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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.245.3416" ID-PMC="PMC3577090" ID-Pensoft-Pub="1313-2970-245-1" ID-Pensoft-UUID="FFFF52503A0AFF882450FFB66D45FF8E" ID-PubMed="23431087" ID-Zenodo-Dep="578462" ModsDocID="1313-2970-245-1" checkinTime="1425642919252" checkinUser="pensoft" docAuthor="Bousquet, Yves" docDate="2012" docId="547DE2B8222E7B1FC316B53D1A12F3BE" docLanguage="en" docName="ZooKeys 245: 1-1722" docOrigin="ZooKeys 245" docPubDate="2012-11-28" docSource="http://dx.doi.org/10.3897/zookeys.245.3416" docTitle="Pelophila borealis" docType="treatment" docVersion="11" id="FFFF52503A0AFF882450FFB66D45FF8E" lastPageNumber="127" masterDocId="FFFF52503A0AFF882450FFB66D45FF8E" masterDocTitle="Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico" masterLastPageNumber="1722" masterPageNumber="1" pageNumber="127" updateTime="1668154792637" updateUser="ExternalLinkService">
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<mods:title id="0A6538BDC6F414AC51B1535A21A87EEE">Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico</mods:title>
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<treatment id="547DE2B8222E7B1FC316B53D1A12F3BE" ID-GBIF-Taxon="127867304" LSID="urn:lsid:plazi:treatment:547DE2B8222E7B1FC316B53D1A12F3BE" httpUri="http://treatment.plazi.org/id/547DE2B8222E7B1FC316B53D1A12F3BE" lastPageNumber="127" pageId="126" pageNumber="127">
<subSubSection id="69A413DEBB70FE1F2D703E1D277E7778" pageId="126" pageNumber="127" type="nomenclature">
<paragraph id="FC306748A5FA09FAA516F8BC203AAA82" pageId="126" pageNumber="127">
<taxonomicName id="AE8B232994DDB837D97EB3F323B7557D" ID-CoL="76CH8" ID-ENA="1483836" LSID="547DE2B8-222E-7B1F-C316-B53D1A12F3BE" authority="(Paykull, 1790)" baseAuthorityName="Paykull" baseAuthorityYear="1790" class="Insecta" family="Carabidae" genus="Pelophila" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Pelophila borealis" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="borealis">Pelophila borealis (Paykull, 1790)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="39476DFABFE9D1C4FA4F817059E4F21D" pageId="126" pageNumber="127" type="reference_group">
<paragraph id="2CD37A717EF73D7BA2C1A15DA417FB0E" pageId="126" pageNumber="127">
<taxonomicName id="07E81F268CE0C2D565BCE20177D42DB8" class="Hexapoda" family="Carabidae" genus="Carabus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="borealis">Carabus borealis</taxonomicName>
Paykull, 1790: 51. Type locality: &quot;Karungi Botniae occidentalis [Norrbotten County, Sweden]&quot; (original citation). Syntype(s) probably in NRSS.
</paragraph>
<paragraph id="7302C1F4A58AE7F1B9F86B76E93B4285" pageId="126" pageNumber="127">
<taxonomicName id="74EA1CD3BB4636276F6F87F7FAE0DF47" authorityName="Mannerheim" authorityYear="1823" class="Insecta" family="Carabidae" genus="Pelophila" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="gebleri">Pelophila gebleri</taxonomicName>
Mannerheim, 1823: 38. Type locality: &quot;Sibiria ad Barnaul [Altai Kray, Russia]&quot; (original citation). One syntype in ZMH (Silfverberg 1987: 17). Synonymy established by Dejean (1833: 22).
</paragraph>
<paragraph id="4E32E567B1BB7977E84061586D40C537" pageId="126" pageNumber="127">
<taxonomicName id="2FC0E8872AF41F82DCE60AE81B87CBBB" authorityName="Mannerheim" authorityYear="1823" class="Insecta" family="Carabidae" genus="Pelophila" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="marginata">Pelophila marginata</taxonomicName>
Mannerheim, 1823: 39. Type locality: &quot;peninsula Kamschatka ad portum St. Petri et Pauli [apparently the town of Petropawlowsk, see Lindroth (1961a: 74)]&quot; (original citation). One syntype in ZMH (Silfverberg 1987: 20). Synonymy established by Dejean (1833: 22).
</paragraph>
<paragraph id="410B3579378D7FDF799C263584D0F577" pageId="126" pageNumber="127">
<taxonomicName id="8EFFE404193A78FBCAF31454233A6266" authorityName="Mannerheim" authorityYear="1823" class="Insecta" family="Carabidae" genus="Pelophila" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="eschscholtzii">Pelophila eschscholtzii</taxonomicName>
Mannerheim, 1823: 40. Type locality: &quot;insula Unalaschka [Aleutian Islands, Alaska]&quot; (original citation). Lectotype (♂), designated by Lindroth (1961a: 57), in ZMH. Synonymy established by Dejean (1833: 22), confirmed by Lindroth (1961a: 57).
</paragraph>
<paragraph id="E8CA978C2D3855BF600470953C81BEE3" pageId="126" pageNumber="127">
<taxonomicName id="63E820CA826061B8C0B084DAE9EB14F7" authorityName="Mannerheim" authorityYear="1823" class="Insecta" family="Carabidae" genus="Pelophila" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="elongata">Pelophila elongata</taxonomicName>
Mannerheim, 1823: 41. Type locality: &quot;peninsula Kamschatka ad portum St. Petri et Pauli [apparently the town of Petropawlowsk, see Lindroth (1961a: 74)]&quot; (original citation). One syntype in ZMH (Silfverberg 1987: 15). Synonymy established by Dejean (1833: 22).
</paragraph>
<paragraph id="3D3786F494DDEA7BD6E082E52FBB924C" pageId="126" pageNumber="127">
<taxonomicName id="10FF504827CFAE7EB1BC91601E83EA8D" baseAuthorityName="Paykull" baseAuthorityYear="1790" class="Insecta" family="Carabidae" genus="Pelophila" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="borealis">Pelophila borealis</taxonomicName>
var.
<emphasis id="F01719613B5AE2A21A44267722FC7B32" italics="true" pageId="126" pageNumber="127">arctica</emphasis>
Dejean, 1826: 265. Type locality not stated. Syntype(s) location unknown (possibly in MHNP). Synonymy established by Dejean (1833: 22).
</paragraph>
<paragraph id="ACA469C5ABDA092048BFBD32DB3F3ADC" pageId="126" pageNumber="127">
<taxonomicName id="D5233B851D4CCDA5AAF5ADFF028CCBD7" baseAuthorityName="Paykull" baseAuthorityYear="1790" class="Insecta" family="Carabidae" genus="Pelophila" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="borealis">Pelophila borealis</taxonomicName>
var.
<emphasis id="38AADAFD7429FBBA1329D61146F1BE66" italics="true" pageId="126" pageNumber="127">dejeanii</emphasis>
Dejean, 1826: 265. Type locality: &quot;environs de Barnaoul,
<normalizedToken id="9F013C0739E523B1F0ECE12FF686F864" originalValue="Sibérie">Siberie</normalizedToken>
[Altai Kray, Russia]&quot; (original citation). Syntype(s) probably in MHNP. Synonymy established by Dejean (1833: 22).
</paragraph>
<paragraph id="7B8F9AE9432B6DDAD832C47B26892CDF" pageId="126" pageNumber="127">
<taxonomicName id="37FB587443D084C06742573A441E60CA" authorityName="Motschulsky" authorityYear="1844" class="Insecta" family="Carabidae" genus="Pelophila" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="laevigata">Pelophila laevigata</taxonomicName>
Motschulsky, 1844: 92. Type locality:
<normalizedToken id="8DA602D1A3C9A7DF3918F07624E88807" originalValue="«près">&quot;pres</normalizedToken>
de la redoute Yamyschevo non loin du fleuve Irtych [Siberia, Russia]&quot; (original citation). Holotype [by monotypy] in ZMMU (Keleinikova 1976: 202). Synonymy established by
<normalizedToken id="3EC6AA069A2B7A3CE08B8BFFE5E5B49E" originalValue="Bänninger">Baenninger</normalizedToken>
(1930: 101).
</paragraph>
<paragraph id="6200F1F5258F4FD888360D90FD5F1D87" pageId="126" pageNumber="127">
<taxonomicName id="2B38A274BE0310E31D6DD85E6F098225" authorityName="Motschulsky" authorityYear="1844" class="Insecta" family="Carabidae" genus="Pelophila" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="californica">Pelophila californica</taxonomicName>
Motschulsky, 1844: 93. Type locality:
<normalizedToken id="66C9CAB3993EA473F78F9CA93ACAE092" originalValue="«Californie»">&quot;Californie&quot;</normalizedToken>
(original citation), which is incorrect (Lindroth 1961a: 57). Syntype(s) location unknown (possibly in ZMMU though not listed in Keleinikova 1976). Synonymy established by
<normalizedToken id="A89D8F646B9345E97EA399049E427440" originalValue="Bänninger">Baenninger</normalizedToken>
(1930: 101).
</paragraph>
<paragraph id="6E19AB32EA982E27788FDC647D9F3175" pageId="126" pageNumber="127">
<taxonomicName id="BD66D3CC426DF7CD75C26D93EB9D9489" authorityName="R. F. Sahlberg" authorityYear="1844" class="Insecta" family="Carabidae" genus="Pelophila" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="ochotica">Pelophila ochotica</taxonomicName>
R.F. Sahlberg, 1844: 17. Type locality: &quot;in monte Morikan [Okhotsk, Khabarovsk Kray, Siberia, Russia]&quot; (original citation). Holotype [by monotypy] location unknown (possibly in ZMUT). Synonymy established by
<normalizedToken id="B07F344F8152D66F2A4710182D9F5D1E" originalValue="Bänninger">Baenninger</normalizedToken>
(1930: 101).
</paragraph>
<paragraph id="908310BA0D7ADE416F2F85DE003D9916" pageId="126" pageNumber="127">
<taxonomicName id="A9910F38D434D8ACC2DD1D2FC43C4285" authorityName="Motschulsky" authorityYear="1860" class="Insecta" family="Carabidae" genus="Pelophila" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="angusticollis">Pelophila angusticollis</taxonomicName>
Motschulsky, 1860: 98. Type locality:
<normalizedToken id="FCC0AFE98AB672B29BF13EB7B8E30466" originalValue="«Kamtschatka»">&quot;Kamtschatka&quot;</normalizedToken>
(original citation). Five syntypes in ZMMU (Keleinikova 1976: 186). Synonymy established by Shilenkov (1994: 9).
</paragraph>
<paragraph id="BE7B7CDF4130C3EA153F13676432C190" pageId="126" pageNumber="127">
<taxonomicName id="A283A082FC957390B2F8D0A2DA728BC5" authorityName="G. Horn" authorityYear="1870" class="Insecta" family="Carabidae" genus="Pelophila" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="ulkei">Pelophila ulkei</taxonomicName>
G.H. Horn, 1870b: 105. Type locality:
<normalizedToken id="663AC7BEF050055E6E3B8E67849B60AE" originalValue="«Hudsons">&quot;Hudson's</normalizedToken>
Bay Territory&quot; (original citation), restricted to &quot;Gillam, Manit[oba]&quot; by Lindroth (1961a: 58). Holotype [by monotypy] (♂) in MCZ (collection LeConte). Synonymy established by Lindroth (1961a: 57).
</paragraph>
<paragraph id="A9E041069F4A07DB74BB5B159C7F194C" pageId="126" pageNumber="127">
<taxonomicName id="C576E657DA31DCA55046653FAA8C84C2" authorityName="Casey" authorityYear="1913" class="Insecta" family="Carabidae" genus="Pelophila" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="shermani">Pelophila shermani</taxonomicName>
Casey, 1913: 45. Type locality: &quot;West S[ain]t Modest[e], Labrador&quot; (original citation). Five syntypes [5 originally cited] in USNM [# 46844]. Synonymy established, under the name
<taxonomicName id="60C799D1CDEE26381198C1F86313A689" class="Insecta" family="Carabidae" genus="Pelophila" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="borealis" subSpecies="ulkei">
<emphasis id="E0CD37B273EB0EBF51D4C25109984611" italics="true" pageId="126" pageNumber="127">Pelophila borealis ulkei</emphasis>
</taxonomicName>
Horn, by
<normalizedToken id="59C0847E44C2A7029F14095A9F188D46" originalValue="Bänninger">Baenninger</normalizedToken>
(1930: 102), confirmed by Lindroth (1961a: 57). Etymology. The specific name was proposed for John Dempster Sherman [1872-1960], dealer of entomological and related books and periodicals.
</paragraph>
</subSubSection>
<subSubSection id="3DB236CD37E1ED87D90E2FEF3F2E59B3" pageId="126" pageNumber="127" type="distribution">
<paragraph id="17BA48B0768EDE95943C170F112CDB94" pageId="126" pageNumber="127">Distribution.</paragraph>
<paragraph id="D0EDD71689E7D8135B29F2B61765D25A" pageId="126" pageNumber="127">
This species is found from northern Europe to the Bering Sea coast (
<normalizedToken id="61B29BC55EE8AA9DCA7BF14674B385C4" originalValue="Farkač">Farkac</normalizedToken>
2003: 98) and from Alaska, including the Aleutian and Kodiak Islands, to Newfoundland, south to the Abitibi region in western Quebec (Larochelle 1975: 98) [see Lindroth 1963a: Fig. 59]. Fossil remnants of this species, dated between about 14,000 and 18,100 years B.P., have been unearthed in southeastern Iowa (Baker et al. 1986: 96; Schwert 1992: 76) and southern Ontario (Morgan and Morgan 1981: 1107).
</paragraph>
</subSubSection>
<subSubSection id="AC30174CEBDBFEB6D82AACBDE2D8B6B8" pageId="126" pageNumber="127" type="records">
<paragraph id="ACF245D23808C7280FCBA16134667355" pageId="126" pageNumber="127">Records.</paragraph>
<paragraph id="F7731F386A8126CE176DE5CAC0218126" pageId="126" pageNumber="127">
<emphasis id="3DEC15A8AA2BD2C417ED39155156438B" bold="true" pageId="126" pageNumber="127">CAN</emphasis>
: AB, BC, LB, MB, NF, NT, NU, ON, QC, SK, YT
<emphasis id="DAF6FF9FE184459A91D75B1ABC1F0C85" bold="true" pageId="126" pageNumber="127">USA</emphasis>
: AK -
<emphasis id="15E943B13EB7FDBC3911FF4303B83E74" bold="true" pageId="126" pageNumber="127">Holarctic</emphasis>
</paragraph>
<caption id="5686E5C26A6C6FB60A37EC07CE2B84B5" httpUri="https://binary.pensoft.net/fig/30907" pageId="126" pageNumber="127" start="Figure 6" startId="F6">
<paragraph id="3228744912620791E5F335301EE97614" pageId="126" pageNumber="127">
<emphasis id="0E3E086034D3AAE7D03F28A2096B47F0" bold="true" pageId="126" pageNumber="127">Figure 6.</emphasis>
<taxonomicName id="C50839E54F30155F69A7BA82F418A561" baseAuthorityName="Paykull" baseAuthorityYear="1790" class="Insecta" family="Carabidae" genus="Pelophila" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="212236E6AC505A748B95A1F8A58E1943" italics="true" pageId="126" pageNumber="127">Pelophila borealis</emphasis>
</taxonomicName>
(Paykull). This pelophiline is one of the 97 species-group taxa which are Holarctic and found naturally in both the Palaearctic and Nearctic Regions. Most of these species are confined to the arctic, subarctic or boreal regions but a few are found mainly in the temperate regions such as
<taxonomicName id="9BCBA2D6B47B48F43E47B1BD89886F40" baseAuthorityName="Dejean" baseAuthorityYear="1825" class="Hexapoda" family="Carabidae" genus="Dyschirius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="politus">
<emphasis id="B691F708A6CC0771621A3868498D781E" italics="true" pageId="126" pageNumber="127">Dyschirius politus</emphasis>
</taxonomicName>
. Most biogeographers agree that these species have spread between the two continents through Beringia during the Quaternary period.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>