,
<bibRefCitation id="E5BAD9E0F89AAFDC3542A8C5BDAF0EB7" author="Grundel" firstAuthor="Grundel" journalOrPublisher="Revue de Paléobiologie" pagination="29-84" refId="B83" refString="Gründel J, Hostettler B, Menkveld-Gfeller U (2022) Die Gastropoden aus der Korallenrifffazies der St-Ursanne-Formation (mittleres Oxfordien) des Schweizer Jura. 3. Die Unterklasse Caenogastropoda Cox, 1960. Revue de Paléobiologie 40: 2984." title="Die Gastropoden aus der Korallenrifffazies der St-Ursanne-Formation (mittleres Oxfordien) des Schweizer Jura. 3. Die Unterklasse Caenogastropoda Cox, 1960." volume="40" year="2022">2022</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETQAE" type="description">
<paragraph id="E1E53981C2E9D0CADF88E49CA4E11C66">
<heading id="18EEB9A99E4AA26A76DDCB6D2835957B" reason="title">Description.</heading>
</paragraph>
<paragraph id="E4921AE069D592209C1B0A9014E69062">
Large specimen has a diameter of
<quantity id="A66F264489FC81D15759AC81B176F764" metricMagnitude="-2" metricUnit="m" metricValue="2.1" unit="mm" value="21.0">21 mm</quantity>
; shell very thin, shallowly cap-shaped in lateral view; trapezoidal (one side of shell slightly narrower than the other side) to subquadratic outline with rounded edges in dorsal view; apex approximately in central position; shell ornamented with irregular concentric bulges.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYQAE" type="remarks">
<paragraph id="B7A5E8C9C77D216AC50C9C8ADB197E90">
<heading id="A0F1F6A8D557AEFC0C637E343303995E" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="911AB09FCABEF737029181DAB2FC2A82">
Species with a similar ornament of bulges (e. g.,
<emphasis id="427D7D1F2DF8C6C132E0118B1F643680" italics="true">P.</emphasis>
?
<emphasis id="1C3E89F19697817115D057E62704A09E" italics="true">ledonii</emphasis>
(
<bibRefCitation id="E02A07A8DD7EE55CEBA0BBE2561DAE30" author="Haber G" journalOrPublisher="Junk, Berlin" refId="B90" refString="Haber G (1932) Gastropoda, Amphineura et Scaphopoda jurassica. In: W. Quenstedt (ed.): Fossilium Catalogus. I, Animalia; pars 53. Junk, Berlin, 304 pp." title="Gastropoda, Amphineura et Scaphopoda jurassica. In: W. Quenstedt (ed.): Fossilium Catalogus. I, Animalia; pars 53." year="1932">Haber, 1932</bibRefCitation>
) sensu
<bibRefCitation id="A5B46C5E02676722FD3DF74DEDC78980" DOI="10.1111/j.1475-4983.2010.00961.x" author="Gatto and Monari" firstAuthor="Gatto" journalOrPublisher="Palaeontology" pagination="771-802" refId="B60" refString="Gatto R, Monari S (2010) Pliensbachian gastropods from Venetian Southern Alps (Italy) and their palaeobiogeographical significance. Palaeontology 53: 771802. https://doi.org/10.1111/j.1475-4983.2010.00961.x" title="Pliensbachian gastropods from Venetian Southern Alps (Italy) and their palaeobiogeographical significance." volume="53" year="2010">Gatto and Monari 2010</bibRefCitation>
,
<taxonomicName id="6C485062EFBE4F2E1893D7B554CBE4DB" authority="Monari et al. 2017" authorityName="Monari" authorityYear="2017" class="Gastropoda" family="Acmaeidae" genus="Fabercapulus" kingdom="Animalia" phylum="Mollusca" rank="species" species="semisculptus">
<emphasis id="6D4AD631B2A0ADFD56C7286D7C444D26" italics="true">Fabercapulus semisculptus</emphasis>
<bibRefCitation id="58A03351616889F56C079D9263EAA8D9" DOI="10.1080/14772019.2017.1312578" author="Monari" etAl="et al." firstAuthor="Monari" issue="6" journalOrPublisher="Journal of Systematic Palaeontology" pagination="449-492" refId="B125" refString="Monari S, Gatto R, Valentini M (2017) Vetigastropoda and Neritimorpha from the Lower Bajocian of Luxembourg and palaeobiogeography of Aalenian-Bajocian (Middle Jurassic) Gastropoda of western Europe. Journal of Systematic Palaeontology 16 (6): 449492. https://doi.org/10.1080/14772019.2017.1312578" title="Vetigastropoda and Neritimorpha from the Lower Bajocian of Luxembourg and palaeobiogeography of Aalenian-Bajocian (Middle Jurassic) Gastropoda of western Europe." volume="16" year="2017">Monari et al. 2017</bibRefCitation>
</taxonomicName>
) have an elliptic outline in dorsal view and a radial furrow. Other species with an oval outline in dorsal view lack a furrow and an ornament of bulges (
<taxonomicName id="E265651317115CCEA766F51999AEF98A" authority="(Sowerby, 1816) sensu Munt" authorityName="Munt" baseAuthorityName="Sowerby" baseAuthorityYear="1816" class="Gastropoda" family="Acmaeidae" genus="Pseudorhytidopilus" kingdom="Animalia" phylum="Mollusca" rank="species" species="latissima">
<emphasis id="BA6232025F04D9DD41E85079B9F01A38" italics="true">P. latissima</emphasis>
(Sowerby, 1816) sensu Munt
</taxonomicName>
in
<bibRefCitation id="B9D924F70E763D5EDFBBDF1D04A9D13A" author="Martill and Etches" firstAuthor="Martill" refId="B122" refString="Martill DM, Etches S (2020) Fossils of the Kimmeridge Clay Formation (Volume 1). Palaeontological Association Field Guide to Fossils 16, 436 pp." year="2020">Martill and Etches 2020</bibRefCitation>
,
<taxonomicName id="1BCD452C7D6F673953754210F66C6EC0" authority="(Eichwald) sensu Gerasimov, 1992" authorityName="sensu Gerasimov" authorityYear="1992" baseAuthorityName="Eichwald" baseAuthorityYear="1840" class="Gastropoda" family="Patellidae" genus="Berlieria" kingdom="Animalia" phylum="Mollusca" rank="species" species="maeotis">
<emphasis id="DFE6BEF94B78C0DF9F6E6F988A649E59" italics="true">Berlieria maeotis</emphasis>
(Eichwald) sensu
<bibRefCitation id="265945E2DB8C5C7148031EB24C59616A" author="Gerasimov PA" refId="B64" refString="Gerasimov PA (1992) Gastropody jurskich i progranichnych nizhnemelovych otlozenij Evropejskoj Rossii. Moskau (Rossiyskaja Akademija Nauk), 190 pp." year="1992">Gerasimov, 1992</bibRefCitation>
</taxonomicName>
and others).
</paragraph>
<paragraph id="9F704EC8A8FA85355BEDE053FC4EBC91">
<taxonomicName id="CD0B1A85C365672CD03ED86B997261F4" authority="(Yokoyama, 1890)" baseAuthorityName="Yokoyama" baseAuthorityYear="1890" class="Magnoliopsida" family="Goodeniaceae" genus="Brunonia" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="annulata">
<emphasis id="07763FAA753677D7C60FF9AE286B701B" italics="true">Brunonia annulata</emphasis>
(Yokoyama, 1890)
</taxonomicName>
from the Cretaceous of
<collectingCountry id="726D2B9F4D20FB062962ED26831268C8" name="Japan">Japan</collectingCountry>
is also cap-shaped and has a similar ornament but its
<typeStatus id="144976CA86A0A614A37B7A45E56182B9">type</typeStatus>
species is much larger and has a rounded, oval outline in dorsal view (
<bibRefCitation id="5E62122ECB5D563E5A7148D934143D70" author="Kase" firstAuthor="Kase" journalOrPublisher="Journal of Paleontology" pagination="766-771" refId="B100" refString="Kase T (1988) Reinterpretation of Brunonia annulata (Yokoyama) as an Early Cretaceous carinariid mesogastropod (Mollusca). Journal of Paleontology 62: 766771." title="Reinterpretation of Brunonia annulata (Yokoyama) as an Early Cretaceous carinariid mesogastropod (Mollusca)." volume="62" year="1988">Kase 1988</bibRefCitation>
;
<bibRefCitation id="C78ACE35679849E10CD524830D601822" author="Dieni" firstAuthor="Dieni" refId="B41" refString="Dieni I (1990) Brunonia annulata (Yokoyama, 1890) (Carinariidae, Mesogastropoda)" year="1990">Dieni 1990</bibRefCitation>
).
<taxonomicName id="CC0EDCB4EAD260AC4E8391CF90F6D67C" authority="Zittel, 1873" authorityName="Zittel" authorityYear="1873" class="Gastropoda" family="Patellidae" genus="Patella" kingdom="Animalia" order="Aplysiida" phylum="Mollusca" rank="species" species="sculptilis" subGenus="Helcion">
<emphasis id="41C51DCF7A79BF72A692D031E022C52E" italics="true">Patella</emphasis>
(
<emphasis id="F47B96A758C81EC2168F9BF089C95104" italics="true">Helcion</emphasis>
)
<emphasis id="E3EB8D0853C7DB20D49DC330870A7708" italics="true">sculptilis</emphasis>
<bibRefCitation id="EFA3E9BE7338261BD29E96A3BE418B4F" author="Zittel K" refId="B183" refString="Zittel K (1873) Die Gastropoden der Stramberger Schichten. Palaeontographica, Supplement 2 / 1: 194 - 373." year="1873">Zittel, 1873</bibRefCitation>
</taxonomicName>
has a straight shell end in lateral view, weak concentric wrinkles and numerous radial ribs.
</paragraph>
</subSubSection>
</treatment>
</document>