treatments-xml/data/54/68/87/54688784633C5F29FF0DFCE5F3368448.xml

<document id="F6511E1B9047DFB6A4F4BB0F13B0B20D" ID-DOI="10.11646/zootaxa.5569.1.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="14703998" ID-ZooBank="EA597E31-AF95-4D31-A14E-9455D6B335B1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1737371943224" checkinUser="plazi" docAuthor="Namayandeh, Armin &amp; Ghaderi, Edris" docDate="2025" docId="54688784633C5F29FF0DFCE5F3368448" docLanguage="en" docName="zootaxa.5569.1.5.pdf" docOrigin="Zootaxa 5569 (1)" docSource="https://doi.org/10.11646/zootaxa.5569.1.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paracricotopus niger sensu Saether 1980" docType="treatment" docVersion="4" lastPageNumber="147" masterDocId="A851FFFC633B5F20FF9AFF84F1478771" masterDocTitle="A review of Paracricotopus (Brundin, 1947), with the description of two new species and redescription of the adult males of Paracricotopus niger (Kieffer, 1913) and Paracricotopus uliginosus (Brundin, 1947) sensu Saether (1980) (Chironomidae: Diptera)" masterLastPageNumber="158" masterPageNumber="138" pageNumber="145" updateTime="1737487211152" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:title id="07C145363DD8D638116BF4F63B2B69BC">A review of Paracricotopus (Brundin, 1947), with the description of two new species and redescription of the adult males of Paracricotopus niger (Kieffer, 1913) and Paracricotopus uliginosus (Brundin, 1947) sensu Saether (1980) (Chironomidae: Diptera)</mods:title>
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<mods:namePart id="4C5F2199A1E2A8DD259A3376FDE58DF8">Namayandeh, Armin</mods:namePart>
<mods:affiliation id="80B2192629584543B78B8823A6ED1029">Department of Environmental and Life Sciences, Trent University, 1600 West Bank Drive, Peterborough, Ontario, Canada.</mods:affiliation>
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<mods:namePart id="5B333AA5551B3287AA8BB74D710E9A52">Ghaderi, Edris</mods:namePart>
<mods:affiliation id="D2F0B891CA65630323E785496A01976A">Department of Fisheries Sciences, Faculty of Natural Resources, University of Kurdistan, Sanandaj, Iran. &amp; Department of Fisheries and Aquatic Ecology, Faculty of Fisheries and Environmental Sciences, Gorgan University of Agricultural Sciences and Natural Resources, Gorgan, Iran.</mods:affiliation>
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<paragraph id="DC7E3692633C5F27FF0DFCE5F223840A" blockId="7.[151,868,865,928]" box="[151,868,865,891]" pageId="7" pageNumber="145">
<heading id="873681FE633C5F27FF0DFCE5F223840A" bold="true" box="[151,868,865,891]" fontSize="11" level="1" pageId="7" pageNumber="145" reason="1">
<taxonomicName id="1BC14D11633C5F27FF0DFCE5F223840A" ID-CoL="4CRJV" authority="(Kieffer, 1913) sensu Saether (1980)" authorityName="sensu Saether" authorityYear="1980" baseAuthorityName="Kieffer" baseAuthorityYear="1913" box="[151,868,865,891]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="145" phylum="Arthropoda" rank="species" species="niger">
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<emphasis id="EEB5EA80633C5F27FF0DFCE5F0DB840A" bold="true" box="[151,412,865,891]" italics="true" pageId="7" pageNumber="145">Paracricotopus niger</emphasis>
(Kieffer, 1913) sensu Saether (1980)
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<paragraph id="DC7E3692633C5F27FF0DFC01F06E84D1" blockId="7.[151,868,865,928]" box="[151,297,901,928]" pageId="7" pageNumber="145">
<figureCitation id="44FA2A17633C5F27FF0DFC01F1A284EE" box="[151,229,901,927]" captionStart="FIGURE 4" captionStartId="8.[151,250,1971,1995]" captionTargetBox="[161,1427,458,1937]" captionTargetId="figure-118@8.[151,1436,447,1948]" captionTargetPageId="8" captionText="FIGURE 4. Paracricotopus niger (Kieffer, 1913) sensu Saether (1980), male (A–D), female (F).A. antenna. B. thorax. C. wing. D. abdominal tergites. E. abdominal tergites. Scale bars are 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14704006" httpUri="https://zenodo.org/record/14704006/files/figure.png" pageId="7" pageNumber="145">Figs. 4</figureCitation>
&amp; 
<figureCitation id="44FA2A17633C5F27FE90FC02F06E84D1" box="[266,297,902,928]" captionStart="FIGURE 7" captionStartId="13.[151,250,1841,1865]" captionTargetBox="[318,1314,159,1806]" captionTargetId="figure-113@13.[310,1322,151,1814]" captionTargetPageId="13" captionText="FIGURE 7. Abdominal tergites I–V of Paracricotopus Brundin, spp. adult males. A. Paracricotopus davoodi Ghaderi et al., 2023. B. Paracricotopus glaber Saether, 1980. C.Paracricotopus insulatus (Bhattacharyay &amp; Chaudhuri, 1987), tergite I missing. D. Paracricotopus mozleyi Steiner, 1983. E. Paracricotopus niger (Kieffer, 1913) sensu Saether (1980). F. Paracricotopus spinicornis Hazra &amp; Chaudhuri, 2002. G. Paracricotopus sturae sp. nov. H. Paracricotopus tamabrevis (Sasa, 1983). I. Paracricotopus togakuroasi (Sasa &amp; Okazawa, 1992). J. Paracricotopus torbjorni sp. nov. K. Paracricotopus uliginosus (Brundin, 1947) sensu Saether (1980)." figureDoi="http://doi.org/10.5281/zenodo.14704014" httpUri="https://zenodo.org/record/14704014/files/figure.png" pageId="7" pageNumber="145">7E</figureCitation>
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<emphasis id="EEB5EA80633C5F27FF0DFC49F0C78496" bold="true" box="[151,384,973,999]" pageId="7" pageNumber="145">Material examined.</emphasis>
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, 
<location id="D91E6049633C5F27FDDCFC49F2078496" LSID="urn:lsid:plazi:treatment:54688784633C5F29FF0DFCE5F3368448:D91E6049633C5F27FDDCFC49F2078496" box="[582,832,973,999]" country="Norway" municipality="Malaisefelle" name="Stor-Elvdal Silbakken" pageId="7" pageNumber="145">Stor-Elvdal Silbakken</location>
, 
<collectingMunicipality id="3C1AACE8633C5F27FCD0FC49F2908496" box="[842,983,973,999]" pageId="7" pageNumber="145">Malaisefelle</collectingMunicipality>
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; 61.74614˚N, 10.74618˚E; 
<date id="A87F1052633C5F27FAC0FC49F05D837A" pageId="7" pageNumber="145" value="2008-06-30" valueMax="2008-07-07" valueMin="2008-06-30">
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; leg. 
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<location id="D91E6049633C5F27FEEEFC75F09E837A" LSID="urn:lsid:plazi:treatment:54688784633C5F29FF0DFCE5F3368448:D91E6049633C5F27FEEEFC75F09E837A" box="[372,473,1009,1035]" country="Norway" name="Hoffstad" pageId="7" pageNumber="145">Hoffstad</location>
; det. 
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, 2015; 
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572; dep. 
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, 
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, 
<location id="D91E6049633C5F27FA80FC75F4DF837A" LSID="urn:lsid:plazi:treatment:54688784633C5F29FF0DFCE5F3368448:D91E6049633C5F27FA80FC75F4DF837A" box="[1306,1432,1009,1035]" country="Norway" county="Moh" municipality="Sommerlystvegen" name="Trondheim" pageId="7" pageNumber="145">Trondheim</location>
, 
<collectingMunicipality id="3C1AACE8633C5F27FF0DFB91F023835E" box="[151,356,1045,1071]" pageId="7" pageNumber="145">Sommerlystvegen</collectingMunicipality>
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, 
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; 63.405˚N, 10.383˚E; 
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<collectingDate id="B83BE9BA633C5F27FCFCFB91F2B6835E" box="[870,1009,1045,1071]" pageId="7" pageNumber="145" value="2014-08-17">17.viii.2014</collectingDate>
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; leg. 
<collectorName id="71345344633C5F27FBB5FB91F532835E" box="[1071,1141,1045,1071]" pageId="7" pageNumber="145">E.Stur</collectorName>
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<location id="D91E6049633C5F27FB20FB91F44F835E" LSID="urn:lsid:plazi:treatment:54688784633C5F29FF0DFCE5F3368448:D91E6049633C5F27FB20FB91F44F835E" box="[1210,1288,1045,1071]" country="Norway" name="Ekrem" pageId="7" pageNumber="145">Ekrem</location>
; det. 
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ID: 
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-D12; dep. 
<collectionCode id="BAD0AE57633C5F27FCDFFBBEF2D48325" box="[837,915,1082,1108]" country="China" httpUri="http://grbio.org/cool/6zr8-87q6" name="National Taiwan Normal University" pageId="7" pageNumber="145">NTNU</collectionCode>
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<location id="D91E6049633C5F27FB0EFBBDF44E8322" LSID="urn:lsid:plazi:treatment:54688784633C5F29FF0DFCE5F3368448:D91E6049633C5F27FB0EFBBDF44E8322" box="[1172,1289,1081,1107]" country="Norway" county="Moh" municipality="Sommerlystvegen" name="Trondelag" pageId="7" pageNumber="145">Trøndelag</location>
, 
<location id="D91E6049633C5F27FA80FBBDF4DF8322" LSID="urn:lsid:plazi:treatment:54688784633C5F29FF0DFCE5F3368448:D91E6049633C5F27FA80FBBDF4DF8322" box="[1306,1432,1081,1107]" country="Norway" county="Moh" municipality="Sommerlystvegen" name="Trondheim" pageId="7" pageNumber="145">Trondheim</location>
, 
<collectingMunicipality id="3C1AACE8633C5F27FF0DFBD9F0238306" box="[151,356,1117,1143]" pageId="7" pageNumber="145">Sommerlystvegen</collectingMunicipality>
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, 
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; 63.405˚N, 10.383˚E; 
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<collectingDate id="B83BE9BA633C5F27FCF0FBD9F2AA8306" box="[874,1005,1117,1143]" pageId="7" pageNumber="145" value="2014-07-20">20.vii.2014</collectingDate>
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; leg. 
<collectorName id="71345344633C5F27FBB6FBD9F5358306" box="[1068,1138,1117,1143]" pageId="7" pageNumber="145">E.Stur</collectorName>
&amp; 
<collectionCode id="BAD0AE57633C5F27FB03FBDAF5EC8309" box="[1177,1195,1118,1144]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="145">T</collectionCode>
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<location id="D91E6049633C5F27FB22FBD9F4418306" LSID="urn:lsid:plazi:treatment:54688784633C5F29FF0DFCE5F3368448:D91E6049633C5F27FB22FBD9F4418306" box="[1208,1286,1117,1143]" country="Norway" name="Ekrem" pageId="7" pageNumber="145">Ekrem</location>
; det. 
<determinerName id="29A2DA56633C5F27FADFFBDAF4DF8306" box="[1349,1432,1117,1144]" pageId="7" pageNumber="145">E. Stur</determinerName>
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ID: 
<specimenCode id="8C679EE9633C5F27FD9FFB05F3ED83ED" box="[517,682,1153,1180]" collectionCode="BIOUG" country="Canada" httpUri="http://grbio.org/cool/5jpr-r1ny" name="Biodiversity Institute of Ontario" pageId="7" pageNumber="145" type="University or college">BIOUG15947</specimenCode>
-C12; dep. 
<collectionCode id="BAD0AE57633C5F27FCBBFB06F22883ED" box="[801,879,1154,1180]" country="China" httpUri="http://grbio.org/cool/6zr8-87q6" name="National Taiwan Normal University" pageId="7" pageNumber="145">NTNU</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="DC7E3692633C5F27FF5DFB21F55E82E9" blockId="7.[151,1437,973,1828]" pageId="7" pageNumber="145">
<emphasis id="EEB5EA80633C5F27FF5DFB21F09383CE" bold="true" box="[199,468,1189,1215]" pageId="7" pageNumber="145">Diagnostic characters.</emphasis>
The adult male of 
<taxonomicName id="1BC14D11633C5F27FD2FFB22F24983CE" authorityName="sensu Saether" authorityYear="1980" baseAuthorityName="Kieffer" baseAuthorityYear="1913" box="[693,782,1190,1215]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="145" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="EEB5EA80633C5F27FD2FFB22F24983CE" box="[693,782,1190,1215]" italics="true" pageId="7" pageNumber="145">P. niger</emphasis>
</taxonomicName>
can be separated from the closely related species by the combination of the following characteristics: AR 0.51–0.73; Haltere brown, 255–256 μm long; wing with costa extension 41–54 μm long; tergites I–VIII with 3 lateral setae; tergite I anterior basal setae, and posterior marginal setae arranged in 3 diagonal rows; tergites 
<collectionCode id="BAD0AE57633C5F27FD18FA96F3D3825D" box="[642,660,1298,1324]" pageId="7" pageNumber="145">II</collectionCode>
– 
<collectionCode id="BAD0AE57633C5F27FD38FA96F3F1825D" box="[674,694,1298,1324]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="145" type="Museum">V</collectionCode>
anterior basal setae and posterior marginal setae in two diagonal rows; tergites 
<collectionCode id="BAD0AE57633C5F27FEADFAB2F0138221" box="[311,340,1334,1360]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="7" pageNumber="145">VI</collectionCode>
– 
<collectionCode id="BAD0AE57633C5F27FEF9FAB2F0CD8221" box="[355,394,1334,1360]" pageId="7" pageNumber="145">VII</collectionCode>
anterior basal setae and posterior marginal setae in straight rows; tergite VIII with 1 row of basal setae and 2 rows of marginal setae; anal point 32–55 μm long; inferior volsella a large lobe slightly curved; gonostylus with large preapical crista dorsalis; anal point bearing 8 lateral setae.
</paragraph>
</subSubSection>
<subSubSection id="94DB6519633C5F29FF5DFA25F5C38464" lastPageId="9" lastPageNumber="147" pageId="7" pageNumber="145" type="description">
<paragraph id="DC7E3692633C5F27FF5DFA25F26782CA" blockId="7.[151,1437,973,1828]" box="[199,800,1441,1468]" pageId="7" pageNumber="145">
<emphasis id="EEB5EA80633C5F27FF5DFA25F03282CA" bold="true" box="[199,373,1441,1467]" pageId="7" pageNumber="145">Redescription.</emphasis>
Male (n = 2, unless otherwise stated).
</paragraph>
<paragraph id="DC7E3692633C5F27FF5DFA41F52082AE" blockId="7.[151,1437,973,1828]" box="[199,1127,1477,1504]" pageId="7" pageNumber="145">
Total length 2.1–2.4, 
<quantity id="1B399B77633C5F27FE2CFA41F3488291" box="[438,527,1477,1504]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="145" unit="mm" value="2.3">2.3 mm</quantity>
. Wing length 
<quantity id="1B399B77633C5F27FD30FA41F2738291" box="[682,820,1477,1504]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="7" pageNumber="145" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.3–1.4 mm</quantity>
. Haltere 255–256 μm long.
</paragraph>
<paragraph id="DC7E3692633C5F27FF5DFA6DF2538159" blockId="7.[151,1437,973,1828]" pageId="7" pageNumber="145">
<emphasis id="EEB5EA80633C5F27FF5DFA6DF0058172" box="[199,322,1513,1539]" italics="true" pageId="7" pageNumber="145">Coloration</emphasis>
. Head, including antenna and thorax, dark brown. Haltere brown. Wing greyish, pale. Legs light brown. Abdominal tergites brown, sternites whitish grey.
</paragraph>
<paragraph id="DC7E3692633C5F27FF5DF9B5F44181AA" blockId="7.[151,1437,973,1828]" pageId="7" pageNumber="145">
<emphasis id="EEB5EA80633C5F27FF5DF9B5F043813A" box="[199,260,1585,1611]" italics="true" pageId="7" pageNumber="145">Head</emphasis>
. Antenna with 13 flagellomere, last flagellomere with 11 sensilla chaetica (
<figureCitation id="44FA2A17633C5F27FBD8F9B5F5DC813A" box="[1090,1179,1585,1611]" captionStart="FIGURE 4" captionStartId="8.[151,250,1971,1995]" captionTargetBox="[161,1427,458,1937]" captionTargetId="figure-118@8.[151,1436,447,1948]" captionTargetPageId="8" captionText="FIGURE 4. Paracricotopus niger (Kieffer, 1913) sensu Saether (1980), male (A–D), female (F).A. antenna. B. thorax. C. wing. D. abdominal tergites. E. abdominal tergites. Scale bars are 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14704006" httpUri="https://zenodo.org/record/14704006/files/figure.png" pageId="7" pageNumber="145">Fig. 4A</figureCitation>
), 2 
<superScript id="2BB49BDA633C5F27FB21F9B5F58C814E" attach="none" box="[1211,1227,1585,1599]" fontSize="6" pageId="7" pageNumber="145">nd</superScript>
–3 
<superScript id="2BB49BDA633C5F27FB72F9B5F5B1814E" attach="left" box="[1256,1270,1585,1599]" fontSize="6" pageId="7" pageNumber="145">rd</superScript>
segments each with 2 sensilla chaetica, groove starts at 4 
<superScript id="2BB49BDA633C5F27FDFDF9D1F3338112" attach="left" box="[615,628,1621,1635]" fontSize="6" pageId="7" pageNumber="145">th</superScript>
segment, AR 0.67. Eyes hairy, without dorsomedial extension, temporal setae 4, including 2 frontals and 2 postoculars (n = 1). Tentorium narrow with tentorial pit close to the apex, tentorium 152–156, 154 μm long. Clypeus rectangular, 64–78, 71 μm long and 91–111, 101 μm wide, bearing 7–12, 10 setae, setae 73–84 μm long. Palpal segments lengths (in μm): 56–60, 58; 51; 68–75, 72; 108–112, 110; 141.
</paragraph>
<paragraph id="DC7E3692633C5F27FF5DF961F3E48055" blockId="7.[151,1437,973,1828]" pageId="7" pageNumber="145">
<emphasis id="EEB5EA80633C5F27FF5DF961F05F818E" box="[199,280,1765,1791]" italics="true" pageId="7" pageNumber="145">Thorax</emphasis>
(
<figureCitation id="44FA2A17633C5F27FEB0F961F0C8818E" box="[298,399,1765,1791]" captionStart="FIGURE 4" captionStartId="8.[151,250,1971,1995]" captionTargetBox="[161,1427,458,1937]" captionTargetId="figure-118@8.[151,1436,447,1948]" captionTargetPageId="8" captionText="FIGURE 4. Paracricotopus niger (Kieffer, 1913) sensu Saether (1980), male (A–D), female (F).A. antenna. B. thorax. C. wing. D. abdominal tergites. E. abdominal tergites. Scale bars are 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14704006" httpUri="https://zenodo.org/record/14704006/files/figure.png" pageId="7" pageNumber="145">Figs. 4B</figureCitation>
). Acrostichals 9; dorsocentrals 8; prealars 3; scutellars 
<quantity id="1B399B77633C5F27FB93F962F570818E" box="[1033,1079,1765,1792]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="7" pageNumber="145" unit="in" value="7.0">7 in</quantity>
single row. Antepronotal lobes developed, with small gap and bearing 5 setae.
</paragraph>
<caption id="88BE661A63335F28FF02FF13F45787DE" box="[152,1296,151,176]" pageId="8" pageNumber="146" startId="8.[152,244,151,175]" targetBox="[163,1364,195,354]" targetIsTable="true" targetPageId="8" targetType="table">
<paragraph id="DC7E369263335F28FF02FF13F45787DE" blockId="8.[152,1296,151,176]" box="[152,1296,151,176]" pageId="8" pageNumber="146">
<emphasis id="EEB5EA8063335F28FF02FF13F04887DE" bold="true" box="[152,271,151,175]" pageId="8" pageNumber="146">TABLE 4.</emphasis>
Lengths (in μm) and proportions of leg segments of 
<taxonomicName id="1BC14D1163335F28FCD9FF1CF58887C1" authority="(Kieffer, 1913)" authorityName="sensu Saether" authorityYear="1980" baseAuthorityName="Kieffer" baseAuthorityYear="1913" box="[835,1231,151,176]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="146" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="EEB5EA8063335F28FCD9FF1CF56087DE" box="[835,1063,152,175]" italics="true" pageId="8" pageNumber="146">Paracricotopus niger</emphasis>
(Kieffer, 1913)
</taxonomicName>
male.
</paragraph>
</caption>
<paragraph id="DC7E369263335F28FE8DFF47F413862C" pageId="8" pageNumber="146">
<table id="AEC1C4326333A0DFFF39FF47F4138613" box="[163,1364,195,354]" gridcols="11" gridrows="4" pageId="8" pageNumber="146">
<tr id="62F134D06333A0DFFF39FF47F4138790" box="[163,1364,195,225]" gridrow="0" pageId="8" pageNumber="146" rowspan-0="1">
<th id="21205DAC6333A0DFFE8DFF47F07A8790" box="[279,317,195,225]" gridcol="1" gridrow="0" pageId="8" pageNumber="146">
<emphasis id="EEB5EA8063335F28FE8DFF47F06C87AA" bold="true" box="[279,299,195,219]" pageId="8" pageNumber="146">fe</emphasis>
</th>
<th id="21205DAC6333A0DFFE16FF47F0F58790" box="[396,434,195,225]" gridcol="2" gridrow="0" pageId="8" pageNumber="146">
<emphasis id="EEB5EA8063335F28FE16FF47F0DC87AA" bold="true" box="[396,411,195,219]" pageId="8" pageNumber="146">ti</emphasis>
</th>
<th id="21205DAC6333A0DFFD9BFF47F3608790" box="[513,551,195,225]" gridcol="3" gridrow="0" pageId="8" pageNumber="146">
<emphasis id="EEB5EA8063335F28FD9BFF40F35187AD" bold="true" box="[513,534,196,220]" pageId="8" pageNumber="146">ta</emphasis>
<emphasis id="EEB5EA8063335F28FD8CFF57F35A8790" bold="true" box="[534,541,211,225]" pageId="8" pageNumber="146">1</emphasis>
</th>
<th id="21205DAC6333A0DFFDECFF47F3DB8790" box="[630,668,195,225]" gridcol="4" gridrow="0" pageId="8" pageNumber="146">
<emphasis id="EEB5EA8063335F28FDECFF40F3CC87AD" bold="true" box="[630,651,196,220]" pageId="8" pageNumber="146">ta</emphasis>
<emphasis id="EEB5EA8063335F28FD11FF57F3D58790" bold="true" box="[651,658,211,225]" pageId="8" pageNumber="146">2</emphasis>
</th>
<th id="21205DAC6333A0DFFD71FF47F2568790" box="[747,785,195,225]" gridcol="5" gridrow="0" pageId="8" pageNumber="146">
<emphasis id="EEB5EA8063335F28FD71FF40F24787AD" bold="true" box="[747,768,196,220]" pageId="8" pageNumber="146">ta</emphasis>
<emphasis id="EEB5EA8063335F28FC9AFF57F2408790" bold="true" box="[768,775,211,225]" pageId="8" pageNumber="146">3</emphasis>
</th>
<th id="21205DAC6333A0DFFCFAFF47F2C18790" box="[864,902,195,225]" gridcol="6" gridrow="0" pageId="8" pageNumber="146">
<emphasis id="EEB5EA8063335F28FCFAFF40F23287AD" bold="true" box="[864,885,196,220]" pageId="8" pageNumber="146">ta</emphasis>
<emphasis id="EEB5EA8063335F28FCEFFF57F23B8790" bold="true" box="[885,892,211,225]" pageId="8" pageNumber="146">4</emphasis>
</th>
<th id="21205DAC6333A0DFFC4FFF47F2B68790" box="[981,1009,195,225]" gridcol="7" gridrow="0" pageId="8" pageNumber="146">
<emphasis id="EEB5EA8063335F28FC4FFF40F2AD87AD" bold="true" box="[981,1002,196,220]" pageId="8" pageNumber="146">ta</emphasis>
<emphasis id="EEB5EA8063335F28FC70FF57F2B68790" bold="true" box="[1002,1009,211,225]" pageId="8" pageNumber="146">5</emphasis>
</th>
<th id="21205DAC6333A0DFFBD0FF47F52A8790" box="[1098,1133,195,225]" gridcol="8" gridrow="0" pageId="8" pageNumber="146">
<emphasis id="EEB5EA8063335F28FBD0FF40F52A87AD" bold="true" box="[1098,1133,196,220]" pageId="8" pageNumber="146">LR</emphasis>
</th>
<th id="21205DAC6333A0DFFB25FF47F5A58790" box="[1215,1250,195,225]" gridcol="9" gridrow="0" pageId="8" pageNumber="146">
<emphasis id="EEB5EA8063335F28FB25FF40F5A587AD" bold="true" box="[1215,1250,196,220]" pageId="8" pageNumber="146">BV</emphasis>
</th>
<th id="21205DAC6333A0DFFAAEFF47F4138790" box="[1332,1364,195,225]" gridcol="10" gridrow="0" pageId="8" pageNumber="146">
<emphasis id="EEB5EA8063335F28FAAEFF47F41387AA" bold="true" box="[1332,1364,195,219]" pageId="8" pageNumber="146">SV</emphasis>
</th>
</tr>
<tr id="62F134D06333A0DFFF39FF74F413867F" box="[163,1364,240,270]" gridrow="1" pageId="8" pageNumber="146">
<th id="21205DAC6333A0DFFF39FF74F1FE867F" box="[163,185,240,270]" gridcol="0" gridrow="1" pageId="8" pageNumber="146">
<emphasis id="EEB5EA8063335F28FF39FF75F1F58678" bold="true" box="[163,178,241,265]" pageId="8" pageNumber="146">P</emphasis>
<emphasis id="EEB5EA8063335F28FF28FE84F1FE867F" bold="true" box="[178,185,256,270]" pageId="8" pageNumber="146">1</emphasis>
</th>
<td id="21205DAC6333A0DFFE8DFF74F07A867F" box="[279,317,240,270]" gridcol="1" gridrow="1" pageId="8" pageNumber="146">488</td>
<td id="21205DAC6333A0DFFE16FF74F0F5867F" box="[396,434,240,270]" gridcol="2" gridrow="1" pageId="8" pageNumber="146">560</td>
<td id="21205DAC6333A0DFFD9BFF74F360867F" box="[513,551,240,270]" gridcol="3" gridrow="1" pageId="8" pageNumber="146">346</td>
<td id="21205DAC6333A0DFFDECFF74F3DB867F" box="[630,668,240,270]" gridcol="4" gridrow="1" pageId="8" pageNumber="146">255</td>
<td id="21205DAC6333A0DFFD71FF74F256867F" box="[747,785,240,270]" gridcol="5" gridrow="1" pageId="8" pageNumber="146">175</td>
<td id="21205DAC6333A0DFFCFAFF74F2C1867F" box="[864,902,240,270]" gridcol="6" gridrow="1" pageId="8" pageNumber="146">123</td>
<td id="21205DAC6333A0DFFC4FFF74F2B6867F" box="[981,1009,240,270]" gridcol="7" gridrow="1" pageId="8" pageNumber="146">82</td>
<td id="21205DAC6333A0DFFBD0FF74F52A867F" box="[1098,1133,240,270]" gridcol="8" gridrow="1" pageId="8" pageNumber="146">0.6</td>
<td id="21205DAC6333A0DFFB25FF74F5A5867F" box="[1215,1250,240,270]" gridcol="9" gridrow="1" pageId="8" pageNumber="146">2.2</td>
<td id="21205DAC6333A0DFFAAEFF74F413867F" box="[1332,1364,240,270]" gridcol="10" gridrow="1" pageId="8" pageNumber="146">3.0</td>
</tr>
<tr id="62F134D06333A0DFFF39FE9EF4138649" box="[163,1364,282,312]" gridrow="2" pageId="8" pageNumber="146">
<th id="21205DAC6333A0DFFF39FE9EF1FE8649" box="[163,185,282,312]" gridcol="0" gridrow="2" pageId="8" pageNumber="146">
<emphasis id="EEB5EA8063335F28FF39FE9FF1F58642" bold="true" box="[163,178,283,307]" pageId="8" pageNumber="146">P</emphasis>
<emphasis id="EEB5EA8063335F28FF28FEAEF1FE8649" bold="true" box="[178,185,298,312]" pageId="8" pageNumber="146">2</emphasis>
</th>
<td id="21205DAC6333A0DFFE8DFE9EF07A8649" box="[279,317,282,312]" gridcol="1" gridrow="2" pageId="8" pageNumber="146">504</td>
<td id="21205DAC6333A0DFFE16FE9EF0F58649" box="[396,434,282,312]" gridcol="2" gridrow="2" pageId="8" pageNumber="146">504</td>
<td id="21205DAC6333A0DFFD9BFE9EF3608649" box="[513,551,282,312]" gridcol="3" gridrow="2" pageId="8" pageNumber="146">231</td>
<td id="21205DAC6333A0DFFDECFE9EF3DB8649" box="[630,668,282,312]" gridcol="4" gridrow="2" pageId="8" pageNumber="146">135</td>
<td id="21205DAC6333A0DFFD71FE9EF2568649" box="[747,785,282,312]" gridcol="5" gridrow="2" pageId="8" pageNumber="146">100</td>
<td id="21205DAC6333A0DFFCFAFE9EF2C18649" box="[864,902,282,312]" gridcol="6" gridrow="2" pageId="8" pageNumber="146">58</td>
<td id="21205DAC6333A0DFFC4FFE9EF2B68649" box="[981,1009,282,312]" gridcol="7" gridrow="2" pageId="8" pageNumber="146">66</td>
<td id="21205DAC6333A0DFFBD0FE9EF52A8649" box="[1098,1133,282,312]" gridcol="8" gridrow="2" pageId="8" pageNumber="146">0.7</td>
<td id="21205DAC6333A0DFFB25FE9EF5A58649" box="[1215,1250,282,312]" gridcol="9" gridrow="2" pageId="8" pageNumber="146">3.9</td>
<td id="21205DAC6333A0DFFAAEFE9EF4138649" box="[1332,1364,282,312]" gridcol="10" gridrow="2" pageId="8" pageNumber="146">4.4</td>
</tr>
<tr id="62F134D06333A0DFFF39FEC0F4138613" box="[163,1364,324,354]" gridrow="3" pageId="8" pageNumber="146">
<th id="21205DAC6333A0DFFF39FEC0F1FE8613" box="[163,185,324,354]" gridcol="0" gridrow="3" pageId="8" pageNumber="146">
<emphasis id="EEB5EA8063335F28FF39FEC1F1F5862C" bold="true" box="[163,178,325,349]" pageId="8" pageNumber="146">P</emphasis>
<emphasis id="EEB5EA8063335F28FF28FED0F1FE8613" bold="true" box="[178,185,340,354]" pageId="8" pageNumber="146">3</emphasis>
</th>
<td id="21205DAC6333A0DFFE8DFEC0F07A8613" box="[279,317,324,354]" gridcol="1" gridrow="3" pageId="8" pageNumber="146">490</td>
<td id="21205DAC6333A0DFFE16FEC0F0F58613" box="[396,434,324,354]" gridcol="2" gridrow="3" pageId="8" pageNumber="146">557</td>
<td id="21205DAC6333A0DFFD9BFEC0F3608613" box="[513,551,324,354]" gridcol="3" gridrow="3" pageId="8" pageNumber="146">312</td>
<td id="21205DAC6333A0DFFDECFEC0F3DB8613" box="[630,668,324,354]" gridcol="4" gridrow="3" pageId="8" pageNumber="146">171</td>
<td id="21205DAC6333A0DFFD71FEC0F2568613" box="[747,785,324,354]" gridcol="5" gridrow="3" pageId="8" pageNumber="146">143</td>
<td id="21205DAC6333A0DFFCFAFEC0F2C18613" box="[864,902,324,354]" gridcol="6" gridrow="3" pageId="8" pageNumber="146">69</td>
<td id="21205DAC6333A0DFFC4FFEC0F2B68613" box="[981,1009,324,354]" gridcol="7" gridrow="3" pageId="8" pageNumber="146">86</td>
<td id="21205DAC6333A0DFFBD0FEC0F52A8613" box="[1098,1133,324,354]" gridcol="8" gridrow="3" pageId="8" pageNumber="146">0.6</td>
<td id="21205DAC6333A0DFFB25FEC0F5A58613" box="[1215,1250,324,354]" gridcol="9" gridrow="3" pageId="8" pageNumber="146">2.9</td>
<td id="21205DAC6333A0DFFAAEFEC0F4138613" box="[1332,1364,324,354]" gridcol="10" gridrow="3" pageId="8" pageNumber="146">3.4</td>
</tr>
</table>
</paragraph>
<caption id="88BE661A63335F28FF0DF837F20C8081" ID-DOI="http://doi.org/10.5281/zenodo.14704006" ID-Zenodo-Dep="14704006" httpUri="https://zenodo.org/record/14704006/files/figure.png" pageId="8" pageNumber="146" startId="8.[151,250,1971,1995]" targetBox="[161,1427,458,1937]" targetPageId="8" targetType="figure">
<paragraph id="DC7E369263335F28FF0DF837F20C8081" blockId="8.[151,1437,1971,2032]" pageId="8" pageNumber="146">
<emphasis id="EEB5EA8063335F28FF0DF837F05580BA" bold="true" box="[151,274,1971,1995]" pageId="8" pageNumber="146">FIGURE 4.</emphasis>
<taxonomicName id="1BC14D1163335F28FE82F830F22480BD" authority="(Kieffer, 1913) sensu Saether (1980)" authorityName="sensu Saether" authorityYear="1980" baseAuthorityName="Kieffer" baseAuthorityYear="1913" box="[280,867,1971,1996]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="146" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="EEB5EA8063335F28FE82F830F0B780BA" box="[280,496,1972,1995]" italics="true" pageId="8" pageNumber="146">Paracricotopus niger</emphasis>
(Kieffer, 1913) sensu Saether (1980)
</taxonomicName>
, male (A–D), female (F). A. antenna. B. thorax. C. wing. D. abdominal tergites. E. abdominal tergites. Scale bars are 100 µm.
</paragraph>
</caption>
<paragraph id="DC7E369263325F29FF5DFF13F44E87A4" blockId="9.[151,1437,150,825]" pageId="9" pageNumber="147">
<emphasis id="EEB5EA8063325F29FF5DFF13F04687C1" box="[199,257,151,176]" italics="true" pageId="9" pageNumber="147">Wing</emphasis>
(
<figureCitation id="44FA2A1763325F29FE95FF13F02087C0" box="[271,359,151,177]" captionStart="FIGURE 4" captionStartId="8.[151,250,1971,1995]" captionTargetBox="[161,1427,458,1937]" captionTargetId="figure-118@8.[151,1436,447,1948]" captionTargetPageId="8" captionText="FIGURE 4. Paracricotopus niger (Kieffer, 1913) sensu Saether (1980), male (A–D), female (F).A. antenna. B. thorax. C. wing. D. abdominal tergites. E. abdominal tergites. Scale bars are 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14704006" httpUri="https://zenodo.org/record/14704006/files/figure.png" pageId="9" pageNumber="147">Fig. 4C</figureCitation>
). Brachiolum with 1 seta. Squama with 6–12, 9 setae. R with 4–6, 5 setae, other veins bare. Costa extension 41–54, 47 μm long. Anal lobe rounded. Microtrichia only visible at&gt; 
<date id="A87F105263325F29FB8BFF3FF51987A4" box="[1041,1118,187,213]" pageId="9" pageNumber="147" value="1000-10">1000 x</date>
magnification.
</paragraph>
<paragraph id="DC7E369263325F29FF5DFF5BF3418630" blockId="9.[151,1437,150,825]" pageId="9" pageNumber="147">
<emphasis id="EEB5EA8063325F29FF5DFF5BF1BB8789" box="[199,252,223,248]" italics="true" pageId="9" pageNumber="147">Legs</emphasis>
. Hind and mid femurs with keel. Pulvilli developed. Fore tibia spur 37–41, 39 μm long, mid tibia spurs 16–17 and 15–17 μm long, hind tibia spurs 36–38, 37 and 19 μm long, hind tibia comb with 12 spines. Lengths and proportions of legs as in 
<tableCitation id="9143032963325F29FE36FEA3F3458630" box="[428,514,295,321]" captionStart="TABLE 4" captionStartId="8.[152,244,151,175]" captionTargetPageId="8" captionText="TABLE 4. Lengths (in μm) and proportions of leg segments of Paracricotopus niger (Kieffer, 1913) male." pageId="9" pageNumber="147">Table 4</tableCitation>
.
</paragraph>
<paragraph id="DC7E369263325F29FF5DFECFF41D86A0" blockId="9.[151,1437,150,825]" pageId="9" pageNumber="147">
<emphasis id="EEB5EA8063325F29FF5DFECFF0BF8615" box="[199,504,331,357]" italics="true" pageId="9" pageNumber="147">Abdominal tergite setation</emphasis>
(
<figureCitation id="44FA2A1763325F29FD91FECFF3348614" box="[523,627,331,357]" captionStart="FIGURE 4" captionStartId="8.[151,250,1971,1995]" captionTargetBox="[161,1427,458,1937]" captionTargetId="figure-118@8.[151,1436,447,1948]" captionTargetPageId="8" captionText="FIGURE 4. Paracricotopus niger (Kieffer, 1913) sensu Saether (1980), male (A–D), female (F).A. antenna. B. thorax. C. wing. D. abdominal tergites. E. abdominal tergites. Scale bars are 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14704006" httpUri="https://zenodo.org/record/14704006/files/figure.png" pageId="9" pageNumber="147">Figs. 4D</figureCitation>
, 
<figureCitation id="44FA2A1763325F29FD1BFECFF3E38614" box="[641,676,331,357]" captionStart="FIGURE 7" captionStartId="13.[151,250,1841,1865]" captionTargetBox="[318,1314,159,1806]" captionTargetId="figure-113@13.[310,1322,151,1814]" captionTargetPageId="13" captionText="FIGURE 7. Abdominal tergites I–V of Paracricotopus Brundin, spp. adult males. A. Paracricotopus davoodi Ghaderi et al., 2023. B. Paracricotopus glaber Saether, 1980. C.Paracricotopus insulatus (Bhattacharyay &amp; Chaudhuri, 1987), tergite I missing. D. Paracricotopus mozleyi Steiner, 1983. E. Paracricotopus niger (Kieffer, 1913) sensu Saether (1980). F. Paracricotopus spinicornis Hazra &amp; Chaudhuri, 2002. G. Paracricotopus sturae sp. nov. H. Paracricotopus tamabrevis (Sasa, 1983). I. Paracricotopus togakuroasi (Sasa &amp; Okazawa, 1992). J. Paracricotopus torbjorni sp. nov. K. Paracricotopus uliginosus (Brundin, 1947) sensu Saether (1980)." figureDoi="http://doi.org/10.5281/zenodo.14704014" httpUri="https://zenodo.org/record/14704014/files/figure.png" pageId="9" pageNumber="147">7E</figureCitation>
). Tergite I with 3 lateral setae, anterior basal setae, and posterior marginal setae arranged in 3 diagonal rows in 2–3/2/3 format. Tergites II–VIII with 3 lateral setae. Tergites II–V anterior basal setae, and posterior marginal setae in two diagonal rows. Tergites VI–VII anterior basal setae and posterior marginal setae in straight rows. Tergite VIII with 1 row of basal setae and 2 rows of marginal setae.
</paragraph>
<paragraph id="DC7E369263325F29FF5DFE5FF2E785F4" blockId="9.[151,1437,150,825]" pageId="9" pageNumber="147">
<emphasis id="EEB5EA8063325F29FF5DFE5FF0118685" box="[199,342,475,500]" italics="true" pageId="9" pageNumber="147">Hypopygium</emphasis>
(
<figureCitation id="44FA2A1763325F29FEFDFE5FF0878684" box="[359,448,475,501]" captionStart="FIGURE 4" captionStartId="8.[151,250,1971,1995]" captionTargetBox="[161,1427,458,1937]" captionTargetId="figure-118@8.[151,1436,447,1948]" captionTargetPageId="8" captionText="FIGURE 4. Paracricotopus niger (Kieffer, 1913) sensu Saether (1980), male (A–D), female (F).A. antenna. B. thorax. C. wing. D. abdominal tergites. E. abdominal tergites. Scale bars are 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14704006" httpUri="https://zenodo.org/record/14704006/files/figure.png" pageId="9" pageNumber="147">Fig. 4E</figureCitation>
). Anal point triangular with sharp apex, bearing 8 lateral setae, 32–55, 44 μm long and 13–19, 16 μm wide. Sternapodeme widely arched with well-reduced oral projections; sternapodeme 65–76, 71 μm long. Phallapodeme 40–54, 47 μm long. Inferior volsella, a large lobe slightly curved, located at the mid-section of gonocoxite. Gonocoxite 160–171, 166 μm long. Gonostylus with large preapical crista dorsalis; gonostylus 78–85, 81 μm long; megaseta 12 μm long. HR 1.9–2.2, 2.0, HV 2.5–3.0, 2.8.
</paragraph>
<paragraph id="DC7E369263325F29FF5DFD0BF03385D8" blockId="9.[151,1437,150,825]" box="[199,372,655,681]" pageId="9" pageNumber="147">Female (n = 1).</paragraph>
<paragraph id="DC7E369263325F29FF5DFD37F2AD85BC" blockId="9.[151,1437,150,825]" box="[199,1002,691,717]" pageId="9" pageNumber="147">Description. As described by Saether 1980 with the following additions:</paragraph>
<paragraph id="DC7E369263325F29FF5DFD53F2518580" blockId="9.[151,1437,150,825]" box="[199,790,727,753]" pageId="9" pageNumber="147">
<emphasis id="EEB5EA8063325F29FF5DFD53F0BB8581" box="[199,508,727,753]" italics="true" pageId="9" pageNumber="147">Abdominal tergites setation</emphasis>
(
<figureCitation id="44FA2A1763325F29FD91FD53F3268580" box="[523,609,727,753]" captionStart="FIGURE 4" captionStartId="8.[151,250,1971,1995]" captionTargetBox="[161,1427,458,1937]" captionTargetId="figure-118@8.[151,1436,447,1948]" captionTargetPageId="8" captionText="FIGURE 4. Paracricotopus niger (Kieffer, 1913) sensu Saether (1980), male (A–D), female (F).A. antenna. B. thorax. C. wing. D. abdominal tergites. E. abdominal tergites. Scale bars are 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14704006" httpUri="https://zenodo.org/record/14704006/files/figure.png" pageId="9" pageNumber="147">Fig. 4F</figureCitation>
). Same as male.
</paragraph>
<paragraph id="DC7E369263325F29FF5DFD7FF5C38464" blockId="9.[151,1437,150,825]" box="[199,1156,763,789]" pageId="9" pageNumber="147">
Immatures. Pupa and larva as described by 
<bibRefCitation id="B8504B6363325F29FD34FD7FF21C8464" author="Saether, O. A." box="[686,859,763,789]" pageId="9" pageNumber="147" pagination="129 - 145" refId="ref15286" refString="Saether, O. A. (1980 b) The Females and immatures of Paracricotopus Thienemann and Harnisch, 1932, with the description of a new species (Diptera: Chironomidae). Aquatic Insects, 2, 129 - 145. https: // doi. org / 10.1080 / 01650428009361018" type="journal article" year="1980">Saether (1980b)</bibRefCitation>
and 
<bibRefCitation id="B8504B6363325F29FC08FD7FF5C78464" author="Ghaderi, E. &amp; Ibrahimi, H. &amp; Mohammadi, H. &amp; Namayandeh, A." box="[914,1152,763,789]" pageId="9" pageNumber="147" pagination="381 - 388" refId="ref14502" refString="Ghaderi, E., Ibrahimi, H., Mohammadi, H. &amp; Namayandeh, A. (2023) Chironomidae (Diptera) of Shalmash Falls, West Azerbaijan, Iran, with the description of Paracricotopus davoodi sp. nov., a new Orthocladiinae. Zootaxa, 5285 (2), 381 - 388. https: // doi. org / 10.11646 / zootaxa. 5285.2.10" type="journal article" year="2023">
Ghaderi 
<emphasis id="EEB5EA8063325F29FC69FD7FF5638464" box="[1011,1060,763,789]" italics="true" pageId="9" pageNumber="147">et al</emphasis>
. (2023)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="94DB651963325F29FF5DFC9AF3368448" box="[199,625,798,825]" pageId="9" pageNumber="147" type="distribution">
<paragraph id="DC7E369263325F29FF5DFC9AF3368448" blockId="9.[151,1437,150,825]" box="[199,625,798,825]" pageId="9" pageNumber="147">
<emphasis id="EEB5EA8063325F29FF5DFC9AF0268449" bold="true" box="[199,353,798,824]" pageId="9" pageNumber="147">Distribution.</emphasis>
Palearctic: Widespread.
</paragraph>
</subSubSection>
</treatment>
</document>