treatments-xml/data/54/BF/FD/54BFFD0035125A8E8C223856B09A20C9.xml

<document id="1668ADC309E32F95F203FF2B69F2AA63" ID-CLB-Dataset="307126" ID-DOI="10.3897/jhr.98.136200" ID-GBIF-Dataset="2f30ca71-e2ba-4645-9a2e-4f8c42de60c0" ID-ZooBank="77736D1C-30F9-4D6C-8310-3A04121CACE2" ID-publisher-id="136200" URI-arpha="CFF2E897-2633-5F4E-8369-35D589D56708" URI-zoobank="http://zoobank.org/77736D1C-30F9-4D6C-8310-3A04121CACE2" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1735917671054" checkinUser="pensoft" docAuthor="Wang, Zhen, Rasnitsyn, Alexandr P., Perkovsky, Evgeny E., Vilhelmsen, Lars &amp; Gao, Taiping" docDate="2025" docId="54BFFD0035125A8E8C223856B09A20C9" docLanguage="en" docName="Journal of Hymenoptera Research 98: 1-18" docOrigin="Journal of Hymenoptera Research 98" docSource="https://jhr.pensoft.net/article/136200/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Siccibythus robustus Wang, Rasnitsyn, Perkovsky &amp; Vilhelmsen 2025, sp. nov." docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="18" masterDocId="CFF2E89726335F4E836935D589D56708" masterDocTitle="New wasps of † Falsiformicidae from mid-Cretaceous Kachin amber" masterLastPageNumber="18" masterPageNumber="1" pageNumber="1" updateTime="1735918735285" updateUser="ExternalLinkService">
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<mods:title id="60E5FB9BBA9FC75A16A28FB5840BFA5E">New wasps of † Falsiformicidae from mid-Cretaceous Kachin amber</mods:title>
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<mods:namePart id="71CBCF86798BB11A2B5000A65AFF3F98">Wang, Zhen</mods:namePart>
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<mods:affiliation id="BC08B4E77B7B3C750317DBC2F96EB230">College of Life Sciences, Capital Normal University, Beijing, 100048, China</mods:affiliation>
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<mods:namePart id="4E16BE07A06115FF64B0105823DF3A64">Rasnitsyn, Alexandr P.</mods:namePart>
<mods:affiliation id="78BCEAE113AF3C5B6E45A423F9B283DC">A. A. Borissiak Palaeontological Institute, Russian Academy of Sciences, Moscow, 117647, Russia</mods:affiliation>
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<mods:namePart id="26A58CACB75666C28ED8688F0980A3B6">Perkovsky, Evgeny E.</mods:namePart>
<mods:nameIdentifier id="F7F5550C4C5464767D168BB68FB03986" type="ORCID">0000-0002-7959-4379</mods:nameIdentifier>
<mods:affiliation id="975B4BD51B3A139A33064B534DCA40AC">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Copenhagen, 2100, Denmark</mods:affiliation>
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<mods:namePart id="C8C90D5C0B4967889B10732925DC677B">Vilhelmsen, Lars</mods:namePart>
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<mods:affiliation id="B446133A981C3217F2BBE27A554828E5">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Copenhagen, 2100, Denmark</mods:affiliation>
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<mods:namePart id="B42D8FA5D11D5DC443EDDC94764A5F07">Gao, Taiping</mods:namePart>
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<mods:affiliation id="ED0528CD0E0004DA7C11777FB91F881D">College of Life Sciences, Capital Normal University, Beijing, 100048, China</mods:affiliation>
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<mods:date id="E1CA4C4E4E73B727369D7D1A8387D899">2025</mods:date>
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<mods:number id="56EE82F42A6D338F0719D8317DFCD04B">2025-01-03</mods:number>
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<mods:number id="D23F6455E1EA04377A006E961E971CB2">98</mods:number>
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<treatment id="54BFFD0035125A8E8C223856B09A20C9" ID-arpha="54BFFD00-3512-5A8E-8C22-3856B09A20C9" ID-zoobank="https://zoobank.org/6F39958F-FB10-4AB2-9752-9AAB01FECDE8" LSID="urn:lsid:plazi:treatment:54BFFD0035125A8E8C223856B09A20C9" LSID-ZBK="urn:lsid:zoobank.org:act:6F39958F-FB10-4AB2-9752-9AAB01FECDE8" httpUri="http://treatment.plazi.org/id/54BFFD0035125A8E8C223856B09A20C9" scope_family="Falsiformicidae" scope_order="Hymenoptera">
<subSubSection id="C4020A10D0A6A399906B71E90D905983" type="nomenclature">
<paragraph id="E43DCE70B61D9382A9FD74079DCA11CE">
<taxonomicName id="98E9670D0E28D688FD65BFB3D130C522" authority="Wang, Rasnitsyn, Perkovsky &amp; Vilhelmsen" authorityName="Wang, Rasnitsyn, Perkovsky &amp; Vilhelmsen" authorityYear="2025" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="robustus" status="sp. nov.">
<emphasis id="A93B2FEEBB724234BF0F665FB771FC30" italics="true">Siccibythus robustus</emphasis>
Wang, Rasnitsyn, Perkovsky &amp; Vilhelmsen
</taxonomicName>
<taxonomicNameLabel id="ADB335495D60C33EB041C8DB77EE196D" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERRAE" type="diagnosis">
<paragraph id="270A5371052612084E86955A94028F69">
<heading id="CF3866132522EE16DED9F6200EFE663B" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="62365BF98B298FA68D53B04A8681B50E">
The length of the scape is less than 0.3 times the length of the flagellum. Fore wing with 2 Rs distal of r-rs several times as long as r-rs; Cu nebulous. Fore tibial spur long, thin and acute apically. Metabasitarsus longer than tarsomeres 2–4 combined. Propodeum outline angular. Anterior outline of 1 
<superScript id="7D273D01FD09119CECCFE3500725BAE9">st</superScript>
metasomal tergum somewhat angular.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZRAE" type="etymology">
<paragraph id="1371E3E9B64602FDDF81B5D881025A6D">
<heading id="25DE3CF6C1E1E6B356E123FA8E47FFB6" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="CD4B1763213D9097C3600F5FB13B6346">
The species is named after the Latin adjective 
<emphasis id="406EC3B919B06489CF788D93B73E0A8C" italics="true">robustus</emphasis>
, meaning “ thick ” and referring to the thick fore femur.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJSAE" type="type material">
<paragraph id="1AB59F69F647F37D93FA52DBD3EC99AB">
<heading id="92318F059220CEDD2BFEC6D42D3B81D2" reason="title">Type material.</heading>
</paragraph>
<paragraph id="152F21776301F101E33D3838F8F14730">
<materialsCitation id="108F657F80ADF3129923FD268D4B5F02" country="Myanmar" location="Kachin" specimenCode="CNU-HYM-MA 2016212" specimenCount="1" specimenCount-female="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="6F81167E535749A0996FB758A6FA3702" bold="true" italics="true">
<typeStatus id="A226A76C6D0DEB41C0EE82E5CC8250A9" type="holotype">Holotype</typeStatus>
</emphasis>
• 
<specimenCount id="9D777D6CFD97C88AA1ECEC0592FE16DE" count="1" type="female">female</specimenCount>
. No. 
<specimenCode id="69CBD4DE5BEED3C71BDE56F2630EFCE9">CNU-HYM-MA 2016212</specimenCode>
. A well-preserved and complete specimen (Figs 
<figureCitation id="F0AE785FF8CCA057DC6D4F6C6168F3A0" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Photographs of Siccibythus robustus sp. nov. (Female holotype; CNU-HYM-MA- 2016212) A habitus in lateral view B head in lateral view C fore wing venation." figureDoi="10.3897/jhr.98.136200.figure1" httpUri="https://binary.pensoft.net/fig/1213364">1</figureCitation>
, 
<figureCitation id="7C07252D79C0AC43468C73734DEA1BA4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photographs of Siccibythus robustus sp. nov. (Female holotype; CNU-HYM-MA- 2016212) A mesosoma in dorsal view B mesosoma in lateral view, arrowheads indicate metapleural-propodeal boundary, black arrow indicates the carinae C hind leg in lateral view D metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure2" httpUri="https://binary.pensoft.net/fig/1213365">2</figureCitation>
)
</materialsCitation>
. 
<materialsCitation id="9883217215A9CEEE3D3ACB67C3C2629E" country="Myanmar" location="Kachin" specimenCode="CNU-HYM-MA 2016213" specimenCount="1" specimenCount-male="1" stateProvince="Kachin" typeStatus="paratype">
<emphasis id="3503CFC9F8C882580586022A6014DF54" bold="true" italics="true">
<typeStatus id="9B95C509049CDFEE7206A0CE3F80490E" type="paratype">Paratype</typeStatus>
</emphasis>
• 
<specimenCount id="C3F7A5ED53AA4E21116FA44A479796D3" count="1" type="male">male</specimenCount>
. No. 
<specimenCode id="2B6032D4CE52C5256D1B5137FE87AA92">CNU-HYM-MA 2016213</specimenCode>
. A well-preserved and complete specimen (Figs 
<figureCitation id="08BF8AB8B5EB3BD09E0BC257DDC4A8A3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Photographs of Siccibythus robustus sp. nov. (Male paratype; CNU-HYM-MA- 2016213) A habitus in lateral view B head in lateral view." figureDoi="10.3897/jhr.98.136200.figure3" httpUri="https://binary.pensoft.net/fig/1213366">3</figureCitation>
, 
<figureCitation id="559707BA08738F3563BA48B368E6872E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Photographs of Siccibythus robustus sp. nov. (Male paratype; CNU-HYM-MA- 2016213) A mesosoma in dorsal view B fore wing venation C hind leg in lateral view D metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure4" httpUri="https://binary.pensoft.net/fig/1213367">4</figureCitation>
)
</materialsCitation>
.
</paragraph>
<caption id="EC9CFF30FB197EF612DFD47327D9A658" ID-DOI="10.3897/jhr.98.136200.figure1" ID-arpha="A42DFDDD-6CFF-5B6F-9EFB-E456B1085D1F" httpUri="https://binary.pensoft.net/fig/1213364" startId="F1">
<paragraph id="DB3C0097E06EBD5B44D90134492C3BF2">
<label id="920DA0E53F8B552A7DF5395BFA19421C">Figure 1.</label>
</paragraph>
<paragraph id="106342625C7D3CA0C9E1BBA31644CC4C">
Photographs of 
<taxonomicName id="9C77CFF93CB280E015B756F3E945C9B6" authorityName="Wang, Rasnitsyn, Perkovsky &amp; Vilhelmsen" authorityYear="2025" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="robustus" status="sp. nov.">
<emphasis id="420C3A988589D2F56145DC71F9302EE7" italics="true">Siccibythus robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4123FBBFE381A12501D9AFEA9F970823" rank="species">sp. nov.</taxonomicNameLabel>
(Female holotype; CNU-HYM-MA- 2016212) 
<emphasis id="C4AFB60CC9C0A6DE552E0A9FB0B96A5B" bold="true">A</emphasis>
habitus in lateral view 
<emphasis id="0871FAE2DA1D69A035EAE9252A283574" bold="true">B</emphasis>
head in lateral view 
<emphasis id="548FE8D4AD72623E639C0882EF0C1D93" bold="true">C</emphasis>
fore wing venation.
</paragraph>
</caption>
<caption id="E3C9E3AFA754ED81DA42B3A4637B606F" ID-DOI="10.3897/jhr.98.136200.figure2" ID-arpha="FFE4DB87-AAE0-5845-8D6E-4EB8D33E1480" httpUri="https://binary.pensoft.net/fig/1213365" startId="F2">
<paragraph id="FC584D3F013CDC035299237C7BC1916C">
<label id="91CBDDF218914FF5BACEDD78431FCFB0">Figure 2.</label>
</paragraph>
<paragraph id="E2AEC1027376027C8C12FA408CB9048A">
Photographs of 
<taxonomicName id="F05CE8D80811E21ACA83F0AB44832331" authorityName="Wang, Rasnitsyn, Perkovsky &amp; Vilhelmsen" authorityYear="2025" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="robustus" status="sp. nov.">
<emphasis id="A503E1BBD672ED0FC4E7E5E453314E85" italics="true">Siccibythus robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70107FF25A110E5BFE0CEB7BBA84E5CE" rank="species">sp. nov.</taxonomicNameLabel>
(Female holotype; CNU-HYM-MA- 2016212) 
<emphasis id="A8D3C9C81F9240040ABA4364953D85D3" bold="true">A</emphasis>
mesosoma in dorsal view 
<emphasis id="76DFD7EC0F82A43A29D1A8EDB94A0379" bold="true">B</emphasis>
mesosoma in lateral view, arrowheads indicate metapleural-propodeal boundary, black arrow indicates the carinae 
<emphasis id="E443B9723F46074361DD0BB078374B78" bold="true">C</emphasis>
hind leg in lateral view 
<emphasis id="71CC2DDF41D4969F168CB2D0E3BD16B6" bold="true">D</emphasis>
metasoma in lateral view.
</paragraph>
</caption>
<caption id="4C041BB5504A7F966761441C0AD37B0F" ID-DOI="10.3897/jhr.98.136200.figure3" ID-arpha="E6E96A44-67D9-5C8C-8AC1-725919698C9E" httpUri="https://binary.pensoft.net/fig/1213366" startId="F3">
<paragraph id="DDC36C3A5C5FE02127F913C635104ECA">
<label id="BD1C3E014A66C632D75D27D8CBB2E79A">Figure 3.</label>
</paragraph>
<paragraph id="9CA13883CDB30320179AFF61F5033F2D">
Photographs of 
<taxonomicName id="15CB95FF3F725A73AA266AF593DFAB5A" authorityName="Wang, Rasnitsyn, Perkovsky &amp; Vilhelmsen" authorityYear="2025" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="robustus" status="sp. nov.">
<emphasis id="5E14813E805812CB554E8842A261BE0B" italics="true">Siccibythus robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EB77FF5F954EEAE645A87C38F074FC44" rank="species">sp. nov.</taxonomicNameLabel>
(Male paratype; CNU-HYM-MA- 2016213) 
<emphasis id="D4C44BA6AC5B7102965D4636117CD1FB" bold="true">A</emphasis>
habitus in lateral view 
<emphasis id="D0522B955611230B0A57E7500D013E7D" bold="true">B</emphasis>
head in lateral view.
</paragraph>
</caption>
<caption id="112A143F703AAA2F521ECC430DEA8218" ID-DOI="10.3897/jhr.98.136200.figure4" ID-arpha="CB887498-BF95-53AC-9F83-9B4FB0C07766" httpUri="https://binary.pensoft.net/fig/1213367" startId="F4">
<paragraph id="758DA2637FFEA198A558E80A4AF73038">
<label id="B1948BF22B9DA31C456EFC1ECC806500">Figure 4.</label>
</paragraph>
<paragraph id="77E61C5C564E9080008AB26A1E21FED3">
Photographs of 
<taxonomicName id="555C3492C6AA0D7FA2C965D139AED29D" authorityName="Wang, Rasnitsyn, Perkovsky &amp; Vilhelmsen" authorityYear="2025" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="robustus" status="sp. nov.">
<emphasis id="C270CDB3F177F3DEE87A001653547A3D" italics="true">Siccibythus robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1CD08DF755BAAC10C1A42BF82C72DD66" rank="species">sp. nov.</taxonomicNameLabel>
(Male paratype; CNU-HYM-MA- 2016213) 
<emphasis id="8142C8CB273495AB206F37096425949E" bold="true">A</emphasis>
mesosoma in dorsal view 
<emphasis id="AE26FDDF69E9F45F0D0AFC3489F6F5E6" bold="true">B</emphasis>
fore wing venation 
<emphasis id="C649A50D860B8C6972188A1392BB56DA" bold="true">C</emphasis>
hind leg in lateral view 
<emphasis id="102EF47FA8868F0233B4E672B3111591" bold="true">D</emphasis>
metasoma in lateral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EBXAE" type="type locality">
<paragraph id="87637630DA09704E8407130B16C2360B">
<heading id="7250C516F78A5B627BD90BC70107DD15" reason="title">Type locality and horizon.</heading>
</paragraph>
<paragraph id="E85ADEF8FE91882A4E270D5B37D31C3E">
The amber specimens were collected from 
<collectingRegion id="9273C124D947FEB65FF68DAD75511F18" country="Myanmar" name="Kachin">Kachin</collectingRegion>
, northern 
<collectingCountry id="761A619A613B55AD812456233696EF4B" name="Myanmar">Myanmar</collectingCountry>
, lowermost Cenomanian, mid-Cretaceous, which was dated at 98.79 ± 0.62 Ma (
<bibRefCitation id="7AB70E8ED6B23D562A687550CCF820CC" DOI="10.1016/S1367-9120(02)00044-5" author="Cruickshank and Ko" firstAuthor="Cruickshank" journalOrPublisher="Journal of Asian Earth Sciences" pagination="441-455" refId="B3" refString="Cruickshank RD, Ko K (2003) Geology of an amber locality in the Hukawng Valley, Northern Myanmar. Journal of Asian Earth Sciences 21: 441–455. https://doi.org/10.1016/S1367-9120 (02) 00044-5" title="Geology of an amber locality in the Hukawng Valley, Northern Myanmar." volume="21" year="2003">Cruickshank and Ko 2003</bibRefCitation>
; 
<bibRefCitation id="AB5387F18AD060B8D1DB155832A8E821" DOI="10.1016/j.cretres.2012.03.014" author="Shi" etAl="et al." firstAuthor="Shi" journalOrPublisher="Cretaceous Research" pagination="155-163" refId="B25" refString="Shi G, Grimaldi DA, Harlow GE, Wang J, Wang J, Yang M, Lei W, Li Q, Li X (2012) Age constraint on Burmese amber based on U – Pb dating of zircons. Cretaceous Research 37: 155–163. https://doi.org/10.1016/j.cretres.2012.03.014" title="Age constraint on Burmese amber based on U - Pb dating of zircons." volume="37" year="2012">Shi et al. 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPXAE" type="description">
<paragraph id="94D6F86FA75D3A644FEA36DFCC5ECE85">
<heading id="FA8765CA507A0ED611BF7849919B01A4" reason="title">Description.</heading>
</paragraph>
<paragraph id="FA8251C47E484315660742FAABDB8C01">
<emphasis id="40ADF2920AD2D12BF606559A45E41602" bold="true">Female.</emphasis>
Head globose, with eyes bulging, occupying most of lateral parts of head (except for wide, convex temples), almost reaching mandibular bases, occipital carina complete, reaching hypostomal carina well behind mandibular base, vertex and frons narrow, longer than wide, ocelli in small acute triangle well distant from eyes, interantennal space flat, clypeus short, simple. Antenna slender, with 12 antennomeres in female (
<quantity id="2C41D53EAEF5D2C76FB7FEE026567228" metricMagnitude="-1" metricUnit="m" metricValue="3.302" unit="in" value="13.0">13 in</quantity>
male), each antennomere is approximately 1–2 times longer than wide, scape slightly longer than pedicel and first flagellomere combined, pedicel longer than first flagellomere, flagellomeres slightly shortening gradually toward apex except for shorter first flagellomere and longer apical segment (Fig. 
<figureCitation id="F699C6D80D7DE1866B99A965E3FF1D70" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Photographs of Siccibythus robustus sp. nov. (Female holotype; CNU-HYM-MA- 2016212) A habitus in lateral view B head in lateral view C fore wing venation." figureDoi="10.3897/jhr.98.136200.figure1" httpUri="https://binary.pensoft.net/fig/1213364">1 B</figureCitation>
). Labrum not visible externally, mandibles small, overlapping, maxillary and labial palps not visible (
<typeStatus id="CC6E94F21DCA027726E016FF8F97042A" type="paratype">paratype</typeStatus>
can be observed mandibles with three teeth, maxillary palp with four palpomeres visible, three apical long, narrow).
</paragraph>
<paragraph id="85AD161937E8B8CD793093CD4EB1DC17">
Pronotum posterior margin extended, with pronotum accounting for 0.4 times the length of mesosoma, pronotum short declivitous part delimited by transverse groove posteriorly. Propleura long, slender, abutting throughout ventrally. Mesoscutum convex between notauli; notauli complete, parapsides and lateral edges incomplete; notauli widely spaced anteriorly, converging to almost meet posteriorly; transscutal articulation distinct; mesoscutellum triangular, narrow, the length is 0.4 times its maximum width, weakly convex, carinate laterally. Meso- and metapleuron smooth, metapleuron delimited from propodeum with incomplete row of pits (Fig. 
<figureCitation id="0C3A0FF577D004AA276D4B0F51FA8E49" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photographs of Siccibythus robustus sp. nov. (Female holotype; CNU-HYM-MA- 2016212) A mesosoma in dorsal view B mesosoma in lateral view, arrowheads indicate metapleural-propodeal boundary, black arrow indicates the carinae C hind leg in lateral view D metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure2" httpUri="https://binary.pensoft.net/fig/1213365">2 A, B</figureCitation>
). Metanotum short, with indistinct metascutellum in female (metanotum short, with distinct metascutellum in male). Propodeum with clear dorsal and posterior surfaces meeting in angle, dorsal with longitudinal and one transverse carinae, posterior surface with lateral longitudinal carinae, sublateral longitudinal carinae and supracoxal transverse carinae (Fig. 
<figureCitation id="96DF58117974BD85ED122880D02D6115" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photographs of Siccibythus robustus sp. nov. (Female holotype; CNU-HYM-MA- 2016212) A mesosoma in dorsal view B mesosoma in lateral view, arrowheads indicate metapleural-propodeal boundary, black arrow indicates the carinae C hind leg in lateral view D metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure2" httpUri="https://binary.pensoft.net/fig/1213365">2 B</figureCitation>
).
</paragraph>
<paragraph id="4D74DB5987C0A293884A2AC0C80596D8">
Fore wing distal of basal veins (1 Rs and 1 M), r-rs and 1 cu-a without tubular veins, except for pterostigma and half section of 2 Rs distal of r-rs, all other distal veins nebulous in female (fore wing distal of basal vein (1 Rs and 1 M) and cu-a with no tubular veins, except for pterostigma, r-rs and most of Rs distal of r-rs, all other veins nebulous in male). Pterostigma prominent, longer than wide; vein 1 Rs straight, shorter than 1 M, arise from Sc + R just proximal to pterostigma; 1 cu-a shorter than 1 M, placed distinctly before it, parallel to 2 cu-a; Cu parallel to A; vein A extending distal to 1 cu-a, and distal part tubular, extending distal to 2 cu-a as nebulous vein, both 1 cu-a and 2 cu-a reaching A; r-rs slightly curved, arising beyond midlength of pterostigma; r-m not present (Fig. 
<figureCitation id="BF173D951FE3C1C7839D1920E5B60A7C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Photographs of Siccibythus robustus sp. nov. (Female holotype; CNU-HYM-MA- 2016212) A habitus in lateral view B head in lateral view C fore wing venation." figureDoi="10.3897/jhr.98.136200.figure1" httpUri="https://binary.pensoft.net/fig/1213364">1 C</figureCitation>
). Hind wing with thin, tubular veins Rs arising from Sc + R; A present; other veins not visible. Seven hamuli present.
</paragraph>
<paragraph id="355B566CBAD0CE561DED1F0FFC8EDEFC">
Fore coxa swollen, triangular, nearly as long as wide; fore trochanter short, fore trochantellus not visible in female (fore trochantellus trapezoidal, the ratio compared to the length of the fore femur is approximately 1: 
<quantity id="0CECB28AB6386D2B07078DD99A5DB1DE" metricMagnitude="-1" metricUnit="m" metricValue="4.318" unit="in" value="17.0">17 in</quantity>
male); fore femur swollen in middle, twice as long as wide (calcar long, thin, gently curved and acute apically in male); fore tibia nearly as long as femur; fore tarsus shorter than fore tibia, basitarsomere shorter than combined length of remaining tarsomeres in female (basitarsomere longer than combined length of remaining tarsomeres in male). Mid coxa slender, smaller than both fore and hind coxa, twice as long as wide; mid trochanter short, mid trochantellus very short and ring-like, approximately 0.1 times as long as femur; mid femur swollen in middle, twice as long as wide; mid tibia slenderer than femur, three times as long as wide, tibia shorter than femur; mid tarsus nearly as long as mid tibia, basitarsomere shorter than combined length of remaining tarsomeres. Hind coxae elongate, swollen, three times as long as wide, with lamella dorsally, basally in female (no lamella in male); hind trochanter short, bulging dorsally, hind trochantellus very short and ring-like (ca. 0.02 as long as femur); hind femur swollen in middle, nearly three times as long as wide in female (nearly six times as long as wide in male); hind tibia very slender, slightly longer than femur; hind tarsus nearly as long as hind tibia, basitarsomere slightly shorter than combined length of remaining tarsomeres (Fig. 
<figureCitation id="71158E2214C902A03F2178B40782DDD7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photographs of Siccibythus robustus sp. nov. (Female holotype; CNU-HYM-MA- 2016212) A mesosoma in dorsal view B mesosoma in lateral view, arrowheads indicate metapleural-propodeal boundary, black arrow indicates the carinae C hind leg in lateral view D metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure2" httpUri="https://binary.pensoft.net/fig/1213365">2 C</figureCitation>
). Claw simple, arolium small. Tibial spur formula 1 / 2 / 2.
</paragraph>
<paragraph id="56B0BBCE11F19F376D53E8523A694C09">
First metasomal segment short, tergum node-like, female with anterior outline, low triangular, second metasomal segment longest and highest, following segments gradually smaller, apical metasomal sternum high, with dorsal sides overlapping (Fig. 
<figureCitation id="FB3C2D48856086166F6CBBB1F47BF0D6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photographs of Siccibythus robustus sp. nov. (Female holotype; CNU-HYM-MA- 2016212) A mesosoma in dorsal view B mesosoma in lateral view, arrowheads indicate metapleural-propodeal boundary, black arrow indicates the carinae C hind leg in lateral view D metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure2" httpUri="https://binary.pensoft.net/fig/1213365">2 D</figureCitation>
) (male first metasomal tergum with fore surface convex, dorsal surface short, rather flat, elevated above hind surface; seventh metasomal sternum with two long upcurved teeth).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXYAE" type="description">
<paragraph id="8EE4CBF29EBF7AFA3D20105DFB7F399A">
<heading id="22C51808E09F262FE0D6F038695042B0" reason="title">Measurements (mm).</heading>
</paragraph>
<paragraph id="6E51014A65FF4706A344F062FE044C61">
<typeStatus id="E0B2232FA29A58944CC703C39E3EE201" type="holotype">Holotype</typeStatus>
female. Total body length (excluding sting and antenna) 3.93; head length 0.47, width 0.61, height 0.53; scape length 0.25, pedicel length 0.13, the first flagellum length 0.08, total flagellum length 0.87; fore wing length 2.22; hind wing length 1.56; fore coxa length 0.30, fore trochanter length 0.57, fore femur length 0.58, fore tibia length 0.55, fore tarsal length 0.32; mid coxa length 0.31, mid trochanter length 0.09, mid trochantellus 0.05, mid femur length 0.50, mid tibia length 0.38, mid tarsal length 0.38; hind coxa length 0.44, hind trochanter length 0.14, hind trochantellus 0.02, hind femur length 0.67, hind tibia length 0.78, hind tarsal length 1.02; the first metasomal segment length 0.28, metasoma length 2.18.
</paragraph>
<paragraph id="174A11C3E7381316AB511D9DB0E4F8C8">
<typeStatus id="CC6EA19B5BB394F69313BF79D8D4DF11" type="paratype">Paratype</typeStatus>
male. Total body length (excluding antenna) 3.54; head length 0.50, width 0.42, height 0.50; scape length 0.25, pedicel length 0.16, the first flagellum length 0.09; total flagellum length 0.99; fore wing length 1.98; hind wing length 1.47; fore coxa length 0.21, fore trochanter length 0.08, fore trochantellus 0.03, fore femur length 0.50, fore tibia length 0.45, fore tarsal length 0.32; mid coxa length 0.12, mid trochanter length 0.10, fore trochantellus 0.02, mid femur length 0.40, mid tibia length 0.30, mid tarsal length 0.34; hind coxa length 0.28, hind trochanter length 0.12, hind femur length 0.55, hind tibia length 0.60, hind tarsal length 0.57; the first metasomal segment length 0.25, metasoma length 1.62.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4YAE" type="remarks">
<paragraph id="FB1D82DF19A11C69D6E8DA02344D0233">
<heading id="D63F8971A268B49E5C26BA4D31D8AA11" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="578CBC58B01488B7356B8689B8AC3322">
The new species is classified in † 
<taxonomicName id="599B0064B45DACD5A3A150E87FC7F15E" authority="Cockx &amp; McKellar, 2016" authorityName="Cockx &amp; McKellar" authorityYear="2016" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
<emphasis id="8F6EE06B86B82173886BC8DBC82FA071" italics="true">Siccibythus</emphasis>
<bibRefCitation id="DE416655AA41DB0EBA1F33A52EBD27BE" DOI="10.1016/j.cretres.2016.06.015" author="Cockx PFD &amp; McKellar RC" journalOrPublisher="Cretaceous Research" pagination="133-139" refId="B2" refString="Cockx PFD, McKellar RC (2016) First record of the family Scolebythidae (Hymenoptera) in mid-Cretaceous amber from Myanmar. Cretaceous Research 67: 133 - 139. https: // doi. org / 10.1016 / j. cretres. 2016.06. 015" title="First record of the family Scolebythidae (Hymenoptera) in mid-Cretaceous amber from Myanmar." volume="67" year="2016">Cockx &amp; McKellar, 2016</bibRefCitation>
</taxonomicName>
based on the fore wing with no cells closed with tubular veins in its middle and distal part (
<bibRefCitation id="78E05863837D24B5350572C242D1F2B3" DOI="10.11646/palaeoentomology.3.6.10" author="Rasnitsyn" etAl="et al." firstAuthor="Rasnitsyn" journalOrPublisher="Palaeoentomology" pagination="582-596" refId="B20" refString="Rasnitsyn AP, Zhang Q, Müller P, Zhang HC (2020) On the identity and limits of Falsiformicidae (Insecta: Hymenoptera, Vespoidea s. l.). Palaeoentomology 3: 582–596. https://doi.org/10.11646/palaeoentomology.3.6.10" title="On the identity and limits of Falsiformicidae (Insecta: Hymenoptera, Vespoidea s. l.)." volume="3" year="2020">Rasnitsyn et al. 2020</bibRefCitation>
). However, the new species is differentiated from 
<taxonomicName id="58BDB0F64D7243F492774E3A309686DB" authorityName="Rasnitsyn, Zhang, Muller &amp; Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="martynovae">
<emphasis id="99BD5F86901C6FDEF1AF17F1D38DF0B8" italics="true">Siccibythus martynovae</emphasis>
</taxonomicName>
, 
<taxonomicName id="0C211A049947632DF234F307CE7D43E8" authorityName="Rasnitsyn, Zhang, Muller &amp; Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="F4968C9B505511E04429EA0A4A0237AD" italics="true">S. pallidus</emphasis>
</taxonomicName>
and 
<taxonomicName id="917667F797B549320B2D7288192A08CB" authorityName="Rasnitsyn, Zhang, Muller &amp; Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="oculatus">
<emphasis id="93C746DA90AF6D27A2761044CD63056E" italics="true">S. oculatus</emphasis>
</taxonomicName>
by fore wing with 2 Rs distal of r-rs several times as long as r-rs, basitarsus longer than tarsomeres 2–4 combined (vs. 2 Rs distal of r-rs at most about as long as r-rs, basitarsus not longer than tarsomeres 2–4 combined in 
<taxonomicName id="09A6C6D38A30EC858B2F37772D232175" authorityName="Rasnitsyn, Zhang, Muller &amp; Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="martynovae">
<emphasis id="1EFDED134CCEC00E6ED3622DC5D74D10" italics="true">S. martynovae</emphasis>
</taxonomicName>
, 
<taxonomicName id="7BC8EEABC15FF4462D8D3A3C886CB067" authorityName="Rasnitsyn, Zhang, Muller &amp; Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="15810D245CC3E615B9402B4900433F25" italics="true">S. pallidus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4CAE7DD5CB3685815926698B0C74BC42" authorityName="Rasnitsyn, Zhang, Muller &amp; Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="oculatus">
<emphasis id="768DF06C37D712CA66A445653A52DE8A" italics="true">S. oculatus</emphasis>
</taxonomicName>
). It is distinguished from 
<taxonomicName id="FF4B8F78C2C47BE809658F28B1330821" authorityName="Cockx &amp; McKellar" authorityYear="2016" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="musculosus">
<emphasis id="AD001712E618813C823F90B3DFDE584C" italics="true">S. musculosus</emphasis>
</taxonomicName>
by the following characters: 1) Cu is nebulous (vs. Cu is tubular); 2) calcar long, thin and acute apical (vs. calcar short, bifid apically). It is differentiated from 
<taxonomicName id="C1F14698F4E18B5199879FBE5C41E758" authorityName="Rasnitsyn, Zhang, Muller &amp; Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="paulus">
<emphasis id="F46570DC92EB65744A6F5D4A3AC765BE" italics="true">S. paulus</emphasis>
</taxonomicName>
by the length of the scape being less than 0.3 times the length of the flagellum (vs. the scape is longer than or equal to 0.3 times of the flagellum). It is distinguished from 
<taxonomicName id="71F0186EAEFBF2C32BFCF0DA9DCE266F" authorityName="Rasnitsyn, Zhang, Muller &amp; Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="ohmkuhnlei">
<emphasis id="01E175FA52540E6714C62DDB1FB830B6" italics="true">S. ohmkuhnlei</emphasis>
</taxonomicName>
by the following characters: 1) anterior outline of 1 
<superScript id="A83D8D7CF8F465D48004A31FEBFDEAD1">st</superScript>
metasomal tergum somewhat angular (vs. straight); 2) propodeum outline angular (vs. more smoothly curved).
</paragraph>
</subSubSection>
</treatment>
</document>