360 lines
57 KiB
XML
360 lines
57 KiB
XML
<document id="2F6EAD23868612B7CF1F9B53F666D5C4" ID-DOI="10.11646/phytotaxa.153.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="5100682" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626239666706" checkinUser="felipe" docAuthor="GRAEFF, C. L., KOCIOLEK, J. P. & RUSHFORTH, S. R." docDate="2013" docId="546B8784F9663964FF3D7C48FEF4652A" docLanguage="en" docName="Phytotaxa.153.1.1-38.pdf" docOrigin="Phytotaxa 153 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.153.1.1" docStyle="DocumentStyle:13F189B31CB4CCCE9197361A4E100104.3:Phytotaxa.2011-2013.journal_article" docStyleId="13F189B31CB4CCCE9197361A4E100104" docStyleName="Phytotaxa.2011-2013.journal_article" docStyleVersion="3" docTitle="Ulnaria tooelensis Graeff, Kociolek & S. R. Rushforth 2013, sp. nov." docType="treatment" docVersion="3" lastPageNumber="12" masterDocId="A852FFFCF96E396FFFAA795FFFDE612E" masterDocTitle="New and Interesting Diatoms (Bacillariophyta) from Blue Lake Warm Springs, Tooele County, Utah" masterLastPageNumber="38" masterPageNumber="1" pageNumber="9" updateTime="1699007293075" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="51C7C4091A26D12E8F951AAC4E3CE50B">New and Interesting Diatoms (Bacillariophyta) from Blue Lake Warm Springs, Tooele County, Utah</mods:title>
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<mods:namePart id="9720C424FF1DC2458953CFF2DC209F33">GRAEFF, C. L.</mods:namePart>
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<mods:namePart id="BC894C8F0AA7E6ADD8D466989EE1C7E8">KOCIOLEK, J. P.</mods:namePart>
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<mods:namePart id="15A5D01EF7CBD40B2535957F577A8666">RUSHFORTH, S. R.</mods:namePart>
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<mods:title id="81EE81E1E6B0CA476022119564103DEC">Phytotaxa</mods:title>
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<mods:part id="1A0C4E838A1A372713F1DD03D769EF83">
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<mods:date id="61FD136E975B3D85C22268258F314361">2013</mods:date>
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<mods:number id="CF06AAA490FAD9E0C2BD79789FE48726">2013-12-13</mods:number>
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<mods:identifier id="AB0CE18F683519B736FE45866FB906C5" type="DOI">10.11646/phytotaxa.153.1.1</mods:identifier>
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<treatment id="546B8784F9663964FF3D7C48FEF4652A" LSID="urn:lsid:plazi:treatment:546B8784F9663964FF3D7C48FEF4652A" httpUri="http://treatment.plazi.org/id/546B8784F9663964FF3D7C48FEF4652A" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9">
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<subSubSection id="94D86519F9663967FF3D7C48FAF26716" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="DC7D3692F9663967FF3D7C48FAB2641D" blockId="8.[151,1436,1303,1816]" box="[151,1388,1303,1332]" pageId="8" pageNumber="9">
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<heading id="873581FEF9663967FF3D7C48FAB2641D" box="[151,1388,1303,1332]" fontSize="12" level="2" pageId="8" pageNumber="9" reason="4">
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<taxonomicName id="1BC24D11F9663967FF3D7C48FC98641D" authority="Graeff, Kociolek & S.R. Rushforth" authorityName="Graeff, Kociolek & S. R. Rushforth" authorityYear="2013" box="[151,838,1303,1332]" class="Bacillariophyceae" family="Fragilariaceae" genus="Ulnaria" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="8" pageNumber="9" phylum="Ochrophyta" rank="species" species="tooelensis" status="sp. nov.">
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<emphasis id="EEB6EA80F9663967FF3D7C48FE5D641A" bold="true" box="[151,387,1303,1332]" italics="true" pageId="8" pageNumber="9">Ulnaria tooelensis</emphasis>
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Graeff, Kociolek & S.R. Rushforth
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</taxonomicName>
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,
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<emphasis id="EEB6EA80F9663967FCF87C47FC6C641A" box="[850,946,1304,1332]" italics="true" pageId="8" pageNumber="9">
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<taxonomicNameLabel id="F58557FBF9663967FCF87C47FC6C641A" box="[850,946,1304,1332]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="44F92A17F9663967FC697C48FB8A641D" box="[963,1108,1303,1331]" captionStart-0="FIGURES 38–48" captionStart-1="FIGURES 49–55" captionStartId-0="9.[151,259,1920,1941]" captionStartId-1="10.[151,259,1715,1736]" captionTargetBox-0="[173,1413,197,1895]" captionTargetBox-1="[169,1417,491,1693]" captionTargetId-0="figure-73@9.[173,1413,197,1895]" captionTargetId-1="figure-303@10.[169,1417,491,1693]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 38–48. LM. Ulnaria tooelensis. Figs 38–46, Valve views showing the size diminution series. Figure 40 is the holotype. Fig. 47, Girdle view showing a cingulum with at least two closed bands, each with a single row of poroids. Fig. 48, Closed girdle band. Scale bars: 10 µm." captionText-1="FIGURES 49–55. SEM. Ulnaria tooelensis. Internal and external valve views of five specimens. Fig. 49, Full external valve view showing rimoportula openings at each end. Fig. 50, Full external valve showing unpaired striae. Fig. 51, Apex showing small, round, depressed external opening of the rimoportula and the ocellimbus. Fig. 52, Large external central area with slightly thickened interstriae and small, unoccluded areolae. Fig. 53, Full internal view showing rimoportulae at each apex. Fig. 54, Small, rounded rimoportula and well-defined apical pore field which extends over a larger area than the external ocellimbus. Fig. 55, Internal view of the central area with unoccluded areolae. Scale bars: Figs 49–50, 53: 10 µm; Figs. 51–52, 54–55: 1 µm." figureDoi-0="http://doi.org/10.5281/zenodo.5100696" figureDoi-1="http://doi.org/10.5281/zenodo.5100700" httpUri-0="https://zenodo.org/record/5100696/files/figure.png" httpUri-1="https://zenodo.org/record/5100700/files/figure.png" pageId="8" pageNumber="9">Figs 38–55</figureCitation>
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; figure 40 =
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<typeStatus id="03798830F9663967FB5E7C48FABA641D" box="[1268,1380,1303,1331]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
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)
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</heading>
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</paragraph>
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<paragraph id="DC7D3692F9663967FF3D7C1EFAF26716" blockId="8.[151,1436,1303,1816]" pageId="8" pageNumber="9">
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Valves linear to linear-elliptical in smaller specimens, tumid on one side of the central area, with apices subcapitate in larger specimens and protracted in smaller specimens (
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<figureCitation id="44F92A17F9663967FC8D7C3EFC766456" box="[807,936,1377,1400]" captionStart="FIGURES 38–48" captionStartId="9.[151,259,1920,1941]" captionTargetBox="[173,1413,197,1895]" captionTargetId="figure-73@9.[173,1413,197,1895]" captionTargetPageId="9" captionText="FIGURES 38–48. LM. Ulnaria tooelensis. Figs 38–46, Valve views showing the size diminution series. Figure 40 is the holotype. Fig. 47, Girdle view showing a cingulum with at least two closed bands, each with a single row of poroids. Fig. 48, Closed girdle band. Scale bars: 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5100696" httpUri="https://zenodo.org/record/5100696/files/figure.png" pageId="8" pageNumber="9">Figs 38–46</figureCitation>
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). Length 14.0–
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<quantity id="1B3A9B77F9663967FBE57C3EFB716456" box="[1103,1199,1377,1400]" metricMagnitude="-2" metricUnit="m" metricValue="9.75" pageId="8" pageNumber="9" unit="mm" value="97.5">97.5 µm</quantity>
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, breadth
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<quantity id="1B3A9B77F9663967FABD7C3EFA466456" box="[1303,1432,1377,1400]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.5" metricValueMin="3.5" pageId="8" pageNumber="9" unit="mm" value="4.5" valueMax="5.5" valueMin="3.5">3.5–5.5 µm</quantity>
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. Central sternum very narrow, straight, widening unilaterally at the center to form a rectangular central area on one side (
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<figureCitation id="44F92A17F9663967FEAE7CFEFE566496" box="[260,392,1441,1464]" captionStart="FIGURES 38–48" captionStartId="9.[151,259,1920,1941]" captionTargetBox="[173,1413,197,1895]" captionTargetId="figure-73@9.[173,1413,197,1895]" captionTargetPageId="9" captionText="FIGURES 38–48. LM. Ulnaria tooelensis. Figs 38–46, Valve views showing the size diminution series. Figure 40 is the holotype. Fig. 47, Girdle view showing a cingulum with at least two closed bands, each with a single row of poroids. Fig. 48, Closed girdle band. Scale bars: 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5100696" httpUri="https://zenodo.org/record/5100696/files/figure.png" pageId="8" pageNumber="9">Figs 40, 43</figureCitation>
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). Striae punctate, more or less opposite and parallel, infrequently with a stray stria without a corresponding stria on the opposite side of the central sternum (
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<figureCitation id="44F92A17F9663967FC237C9EFBD564F6" box="[905,1035,1473,1496]" captionStart="FIGURES 38–48" captionStartId="9.[151,259,1920,1941]" captionTargetBox="[173,1413,197,1895]" captionTargetId="figure-73@9.[173,1413,197,1895]" captionTargetPageId="9" captionText="FIGURES 38–48. LM. Ulnaria tooelensis. Figs 38–46, Valve views showing the size diminution series. Figure 40 is the holotype. Fig. 47, Girdle view showing a cingulum with at least two closed bands, each with a single row of poroids. Fig. 48, Closed girdle band. Scale bars: 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5100696" httpUri="https://zenodo.org/record/5100696/files/figure.png" pageId="8" pageNumber="9">Figs 41, 44</figureCitation>
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). Striae number 11–15/
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<quantity id="1B3A9B77F9663967FAA67C9EFA8A64F6" box="[1292,1364,1473,1496]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="9" unit="mm" value="10.0">10 µm</quantity>
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in the center of the valve, 13–16/
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<quantity id="1B3A9B77F9663967FE4D7CBEFDF364D6" box="[487,557,1505,1528]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="9" unit="mm" value="10.0">10 µm</quantity>
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at the ends. A labiate process (rimoportula) is positioned at each end of the valve (
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<figureCitation id="44F92A17F9663967FF647F5EFE8E6736" box="[206,336,1537,1560]" captionStart="FIGURES 38–48" captionStartId="9.[151,259,1920,1941]" captionTargetBox="[173,1413,197,1895]" captionTargetId="figure-73@9.[173,1413,197,1895]" captionTargetPageId="9" captionText="FIGURES 38–48. LM. Ulnaria tooelensis. Figs 38–46, Valve views showing the size diminution series. Figure 40 is the holotype. Fig. 47, Girdle view showing a cingulum with at least two closed bands, each with a single row of poroids. Fig. 48, Closed girdle band. Scale bars: 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5100696" httpUri="https://zenodo.org/record/5100696/files/figure.png" pageId="8" pageNumber="9">Figs 40–41</figureCitation>
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). A pore field is present at the very terminus of the valve on each end (
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<figureCitation id="44F92A17F9663967FB2D7F5EFA836736" box="[1159,1373,1537,1560]" captionStart="FIGURES 38–48" captionStartId="9.[151,259,1920,1941]" captionTargetBox="[173,1413,197,1895]" captionTargetId="figure-73@9.[173,1413,197,1895]" captionTargetPageId="9" captionText="FIGURES 38–48. LM. Ulnaria tooelensis. Figs 38–46, Valve views showing the size diminution series. Figure 40 is the holotype. Fig. 47, Girdle view showing a cingulum with at least two closed bands, each with a single row of poroids. Fig. 48, Closed girdle band. Scale bars: 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5100696" httpUri="https://zenodo.org/record/5100696/files/figure.png" pageId="8" pageNumber="9">Figs 39–41, 45–46</figureCitation>
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). The cingulum is comprised of at least two closed girdle bands each with a single row of poroids (
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<figureCitation id="44F92A17F9663967FB097F7EFAFE6716" box="[1187,1312,1569,1592]" captionStart="FIGURES 38–48" captionStartId="9.[151,259,1920,1941]" captionTargetBox="[173,1413,197,1895]" captionTargetId="figure-73@9.[173,1413,197,1895]" captionTargetPageId="9" captionText="FIGURES 38–48. LM. Ulnaria tooelensis. Figs 38–46, Valve views showing the size diminution series. Figure 40 is the holotype. Fig. 47, Girdle view showing a cingulum with at least two closed bands, each with a single row of poroids. Fig. 48, Closed girdle band. Scale bars: 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5100696" httpUri="https://zenodo.org/record/5100696/files/figure.png" pageId="8" pageNumber="9">Figs 47–48</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="94D86519F9663967FF3D7F1EFCB16636" pageId="8" pageNumber="9" type="description">
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<paragraph id="DC7D3692F9663967FF3D7F1EFCB16636" blockId="8.[151,1436,1303,1816]" pageId="8" pageNumber="9">
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In the SEM, the valve has small, rounded, unoccluded areolae comprising the striae (
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<figureCitation id="44F92A17F9663967FB837F1EFB296776" box="[1065,1271,1601,1624]" captionStart="FIGURES 49–55" captionStartId="10.[151,259,1715,1736]" captionTargetBox="[169,1417,491,1693]" captionTargetId="figure-303@10.[169,1417,491,1693]" captionTargetPageId="10" captionText="FIGURES 49–55. SEM. Ulnaria tooelensis. Internal and external valve views of five specimens. Fig. 49, Full external valve view showing rimoportula openings at each end. Fig. 50, Full external valve showing unpaired striae. Fig. 51, Apex showing small, round, depressed external opening of the rimoportula and the ocellimbus. Fig. 52, Large external central area with slightly thickened interstriae and small, unoccluded areolae. Fig. 53, Full internal view showing rimoportulae at each apex. Fig. 54, Small, rounded rimoportula and well-defined apical pore field which extends over a larger area than the external ocellimbus. Fig. 55, Internal view of the central area with unoccluded areolae. Scale bars: Figs 49–50, 53: 10 µm; Figs. 51–52, 54–55: 1 µm." figureDoi="http://doi.org/10.5281/zenodo.5100700" httpUri="https://zenodo.org/record/5100700/files/figure.png" pageId="8" pageNumber="9">Figs 51–52, 54–55</figureCitation>
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). The striae are more or less aligned, though that arrangement breaks down in places along the valve (
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<figureCitation id="44F92A17F9663967FBF47F3EFB026756" box="[1118,1244,1633,1656]" captionStart="FIGURES 49–55" captionStartId="10.[151,259,1715,1736]" captionTargetBox="[169,1417,491,1693]" captionTargetId="figure-303@10.[169,1417,491,1693]" captionTargetPageId="10" captionText="FIGURES 49–55. SEM. Ulnaria tooelensis. Internal and external valve views of five specimens. Fig. 49, Full external valve view showing rimoportula openings at each end. Fig. 50, Full external valve showing unpaired striae. Fig. 51, Apex showing small, round, depressed external opening of the rimoportula and the ocellimbus. Fig. 52, Large external central area with slightly thickened interstriae and small, unoccluded areolae. Fig. 53, Full internal view showing rimoportulae at each apex. Fig. 54, Small, rounded rimoportula and well-defined apical pore field which extends over a larger area than the external ocellimbus. Fig. 55, Internal view of the central area with unoccluded areolae. Scale bars: Figs 49–50, 53: 10 µm; Figs. 51–52, 54–55: 1 µm." figureDoi="http://doi.org/10.5281/zenodo.5100700" httpUri="https://zenodo.org/record/5100700/files/figure.png" pageId="8" pageNumber="9">Figs 49–50</figureCitation>
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). The rimoportula opening is round, similar in size to the areolae, but located within a small depression (
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<figureCitation id="44F92A17F9663967FBFC7FDEFB7867B6" box="[1110,1190,1665,1688]" captionStart="FIGURES 49–55" captionStartId="10.[151,259,1715,1736]" captionTargetBox="[169,1417,491,1693]" captionTargetId="figure-303@10.[169,1417,491,1693]" captionTargetPageId="10" captionText="FIGURES 49–55. SEM. Ulnaria tooelensis. Internal and external valve views of five specimens. Fig. 49, Full external valve view showing rimoportula openings at each end. Fig. 50, Full external valve showing unpaired striae. Fig. 51, Apex showing small, round, depressed external opening of the rimoportula and the ocellimbus. Fig. 52, Large external central area with slightly thickened interstriae and small, unoccluded areolae. Fig. 53, Full internal view showing rimoportulae at each apex. Fig. 54, Small, rounded rimoportula and well-defined apical pore field which extends over a larger area than the external ocellimbus. Fig. 55, Internal view of the central area with unoccluded areolae. Scale bars: Figs 49–50, 53: 10 µm; Figs. 51–52, 54–55: 1 µm." figureDoi="http://doi.org/10.5281/zenodo.5100700" httpUri="https://zenodo.org/record/5100700/files/figure.png" pageId="8" pageNumber="9">Fig. 51</figureCitation>
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). The unilateral central area has some thickened interstriae present (
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<figureCitation id="44F92A17F9663967FD357FFEFD316796" box="[671,751,1697,1720]" captionStart="FIGURES 49–55" captionStartId="10.[151,259,1715,1736]" captionTargetBox="[169,1417,491,1693]" captionTargetId="figure-303@10.[169,1417,491,1693]" captionTargetPageId="10" captionText="FIGURES 49–55. SEM. Ulnaria tooelensis. Internal and external valve views of five specimens. Fig. 49, Full external valve view showing rimoportula openings at each end. Fig. 50, Full external valve showing unpaired striae. Fig. 51, Apex showing small, round, depressed external opening of the rimoportula and the ocellimbus. Fig. 52, Large external central area with slightly thickened interstriae and small, unoccluded areolae. Fig. 53, Full internal view showing rimoportulae at each apex. Fig. 54, Small, rounded rimoportula and well-defined apical pore field which extends over a larger area than the external ocellimbus. Fig. 55, Internal view of the central area with unoccluded areolae. Scale bars: Figs 49–50, 53: 10 µm; Figs. 51–52, 54–55: 1 µm." figureDoi="http://doi.org/10.5281/zenodo.5100700" httpUri="https://zenodo.org/record/5100700/files/figure.png" pageId="8" pageNumber="9">Fig. 52</figureCitation>
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). Ocellimbi are evident on the margin at the poles (
|
||
<figureCitation id="44F92A17F9663967FABE7FFEFA4E6796" box="[1300,1424,1697,1720]" captionStart="FIGURES 49–55" captionStartId="10.[151,259,1715,1736]" captionTargetBox="[169,1417,491,1693]" captionTargetId="figure-303@10.[169,1417,491,1693]" captionTargetPageId="10" captionText="FIGURES 49–55. SEM. Ulnaria tooelensis. Internal and external valve views of five specimens. Fig. 49, Full external valve view showing rimoportula openings at each end. Fig. 50, Full external valve showing unpaired striae. Fig. 51, Apex showing small, round, depressed external opening of the rimoportula and the ocellimbus. Fig. 52, Large external central area with slightly thickened interstriae and small, unoccluded areolae. Fig. 53, Full internal view showing rimoportulae at each apex. Fig. 54, Small, rounded rimoportula and well-defined apical pore field which extends over a larger area than the external ocellimbus. Fig. 55, Internal view of the central area with unoccluded areolae. Scale bars: Figs 49–50, 53: 10 µm; Figs. 51–52, 54–55: 1 µm." figureDoi="http://doi.org/10.5281/zenodo.5100700" httpUri="https://zenodo.org/record/5100700/files/figure.png" pageId="8" pageNumber="9">Figs 49–51</figureCitation>
|
||
). Internally, the unornamented central area is evident, and the central sternum is narrow (
|
||
<figureCitation id="44F92A17F9663967FBCB7F9EFB0267F6" box="[1121,1244,1729,1752]" captionStart="FIGURES 49–55" captionStartId="10.[151,259,1715,1736]" captionTargetBox="[169,1417,491,1693]" captionTargetId="figure-303@10.[169,1417,491,1693]" captionTargetPageId="10" captionText="FIGURES 49–55. SEM. Ulnaria tooelensis. Internal and external valve views of five specimens. Fig. 49, Full external valve view showing rimoportula openings at each end. Fig. 50, Full external valve showing unpaired striae. Fig. 51, Apex showing small, round, depressed external opening of the rimoportula and the ocellimbus. Fig. 52, Large external central area with slightly thickened interstriae and small, unoccluded areolae. Fig. 53, Full internal view showing rimoportulae at each apex. Fig. 54, Small, rounded rimoportula and well-defined apical pore field which extends over a larger area than the external ocellimbus. Fig. 55, Internal view of the central area with unoccluded areolae. Scale bars: Figs 49–50, 53: 10 µm; Figs. 51–52, 54–55: 1 µm." figureDoi="http://doi.org/10.5281/zenodo.5100700" httpUri="https://zenodo.org/record/5100700/files/figure.png" pageId="8" pageNumber="9">Figs 53, 55</figureCitation>
|
||
). The rimoportula is small, elliptical and bordered by narrow “lips” (
|
||
<figureCitation id="44F92A17F9663967FD5F7FBEFC9667D6" box="[757,840,1761,1784]" captionStart="FIGURES 49–55" captionStartId="10.[151,259,1715,1736]" captionTargetBox="[169,1417,491,1693]" captionTargetId="figure-303@10.[169,1417,491,1693]" captionTargetPageId="10" captionText="FIGURES 49–55. SEM. Ulnaria tooelensis. Internal and external valve views of five specimens. Fig. 49, Full external valve view showing rimoportula openings at each end. Fig. 50, Full external valve showing unpaired striae. Fig. 51, Apex showing small, round, depressed external opening of the rimoportula and the ocellimbus. Fig. 52, Large external central area with slightly thickened interstriae and small, unoccluded areolae. Fig. 53, Full internal view showing rimoportulae at each apex. Fig. 54, Small, rounded rimoportula and well-defined apical pore field which extends over a larger area than the external ocellimbus. Fig. 55, Internal view of the central area with unoccluded areolae. Scale bars: Figs 49–50, 53: 10 µm; Figs. 51–52, 54–55: 1 µm." figureDoi="http://doi.org/10.5281/zenodo.5100700" httpUri="https://zenodo.org/record/5100700/files/figure.png" pageId="8" pageNumber="9">Fig. 54</figureCitation>
|
||
). Internally, the ocellimbi are distinct with a fine grid pattern positioned at the valve terminus on the mantle (
|
||
<figureCitation id="44F92A17F9663967FCB87E5EFCBC6636" box="[786,866,1793,1816]" captionStart="FIGURES 49–55" captionStartId="10.[151,259,1715,1736]" captionTargetBox="[169,1417,491,1693]" captionTargetId="figure-303@10.[169,1417,491,1693]" captionTargetPageId="10" captionText="FIGURES 49–55. SEM. Ulnaria tooelensis. Internal and external valve views of five specimens. Fig. 49, Full external valve view showing rimoportula openings at each end. Fig. 50, Full external valve showing unpaired striae. Fig. 51, Apex showing small, round, depressed external opening of the rimoportula and the ocellimbus. Fig. 52, Large external central area with slightly thickened interstriae and small, unoccluded areolae. Fig. 53, Full internal view showing rimoportulae at each apex. Fig. 54, Small, rounded rimoportula and well-defined apical pore field which extends over a larger area than the external ocellimbus. Fig. 55, Internal view of the central area with unoccluded areolae. Scale bars: Figs 49–50, 53: 10 µm; Figs. 51–52, 54–55: 1 µm." figureDoi="http://doi.org/10.5281/zenodo.5100700" httpUri="https://zenodo.org/record/5100700/files/figure.png" pageId="8" pageNumber="9">Fig. 54</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="94D86519F9663967FF3D7E1EFE3B66AC" pageId="8" pageNumber="9" type="materials_examined">
|
||
<paragraph id="DC7D3692F9663967FF3D7E1EFE3B66AC" blockId="8.[151,1436,1857,1999]" pageId="8" pageNumber="9">
|
||
<emphasis id="EEB6EA80F9663967FF3D7E1EFF0A6675" bold="true" box="[151,212,1857,1883]" pageId="8" pageNumber="9">Type</emphasis>
|
||
:—
|
||
<materialsCitation id="6CAA3CCFF9663967FF537E1EFE3B66AC" accessionNumber="BM 101677" collectionCode="BM" country="United States of America" county="Tooele Co." location="Blue Lake" pageId="8" pageNumber="9" specimenCode="COLO 439033, COLO 8505" specimenCount="1" stateProvince="Utah" typeStatus="holotype">
|
||
<collectingCountry id="A4D57602F9663967FF537E1EFEE76675" box="[249,313,1857,1883]" name="United States of America" pageId="8" pageNumber="9">USA</collectingCountry>
|
||
.
|
||
<location id="D91D6049F9663967FEEE7E1EFE1C6675" LSID="urn:lsid:plazi:treatment:546B8784F9663964FF3D7C48FEF4652A:D91D6049F9663967FEEE7E1EFE1C6675" box="[324,450,1857,1883]" country="United States of America" county="Tooele Co." name="Blue Lake" pageId="8" pageNumber="9" stateProvince="Utah">Blue Lake</location>
|
||
,
|
||
<collectingCounty id="351C4E1EF9663967FE647E1EFD8C6675" box="[462,594,1857,1883]" pageId="8" pageNumber="9">Tooele Co.</collectingCounty>
|
||
,
|
||
<collectingRegion id="1E06F870F9663967FDF57E1EFD426675" box="[607,668,1857,1883]" country="United States of America" name="Utah" pageId="8" pageNumber="9">Utah</collectingRegion>
|
||
. (
|
||
<specimenCode id="8C649EE9F9663967FD1A7E1EFCBB6675" box="[688,869,1857,1883]" collectionCode="COLO" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15540" lsid="urn:lsid:biocol.org:col:15540" name="University of Colorado Herbarium" pageId="8" pageNumber="9" type="Herbarium">COLO 439033</specimenCode>
|
||
,
|
||
<typeStatus id="03798830F9663967FCDB7E1EFC016675" box="[881,991,1857,1883]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
! (=
|
||
<figureCitation id="44F92A17F9663967FBBF7E1EFBB36675" box="[1045,1133,1857,1883]" captionStart="FIGURES 38–48" captionStartId="9.[151,259,1920,1941]" captionTargetBox="[173,1413,197,1895]" captionTargetId="figure-73@9.[173,1413,197,1895]" captionTargetPageId="9" captionText="FIGURES 38–48. LM. Ulnaria tooelensis. Figs 38–46, Valve views showing the size diminution series. Figure 40 is the holotype. Fig. 47, Girdle view showing a cingulum with at least two closed bands, each with a single row of poroids. Fig. 48, Closed girdle band. Scale bars: 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5100696" httpUri="https://zenodo.org/record/5100696/files/figure.png" pageId="8" pageNumber="9">Fig. 40</figureCitation>
|
||
), designated here;
|
||
<specimenCode id="8C649EE9F9663967FAE17E1EFF0A66AC" collectionCode="COLO" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15540" lsid="urn:lsid:biocol.org:col:15540" name="University of Colorado Herbarium" pageId="8" pageNumber="9" type="Herbarium">COLO 8505</specimenCode>
|
||
,
|
||
<collectionCode id="BAD3AE57F9663967FF4A7E37FED066AC" box="[224,270,1896,1922]" country="United Kingdom" name="Bristol Museum" pageId="8" pageNumber="9">BM</collectionCode>
|
||
101677, isotypes)
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="94D86519F9663967FF6C7ED1FCB86686" box="[198,870,1934,1960]" pageId="8" pageNumber="9" type="etymology">
|
||
<paragraph id="DC7D3692F9663967FF6C7ED1FCB86686" blockId="8.[151,1436,1857,1999]" box="[198,870,1934,1960]" pageId="8" pageNumber="9">
|
||
<emphasis id="EEB6EA80F9663967FF6C7ED1FE936686" bold="true" box="[198,333,1934,1960]" pageId="8" pageNumber="9">Etymology</emphasis>
|
||
:—Named for the county in which it is found.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="94D86519F9663964FF6C7EEAFEF4652A" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9" type="distribution">
|
||
<paragraph id="DC7D3692F9663967FF6C7EEAFA4266E1" blockId="8.[151,1436,1857,1999]" box="[198,1436,1973,1999]" pageId="8" pageNumber="9">
|
||
<emphasis id="EEB6EA80F9663967FF6C7EEAFE8166E1" bold="true" box="[198,351,1973,1999]" pageId="8" pageNumber="9">Distribution</emphasis>
|
||
:—
|
||
<taxonomicName id="1BC24D11F9663967FE2E7EEAFD8066E1" authorityName="Graeff, Kociolek & S. R. Rushforth" authorityYear="2013" box="[388,606,1973,1999]" class="Bacillariophyceae" family="Fragilariaceae" genus="Ulnaria" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="8" pageNumber="9" phylum="Ochrophyta" rank="species" species="tooelensis">
|
||
<emphasis id="EEB6EA80F9663967FE2E7EEAFD8066E1" box="[388,606,1973,1999]" italics="true" pageId="8" pageNumber="9">Ulnaria tooelensis</emphasis>
|
||
</taxonomicName>
|
||
is common in the main basin of Blue Lake and the surrounding marsh.
|
||
</paragraph>
|
||
<caption id="88BD661AF9673966FF3D7EDFFE9466FE" ID-DOI="http://doi.org/10.5281/zenodo.5100696" ID-Zenodo-Dep="5100696" httpUri="https://zenodo.org/record/5100696/files/figure.png" pageId="9" pageNumber="10" startId="9.[151,259,1920,1941]" targetBox="[173,1413,197,1895]" targetPageId="9">
|
||
<paragraph id="DC7D3692F9673966FF3D7EDFFE9466FE" blockId="9.[151,1437,1920,2000]" pageId="9" pageNumber="10">
|
||
<emphasis id="EEB6EA80F9673966FF3D7EDFFE9866BB" bold="true" box="[151,326,1920,1941]" pageId="9" pageNumber="10">FIGURES 38–48</emphasis>
|
||
. LM.
|
||
<taxonomicName id="1BC24D11F9673966FE2E7EDFFDE266B8" authorityName="Graeff, Kociolek & S. R. Rushforth" authorityYear="2013" box="[388,572,1920,1942]" class="Bacillariophyceae" family="Fragilariaceae" genus="Ulnaria" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="9" pageNumber="10" phylum="Ochrophyta" rank="species" species="tooelensis">
|
||
<emphasis id="EEB6EA80F9673966FE2E7EDFFDE266B8" bold="true" box="[388,572,1920,1942]" italics="true" pageId="9" pageNumber="10">Ulnaria tooelensis</emphasis>
|
||
</taxonomicName>
|
||
. Figs 38–46, Valve views showing the size diminution series. Figure 40 is the holotype. Fig. 47, Girdle view showing a cingulum with at least two closed bands, each with a single row of poroids. Fig. 48, Closed girdle band. Scale bars: 10 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DC7D3692F9643965FF6C79C7FD2960B4" blockId="10.[151,1436,152,410]" pageId="10" pageNumber="11">
|
||
<emphasis id="EEB6EA80F9643965FF6C79C7FEB2619C" bold="true" box="[198,364,152,178]" pageId="10" pageNumber="11">Observations</emphasis>
|
||
:—This diatom was identified by Kaczmarska and Rushforth (1983) as
|
||
<taxonomicName id="1BC24D11F9643965FA8A79C7FD4D61F6" authority="(Kutzing) Petersen (1938: 167)" authorityName="Petersen" authorityPageNumber="167" authorityYear="1938" baseAuthorityName="Kutzing" class="Bacillariophyceae" family="Fragilariaceae" genus="Fragilaria" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="species" species="vaucheriae">
|
||
<emphasis id="EEB6EA80F9643965FA8A79C7FEC761F6" italics="true" pageId="10" pageNumber="11">Fragilaria vaucheriae</emphasis>
|
||
(Kützing)
|
||
<bibRefCitation id="B8534B63F9643965FE3179E1FD4D61F6" author="Petersen, J. B." box="[411,659,190,216]" pageId="10" pageNumber="11" pagination="164 - 170" refId="ref18896" refString="Petersen, J. B. (1938) Fragilaria intermedia-Synedra vaucheriae? Botaniska Notiser 1938: 164 - 170." type="journal article" year="1938">Petersen (1938: 167)</bibRefCitation>
|
||
</taxonomicName>
|
||
. These two diatoms are morphologically distinct, as illustrated by Krammer & Lange-Bertalot (1991: pl. 108, figs 10–15). They examined the
|
||
<typeStatus id="03798830F9643965FBC879BAFB4961D1" box="[1122,1175,229,255]" pageId="10" pageNumber="11">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="1BC24D11F9643965FA9F79BAFDDD6008" authority="Kutzing (1833: 32)" authorityName="Kutzing" authorityPageNumber="32" authorityYear="1833" class="Bacillariophyceae" family="Licmophoraceae" genus="Exilaria" higherTaxonomySource="GBIF" kingdom="Chromista" order="Licmophorales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="species" species="vaucheriae">
|
||
<emphasis id="EEB6EA80F9643965FA9F79BAFEC76008" italics="true" pageId="10" pageNumber="11">Exilaria vaucheriae</emphasis>
|
||
<bibRefCitation id="B8534B63F9643965FE8A7853FDDD6008" author="Kutzing, F. T." box="[288,515,268,294]" pageId="10" pageNumber="11" pagination="1 - 4" refId="ref17830" refString="Kutzing, F. T. (1833) Algarum aquae dulcis Germanicarum. Decas I. Collegit Fridericus Traugott Kutzing, Cocit Botanischen. Ratisbon Sodalis. Halis Saxonum in Commissis C. A. Schwetschkii et Fil. 1: 1 - 4." type="journal article" year="1833">Kützing (1833: 32)</bibRefCitation>
|
||
</taxonomicName>
|
||
, the basionym of
|
||
<taxonomicName id="1BC24D11F9643965FD797853FCAC6008" authorityName="Kutzing" authorityYear="1833" box="[723,882,268,294]" class="Bacillariophyceae" family="Licmophoraceae" genus="Exilaria" higherTaxonomySource="GBIF" kingdom="Chromista" order="Licmophorales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="species" species="vaucheriae">
|
||
<emphasis id="EEB6EA80F9643965FD797853FCAC6008" box="[723,882,268,294]" italics="true" pageId="10" pageNumber="11">F. vaucheriae</emphasis>
|
||
</taxonomicName>
|
||
. The valve margins of
|
||
<taxonomicName id="1BC24D11F9643965FB2B7853FA846008" authorityName="Graeff, Kociolek & S. R. Rushforth" authorityYear="2013" box="[1153,1370,268,294]" class="Bacillariophyceae" family="Fragilariaceae" genus="Ulnaria" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="species" species="tooelensis">
|
||
<emphasis id="EEB6EA80F9643965FB2B7853FA846008" box="[1153,1370,268,294]" italics="true" pageId="10" pageNumber="11">Ulnaria tooelensis</emphasis>
|
||
</taxonomicName>
|
||
taper gradually toward the apices while they are relatively parallel in
|
||
<emphasis id="EEB6EA80F9643965FC2E786DFBF56062" box="[900,1067,306,332]" italics="true" pageId="10" pageNumber="11">
|
||
<taxonomicName id="1BC24D11F9643965FC2E786DFBF96062" authorityName="Kutzing" authorityYear="1833" box="[900,1063,306,332]" class="Bacillariophyceae" family="Licmophoraceae" genus="Exilaria" higherTaxonomySource="GBIF" kingdom="Chromista" order="Licmophorales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="species" species="vaucheriae">F. vaucheriae</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The apices of
|
||
<taxonomicName id="1BC24D11F9643965FB7D786DFAAF6062" authorityName="Graeff, Kociolek & S. R. Rushforth" authorityYear="2013" box="[1239,1393,306,332]" class="Bacillariophyceae" family="Fragilariaceae" genus="Ulnaria" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="species" species="tooelensis">
|
||
<emphasis id="EEB6EA80F9643965FB7D786DFAAF6062" box="[1239,1393,306,332]" italics="true" pageId="10" pageNumber="11">U. tooelensis</emphasis>
|
||
</taxonomicName>
|
||
are broader than those in in
|
||
<taxonomicName id="1BC24D11F9643965FE157806FDBD605D" authorityName="Kutzing" authorityYear="1833" box="[447,611,345,371]" class="Bacillariophyceae" family="Licmophoraceae" genus="Exilaria" higherTaxonomySource="GBIF" kingdom="Chromista" order="Licmophorales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="species" species="vaucheriae">
|
||
<emphasis id="EEB6EA80F9643965FE157806FDBD605D" box="[447,611,345,371]" italics="true" pageId="10" pageNumber="11">F. vaucheriae</emphasis>
|
||
</taxonomicName>
|
||
. However, the valve breadths and stria densities of the two taxa are congruent (Krammer & Lange–Bertalot 1991: 124).
|
||
</paragraph>
|
||
<caption id="88BD661AF9643965FF3D7FECFBA66674" ID-DOI="http://doi.org/10.5281/zenodo.5100700" ID-Zenodo-Dep="5100700" httpUri="https://zenodo.org/record/5100700/files/figure.png" pageId="10" pageNumber="11" startId="10.[151,259,1715,1736]" targetBox="[169,1417,491,1693]" targetPageId="10">
|
||
<paragraph id="DC7D3692F9643965FF3D7FECFBA66674" blockId="10.[151,1437,1714,1882]" pageId="10" pageNumber="11">
|
||
<emphasis id="EEB6EA80F9643965FF3D7FECFE9867E6" bold="true" box="[151,326,1715,1736]" pageId="10" pageNumber="11">FIGURES 49–55</emphasis>
|
||
. SEM.
|
||
<taxonomicName id="1BC24D11F9643965FE3B7FEDFD9767E6" authorityName="Graeff, Kociolek & S. R. Rushforth" authorityYear="2013" box="[401,585,1714,1736]" class="Bacillariophyceae" family="Fragilariaceae" genus="Ulnaria" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="species" species="tooelensis">
|
||
<emphasis id="EEB6EA80F9643965FE3B7FEDFD9767E6" bold="true" box="[401,585,1714,1736]" italics="true" pageId="10" pageNumber="11">Ulnaria tooelensis</emphasis>
|
||
</taxonomicName>
|
||
. Internal and external valve views of five specimens. Fig. 49, Full external valve view showing rimoportula openings at each end. Fig. 50, Full external valve showing unpaired striae. Fig. 51, Apex showing small, round, depressed external opening of the rimoportula and the ocellimbus. Fig. 52, Large external central area with slightly thickened interstriae and small, unoccluded areolae. Fig. 53, Full internal view showing rimoportulae at each apex. Fig. 54, Small, rounded rimoportula and well-defined apical pore field which extends over a larger area than the external ocellimbus. Fig. 55, Internal view of the central area with unoccluded areolae. Scale bars: Figs 49–50, 53: 10 µm; Figs. 51–52, 54–55: 1 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DC7D3692F9643964FF6C7ED5FCC260C9" blockId="10.[151,1436,1930,2033]" lastBlockId="11.[151,1436,152,1028]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
|
||
Other species that appear similar to
|
||
<taxonomicName id="1BC24D11F9643965FD297ED5FCC1668A" authorityName="Graeff, Kociolek & S. R. Rushforth" authorityYear="2013" box="[643,799,1930,1956]" class="Bacillariophyceae" family="Fragilariaceae" genus="Ulnaria" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="species" species="tooelensis">
|
||
<emphasis id="EEB6EA80F9643965FD297ED5FCC1668A" box="[643,799,1930,1956]" italics="true" pageId="10" pageNumber="11">U. tooelensis</emphasis>
|
||
</taxonomicName>
|
||
include
|
||
<taxonomicName id="1BC24D11F9643965FC267ED5FB01668A" authority="Grunow" authorityName="Grunow" authorityYear="1881" box="[908,1247,1930,1956]" class="Bacillariophyceae" family="Fragilariaceae" genus="Synedra" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="species" species="capitellata">
|
||
<emphasis id="EEB6EA80F9643965FC267ED5FBAA668A" box="[908,1140,1930,1956]" italics="true" pageId="10" pageNumber="11">Synedra capitellata</emphasis>
|
||
Grunow
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="B8534B63F9643965FAA67ED5FF0166E4" author="Van Heurck, H." pageId="10" pageNumber="11" pagination="31 - 77" refId="ref20141" refString="Van Heurck, H. (1881) Synopsis des Diatomees de Belgique. Atlas. Ducaju & Cie., Anvers, pls. 31 - 77." type="book chapter" year="1881">Van Heurck (1881</bibRefCitation>
|
||
: plate 40, fig. 26),
|
||
<taxonomicName id="1BC24D11F9643965FE127EEFFCCC66E4" authority="Grunow" authorityName="Grunow" authorityYear="1881" box="[440,786,1968,1994]" class="Bacillariophyceae" family="Fragilariaceae" form="major" genus="Synedra" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="form" species="familiaris">
|
||
<emphasis id="EEB6EA80F9643965FE127EEFFD9866E4" box="[440,582,1968,1994]" italics="true" pageId="10" pageNumber="11">S. familiaris</emphasis>
|
||
f.
|
||
<emphasis id="EEB6EA80F9643965FDCF7EEFFD7566E4" box="[613,683,1968,1994]" italics="true" pageId="10" pageNumber="11">major</emphasis>
|
||
Grunow
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="B8534B63F9643965FC9C7EEFFBD166E4" author="Van Heurck, H." box="[822,1039,1968,1994]" pageId="10" pageNumber="11" pagination="31 - 77" refId="ref20141" refString="Van Heurck, H. (1881) Synopsis des Diatomees de Belgique. Atlas. Ducaju & Cie., Anvers, pls. 31 - 77." type="book chapter" year="1881">Van Heurck (1881</bibRefCitation>
|
||
: plate 40, fig. 16),
|
||
<taxonomicName id="1BC24D11F9643965FB437EEFFE8B66DF" authority="Grunow" authorityName="Grunow" authorityYear="1881" class="Bacillariophyceae" family="Fragilariaceae" genus="Synedra" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="variety" species="rumpens" variety="scotica">
|
||
<emphasis id="EEB6EA80F9643965FB437EEFFAB466E4" box="[1257,1386,1968,1994]" italics="true" pageId="10" pageNumber="11">S. rumpens</emphasis>
|
||
var.
|
||
<emphasis id="EEB6EA80F9643965FF3D7E88FF3266DF" box="[151,236,2007,2033]" italics="true" pageId="10" pageNumber="11">scotica</emphasis>
|
||
Grunow
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="B8534B63F9643965FED47E88FD8066DF" author="Van Heurck, H." box="[382,606,2007,2033]" pageId="10" pageNumber="11" pagination="31 - 77" refId="ref20141" refString="Van Heurck, H. (1881) Synopsis des Diatomees de Belgique. Atlas. Ducaju & Cie., Anvers, pls. 31 - 77." type="book chapter" year="1881">Van Heurck (1881</bibRefCitation>
|
||
: plate 40, fig. 11) and
|
||
<taxonomicName id="1BC24D11F9643965FCDA7E88FA9B66DF" authority="Grunow" authorityName="Grunow" authorityYear="1881" box="[880,1349,2007,2033]" class="Bacillariophyceae" family="Fragilariaceae" genus="Synedra" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="variety" species="rumpens" variety="meneghiniana">
|
||
<emphasis id="EEB6EA80F9643965FCDA7E88FC2A66DF" box="[880,1012,2007,2033]" italics="true" pageId="10" pageNumber="11">S. rumpens</emphasis>
|
||
var.
|
||
<emphasis id="EEB6EA80F9643965FB987E88FB0566DF" box="[1074,1243,2007,2033]" italics="true" pageId="10" pageNumber="11">meneghiniana</emphasis>
|
||
Grunow
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="B8534B63F9643964FAC77E88FEE1619C" author="Van Heurck, H." lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" pagination="31 - 77" refId="ref20141" refString="Van Heurck, H. (1881) Synopsis des Diatomees de Belgique. Atlas. Ducaju & Cie., Anvers, pls. 31 - 77." type="book chapter" year="1881">Van Heurck (1881</bibRefCitation>
|
||
: plate 40, fig. 13). All of these taxa are linear, with barely to distinctly capitate headpoles and coarse striae. They are all much smaller (Grunow in
|
||
<bibRefCitation id="B8534B63F9653964FC8E79E1FC2361F6" author="Van Heurck, H." box="[804,1021,190,216]" pageId="11" pageNumber="12" pagination="31 - 77" refId="ref20141" refString="Van Heurck, H. (1881) Synopsis des Diatomees de Belgique. Atlas. Ducaju & Cie., Anvers, pls. 31 - 77." type="book chapter" year="1881">Van Heurck 1881</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="B8534B63F9653964FBA679E1FAF361F6" author="Patrick, R. M. & Reimer, C. W." box="[1036,1325,190,216]" pageId="11" pageNumber="12" refId="ref18860" refString="Patrick, R. M. & Reimer, C. W. (1966) The Diatoms of the United States. Monograph 13. Academy of Natural Sciences, Philadelphia, 688 pp." type="book" year="1966">Patrick & Reimer 1966</bibRefCitation>
|
||
, also see Krammer & Lange-Bertalot 1991 under names within the genus
|
||
<taxonomicName id="1BC24D11F9653964FC3E79BAFBCE61D1" authorityName="Lyngbye" authorityYear="1819" box="[916,1040,229,255]" class="Bacillariophyceae" family="Fragilariaceae" genus="Fragilaria" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="11" pageNumber="12" phylum="Ochrophyta" rank="genus">
|
||
<emphasis id="EEB6EA80F9653964FC3E79BAFBCE61D1" box="[916,1040,229,255]" italics="true" pageId="11" pageNumber="12">Fragilaria</emphasis>
|
||
</taxonomicName>
|
||
) than what we have observed for
|
||
<taxonomicName id="1BC24D11F9653964FF3D7853FEEF6008" authorityName="Graeff, Kociolek & S. R. Rushforth" authorityYear="2013" box="[151,305,268,294]" class="Bacillariophyceae" family="Fragilariaceae" genus="Ulnaria" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="11" pageNumber="12" phylum="Ochrophyta" rank="species" species="tooelensis">
|
||
<emphasis id="EEB6EA80F9653964FF3D7853FEEF6008" box="[151,305,268,294]" italics="true" pageId="11" pageNumber="12">U. tooelensis</emphasis>
|
||
</taxonomicName>
|
||
, and the valves of our new taxon that are in the same size range as these are shaped differently and have striae that are much more disorganized. A review of the nomenclature,
|
||
<typeStatus id="03798830F9653964FBCB786DFB7E6062" box="[1121,1184,306,332]" pageId="11" pageNumber="12">types</typeStatus>
|
||
and ultrastructure of these species is presented in
|
||
<bibRefCitation id="B8534B63F9653964FE497806FD31605D" author="Tuji, A. & Williams, D. M." box="[483,751,345,371]" pageId="11" pageNumber="12" pagination="99 - 103" refId="ref19914" refString="Tuji, A. & Williams, D. M. (2006) Examination of the type material of Synedra rumpens Fragilaria rumpens, Bacillariophyceae. Phycological Research 54: 99 - 103. http: // dx. doi. org / 10.1111 / j. 1440 - 1835.2006.00414. x" type="journal article" year="2006">Tuji & Williams (2006</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="B8534B63F9653964FD507806FCE7605D" author="Tuji, A. & Williams, D. M." box="[762,825,345,371]" pageId="11" pageNumber="12" pagination="1 - 9" refId="ref20017" refString="Tuji, A. & Williams, D. M. (2013) Examination of types in the Fragilaria vaucheriae-intermedia species complex. Bulletin of the National Museum of Nature and Science Series B (Botany) 39: 1 - 9." type="journal article" year="2013">2013</bibRefCitation>
|
||
). Further work on the ultrastructure and systematics of this complex of species, including the
|
||
<taxonomicName id="1BC24D11F9653964FDD778DFFCE960B4" box="[637,823,384,410]" class="Bacillariophyceae" family="Fragilariaceae" genus="Synedra" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="11" pageNumber="12" phylum="Ochrophyta" rank="species" species="delicatissima">
|
||
<emphasis id="EEB6EA80F9653964FDD778DFFCE960B4" box="[637,823,384,410]" italics="true" pageId="11" pageNumber="12">S. delicatissima</emphasis>
|
||
</taxonomicName>
|
||
W.
|
||
<bibRefCitation id="B8534B63F9653964FCCC78DFFBEE60B4" author="Smith, W." box="[870,1072,384,410]" pageId="11" pageNumber="12" refId="ref19359" refString="Smith, W. 1853. Synopsis of the British Diatomaceae. 1. John Van Voorst, London. 89 pp." type="book" year="1853">
|
||
<collectingRegion id="1E06F870F9653964FCCC78DFFC7260B4" box="[870,940,384,410]" country="Bermuda" name="Smith's" pageId="11" pageNumber="12">Smith</collectingRegion>
|
||
(1853: 72)
|
||
</bibRefCitation>
|
||
group (
|
||
<bibRefCitation id="B8534B63F9653964FB2678DFFA4D60B4" author="Tuji, A. & Williams, D. M." box="[1164,1427,384,410]" pageId="11" pageNumber="12" pagination="69 - 79" refId="ref19967" refString="Tuji, A. & Williams, D. M. (2007) Type examination of Japanese diatoms described by Friedrich Meister (1913) from Lake Suwa. Bulletin of the National Museum of Nature and Science Series B (Botany) 33: 69 - 79." type="journal article" year="2007">Tuji & Williams 2007</bibRefCitation>
|
||
) and
|
||
<taxonomicName id="1BC24D11F9653964FF7978F9FD0960EE" authority="Ostrup (1910: 190)" authorityName="Ostrup" authorityPageNumber="190" authorityYear="1910" box="[211,727,422,448]" class="Bacillariophyceae" family="Fragilariaceae" genus="Fragilaria" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="11" pageNumber="12" phylum="Ochrophyta" rank="species" species="gracilis">
|
||
<emphasis id="EEB6EA80F9653964FF7978F9FE9060EE" box="[211,334,422,448]" italics="true" pageId="11" pageNumber="12">F. gracilis</emphasis>
|
||
Østrup (1910: 190) (Tuji 2007)
|
||
</taxonomicName>
|
||
, will go far to help elucidate circumscription of these two genera and the placement of these species within them.
|
||
</paragraph>
|
||
<paragraph id="DC7D3692F9653964FF6C78ABFC5E63D8" blockId="11.[151,1436,152,1028]" pageId="11" pageNumber="12">
|
||
Smaller specimens of this species also bear a superficial resemblance to
|
||
<taxonomicName id="1BC24D11F9653964FB9E78ABFEAA631A" authority="Levkov & Williams (2011: 8)" authorityName="Levkov & Williams" authorityPageNumber="8" authorityYear="2011" class="Bacillariophyceae" family="Fragilariaceae" genus="Staurosirella" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="11" pageNumber="12" phylum="Ochrophyta" rank="species" species="micra">
|
||
<emphasis id="EEB6EA80F9653964FB9E78ABFAC96320" box="[1076,1303,500,526]" italics="true" pageId="11" pageNumber="12">Staurosirella micra</emphasis>
|
||
<bibRefCitation id="B8534B63F9653964FA8878ABFEAA631A" author="Levkov, Z. & Williams, D. M." pageId="11" pageNumber="12" pagination="1 - 41" refId="ref17973" refString="Levkov, Z. & Williams, D. M. (2011) Fifteen new diatom (Bacillariophyta) species from Lake Ohrid, Macedonia. Phytotaxa 30: 1 - 41." type="journal article" year="2011">Levkov & Williams (2011: 8)</bibRefCitation>
|
||
</taxonomicName>
|
||
, described recently from Lake
|
||
<collectingRegion id="1E06F870F9653964FD4A7B45FCFA631A" box="[736,804,538,564]" country="Macedonia" name="Ohrid" pageId="11" pageNumber="12">Ohrid</collectingRegion>
|
||
(
|
||
<bibRefCitation id="B8534B63F9653964FC9E7B45FBBD631A" author="Levkov, Z. & Williams, D. M." box="[820,1123,538,564]" pageId="11" pageNumber="12" pagination="1 - 41" refId="ref17973" refString="Levkov, Z. & Williams, D. M. (2011) Fifteen new diatom (Bacillariophyta) species from Lake Ohrid, Macedonia. Phytotaxa 30: 1 - 41." type="journal article" year="2011">Levkov & Williams 2011</bibRefCitation>
|
||
). This
|
||
<collectingRegion id="1E06F870F9653964FB107B45FB20631A" box="[1210,1278,538,564]" country="Macedonia" name="Ohrid" pageId="11" pageNumber="12">Ohrid</collectingRegion>
|
||
species has a tumid central area that is has one side without ornamentation. Like
|
||
<taxonomicName id="1BC24D11F9653964FC1F7B1EFB916375" authorityName="Graeff, Kociolek & S. R. Rushforth" authorityYear="2013" box="[949,1103,577,603]" class="Bacillariophyceae" family="Fragilariaceae" genus="Ulnaria" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="11" pageNumber="12" phylum="Ochrophyta" rank="species" species="tooelensis">
|
||
<emphasis id="EEB6EA80F9653964FC1F7B1EFB916375" box="[949,1103,577,603]" italics="true" pageId="11" pageNumber="12">U. tooelensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="1BC24D11F9653964FBF47B1EFAEB6375" box="[1118,1333,577,603]" class="Bacillariophyceae" family="Fragilariaceae" genus="Staurosirella" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="11" pageNumber="12" phylum="Ochrophyta" rank="species" species="mica">
|
||
<emphasis id="EEB6EA80F9653964FBF47B1EFAEB6375" box="[1118,1333,577,603]" italics="true" pageId="11" pageNumber="12">Staurosirella mica</emphasis>
|
||
</taxonomicName>
|
||
also has pore fields at both apices and striae composed of uniseriate rows of round to elliptical areolae (
|
||
<bibRefCitation id="B8534B63F9653964FAB57B37FE956386" author="Levkov, Z. & Williams, D. M." pageId="11" pageNumber="12" pagination="1 - 41" refId="ref17973" refString="Levkov, Z. & Williams, D. M. (2011) Fifteen new diatom (Bacillariophyta) species from Lake Ohrid, Macedonia. Phytotaxa 30: 1 - 41." type="journal article" year="2011">Levkov & Williams 2011</bibRefCitation>
|
||
: figs 39–56). The two species differ in the structure of the pore fields (an ocellimbus in
|
||
<taxonomicName id="1BC24D11F9653964FAD57BD1FECD63E1" authorityName="Graeff, Kociolek & S. R. Rushforth" authorityYear="2013" class="Bacillariophyceae" family="Fragilariaceae" genus="Ulnaria" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="11" pageNumber="12" phylum="Ochrophyta" rank="species" species="tooelensis">
|
||
<emphasis id="EEB6EA80F9653964FAD57BD1FECD63E1" italics="true" pageId="11" pageNumber="12">U. tooelensis</emphasis>
|
||
</taxonomicName>
|
||
) and areolae (simple rounded pores both internally and externally in
|
||
<taxonomicName id="1BC24D11F9653964FB2B7BEAFAFD63E1" authorityName="Graeff, Kociolek & S. R. Rushforth" authorityYear="2013" box="[1153,1315,693,719]" class="Bacillariophyceae" family="Fragilariaceae" genus="Ulnaria" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="11" pageNumber="12" phylum="Ochrophyta" rank="species" species="tooelensis">
|
||
<emphasis id="EEB6EA80F9653964FB2B7BEAFAFD63E1" box="[1153,1315,693,719]" italics="true" pageId="11" pageNumber="12">U. tooelensis</emphasis>
|
||
</taxonomicName>
|
||
), and the presence of two rimoportulae, one at each end, in
|
||
<emphasis id="EEB6EA80F9653964FD4A7B83FC5E63D8" box="[736,896,732,758]" italics="true" pageId="11" pageNumber="12">
|
||
<taxonomicName id="1BC24D11F9653964FD4A7B83FCA363D8" authorityName="Graeff, Kociolek & S. R. Rushforth" authorityYear="2013" box="[736,893,732,758]" class="Bacillariophyceae" family="Fragilariaceae" genus="Ulnaria" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="11" pageNumber="12" phylum="Ochrophyta" rank="species" species="tooelensis">U. tooelensis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="DC7D3692F9653964FF6C7A5DFEF4652A" blockId="11.[151,1436,152,1028]" pageId="11" pageNumber="12">
|
||
Because
|
||
<taxonomicName id="1BC24D11F9653964FE987A5DFE136232" authorityName="Graeff, Kociolek & S. R. Rushforth" authorityYear="2013" box="[306,461,770,796]" class="Bacillariophyceae" family="Fragilariaceae" genus="Ulnaria" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="11" pageNumber="12" phylum="Ochrophyta" rank="species" species="tooelensis">
|
||
<emphasis id="EEB6EA80F9653964FE987A5DFE136232" box="[306,461,770,796]" italics="true" pageId="11" pageNumber="12">U. tooelensis</emphasis>
|
||
</taxonomicName>
|
||
has a rimoportula and ocellulimbus at each apex (
|
||
<bibRefCitation id="B8534B63F9653964FB847A5DFAD56232" author="Round, F. E. & Crawford, R. M. & Mann, D. G." box="[1070,1291,770,796]" pageId="11" pageNumber="12" refId="ref19074" refString="Round, F. E., Crawford, R. M. & Mann, D. G. (1990) The Diatoms. Biology and Morphology of the Genera. Cambridge University Press, Cambridge, 747 pp." type="book" year="1990">
|
||
Round
|
||
<emphasis id="EEB6EA80F9653964FB227A5DFB606232" box="[1160,1214,770,796]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
|
||
. 1990
|
||
</bibRefCitation>
|
||
) and closed girdle bands with a row of poroids, the taxon is excluded from the genus
|
||
<taxonomicName id="1BC24D11F9653964FC407A76FBB8626D" authorityName="Lyngbye" authorityYear="1819" box="[1002,1126,809,835]" class="Bacillariophyceae" family="Fragilariaceae" genus="Fragilaria" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="11" pageNumber="12" phylum="Ochrophyta" rank="genus">
|
||
<emphasis id="EEB6EA80F9653964FC407A76FBB8626D" box="[1002,1126,809,835]" italics="true" pageId="11" pageNumber="12">Fragilaria</emphasis>
|
||
</taxonomicName>
|
||
(Williams 1986;
|
||
<bibRefCitation id="B8534B63F9653964FA987A76FE926244" author="Williams, D. M. & Round, F. E." pageId="11" pageNumber="12" pagination="313 - 339" refId="ref20447" refString="Williams, D. M. & Round, F. E. (1986) Revision of the genus Synedra Ehrenb. Diatom Research 1: 313 - 339. http: // dx. doi. org / 10.1080 / 0269249 X. 1986.9704976" type="journal article" year="1986">Williams & Round 1986</bibRefCitation>
|
||
). Our species lacks the external areolar occlusions of
|
||
<taxonomicName id="1BC24D11F9653964FC617A0FFF0E62BE" authority="Williams & Round (1986: 314)" authorityName="Williams & Round" authorityPageNumber="314" authorityYear="1986" class="Bacillariophyceae" family="Fragilariaceae" genus="Catacombas" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="11" pageNumber="12" phylum="Ochrophyta" rank="genus">
|
||
<emphasis id="EEB6EA80F9653964FC617A0FFB806244" box="[971,1118,848,874]" italics="true" pageId="11" pageNumber="12">Catacombas</emphasis>
|
||
<bibRefCitation id="B8534B63F9653964FBCC7A0FFF0E62BE" author="Williams, D. M. & Round, F. E." pageId="11" pageNumber="12" pagination="313 - 339" refId="ref20447" refString="Williams, D. M. & Round, F. E. (1986) Revision of the genus Synedra Ehrenb. Diatom Research 1: 313 - 339. http: // dx. doi. org / 10.1080 / 0269249 X. 1986.9704976" type="journal article" year="1986">Williams & Round (1986: 314)</bibRefCitation>
|
||
</taxonomicName>
|
||
, leaving us with placement of this species in
|
||
<taxonomicName id="1BC24D11F9653964FD487A29FB6162BE" authority="(Kutzing) Compere (2001: 100)" authorityName="Compere" authorityPageNumber="100" authorityYear="2001" baseAuthorityName="Kutzing" box="[738,1215,886,912]" class="Bacillariophyceae" family="Fragilariaceae" genus="Ulnaria" higherTaxonomySource="GBIF" kingdom="Chromista" order="Fragilariales" pageId="11" pageNumber="12" phylum="Ochrophyta" rank="genus">
|
||
<emphasis id="EEB6EA80F9653964FD487A29FCE162BE" box="[738,831,886,912]" italics="true" pageId="11" pageNumber="12">Ulnaria</emphasis>
|
||
(Kützing)
|
||
<bibRefCitation id="B8534B63F9653964FC157A29FB6162BE" author="Compere, P." box="[959,1215,886,912]" pageId="11" pageNumber="12" pagination="97 - 102" refId="ref15125" refString="Compere, P. (2001) Ulnaria (Kutzing) Compere, a new genus name for Fragilaria subgen. Altersynedra Lange-Bertalot with comments on the typification of Synedra Ehrenberg. In: R. Jahn, J. P. Kociolek, A. Witkowski & P. Compere (Eds), Lange-Bertalot Festschrift, Studies on Diatoms. A. R. G. Gantner, Ruggell, pp. 97 - 102." type="book chapter" year="2001">Compére (2001: 100)</bibRefCitation>
|
||
</taxonomicName>
|
||
. This taxon and its interesting suite of character states provides additional information for a future review of the phylogenetic relationships of this interesting group of diatoms, and the taxonomic implications such a review and analysis would yield.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |