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<document id="45EB2DF28AD9CAE822DC6E74ED39B8D4" ID-DOI="10.11646/zootaxa.5315.6.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8142503" ID-ZooBank="33189294-DC74-4CFA-8213-2600B8459040" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1689237480792" checkinUser="plazi" docAuthor="Sassi, Davide" docDate="2023" docId="546A6413FFA2FF8DFF5FFBFBFBE5F999" docLanguage="en" docName="zootaxa.5315.6.1.pdf" docOrigin="Zootaxa 5315 (6)" docSource="http://dx.doi.org/10.11646/zootaxa.5315.6.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Griburius boucardi" docType="treatment" docVersion="2" lastPageNumber="518" masterDocId="A8531C6BFFADFF9CFFC8FFC5FF89FFC8" masterDocTitle="Revision of the Griburius scutellaris (Fabricius, 1801) species group (Coleoptera: Chrysomelidae: Cryptocephalinae)" masterLastPageNumber="548" masterPageNumber="501" pageNumber="516" updateTime="1689238041822" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="21576253CA0749626D57FF786D77D70B">Revision of the Griburius scutellaris (Fabricius, 1801) species group (Coleoptera: Chrysomelidae: Cryptocephalinae)</mods:title>
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<mods:namePart id="ECE608597974AD526219368B457D8BF4">Sassi, Davide</mods:namePart>
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<mods:title id="92180D3B08215929EB06BA8B9D132472">Zootaxa</mods:title>
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<mods:date id="BAA8C4F75609DEE635CECE5AD4D2281C">2023</mods:date>
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<mods:number id="A74C967F4A329D2E5028664E5BE522B1">2023-07-13</mods:number>
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<mods:number id="53D87C0CD5B8672437BA3B49D1F24DA2">5315</mods:number>
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<mods:number id="E9B2F24DB9F43DDD027437D9BEFCBB17">6</mods:number>
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<mods:classification id="237275FFAFAF53715862216BFF743C19">journal article</mods:classification>
<mods:identifier id="F34478881777A3A1A25DE6CA598577AB" type="DOI">10.11646/zootaxa.5315.6.1</mods:identifier>
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</mods:mods>
<treatment id="546A6413FFA2FF8DFF5FFBFBFBE5F999" LSID="urn:lsid:plazi:treatment:546A6413FFA2FF8DFF5FFBFBFBE5F999" httpUri="http://treatment.plazi.org/id/546A6413FFA2FF8DFF5FFBFBFBE5F999" lastPageId="17" lastPageNumber="518" pageId="15" pageNumber="516">
<subSubSection id="94D9868EFFA2FF93FF5FFBFBFDB2FB90" box="[151,571,1086,1113]" pageId="15" pageNumber="516" type="nomenclature">
<paragraph id="DC7CD505FFA2FF93FF5FFBFBFDB2FB90" blockId="15.[151,571,1086,1149]" box="[151,571,1086,1113]" pageId="15" pageNumber="516">
<heading id="87346269FFA2FF93FF5FFBFBFDB2FB90" bold="true" box="[151,571,1086,1113]" fontSize="11" level="1" pageId="15" pageNumber="516" reason="1">
<taxonomicName id="1BC3AE86FFA2FF93FF5FFBFBFDB2FB90" authority="(Jacoby, 1880)" baseAuthorityName="Jacoby" baseAuthorityYear="1880" box="[151,571,1086,1113]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="boucardi">
<emphasis id="EEB70917FFA2FF93FF5FFBFBFDB2FB90" bold="true" box="[151,571,1086,1113]" pageId="15" pageNumber="516">
<emphasis id="EEB70917FFA2FF93FF5FFBFBFE0BFB91" bold="true" box="[151,386,1086,1113]" italics="true" pageId="15" pageNumber="516">Griburius boucardi</emphasis>
(
<bibRefCitation id="B852A8F4FFA2FF93FE58FBFBFDBAFB90" author="Jacoby, M." box="[400,563,1086,1112]" pageId="15" pageNumber="516" pagination="1 - 73" refId="ref44547" refString="Jacoby, M. (1880) s. n. In: Biologia Centrali-Americana. Insecta. Coleoptera. Vol. VI. Part 1. Phytophaga (part). Published for the editors by R. H. Porter, London, pp. 1 - 73." type="book chapter" year="1880">Jacoby, 1880</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="94D9868EFFA2FF93FF5FFBA6FEAFFBB5" box="[151,294,1123,1149]" pageId="15" pageNumber="516" type="description">
<paragraph id="DC7CD505FFA2FF93FF5FFBA6FEAFFBB5" blockId="15.[151,571,1086,1149]" box="[151,294,1123,1149]" pageId="15" pageNumber="516">
(
<figureCitation id="44F8C980FFA2FF93FF57FBA6FF60FBB5" box="[159,233,1123,1149]" captionStart="FIGURE 4" captionStartId="16.[151,260,1595,1621]" captionTargetBox="[186,1401,276,1564]" captionTargetId="figure-68@16.[180,1408,270,1570]" captionTargetPageId="16" captionText="FIGURE 4. G. boucardi. a–b: habitus, dorsal and lateral view (HT); c: frontal view, female (HT); d: frontal view, male (Varaguas, Panama); e–f: colour variations (e: male, Varaguas, Panama; f: female, Varaguas, Panama); g–k: median lobe of aedeagus in ventral (g), dorsal (h) and lateral (k) view [0.5 mm]; m: antenna [1.3 mm]; n: spermatheca [0.3 mm]. hd: setose depressions. Square brackets: length of scale segment. Photos a, b and c thanks to the courtesy of Charles Farnum (MCZ)." figureDoi="http://doi.org/10.5281/zenodo.8142511" httpUri="https://zenodo.org/record/8142511/files/figure.png" pageId="15" pageNumber="516">Figs 4</figureCitation>
;
<figureCitation id="44F8C980FFA2FF93FF3DFBA6FE97FBB5" box="[245,286,1123,1149]" captionStart="FIGURE 12" captionStartId="42.[151,260,1780,1806]" captionTargetBox="[156,1430,185,1751]" captionTargetId="figure-36@42.[151,1436,179,1756]" captionTargetPageId="42" captionText="FIGURE 12. Distribution maps for the Griburius scutellaris species-group. a: G. albilabris; b: G. biverrucatus; c: G. boucardi; d: G. decoratus; e: G. errans; f: G. larvatus. Black markings: data from specimens analyzed during the study. Red markings: data from the public database Global Biodiversity Information Facility (GBIF.com)." figureDoi="http://doi.org/10.5281/zenodo.8142531" httpUri="https://zenodo.org/record/8142531/files/figure.png" pageId="15" pageNumber="516">12c</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="94D9868EFFA2FF93FF5FFB6FFB5FFA55" pageId="15" pageNumber="516" type="reference_group">
<paragraph id="DC7CD505FFA2FF93FF5FFB6FFCCFFB0D" blockId="15.[151,1437,1194,2013]" box="[151,838,1194,1221]" pageId="15" pageNumber="516">
<treatmentCitationGroup id="FCD3F22BFFA2FF93FF5FFB6FFCCFFB0D" box="[151,838,1194,1221]" pageId="15" pageNumber="516">
<treatmentCitation id="5D62F314FFA2FF93FF5FFB6FFDC3FB0D" author="Jacoby, M." box="[151,586,1194,1221]" page="63" pageId="15" pageNumber="516" year="1880">
<taxonomicName id="1BC3AE86FFA2FF93FF5FFB6FFDC3FB0D" authority="Jacoby, 1880: 63" authorityName="Jacoby" authorityPageNumber="63" authorityYear="1880" box="[151,586,1194,1221]" class="Insecta" family="Chrysomelidae" genus="Scolochrus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="boucardi">
<emphasis id="EEB70917FFA2FF93FF5FFB6FFE0FFB0D" bold="true" box="[151,390,1194,1221]" italics="true" pageId="15" pageNumber="516">Scolochrus boucardi</emphasis>
<bibRefCitation id="B852A8F4FFA2FF93FE45FB6EFDC3FB0D" author="Jacoby, M." box="[397,586,1195,1221]" pageId="15" pageNumber="516" pagination="1 - 73" refId="ref44547" refString="Jacoby, M. (1880) s. n. In: Biologia Centrali-Americana. Insecta. Coleoptera. Vol. VI. Part 1. Phytophaga (part). Published for the editors by R. H. Porter, London, pp. 1 - 73." type="book chapter" year="1880">Jacoby, 1880: 63</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
</treatmentCitationGroup>
</paragraph>
<paragraph id="DC7CD505FFA2FF93FF0FFB0BFB33FB21" blockId="15.[151,1437,1194,2013]" box="[199,1210,1230,1257]" pageId="15" pageNumber="516">
<treatmentCitationGroup id="FCD3F22BFFA2FF93FF0FFB0BFB33FB21" box="[199,1210,1230,1257]" pageId="15" pageNumber="516">
Giburius
<treatmentCitation id="5D62F314FFA2FF93FEFDFB0BFD07FB21" author="Clavareau, C. H." box="[309,654,1230,1257]" page="89" pageId="15" pageNumber="516" year="1913">
<taxonomicName id="1BC3AE86FFA2FF93FEFDFB0BFD07FB21" authority=": Clavareau, 1913: 89" authorityName=": Clavareau" authorityPageNumber="89" authorityYear="1913" box="[309,654,1230,1257]" class="Insecta" family="Chrysomelidae" genus="Scolochrus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="boucardi">
<emphasis id="EEB70917FFA2FF93FEFDFB0BFE2CFB20" box="[309,421,1230,1257]" italics="true" pageId="15" pageNumber="516">
<emphasis id="EEB70917FFA2FF93FEFDFB0BFE15FB21" bold="true" box="[309,412,1230,1257]" italics="true" pageId="15" pageNumber="516">boucardi</emphasis>
:
</emphasis>
<bibRefCitation id="B852A8F4FFA2FF93FE64FB0AFD07FB21" author="Clavareau, C. H." box="[428,654,1231,1257]" pageId="15" pageNumber="516" pagination="1 - 278" refId="ref44237" refString="Clavareau, C. H. (1913) Pars 53: Chrysomelidae: Megascelinae, Megalopodinae, Clytrinae, Cryptocephalinae, Chlamydinae, Lamprosominae. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. W. Junk, Berlin, pp. 1 - 278." type="book chapter" year="1913">Clavareau, 1913: 89</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(catalogue);
<treatmentCitation id="5D62F314FFA2FF93FCEAFB0AFBA4FB21" author="Blackwelder, R. E." box="[802,1069,1231,1257]" page="639" pageId="15" pageNumber="516" year="1946">
<bibRefCitation id="B852A8F4FFA2FF93FCEAFB0AFBA4FB21" author="Blackwelder, R. E." box="[802,1069,1231,1257]" pageId="15" pageNumber="516" pagination="551 - 763" refId="ref43906" refString="Blackwelder, R. E. (1946) Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America. Part 4. Bulletin of the United States National Museum, 185, 551 - 763. https: // doi. org / 10.5479 / si. 03629236.185.4" type="journal article" year="1946">Blackwelder, 1946: 639</bibRefCitation>
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
<paragraph id="DC7CD505FFA2FF93FF0FFB36FB5FFA55" blockId="15.[151,1437,1194,2013]" pageId="15" pageNumber="516">
<emphasis id="EEB70917FFA2FF93FF0FFB36FE99FAC5" bold="true" box="[199,272,1267,1293]" italics="true" pageId="15" pageNumber="516">Types.</emphasis>
Jacoby reported that only a single specimen of the species was available for the study. The single female housed in MCZ matches the original description, therefore it is here accepted as the
<typeStatus id="03786BA7FFA2FF93FB5DFAD2FB7EFAF9" box="[1173,1271,1303,1329]" pageId="15" pageNumber="516">holotype</typeStatus>
of the species.
<typeStatus id="03786BA7FFA2FF93FF5FFAFEFEA6FA9D" box="[151,303,1339,1365]" pageId="15" pageNumber="516">HOLOTYPE</typeStatus>
:
<specimenCount id="CAC51E8CFFA2FF93FEF5FAFEFEDBFA9D" box="[317,338,1339,1365]" pageId="15" pageNumber="516" type="female">♀</specimenCount>
, pinned, // “
<collectingCountry id="A4D49595FFA2FF93FE2BFAFEFDB7FA9D" box="[483,574,1339,1365]" name="Panama" pageId="15" pageNumber="516">Panama</collectingCountry>
Bouchard.” [white label, handwritten] // “Type Sp. Figured.” [white label, handwritten] // “1st Jacoby Coll.” [white label, printed] // “Type 8642” [red label, partly printed] “
<taxonomicName id="1BC3AE86FFA2FF93FB0DFA9AFEB5FA55" authority="(Jacoby, 1880)" baseAuthorityName="Jacoby" baseAuthorityYear="1880" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="boucardi">
<emphasis id="EEB70917FFA2FF93FB0DFA9AFA15FAB1" box="[1221,1436,1375,1401]" italics="true" pageId="15" pageNumber="516">Griburius boucardi</emphasis>
(
<bibRefCitation id="B852A8F4FFA2FF93FF57FA46FEBAFA55" author="Jacoby, M." box="[159,307,1411,1437]" pageId="15" pageNumber="516" pagination="1 - 73" refId="ref44547" refString="Jacoby, M. (1880) s. n. In: Biologia Centrali-Americana. Insecta. Coleoptera. Vol. VI. Part 1. Phytophaga (part). Published for the editors by R. H. Porter, London, pp. 1 - 73." type="book chapter" year="1880">Jacoby, 1880</bibRefCitation>
)
</taxonomicName>
(
<taxonomicName id="1BC3AE86FFA2FF93FE84FA46FDBDFA55" authorityName=": Clavareau" authorityYear="1913" box="[332,564,1411,1437]" class="Insecta" family="Chrysomelidae" genus="Scolochrus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="boucardi">
<emphasis id="EEB70917FFA2FF93FE84FA46FDBDFA55" box="[332,564,1411,1437]" italics="true" pageId="15" pageNumber="516">Scolochrus boucardi</emphasis>
</taxonomicName>
)
<typeStatus id="03786BA7FFA2FF93FD8CFA46FD63FA55" box="[580,746,1411,1437]" pageId="15" pageNumber="516">HOLOTYPUS</typeStatus>
D. Sassi det.” [red label, printed] // (MCZ).
</paragraph>
</subSubSection>
<subSubSection id="94D9868EFFA2FF93FF0FFA62FE2BF999" pageId="15" pageNumber="516" type="materials_examined">
<paragraph id="DC7CD505FFA2FF93FF0FFA62FE56FA09" blockId="15.[151,1437,1194,2013]" box="[199,479,1446,1473]" pageId="15" pageNumber="516">
<materialsCitation id="6CABDF58FFA2FF93FF0FFA62FE52FA09" box="[199,475,1446,1473]" country="Panama" location="Panama" pageId="15" pageNumber="516" specimenCount="1" typeStatus="holotype">
<emphasis id="EEB70917FFA2FF93FF0FFA62FED6FA09" bold="true" box="[199,351,1446,1473]" italics="true" pageId="15" pageNumber="516">
<typeStatus id="03786BA7FFA2FF93FF0FFA62FF77FA09" box="[199,254,1447,1473]" pageId="15" pageNumber="516">Type</typeStatus>
locality.
</emphasis>
“
<collectingCountry id="A4D49595FFA2FF93FEB8FA62FE58FA09" box="[368,465,1447,1473]" name="Panama" pageId="15" pageNumber="516">Panama</collectingCountry>
”
</materialsCitation>
.
</paragraph>
<paragraph id="DC7CD505FFA2FF93FF0FFA0FFE2BF999" blockId="15.[151,1437,1194,2013]" pageId="15" pageNumber="516">
<emphasis id="EEB70917FFA2FF93FF0FFA0FFDA8FA2D" bold="true" box="[199,545,1482,1509]" italics="true" pageId="15" pageNumber="516">Additional material examined</emphasis>
.
<materialsCitation id="6CABDF58FFA2FF93FDF8FA0FFF7CF9C1" collectingDate="1971-06-19" collectionCode="TAMU" country="Panama" elevation="450" latitude="8.666667" location="Cerro Campana" longLatPrecision="1300" longitude="-79.933334" pageId="15" pageNumber="516" specimenCount="1" stateProvince="Panama">
<emphasis id="EEB70917FFA2FF93FDF8FA0FFD23FA2C" bold="true" box="[560,682,1482,1508]" pageId="15" pageNumber="516">
<collectingCountry id="A4D49595FFA2FF93FDF8FA0FFD23FA2C" box="[560,682,1482,1508]" name="Panama" pageId="15" pageNumber="516">PANAMA</collectingCountry>
</emphasis>
:
<emphasis id="EEB70917FFA2FF93FD72FA0EFC95FA2D" bold="true" box="[698,796,1483,1509]" pageId="15" pageNumber="516">
<collectingRegion id="1E071BE7FFA2FF93FD72FA0EFC95FA2D" box="[698,796,1483,1509]" country="Panama" name="Panama" pageId="15" pageNumber="516">Panama</collectingRegion>
</emphasis>
:
<location id="D91C83DEFFA2FF93FCE4FA0EFC57FA2D" LSID="urn:lsid:plazi:treatment:546A6413FFA2FF8DFF5FFBFBFBE5F999:D91C83DEFFA2FF93FCE4FA0EFC57FA2D" box="[812,990,1483,1509]" country="Panama" latitude="8.666667" longLatPrecision="1300" longitude="-79.933334" name="Cerro Campana" pageId="15" pageNumber="516" stateProvince="Panama">Cerro Campana</location>
<quantity id="1B3B78E0FFA2FF93FC2DFA0FFBACFA2D" box="[997,1061,1482,1509]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="15" pageNumber="516" unit="m" value="450.0">
<elevation id="57EE3236FFA2FF93FC2DFA0FFBACFA2D" box="[997,1061,1482,1509]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="15" pageNumber="516" unit="m" value="450.0">450m</elevation>
</quantity>
<geoCoordinate id="B9F7B3C2FFA2FF93FBE5FA0EFB09FA2D" box="[1069,1152,1483,1509]" degrees="8" direction="north" minutes="40" orientation="latitude" pageId="15" pageNumber="516" precision="925" value="8.666667">8°40’N</geoCoordinate>
<geoCoordinate id="B9F7B3C2FFA2FF93FB40FA0FFB67FA2D" box="[1160,1262,1482,1509]" degrees="79" direction="west" minutes="56" orientation="longitude" pageId="15" pageNumber="516" precision="925" value="-79.933334">79°56’W</geoCoordinate>
<date id="A87DF3C5FFA2FF93FB3EFA0EFAFFFA2D" box="[1270,1398,1483,1509]" pageId="15" pageNumber="516" value="1971-06-19">
<collectingDate id="B8390A2DFFA2FF93FB3EFA0EFAFFFA2D" box="[1270,1398,1483,1509]" pageId="15" pageNumber="516" value="1971-06-19">19.VI.1971</collectingDate>
</date>
(1,
<collectionCode id="BAD24DC0FFA2FF93FF5FFA2AFF64F9C1" box="[151,237,1519,1545]" pageId="15" pageNumber="516">TAMU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6CABDF58FFA2FF93FEC8FA2AFD23F9C1" box="[256,682,1519,1545]" collectingDate="1980-05-11" collectionCode="ERPC" collectorName="Cerro Campana" country="Panama" pageId="15" pageNumber="516" specimenCount="1" stateProvince="Panama">
<collectorName id="7136B0D3FFA2FF93FEC8FA2AFE38F9C1" box="[256,433,1519,1545]" pageId="15" pageNumber="516">Cerro Campana</collectorName>
<date id="A87DF3C5FFA2FF93FE70FA2AFDA0F9C1" box="[440,553,1519,1545]" pageId="15" pageNumber="516" value="1980-05-11">
<collectingDate id="B8390A2DFFA2FF93FE70FA2AFDA0F9C1" box="[440,553,1519,1545]" pageId="15" pageNumber="516" value="1980-05-11">11.V.1980</collectingDate>
</date>
(1,
<collectionCode id="BAD24DC0FFA2FF93FD9FFA2AFD2BF9C1" box="[599,674,1519,1545]" pageId="15" pageNumber="516">ERPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6CABDF58FFA2FF93FD7EFA2AFB22F9C1" box="[694,1195,1519,1545]" collectingDate="1976-07-18" collectionCode="USNMHN" collectorName="Cerro Campana" country="Panama" pageId="15" pageNumber="516" specimenCount="1" stateProvince="Panama">
<collectorName id="7136B0D3FFA2FF93FD7EFA2AFCEEF9C1" box="[694,871,1519,1545]" pageId="15" pageNumber="516">Cerro Campana</collectorName>
<date id="A87DF3C5FFA2FF93FCA6FA2AFC7FF9C1" box="[878,1014,1519,1545]" pageId="15" pageNumber="516" value="1976-07-18">
<collectingDate id="B8390A2DFFA2FF93FCA6FA2AFC7FF9C1" box="[878,1014,1519,1545]" pageId="15" pageNumber="516" value="1976-07-18">18.VII.1976</collectingDate>
</date>
(1,
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)
</materialsCitation>
;
<materialsCitation id="6CABDF58FFA2FF93FB7EFA2AFE7FF9E5" collectingDate="1980-05-26" collectionCode="ERPC" country="Panama" location="20 km W Santiago" pageId="15" pageNumber="516" specimenCount="1" stateProvince="Veraguas">
<emphasis id="EEB70917FFA2FF93FB7EFA2AFAACF9C1" bold="true" box="[1206,1317,1519,1545]" pageId="15" pageNumber="516">
<collectingRegion id="1E071BE7FFA2FF93FB7EFA2AFAACF9C1" box="[1206,1317,1519,1545]" country="Panama" name="Veraguas" pageId="15" pageNumber="516">Veraguas</collectingRegion>
</emphasis>
:
<location id="D91C83DEFFA2FF93FAFCFA2AFF70F9E5" LSID="urn:lsid:plazi:treatment:546A6413FFA2FF8DFF5FFBFBFBE5F999:D91C83DEFFA2FF93FAFCFA2AFF70F9E5" country="Panama" name="20 km W Santiago" pageId="15" pageNumber="516" stateProvince="Veraguas">
<quantity id="1B3B78E0FFA2FF93FAFCFA2AFAF5F9C1" box="[1332,1404,1519,1545]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="516" unit="km" value="20.0">20 km</quantity>
W Santiago
</location>
<date id="A87DF3C5FFA2FF93FEC9F9D6FEFAF9E5" box="[257,371,1555,1581]" pageId="15" pageNumber="516" value="1980-05-26">
<collectingDate id="B8390A2DFFA2FF93FEC9F9D6FEFAF9E5" box="[257,371,1555,1581]" pageId="15" pageNumber="516" value="1980-05-26">26.V.1980</collectingDate>
</date>
(6,
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)
</materialsCitation>
;
<materialsCitation id="6CABDF58FFA2FF93FDCAF9D6FBC5F9E5" box="[514,1100,1555,1581]" collectingDate="2021-05-03" collectionCode="GBIF" collectorName="La Esmeralda" country="Panama" location="Canto del Llano" pageId="15" pageNumber="516" specimenCount="1" stateProvince="Veraguas">
<collectorName id="7136B0D3FFA2FF93FDCAF9D6FD28F9E5" box="[514,673,1555,1581]" pageId="15" pageNumber="516">La Esmeralda</collectorName>
,
<location id="D91C83DEFFA2FF93FD65F9D6FCEDF9E5" LSID="urn:lsid:plazi:treatment:546A6413FFA2FF8DFF5FFBFBFBE5F999:D91C83DEFFA2FF93FD65F9D6FCEDF9E5" box="[685,868,1555,1581]" country="Panama" name="Canto del Llano" pageId="15" pageNumber="516" stateProvince="Veraguas">Canto del Llano</location>
<date id="A87DF3C5FFA2FF93FCA4F9D6FC59F9E5" box="[876,976,1555,1581]" pageId="15" pageNumber="516" value="2021-05-03">
<collectingDate id="B8390A2DFFA2FF93FCA4F9D6FC59F9E5" box="[876,976,1555,1581]" pageId="15" pageNumber="516" value="2021-05-03">3.V.2021</collectingDate>
</date>
(1,
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)
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;
<materialsCitation id="6CABDF58FFA2FF93FB90F9D7FE17F999" collectingDate="28. V. 2020 &amp; 5. VI. 2020" collectionCode="GBIF" collectorName="Penonome" country="Panama" location="Cocle" pageId="15" pageNumber="516" specimenCount="1" stateProvince="Cocle">
<emphasis id="EEB70917FFA2FF93FB90F9D7FB12F9E5" bold="true" box="[1112,1179,1554,1581]" pageId="15" pageNumber="516">
<collectingRegion id="1E071BE7FFA2FF93FB90F9D7FB12F9E5" box="[1112,1179,1554,1581]" country="Panama" name="Cocle" pageId="15" pageNumber="516">Coclé</collectingRegion>
</emphasis>
:
<collectorName id="7136B0D3FFA2FF93FB63F9D6FAA8F9E5" box="[1195,1313,1555,1581]" pageId="15" pageNumber="516">Penonomé</collectorName>
<collectingDate id="B8390A2DFFA2FF93FAE2F9D6FEAFF999" isEnumeration="true" pageId="15" pageNumber="516">
<date id="A87DF3C5FFA2FF93FAE2F9D6FA14F9E5" box="[1322,1437,1555,1581]" pageId="15" pageNumber="516" value="2020-05-28">28.V.2020</date>
&amp;
<date id="A87DF3C5FFA2FF93FF7CF9F3FEAFF999" box="[180,294,1590,1617]" pageId="15" pageNumber="516" value="2020-06-05">5.VI.2020</date>
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(3,
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="94D9868EFFA2FF93FF0FF99FFC21F9BD" box="[199,936,1626,1653]" pageId="15" pageNumber="516" type="distribution">
<paragraph id="DC7CD505FFA2FF93FF0FF99FFC21F9BD" blockId="15.[151,1437,1194,2013]" box="[199,936,1626,1653]" pageId="15" pageNumber="516">
<emphasis id="EEB70917FFA2FF93FF0FF99FFED3F9BD" bold="true" box="[199,346,1626,1653]" italics="true" pageId="15" pageNumber="516">Distribution.</emphasis>
At present, the species is known only from
<collectingCountry id="A4D49595FFA2FF93FC8FF99EFC2AF9BD" box="[839,931,1627,1653]" name="Panama" pageId="15" pageNumber="516">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="94D9868EFFA2FF93FF0FF9BAFA04F975" pageId="15" pageNumber="516" type="diagnosis">
<paragraph id="DC7CD505FFA2FF93FF0FF9BAFA04F975" blockId="15.[151,1437,1194,2013]" pageId="15" pageNumber="516">
<emphasis id="EEB70917FFA2FF93FF0FF9BAFEB3F951" bold="true" box="[199,314,1663,1689]" italics="true" pageId="15" pageNumber="516">Diagnosis</emphasis>
. Even though the dorsal colouration is generally highly variable within the
<taxonomicName id="1BC3AE86FFA2FF93FB60F9BAFACBF951" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[1192,1346,1663,1689]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="scutellaris">
<emphasis id="EEB70917FFA2FF93FB60F9BAFACBF951" box="[1192,1346,1663,1689]" italics="true" pageId="15" pageNumber="516">G. scutellaris</emphasis>
</taxonomicName>
species group,
<taxonomicName id="1BC3AE86FFA2FF93FF2FF966FEE4F975" authorityName=": Clavareau" authorityYear="1913" box="[231,365,1699,1725]" class="Insecta" family="Chrysomelidae" genus="Scolochrus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="boucardi">
<emphasis id="EEB70917FFA2FF93FF2FF966FEE4F975" box="[231,365,1699,1725]" italics="true" pageId="15" pageNumber="516">G. boucardi</emphasis>
</taxonomicName>
seems to be quite constant in its peculiar black elytral pattern, that resembles a large H-shape.
</paragraph>
</subSubSection>
<subSubSection id="94D9868EFFA2FF8DFF0FF902FBE5F999" lastPageId="17" lastPageNumber="518" pageId="15" pageNumber="516" type="description">
<paragraph id="DC7CD505FFA2FF93FF0FF902FE34F8CD" blockId="15.[151,1437,1194,2013]" pageId="15" pageNumber="516">
<emphasis id="EEB70917FFA2FF93FF0FF902FE3BF929" bold="true" box="[199,434,1734,1761]" italics="true" pageId="15" pageNumber="516">Description of male.</emphasis>
BL = 4.3–5.0 mm, BW =
<quantity id="1B3B78E0FFA2FF93FD13F902FCECF929" box="[731,869,1735,1761]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.2" metricValueMin="2.7" pageId="15" pageNumber="516" unit="mm" value="2.95" valueMax="3.2" valueMin="2.7">2.7–3.2 mm</quantity>
, PL = 1.6–2.0 mm, PW = 2.5–3.0 mm. Interocular distance 4.0–4.7 % of BL.
</paragraph>
<paragraph id="DC7CD505FFA2FF8CFF0FF8CAFE15FF31" blockId="15.[151,1437,1194,2013]" lastBlockId="16.[151,1436,151,249]" lastPageId="16" lastPageNumber="517" pageId="15" pageNumber="516">
Head (
<figureCitation id="44F8C980FFA2FF93FEDEF8CAFEE5F8E1" box="[278,364,1807,1833]" captionStart="FIGURE 4" captionStartId="16.[151,260,1595,1621]" captionTargetBox="[186,1401,276,1564]" captionTargetId="figure-68@16.[180,1408,270,1570]" captionTargetPageId="16" captionText="FIGURE 4. G. boucardi. a–b: habitus, dorsal and lateral view (HT); c: frontal view, female (HT); d: frontal view, male (Varaguas, Panama); e–f: colour variations (e: male, Varaguas, Panama; f: female, Varaguas, Panama); g–k: median lobe of aedeagus in ventral (g), dorsal (h) and lateral (k) view [0.5 mm]; m: antenna [1.3 mm]; n: spermatheca [0.3 mm]. hd: setose depressions. Square brackets: length of scale segment. Photos a, b and c thanks to the courtesy of Charles Farnum (MCZ)." figureDoi="http://doi.org/10.5281/zenodo.8142511" httpUri="https://zenodo.org/record/8142511/files/figure.png" pageId="15" pageNumber="516">Fig. 4d</figureCitation>
) yellow with vertex, surface between upper lobes of eyes and lower part of clypeus black. Labrum brownish. Vertex sparsely and shallowly punctured with very short, recumbent, whitish setae. Surface of frontoclypeal area with sparse, weakly impressed punctation and scattered setae. Longer setae only along inner eye rims and on mouth parts. Mid-cranial suture well detectable between upper lobes of eyes. Upper lobes of eyes close to each other along midline, but always separated by distinct strip of frons space. Ocular lines narrow, marked by row of weakly impressed punctures, strictly adhering to ocular rim up to ocular canthus. Ocular canthus large not differentiated in punctation and setosity from the remainder of frontoclypeal surface. Antennae (
<figureCitation id="44F8C980FFBDFF8CFB0DFF52FA97FF79" box="[1221,1310,151,177]" captionStart="FIGURE 4" captionStartId="16.[151,260,1595,1621]" captionTargetBox="[186,1401,276,1564]" captionTargetId="figure-68@16.[180,1408,270,1570]" captionTargetPageId="16" captionText="FIGURE 4. G. boucardi. a–b: habitus, dorsal and lateral view (HT); c: frontal view, female (HT); d: frontal view, male (Varaguas, Panama); e–f: colour variations (e: male, Varaguas, Panama; f: female, Varaguas, Panama); g–k: median lobe of aedeagus in ventral (g), dorsal (h) and lateral (k) view [0.5 mm]; m: antenna [1.3 mm]; n: spermatheca [0.3 mm]. hd: setose depressions. Square brackets: length of scale segment. Photos a, b and c thanks to the courtesy of Charles Farnum (MCZ)." figureDoi="http://doi.org/10.5281/zenodo.8142511" httpUri="https://zenodo.org/record/8142511/files/figure.png" pageId="16" pageNumber="517">
Fig.
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</figureCitation>
) uniformly brownish rather short with antennomeres 3–5 bright, subcylindrical; antennomeres 6–11, dull, more flattened and more diffusedly setose.
</paragraph>
<caption id="88BC858DFFBDFF8CFF5FF9FEFE4EF92E" ID-DOI="http://doi.org/10.5281/zenodo.8142511" ID-Zenodo-Dep="8142511" httpUri="https://zenodo.org/record/8142511/files/figure.png" pageId="16" pageNumber="517" startId="16.[151,260,1595,1621]" targetBox="[186,1401,276,1564]" targetPageId="16" targetType="figure">
<paragraph id="DC7CD505FFBDFF8CFF5FF9FEFE4EF92E" blockId="16.[151,1437,1595,1766]" pageId="16" pageNumber="517">
<emphasis id="EEB70917FFBDFF8CFF5FF9FEFEA8F99E" bold="true" box="[151,289,1595,1622]" pageId="16" pageNumber="517">FIGURE 4.</emphasis>
<emphasis id="EEB70917FFBDFF8CFEE0F9F9FE3EF99E" box="[296,439,1596,1622]" italics="true" pageId="16" pageNumber="517">
<taxonomicName id="1BC3AE86FFBDFF8CFEE0F9F9FE3AF99E" authorityName=": Clavareau" authorityYear="1913" box="[296,435,1596,1622]" class="Insecta" family="Chrysomelidae" genus="Scolochrus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="517" phylum="Arthropoda" rank="species" species="boucardi">G. boucardi</taxonomicName>
.
</emphasis>
a–b: habitus, dorsal and lateral view (HT); c: frontal view, female (HT); d: frontal view, male (Varaguas, Panama); e–f: colour variations (e: male, Varaguas, Panama; f: female, Varaguas, Panama); g–k: median lobe of aedeagus in ventral (g), dorsal (h) and lateral (k) view [0.5 mm]; m: antenna [1.3 mm]; n: spermatheca [0.3 mm]. hd: setose depressions. Square brackets: length of scale segment. Photos a, b and c thanks to the courtesy of Charles Farnum (MCZ).
</paragraph>
</caption>
<paragraph id="DC7CD505FFBDFF8DFF0FF8EEFD80FF31" blockId="16.[151,1437,1834,2041]" lastBlockId="17.[151,1437,151,1617]" lastPageId="17" lastPageNumber="518" pageId="16" pageNumber="517">Pronotum black with anterior and lateral margins yellow. Sometimes anterior margin yellow only along sides. Yellow median longitudinal stripe from anterior margin to middle of disc. At times, two further yellow spots on posterior part of disc, close to midline, reaching posterior margin. Posterior margin narrowly light yellow just in front to scutellum. Pronotal shape roughly elliptical, moderately transverse, regularly convex. Lateral margins narrow, not visible from above, regularly curved so that maximum width nearly at middle. Surface moderately shiny with scattered, fine to almost obliterated punctation at middle of disc. Punctures coarser and more deeply impressed along sides and in proximity of posterolateral impressions. Posterolateral impressions well impressed and obliquely arranged, marked by strong punctures at bottom of depression. Pronotal posterior margin thickened along posterolateral impressions.</paragraph>
<paragraph id="DC7CD505FFBCFF8DFF0FFEC6FBCCFE89" blockId="17.[151,1437,151,1617]" pageId="17" pageNumber="518">Scutellum black, sometimes vaguely tinged with brownish at center, subtriangular with apex truncated in almost straight line. Surface minutely and sparsely punctured, with scarce, very short setae.</paragraph>
<paragraph id="DC7CD505FFBCFF8DFF0FFE8EFCF0FDF5" blockId="17.[151,1437,151,1617]" pageId="17" pageNumber="518">Elytron yellow with large subquadrate common black patch with concave sides on disc. Such patch not including scutellar area, which is yellow. Often such black pattern spotted with yellow along suture just behind middle. Suture, lateral margin and humeral callus black. Epipleuron yellow. Elytral outline short with sides almost straight and convergent posteriorly. Lateral margins narrow, simultaneously visible from above only along posterior half. Elytral surface slightly flattened on disc, moderately shiny with strong, well-impressed punctation arranged in almost regular rows, distinct up to posterior clivus. Intervals flat. Postscutellar area fairly raised. Humeral callus prominent, impunctate. Epipleuron smooth, impunctate, with convex surface.</paragraph>
<paragraph id="DC7CD505FFBCFF8DFF0FFD82FE00FD4D" blockId="17.[151,1437,151,1617]" pageId="17" pageNumber="518">Pygidium yellow, at times with brownish margins. Surface matt, covered with close shallow punctures and appressed, pale setae.</paragraph>
<paragraph id="DC7CD505FFBCFF8DFF0FFD4AFE1CFC6D" blockId="17.[151,1437,151,1617]" pageId="17" pageNumber="518">Ventral parts of thorax black with prosternal process sometimes brownish. Abdominal ventrites 1–4 black with yellow margins. Fifth abdominal ventrite almost completely yellow. Medial projection of ventrite 1 usually yellow. Hypomera, mesepimera and mesepisterna almost bare, shiny, with scattered punctures. Remainder of ventral surface of thorax matt, covered with thick, short, regularly distributed setae and fine, shallow punctures. Prosternal process large, with sides almost straight between anterior coxae, then weakly converging in large, round apex; surface anteriorly almost flat, then slightly concave toward apex, covered with well-impressed punctures and sparse, long, semi-erect setae. Anterior legs brownish with yellow patches, median and posterior legs yellow with tibiae and tarsi more or less darkened.</paragraph>
<paragraph id="DC7CD505FFBCFF8DFF0FFC6AFCEBFB91" blockId="17.[151,1437,151,1617]" pageId="17" pageNumber="518">
Median depression on fifth abdominal ventrite hardly detectable, evenly covered with short setae and small punctures as the remainder of ventrite surface. Ventrite posterior margin straight. Median lobe of aedeagus (
<figureCitation id="44F8C980FFBCFF8DFAF7FC16FF2FFBD9" captionStart="FIGURE 4" captionStartId="16.[151,260,1595,1621]" captionTargetBox="[186,1401,276,1564]" captionTargetId="figure-68@16.[180,1408,270,1570]" captionTargetPageId="16" captionText="FIGURE 4. G. boucardi. a–b: habitus, dorsal and lateral view (HT); c: frontal view, female (HT); d: frontal view, male (Varaguas, Panama); e–f: colour variations (e: male, Varaguas, Panama; f: female, Varaguas, Panama); g–k: median lobe of aedeagus in ventral (g), dorsal (h) and lateral (k) view [0.5 mm]; m: antenna [1.3 mm]; n: spermatheca [0.3 mm]. hd: setose depressions. Square brackets: length of scale segment. Photos a, b and c thanks to the courtesy of Charles Farnum (MCZ)." figureDoi="http://doi.org/10.5281/zenodo.8142511" httpUri="https://zenodo.org/record/8142511/files/figure.png" pageId="17" pageNumber="518">
Fig.
<quantity id="1B3B78E0FFBCFF8DFABAFC16FA04FC25" box="[1394,1421,979,1005]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="17" pageNumber="518" unit="g" value="4.0">4g</quantity>
– k
</figureCitation>
) with apex terminated by well differentiated median, blunt denticle. In lateral view apex almost straight. Ventral outline marked with arcuate, median carina along apical half. Setose depressions large, shallow, barely delimited, with surface covered by small punctures and long arcuate setae.
</paragraph>
<paragraph id="DC7CD505FFBCFF8DFF0FFBA6FDC4FB69" blockId="17.[151,1437,151,1617]" pageId="17" pageNumber="518">
<emphasis id="EEB70917FFBCFF8DFF0FFBA6FE92FBB5" box="[199,283,1123,1149]" italics="true" pageId="17" pageNumber="518">Female</emphasis>
. Habitus in
<figureCitation id="44F8C980FFBCFF8DFE68FBA6FD80FBB5" box="[416,521,1123,1149]" captionStart="FIGURE 4" captionStartId="16.[151,260,1595,1621]" captionTargetBox="[186,1401,276,1564]" captionTargetId="figure-68@16.[180,1408,270,1570]" captionTargetPageId="16" captionText="FIGURE 4. G. boucardi. a–b: habitus, dorsal and lateral view (HT); c: frontal view, female (HT); d: frontal view, male (Varaguas, Panama); e–f: colour variations (e: male, Varaguas, Panama; f: female, Varaguas, Panama); g–k: median lobe of aedeagus in ventral (g), dorsal (h) and lateral (k) view [0.5 mm]; m: antenna [1.3 mm]; n: spermatheca [0.3 mm]. hd: setose depressions. Square brackets: length of scale segment. Photos a, b and c thanks to the courtesy of Charles Farnum (MCZ)." figureDoi="http://doi.org/10.5281/zenodo.8142511" httpUri="https://zenodo.org/record/8142511/files/figure.png" pageId="17" pageNumber="518">Fig. 4a–b</figureCitation>
(HT). BL = 5.0–
<quantity id="1B3B78E0FFBCFF8DFD0CFBA7FC90FBB5" box="[708,793,1122,1149]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="17" pageNumber="518" unit="mm" value="5.3">5.3 mm</quantity>
, BW =
<quantity id="1B3B78E0FFBCFF8DFCA4FBA7FC7DFBB5" box="[876,1012,1122,1149]" metricMagnitude="-3" metricUnit="m" metricValue="3.35" metricValueMax="3.5" metricValueMin="3.2" pageId="17" pageNumber="518" unit="mm" value="3.35" valueMax="3.5" valueMin="3.2">3.2–3.5 mm</quantity>
, PL = 1.9–2.0 mm, PW =
<quantity id="1B3B78E0FFBCFF8DFAD8FBA6FA10FBB5" box="[1296,1433,1123,1149]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="17" pageNumber="518" unit="mm" value="3.0" valueMax="3.1" valueMin="2.9">2.9–3.1 mm</quantity>
. Interocular distance 9.4–10.0 % of BL.
</paragraph>
<paragraph id="DC7CD505FFBCFF8DFF0FFB6EFF77FA9D" blockId="17.[151,1437,151,1617]" pageId="17" pageNumber="518">
In females, the eyes (
<figureCitation id="44F8C980FFBCFF8DFE73FB6EFD84FB0D" box="[443,525,1195,1221]" captionStart="FIGURE 4" captionStartId="16.[151,260,1595,1621]" captionTargetBox="[186,1401,276,1564]" captionTargetId="figure-68@16.[180,1408,270,1570]" captionTargetPageId="16" captionText="FIGURE 4. G. boucardi. a–b: habitus, dorsal and lateral view (HT); c: frontal view, female (HT); d: frontal view, male (Varaguas, Panama); e–f: colour variations (e: male, Varaguas, Panama; f: female, Varaguas, Panama); g–k: median lobe of aedeagus in ventral (g), dorsal (h) and lateral (k) view [0.5 mm]; m: antenna [1.3 mm]; n: spermatheca [0.3 mm]. hd: setose depressions. Square brackets: length of scale segment. Photos a, b and c thanks to the courtesy of Charles Farnum (MCZ)." figureDoi="http://doi.org/10.5281/zenodo.8142511" httpUri="https://zenodo.org/record/8142511/files/figure.png" pageId="17" pageNumber="518">Fig. 4c</figureCitation>
) are smaller and more separated along midline, and the head surface is covered with rather coarse punctation. The head surface is black, with more or less extended yellow patch in the middle; sometimes (like in the HT,
<figureCitation id="44F8C980FFBCFF8DFE1CFB36FDAFFAC5" box="[468,550,1267,1293]" captionStart="FIGURE 4" captionStartId="16.[151,260,1595,1621]" captionTargetBox="[186,1401,276,1564]" captionTargetId="figure-68@16.[180,1408,270,1570]" captionTargetPageId="16" captionText="FIGURE 4. G. boucardi. a–b: habitus, dorsal and lateral view (HT); c: frontal view, female (HT); d: frontal view, male (Varaguas, Panama); e–f: colour variations (e: male, Varaguas, Panama; f: female, Varaguas, Panama); g–k: median lobe of aedeagus in ventral (g), dorsal (h) and lateral (k) view [0.5 mm]; m: antenna [1.3 mm]; n: spermatheca [0.3 mm]. hd: setose depressions. Square brackets: length of scale segment. Photos a, b and c thanks to the courtesy of Charles Farnum (MCZ)." figureDoi="http://doi.org/10.5281/zenodo.8142511" httpUri="https://zenodo.org/record/8142511/files/figure.png" pageId="17" pageNumber="518">Fig. 4c</figureCitation>
) the yellow pattern prevails on the frons In the
<typeStatus id="03786BA7FFBCFF8DFB9AFB36FB3DFAC5" box="[1106,1204,1267,1293]" pageId="17" pageNumber="518">holotype</typeStatus>
the pronotal yellow pattern is more extended to form a roughly anchor-shaped marking. Remainder of colour pattern is similar to that of males.
</paragraph>
<paragraph id="DC7CD505FFBCFF8DFF0FFA9AFBE5F999" blockId="17.[151,1437,151,1617]" pageId="17" pageNumber="518">
The fifth abdominal ventrite in females has a quite large, rounded and deep pit. The bottom of the pit is glabrous, matt, impunctate but covered by tiny wrinkles. The vasculum of the spermatheca (
<figureCitation id="44F8C980FFBCFF8DFB05FA46FAABFA55" box="[1229,1314,1411,1437]" captionStart="FIGURE 4" captionStartId="16.[151,260,1595,1621]" captionTargetBox="[186,1401,276,1564]" captionTargetId="figure-68@16.[180,1408,270,1570]" captionTargetPageId="16" captionText="FIGURE 4. G. boucardi. a–b: habitus, dorsal and lateral view (HT); c: frontal view, female (HT); d: frontal view, male (Varaguas, Panama); e–f: colour variations (e: male, Varaguas, Panama; f: female, Varaguas, Panama); g–k: median lobe of aedeagus in ventral (g), dorsal (h) and lateral (k) view [0.5 mm]; m: antenna [1.3 mm]; n: spermatheca [0.3 mm]. hd: setose depressions. Square brackets: length of scale segment. Photos a, b and c thanks to the courtesy of Charles Farnum (MCZ)." figureDoi="http://doi.org/10.5281/zenodo.8142511" httpUri="https://zenodo.org/record/8142511/files/figure.png" pageId="17" pageNumber="518">Fig. 4n</figureCitation>
) is sickle-shaped, sometimes weakly pigmented along its not-swollen proximal lobe. The distal lobe is long, slender, slightly tapered with a rather blunt apex fairly bent downwards. The ampulla is short, not or slightly pigmented. The duct insertion on the ampulla is very short. The sperm gland insertion is longer and slightly bent upwards. The duct is uniform in size, quite rigid, not coiled but with a variable number (2–5) of turns near the vasculum, then almost straight. The insertion on the bursa copulatrix is simple, neither swollen nor pigmented.
</paragraph>
</subSubSection>
</treatment>
</document>
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