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<mods:title id="C665590E50F7EB21D44C0C5C80DE90D5">Contributions to the taxonomy of the ant genus Echinopla Smith, 1857 (Hymenoptera, Formicidae)</mods:title>
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<mods:namePart id="5C5EB6C4539544E3B6F702EF32650203">Laciny, Alice</mods:namePart>
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<treatment id="548947155CEA5B18ADB5705EA174E7F1" ID-GBIF-Taxon="127900112" LSID="urn:lsid:plazi:treatment:548947155CEA5B18ADB5705EA174E7F1" httpUri="http://treatment.plazi.org/id/548947155CEA5B18ADB5705EA174E7F1" lastPageId="15" lastPageNumber="115" pageId="13" pageNumber="114">
<subSubSection id="9560D2811FAF12220D64349A9A600BA9" pageId="13" pageNumber="114" type="multiple">
<paragraph id="A9CDFBFB3892444B7FE9089540E817D9" pageId="13" pageNumber="114">Taxon classification Animalia Hymenoptera Formicidae</paragraph>
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<subSubSection id="0C59415D7146A7AA4A080F0BB03C5A65" pageId="13" pageNumber="114" type="nomenclature">
<paragraph id="D51EA6D84AF814597E48D23ACFB479F9" pageId="13" pageNumber="114">
<taxonomicName id="6E415D799497B052870867E7066EDB51" ID-CoL="38JSR" authority="Mayr, 1862" authorityName="Mayr" authorityYear="1862" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla lineata" order="Hymenoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="lineata">Echinopla lineata Mayr, 1862</taxonomicName>
Figs 35-38, 43
</paragraph>
</subSubSection>
<subSubSection id="96E6435E4E9554DC94696F03A18EAA18" pageId="13" pageNumber="114" type="type material examined">
<paragraph id="4B91B3BECA50D7AD484FFAFC7556DD9F" pageId="13" pageNumber="114">Type material examined.</paragraph>
<paragraph id="B3EC7EA3802CEBA45202CDDCCC33DEC6" pageId="13" pageNumber="114">
Lectotype (worker, present designation, NHMW) from &quot;Batavia (Novara)&quot; (
<bibRefCitation id="6516F6849386763BB326742F63F9C4F8" author="Mayr, G" journalOrPublisher="Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien" pageId="18" pageNumber="119" pagination="649 - 776" title="Myrmecologische Studien." volume="12" year="1862">Mayr 1862</bibRefCitation>
); labels see Figure 26. Paralectotypes (2 workers, NHMW), from the same locality.
</paragraph>
</subSubSection>
<subSubSection id="D3CEF8DFA973CCCCCAD2BEE3DE193F0E" lastPageId="14" lastPageNumber="115" pageId="13" pageNumber="114" type="additional material examined">
<paragraph id="B32FD6830BF7161FE80ECD6349421A5A" pageId="13" pageNumber="114">Additional material examined.</paragraph>
<paragraph id="AA2F20CB3D446C2DB9A52733898FDC67" lastPageId="14" lastPageNumber="115" pageId="13" pageNumber="114">
1 worker (NHMW) of unknown origin, leg. Ida Pfeiffer, det. G. Mayr; 1 worker (NHMW) from Java, leg. Vollenhoven, det. G. Mayr; 5 workers (ZCW) from Singapore, Singapore Zoo, 2.XI.2003, leg. H. Zettel; 8 workers (SCV, ZCW)
<pageBreakToken id="BD0380D228193316984EA9FA6C4A8463" pageId="14" pageNumber="115" start="start">from</pageBreakToken>
Borneo, Sarawak, Mulu National Park, Kerangas Forest behind Mulu Airport, 21.VII.2010, leg. D.M. Sorger; 1 worker (WCD) from Borneo, Sabah, Danum Valley Field Centre,
<geoCoordinate id="9DD1A7938E4FA3F5580FE8137A192C3C" direction="north" orientation="latitude" precision="1" value="4.96462">4.96462°N</geoCoordinate>
;
<geoCoordinate id="8115ADB59F8DDAA771CFCC912571D87F" direction="east" orientation="longitude" precision="1" value="117.804">117.804000°E</geoCoordinate>
<normalizedToken id="A333E42599A925318B2C113DDEAB0650" originalValue="±">+/-</normalizedToken>
70 m, 180 m a.s.l., 24.VIII.2010, leg. P.S. Ward (#16476); 5 workers (CAS), from same locality, 525 m a.s.l.,
<geoCoordinate id="7056905ACBB7375A3EA3A0CFF555E2C8" direction="north" orientation="latitude" precision="1" value="4.965611">4.965611°N</geoCoordinate>
;
<geoCoordinate id="D19E1992C10283EBC0E08E512CCA7496" direction="east" orientation="longitude" precision="1" value="117.799805">117.799806°E</geoCoordinate>
, 16-26.VIII.2010, rainforest, Ant Course 2010, various collectors; 1 workers (CAS), from same area, Danum Valley, West Trail, 225 m a.s.l.,
<geoCoordinate id="F0CF276005C046AED1DE9E45923773D8" direction="north" orientation="latitude" precision="1" value="4.963056">4.963056°N</geoCoordinate>
;
<geoCoordinate id="5116A4C27A5080C63F3A2C646FC23EAD" direction="east" orientation="longitude" precision="1" value="117.8028">117.802806°E</geoCoordinate>
, 16-26.VIII.2010, rainforest, Ant Course 2010, unknown collector.
</paragraph>
</subSubSection>
<subSubSection id="18033E602AE44022731DC70D323D2093" pageId="14" pageNumber="115" type="measurements of the lectotype">
<paragraph id="D02ECA56B400AD88C6EA9C5DAE6204EC" pageId="14" pageNumber="115">Measurements of the lectotype.</paragraph>
<paragraph id="FE123A30BA07DB3B662BECFACF4ACC0F" pageId="14" pageNumber="115">TL 6.3; HW1 1.66; HW2 1.58; HL 1.57; EL 0.36; SL 1.39; SW 0.18; HaL 0.48; PML 1.17; PMW 1.50; PpL 1.04; PpW 1.20; PH 0.47; PL 0.51; PW 1.23; GL 1.65; GW 1.80. Indices: CI 106; SI 84; MI 148.</paragraph>
</subSubSection>
<subSubSection id="844884B641E9A67018D3CACCE288B20C" lastPageId="15" lastPageNumber="116" pageId="14" pageNumber="115" type="notes">
<paragraph id="4F59B046EAC2C1836DC5C68675CD656C" pageId="14" pageNumber="115">Notes.</paragraph>
<paragraph id="AA95C49FCCE2BAC66AF036CBBBE11EF4" pageId="14" pageNumber="115">
<taxonomicName id="AAB1F89777F68A7CB9BEC5BC5F2D731E" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla lineata" order="Hymenoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="lineata">Echinopla lineata</taxonomicName>
has some similarities with
<taxonomicName id="9B5BFBDD9CCB0BB0D49FFA31789BEE59" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla striata" order="Hymenoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="striata">Echinopla striata</taxonomicName>
, especially in sculpture, but can be recognized by the combination of black colour (without metallic lustre), coarse longitudinal striation on head and mesosoma, fine longitudinal striation on gaster, long white setae, and relatively abundant short appressed hair that gives the specimens a dull appearance. See also notes on the similar
<taxonomicName id="B29A61BE5946F25C5F6EAE43FBDDC67C" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla senilis" order="Hymenoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="senilis">Echinopla senilis</taxonomicName>
.
</paragraph>
<paragraph id="5B5FF724806C408B8A796E13923CF21B" pageId="14" pageNumber="115">
The collection of NHMW contains five historical specimens of
<taxonomicName id="9D4B17858F87AE662899E198AA3F640E" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla lineata" order="Hymenoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="lineata">Echinopla lineata</taxonomicName>
, one of which was erroneously labelled as
<taxonomicName id="2F1506BFB384C20ADF77AFD6DFE01D97" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla senilis" order="Hymenoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="senilis">Echinopla senilis</taxonomicName>
. Three specimens are considered as types, whereas one gyne from Java (see
<bibRefCitation id="37D39B2349F8A29BCEEAFFE8DD545A0E" author="Mayr, G" journalOrPublisher="Annali del Museo Civico di Storia Naturale" pageId="18" pageNumber="119" pagination="133 - 155" title="Formicidae Borneenses collectae a J. Doria et O. Beccari in territorio Sarawak annis 1865 - 1867." volume="2" year="1872">Mayr 1872</bibRefCitation>
) and one worker collected by Ida Pfeiffer from an unknown locality are non-type specimens. The worker (paralectotype) labelled
<normalizedToken id="6FFA1DDAA5298C1178E1F164B86AF472" originalValue="“Batavia”">&quot;Batavia&quot;</normalizedToken>
(now Jakarta, Java, Indonesia) is missing its head. One worker (paralectotype) bears the same locality code label Q and the same Novara expedition label as the lectotype (see Fig. 38), but a determination label &quot;
<taxonomicName id="9238872C3BD6B5DC85E5BE9DBBFEC3E3" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla senilis" order="Hymenoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="senilis">Echinopla senilis</taxonomicName>
det. G. Mayr&quot; (in Anton
<normalizedToken id="9EFC01555CD87A65497F07187FB0CAA3" originalValue="Handlirsch’s">Handlirsch's</normalizedToken>
handwriting!) which is probably a curatorial error (see notes for
<taxonomicName id="4E1C2BDD7C642A13F60968697AA17E02" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla senilis" order="Hymenoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="senilis">Echinopla senilis</taxonomicName>
). This slightly damaged specimen clearly belongs to
<taxonomicName id="6FD3D9CFF921BED911086F1230782B81" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla lineata" order="Hymenoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="lineata">Echinopla lineata</taxonomicName>
. The third type specimen, which is in a very good condition (see Figs 35-37), was selected as the lectotype.
</paragraph>
<caption id="29D7C50418107D785007D013DFF37BD8" pageId="14" pageNumber="115">
<paragraph id="7AE713813B489AEB5E4741F0D8F93CB7" pageId="14" pageNumber="115">
Figures 35-38.
<taxonomicName id="756450DF7779A00B88BCAF0A1268437F" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla lineata" order="Hymenoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="lineata">Echinopla lineata</taxonomicName>
, lectotype: (35) Head, full face view. (36) Habitus, lateral view. (37) Habitus, dorsal view. (38) Labels.
</paragraph>
</caption>
<paragraph id="61B724CE74E5FDADE4D95E31EFB3C5F0" lastPageId="15" lastPageNumber="116" pageId="14" pageNumber="115">
The species was originally described from Java (
<bibRefCitation id="C35092572A7CAE163B980AF049FD7894" author="Mayr, G" journalOrPublisher="Verhandlungen der Kaiserlich-Koeniglichen Zoologisch-Botanischen Gesellschaft in Wien" pageId="18" pageNumber="119" pagination="649 - 776" title="Myrmecologische Studien." volume="12" year="1862">Mayr 1862</bibRefCitation>
). This first description is very short, but
<bibRefCitation id="E326F2F83D82DD91298389CE9BA30DB5" author="Mayr, G" journalOrPublisher="K. Gerold's Sohn, Vienna" pageId="18" pageNumber="119" title="Formicidae. In: Reise der Oesterreichischen Fregatte &quot; Novara &quot; um die Erde in den Jahren 1857, 1858, 1859. Zoologischer Theil" year="1865">Mayr (1865)</bibRefCitation>
presents illustrations (Fig. 43) and a much more detailed description.
<bibRefCitation id="28C5B6B502EA23F3F3B855A09A5043C7" author="Emery, C" journalOrPublisher="Annali del Museo Civico di Storia Naturale" pageId="18" pageNumber="119" pagination="661 - 722" title="Formiche raccolte da Elio Modigliani in Sumatra, Engano e Mentawei." volume="40" year="1900">Emery (1900)</bibRefCitation>
recorded
<taxonomicName id="8363097B12D688C23EADFB57B8A7FB60" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla lineata" order="Hymenoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="lineata">
<pageBreakToken id="3854C0783C7D5E313044318B6E71837E" pageId="15" pageNumber="116" start="start">Echinopla</pageBreakToken>
lineata
</taxonomicName>
from Sumatra.
<taxonomicName id="0CF34E79BD3A8E6AAF862434657EFE82" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla sucki" order="Hymenoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="sucki">Echinopla sucki</taxonomicName>
was described by
<bibRefCitation id="5BC974DD90EFDAEA5BF303F710D0569D" author="Forel, A" journalOrPublisher="Mitteilungen aus dem Naturhistorischen Museum in Hamburg" pageId="18" pageNumber="119" pagination="43 - 82" title="Formiciden des Naturhistorischen Museums zu Hamburg. Neue Calyptomyrmex-, Dacryon-, Podomyrma- und Echinopla-Arten." url="10.5281/zenodo.14254" volume="18" year="1901">Forel (1901)</bibRefCitation>
from Sarawak, Borneo, and synonymised by
<bibRefCitation id="249B4F22F6AAE2C778345C315CC4975B" author="Emery, C" journalOrPublisher="Genera Insectorum" pageId="18" pageNumber="119" pagination="1 - 302" title="Hymenoptera. Fam. Formicidae. Subfam. Formicinae." volume="183" year="1925">Emery (1925)</bibRefCitation>
. After having studied the pictures of a syntype of
<taxonomicName id="18B22FA8E161890F0EE4C8BDE48A719A" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla sucki" order="Hymenoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="sucki">Echinopla sucki</taxonomicName>
in
<bibRefCitation id="54F7B27FFB80ED1969DB8488C12B2DB3" author="Andre, E" journalOrPublisher="Memoires de la Societe Zoologique de France" pageId="18" pageNumber="119" url="http://www.antweb.org" year="2015">Antweb (2015)</bibRefCitation>
and non-type specimens from Sarawak, we affirm this synonymy. We also confirm the presence of
<taxonomicName id="D6C3506B10E92A3854CBEB8574984C15" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla lineata" order="Hymenoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="lineata">Echinopla lineata</taxonomicName>
in Sabah, northern Borneo, based on examined worker specimens in the P.S. Ward Collection and in the California Academy of Sciences.
<bibRefCitation id="384A2880E10FF8FFB92F018738CA025E" author="Andre, E" journalOrPublisher="Memoires de la Societe Zoologique de France" pageId="18" pageNumber="119" url="http://www.antweb.org" year="2015">Antweb (2015)</bibRefCitation>
records two specimens from Sabah (not examined), but the illustrated one from the Maliau Basin is not
<taxonomicName id="37D9AB1ADE8A364933D64998B6BE13ED" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla lineata" order="Hymenoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="lineata">Echinopla lineata</taxonomicName>
, because it differs strongly by a fine sculpture of head and thoracic nota, by long black pilosity and the almost absent short hair on the thoracic nota; except for its non-metallic colour it resembles
<taxonomicName id="63C46743F4EBC7B711B71FCEF45FA873" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla striata" order="Hymenoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="striata">Echinopla striata</taxonomicName>
. Also the
<bibRefCitation id="AA83F04EF3D203918029B1FC0F919A4B" author="Andre, E" journalOrPublisher="Memoires de la Societe Zoologique de France" pageId="18" pageNumber="119" url="http://www.antweb.org" year="2015">Antweb (2015)</bibRefCitation>
record of
<taxonomicName id="92BC684F5C24615FE90AB993F68ABE94" class="Insecta" family="Formicidae" genus="Echinopla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Echinopla lineata" order="Hymenoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="lineata">Echinopla lineata</taxonomicName>
in Singapore can be confirmed after having studied the illustrations of this specimen and additionally collected specimens.
<bibRefCitation id="51164BAF82C4AE88D3F3F0B0AF6C83B7" author="Andre, E" journalOrPublisher="Memoires de la Societe Zoologique de France" pageId="18" pageNumber="119" url="http://www.antbase.net" year="2015">Antbase (2015)</bibRefCitation>
illustrates a correctly identified worker from Kuala Lumpur, West Malaysia, in the Natural History Museum in London.
</paragraph>
</subSubSection>
</treatment>
</document>
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