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<document id="9F5F163409218BFBCD8308AFC45A8025" ID-DOI="10.11646/zootaxa.5569.1.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="14703998" ID-ZooBank="EA597E31-AF95-4D31-A14E-9455D6B335B1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1737371943224" checkinUser="plazi" docAuthor="Namayandeh, Armin &amp; Ghaderi, Edris" docDate="2025" docId="5468878463355F30FF0DF98BF089827C" docLanguage="en" docName="zootaxa.5569.1.5.pdf" docOrigin="Zootaxa 5569 (1)" docSource="https://doi.org/10.11646/zootaxa.5569.1.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paracricotopus togakuroasi" docType="treatment" docVersion="4" lastPageNumber="154" masterDocId="A851FFFC633B5F20FF9AFF84F1478771" masterDocTitle="A review of Paracricotopus (Brundin, 1947), with the description of two new species and redescription of the adult males of Paracricotopus niger (Kieffer, 1913) and Paracricotopus uliginosus (Brundin, 1947) sensu Saether (1980) (Chironomidae: Diptera)" masterLastPageNumber="158" masterPageNumber="138" pageNumber="152" updateTime="1737487211152" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:title id="19A191586D5E5086E5D8F0A253C93131">A review of Paracricotopus (Brundin, 1947), with the description of two new species and redescription of the adult males of Paracricotopus niger (Kieffer, 1913) and Paracricotopus uliginosus (Brundin, 1947) sensu Saether (1980) (Chironomidae: Diptera)</mods:title>
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<mods:namePart id="6C6E9E32D65BF9EBF0A373A12DDA4F0D">Namayandeh, Armin</mods:namePart>
<mods:affiliation id="C42690F6AE4611A7498F34EC1A44025E">Department of Environmental and Life Sciences, Trent University, 1600 West Bank Drive, Peterborough, Ontario, Canada.</mods:affiliation>
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<mods:namePart id="662E0F9382C89E17D189FA12C0C9797A">Ghaderi, Edris</mods:namePart>
<mods:affiliation id="90E1A371ABD2488F88A74AF739C8C089">Department of Fisheries Sciences, Faculty of Natural Resources, University of Kurdistan, Sanandaj, Iran. &amp; Department of Fisheries and Aquatic Ecology, Faculty of Fisheries and Environmental Sciences, Gorgan University of Agricultural Sciences and Natural Resources, Gorgan, Iran.</mods:affiliation>
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<treatment id="5468878463355F30FF0DF98BF089827C" LSID="urn:lsid:plazi:treatment:5468878463355F30FF0DF98BF089827C" httpUri="http://treatment.plazi.org/id/5468878463355F30FF0DF98BF089827C" lastPageId="16" lastPageNumber="154" pageId="14" pageNumber="152">
<subSubSection id="94DB651963355F2EFF0DF98BF2678158" box="[151,800,1551,1578]" pageId="14" pageNumber="152" type="nomenclature">
<paragraph id="DC7E369263355F2EFF0DF98BF2678158" blockId="14.[151,800,1551,1578]" box="[151,800,1551,1578]" pageId="14" pageNumber="152">
<heading id="873681FE63355F2EFF0DF98BF2678158" bold="true" box="[151,800,1551,1578]" fontSize="11" level="1" pageId="14" pageNumber="152" reason="1">
<taxonomicName id="1BC14D1163355F2EFF0DF98BF2678158" ID-CoL="9HW5T" authority="(Sasa &amp; Okazawa, 1992)" baseAuthorityName="Sasa &amp; Okazawa" baseAuthorityYear="1992" box="[151,800,1551,1578]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="152" phylum="Arthropoda" rank="species" species="togakuroasi">
<emphasis id="EEB5EA8063355F2EFF0DF98BF2678158" bold="true" box="[151,800,1551,1578]" pageId="14" pageNumber="152">
<emphasis id="EEB5EA8063355F2EFF0DF98BF0A1815B" bold="true" box="[151,486,1551,1578]" italics="true" pageId="14" pageNumber="152">Paracricotopus togakuroasi</emphasis>
(
<bibRefCitation id="B8504B6363355F2EFE6FF98BF2508158" author="Sasa, M. &amp; Okazawa, T." box="[501,791,1551,1577]" pageId="14" pageNumber="152" pagination="40 - 91" refId="ref15460" refString="Sasa, M. &amp; Okazawa, T. (1992) Studies on the chironomid midges (yusurika) of Kurobe River. In: Some Characteristics of Nature Conservation within the Chief Rivers in Toyama Prefecture (The upper reach of Kurobe River). Research Report from Toyama Prefectural Environmental Pollution Research Center, pp. 40 - 91." type="book chapter" year="1992">Sasa &amp; Okazawa, 1992</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="94DB651963355F2EFF0DF9D3F2678073" pageId="14" pageNumber="152" type="description">
<paragraph id="DC7E369263355F2EFF0DF9D3F5A88100" blockId="14.[151,1437,1623,1830]" box="[151,1263,1623,1650]" pageId="14" pageNumber="152">
<emphasis id="EEB5EA8063355F2EFF0DF9D3F06E8100" bold="true" box="[151,297,1623,1649]" pageId="14" pageNumber="152">Description.</emphasis>
The adult male as described by
<bibRefCitation id="B8504B6363355F2EFD08F9D3F2E48100" author="Sasa, M. &amp; Okazawa, T." box="[658,931,1623,1649]" pageId="14" pageNumber="152" pagination="40 - 91" refId="ref15460" refString="Sasa, M. &amp; Okazawa, T. (1992) Studies on the chironomid midges (yusurika) of Kurobe River. In: Some Characteristics of Nature Conservation within the Chief Rivers in Toyama Prefecture (The upper reach of Kurobe River). Research Report from Toyama Prefectural Environmental Pollution Research Center, pp. 40 - 91." type="book chapter" year="1992">Sasa &amp; Okazawa (1992)</bibRefCitation>
with the following additions:
</paragraph>
<paragraph id="DC7E369263355F2EFF5DF9FFF2678073" blockId="14.[151,1437,1623,1830]" pageId="14" pageNumber="152">
<emphasis id="EEB5EA8063355F2EFF5DF9FFF0B481E4" box="[199,499,1659,1685]" italics="true" pageId="14" pageNumber="152">Abdominal tergite setation</emphasis>
(
<figureCitation id="44FA2A1763355F2EFD99F9FFF31481E4" box="[515,595,1659,1685]" captionStart="FIGURE 7" captionStartId="13.[151,250,1841,1865]" captionTargetBox="[318,1314,159,1806]" captionTargetId="figure-113@13.[310,1322,151,1814]" captionTargetPageId="13" captionText="FIGURE 7. Abdominal tergites I–V of Paracricotopus Brundin, spp. adult males. A. Paracricotopus davoodi Ghaderi et al., 2023. B. Paracricotopus glaber Saether, 1980. C.Paracricotopus insulatus (Bhattacharyay &amp; Chaudhuri, 1987), tergite I missing. D. Paracricotopus mozleyi Steiner, 1983. E. Paracricotopus niger (Kieffer, 1913) sensu Saether (1980). F. Paracricotopus spinicornis Hazra &amp; Chaudhuri, 2002. G. Paracricotopus sturae sp. nov. H. Paracricotopus tamabrevis (Sasa, 1983). I. Paracricotopus togakuroasi (Sasa &amp; Okazawa, 1992). J. Paracricotopus torbjorni sp. nov. K. Paracricotopus uliginosus (Brundin, 1947) sensu Saether (1980)." figureDoi="http://doi.org/10.5281/zenodo.14704014" httpUri="https://zenodo.org/record/14704014/files/figure.png" pageId="14" pageNumber="152">Fig. 7I</figureCitation>
). Tergite I with 2 lateral setae, anterior basal setae, and posterior marginal setae arranged in 3 diagonal rows in 2/1/2 format. Tergites II with 3 lateral setae, anterior basal setae, and posterior marginal setae in two continuous diagonal rows. Tergite III-VIII with 4 lateral setae, anterior basal setae, and posterior marginal setae in two continuous diagonal rows.
</paragraph>
</subSubSection>
<subSubSection id="94DB651963355F2EFF5DF88FF36B8057" box="[199,556,1803,1830]" pageId="14" pageNumber="152" type="distribution">
<paragraph id="DC7E369263355F2EFF5DF88FF36B8057" blockId="14.[151,1437,1623,1830]" box="[199,556,1803,1830]" pageId="14" pageNumber="152">
<emphasis id="EEB5EA8063355F2EFF5DF88FF0268054" bold="true" box="[199,353,1803,1829]" pageId="14" pageNumber="152">Distribution.</emphasis>
Palearctic:
<collectingCountry id="A4D6760263355F2EFE7FF888F36F8057" box="[485,552,1804,1830]" name="Japan" pageId="14" pageNumber="152">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="94DB651963355F30FF0DF8F7F089827C" lastPageId="16" lastPageNumber="154" pageId="14" pageNumber="152" type="discussion">
<paragraph id="DC7E369263355F2EFF0DF8F7F04A80FC" blockId="14.[151,269,1907,1933]" box="[151,269,1907,1933]" pageId="14" pageNumber="152">
<emphasis id="EEB5EA8063355F2EFF0DF8F7F04A80FC" bold="true" box="[151,269,1907,1933]" pageId="14" pageNumber="152">Remarks</emphasis>
</paragraph>
<paragraph id="DC7E369263355F30FF0DF83FF3F986DC" blockId="14.[151,1437,1979,2042]" lastBlockId="16.[151,1437,151,1293]" lastPageId="16" lastPageNumber="154" pageId="14" pageNumber="152">
The adult males of
<taxonomicName id="1BC14D1163355F2EFEF5F838F09780A4" authorityName="Namayandeh &amp; Ghaderi" authorityYear="2025" box="[367,464,1980,2005]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="152" phylum="Arthropoda" rank="species" species="sturae" status="sp. nov.">
<emphasis id="EEB5EA8063355F2EFEF5F838F09780A4" box="[367,464,1980,2005]" italics="true" pageId="14" pageNumber="152">P. sturae</emphasis>
</taxonomicName>
<emphasis id="EEB5EA8063355F2EFE4DF83FF37780A4" bold="true" box="[471,560,1979,2005]" pageId="14" pageNumber="152">
<taxonomicNameLabel id="F58657FB63355F2EFE4DF83FF37780A4" box="[471,560,1979,2005]" pageId="14" pageNumber="152" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="1BC14D1163355F2EFDFDF838F3A080A4" authorityName="Namayandeh &amp; Ghaderi" authorityYear="2025" box="[615,743,1979,2005]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="152" phylum="Arthropoda" rank="species" species="torbjorni" status="sp. nov.">
<emphasis id="EEB5EA8063355F2EFDFDF838F3A080A4" box="[615,743,1979,2005]" italics="true" pageId="14" pageNumber="152">P. torbjorni</emphasis>
</taxonomicName>
<emphasis id="EEB5EA8063355F2EFD74F83FF20080A4" bold="true" box="[750,839,1979,2005]" pageId="14" pageNumber="152">
<taxonomicNameLabel id="F58657FB63355F2EFD74F83FF20080A4" box="[750,839,1979,2005]" pageId="14" pageNumber="152" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are both similar to
<taxonomicName id="1BC14D1163355F2EFBBBF838F53080A4" authorityName="sensu Saether" authorityYear="1980" baseAuthorityName="Kieffer" baseAuthorityYear="1913" box="[1057,1143,1980,2005]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="152" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="EEB5EA8063355F2EFBBBF838F53080A4" box="[1057,1143,1980,2005]" italics="true" pageId="14" pageNumber="152">P. niger</emphasis>
</taxonomicName>
. A combination of shorter anal points, numbers, and patterns of the tergite setation separates adult males of both species from
<taxonomicName id="1BC14D1163355F2EFA84F864F4308088" authorityName="sensu Saether" authorityYear="1980" baseAuthorityName="Kieffer" baseAuthorityYear="1913" box="[1310,1399,2016,2041]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="152" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="EEB5EA8063355F2EFA84F864F4308088" box="[1310,1399,2016,2041]" italics="true" pageId="14" pageNumber="152">P. niger</emphasis>
</taxonomicName>
.
<taxonomicName id="1BC14D1163355F30FA12F864F1BB87C0" authorityName="Namayandeh &amp; Ghaderi" authorityYear="2025" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" lastPageId="16" lastPageNumber="154" order="Diptera" pageId="14" pageNumber="152" phylum="Arthropoda" rank="species" species="torbjorni" status="sp. nov.">
<emphasis id="EEB5EA8063355F30FA12F864F1BB87C0" italics="true" lastPageId="16" lastPageNumber="154" pageId="14" pageNumber="152">P. torbjorni</emphasis>
</taxonomicName>
<emphasis id="EEB5EA80632B5F30FE9EFF13F01887C0" bold="true" box="[260,351,151,177]" pageId="16" pageNumber="154">
<taxonomicNameLabel id="F58657FB632B5F30FE9EFF13F01887C0" box="[260,351,151,177]" pageId="16" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
adult male has a much smaller anal point than
<taxonomicName id="1BC14D11632B5F30FCE2FF13F29C87C1" authorityName="Namayandeh &amp; Ghaderi" authorityYear="2025" box="[888,987,151,176]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="species" species="sturae" status="sp. nov.">
<emphasis id="EEB5EA80632B5F30FCE2FF13F29C87C1" box="[888,987,151,176]" italics="true" pageId="16" pageNumber="154">P. sturae</emphasis>
</taxonomicName>
<emphasis id="EEB5EA80632B5F30FC79FF13F57987C0" bold="true" box="[995,1086,151,177]" pageId="16" pageNumber="154">
<taxonomicNameLabel id="F58657FB632B5F30FC79FF13F57987C0" box="[995,1086,151,177]" pageId="16" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="1BC14D11632B5F30FBECFF13F58287C1" authorityName="sensu Saether" authorityYear="1980" baseAuthorityName="Kieffer" baseAuthorityYear="1913" box="[1142,1221,151,176]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="EEB5EA80632B5F30FBECFF13F58287C1" box="[1142,1221,151,176]" italics="true" pageId="16" pageNumber="154">P.niger</emphasis>
</taxonomicName>
, which can further separate it from the latter two species. The single
<taxonomicName id="1BC14D11632B5F30FD5FFF3FF20187A4" authorityName="Namayandeh &amp; Ghaderi" authorityYear="2025" box="[709,838,187,213]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="species" species="torbjorni" status="sp. nov.">
<emphasis id="EEB5EA80632B5F30FD5FFF3FF20187A4" box="[709,838,187,213]" italics="true" pageId="16" pageNumber="154">P. torbjorni</emphasis>
</taxonomicName>
<emphasis id="EEB5EA80632B5F30FCD7FF3FF2E087A4" bold="true" box="[845,935,187,213]" pageId="16" pageNumber="154">
<taxonomicNameLabel id="F58657FB632B5F30FCD7FF3FF2E087A4" box="[845,935,187,213]" pageId="16" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
adult male we examined lacked lateral setae on its anal point, similar to
<taxonomicName id="1BC14D11632B5F30FE5CFF5BF3718788" authorityName="Steiner" authorityYear="1983" box="[454,566,223,249]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="species" species="mozleyi">
<emphasis id="EEB5EA80632B5F30FE5CFF5BF3718788" box="[454,566,223,249]" italics="true" pageId="16" pageNumber="154">P. mozleyi</emphasis>
</taxonomicName>
a
<emphasis id="EEB5EA80632B5F30FDD2FF5BF2308788" box="[584,887,223,249]" italics="true" pageId="16" pageNumber="154">
nd
<taxonomicName id="1BC14D11632B5F30FDF1FF5BF3A98788" baseAuthorityName="Bhattacharyay &amp; Chaudhuri" baseAuthorityYear="1987" box="[619,750,223,249]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="species" species="insulatus">P. insulatus</taxonomicName>
.
<taxonomicName id="1BC14D11632B5F30FD62FF5BF2308788" baseAuthorityName="Bhattacharyay &amp; Chaudhuri" baseAuthorityYear="1987" box="[760,887,223,249]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="species" species="insulatus">P. insulatus</taxonomicName>
</emphasis>
has a much longer anal point and costa extension compared to
<taxonomicName id="1BC14D11632B5F30FEB3FE87F0ED866C" authorityName="Namayandeh &amp; Ghaderi" authorityYear="2025" box="[297,426,259,285]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="species" species="torbjorni" status="sp. nov.">
<emphasis id="EEB5EA80632B5F30FEB3FE87F0ED866C" box="[297,426,259,285]" italics="true" pageId="16" pageNumber="154">P. torbjorni</emphasis>
</taxonomicName>
<emphasis id="EEB5EA80632B5F30FE2BFE87F34C866C" bold="true" box="[433,523,259,285]" pageId="16" pageNumber="154">
<taxonomicNameLabel id="F58657FB632B5F30FE2BFE87F34C866C" box="[433,523,259,285]" pageId="16" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="1BC14D11632B5F30FDD2FE87F3FE866C" authorityName="Steiner" authorityYear="1983" box="[584,697,259,285]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="species" species="mozleyi">
<emphasis id="EEB5EA80632B5F30FDD2FE87F3FE866C" box="[584,697,259,285]" italics="true" pageId="16" pageNumber="154">P. mozleyi</emphasis>
</taxonomicName>
has a gonostylus without crista dorsalis; whereas gonostylus of
<taxonomicName id="1BC14D11632B5F30FA12FE87F1BB8630" authorityName="Namayandeh &amp; Ghaderi" authorityYear="2025" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="species" species="torbjorni" status="sp. nov.">
<emphasis id="EEB5EA80632B5F30FA12FE87F1BB8630" italics="true" pageId="16" pageNumber="154">P. torbjorni</emphasis>
</taxonomicName>
<emphasis id="EEB5EA80632B5F30FE99FEA3F01A8630" bold="true" box="[259,349,295,321]" pageId="16" pageNumber="154">
<taxonomicNameLabel id="F58657FB632B5F30FE99FEA3F01A8630" box="[259,349,295,321]" pageId="16" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a prominent crista dorsalis.
<tableCitation id="91430329632B5F30FD5FFEA3F2508630" box="[709,791,295,321]" captionStart="TABLE 7" captionStartId="15.[151,239,569,595]" captionTargetPageId="15" captionText="TABLE 7. Adult male of Paracricotopus Brundin spp., comparison of some relevant characters. Ac = Acrostichals; AnP = Anal point; AR = Antennal Ratio; Dc = Dorsocentrals; ext. = extension; H = Head; Hal = Haltere; L = Length; LS = Lateral Setae; Sq = Squama; TL = Total Length; WL = Wing Lenght." pageId="16" pageNumber="154">Table 7</tableCitation>
provides additional characteristics that can separate adult males of
<taxonomicName id="1BC14D11632B5F30FE99FECFF0F68615" authorityName="Brundin" authorityYear="1956" box="[259,433,331,356]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="EEB5EA80632B5F30FE99FECFF0F68615" box="[259,433,331,356]" italics="true" pageId="16" pageNumber="154">Paracricotopus</emphasis>
</taxonomicName>
. A combination of longer notum, smaller seminal capsule, and pattern of tergites setae separates the female of
<taxonomicName id="1BC14D11632B5F30FE38FEEBF34386F9" authorityName="Namayandeh &amp; Ghaderi" authorityYear="2025" box="[418,516,367,392]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="species" species="sturae" status="sp. nov.">
<emphasis id="EEB5EA80632B5F30FE38FEEBF34386F9" box="[418,516,367,392]" italics="true" pageId="16" pageNumber="154">P. sturae</emphasis>
</taxonomicName>
<emphasis id="EEB5EA80632B5F30FD96FEEBF32186F8" bold="true" box="[524,614,367,393]" pageId="16" pageNumber="154">
<taxonomicNameLabel id="F58657FB632B5F30FD96FEEBF32186F8" box="[524,614,367,393]" pageId="16" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from that of
<taxonomicName id="1BC14D11632B5F30FD66FEEBF21486F9" authorityName="sensu Saether" authorityYear="1980" baseAuthorityName="Kieffer" baseAuthorityYear="1913" box="[764,851,367,392]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="EEB5EA80632B5F30FD66FEEBF21486F9" box="[764,851,367,392]" italics="true" pageId="16" pageNumber="154">P. niger</emphasis>
</taxonomicName>
.
<tableCitation id="91430329632B5F30FCFBFEEBF2F286F8" box="[865,949,367,393]" captionStart="TABLE 8" captionStartId="16.[151,239,1340,1364]" captionTargetPageId="16" captionText="TABLE 8. Adult female of Paracricotopus Brundin spp., comparison of some relevant characters. AR = Antennal Ratio, L = Length; No. = Number, TL = Total Length; WL = Wing Length." pageId="16" pageNumber="154">Table 8</tableCitation>
provides additional characteristics that can separate known adult females of
<taxonomicName id="1BC14D11632B5F30FD92FE17F3F186DD" authorityName="Brundin" authorityYear="1956" box="[520,694,403,428]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="EEB5EA80632B5F30FD92FE17F3F186DD" box="[520,694,403,428]" italics="true" pageId="16" pageNumber="154">Paracricotopus</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="88BE661A63345F2FFF0DFDBDF7428506" pageId="15" pageNumber="153" startId="15.[151,239,569,595]" targetBox="[162,1881,682,1620]" targetIsTable="true" targetPageId="15" targetType="table">
<paragraph id="DC7E369263345F2FFF0DFDBDF7428506" blockId="15.[151,2037,569,631]" pageId="15" pageNumber="153">
<emphasis id="EEB5EA8063345F2FFF0DFDBDF04E8522" bold="true" box="[151,265,569,595]" pageId="15" pageNumber="153">TABLE 7.</emphasis>
Adult male of
<taxonomicName id="1BC14D1163345F2FFE32FDBDF3A68522" box="[424,737,569,595]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="153" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="EEB5EA8063345F2FFE32FDBDF3098522" box="[424,590,569,595]" italics="true" pageId="15" pageNumber="153">Paracricotopus</emphasis>
Brundin spp.
</taxonomicName>
, comparison of some relevant characters. Ac = Acrostichals; AnP = Anal point; AR = Antennal Ratio; Dc = Dorsocentrals; ext. = extension; H = Head; Hal = Haltere; L = Length; LS = Lateral Setae; Sq = Squama; TL = Total Length; WL = Wing Lenght.
</paragraph>
</caption>
<paragraph id="DC7E369263345F2FFF38FD38F6558133" pageId="15" pageNumber="153">
<table id="AEC1C4326334A0DFFF38FD2EF61E8125" box="[162,1881,682,1620]" gridcols="12" gridrows="16" pageId="15" pageNumber="153">
<tr id="62F134D06334A0DFFF38FD2EF61E8596" box="[162,1881,682,743]" gridrow="0" pageId="15" pageNumber="153">
<th id="21205DAC6334A0DFFF38FD2EF0288596" box="[162,367,682,743]" gridcol="0" gridrow="0" pageId="15" pageNumber="153">
<emphasis id="EEB5EA8063345F2FFF38FD38F1B685A4" bold="true" box="[162,241,700,725]" pageId="15" pageNumber="153">Species</emphasis>
</th>
<th id="21205DAC6334A0DFFE3EFD2EF3418596" box="[420,518,682,743]" gridcol="1" gridrow="0" pageId="15" pageNumber="153">
<emphasis id="EEB5EA8063345F2FFE3EFD38F34185A4" bold="true" box="[420,518,700,725]" pageId="15" pageNumber="153">TL (mm)</emphasis>
</th>
<th id="21205DAC6334A0DFFDBDFD2EF3D68596" box="[551,657,682,743]" gridcol="2" gridrow="0" pageId="15" pageNumber="153">
<emphasis id="EEB5EA8063345F2FFDBDFD38F3D685A4" bold="true" box="[551,657,700,725]" pageId="15" pageNumber="153">WL (mm)</emphasis>
</th>
<th id="21205DAC6334A0DFFD33FD2EF2458596" box="[681,770,682,743]" gridcol="3" gridrow="0" pageId="15" pageNumber="153">
<emphasis id="EEB5EA8063345F2FFD33FD38F38985A4" bold="true" box="[681,718,700,725]" pageId="15" pageNumber="153">HL</emphasis>
</th>
<th id="21205DAC6334A0DFFCA4FD2EF2AC8596" box="[830,1003,682,743]" gridcol="4" gridrow="0" pageId="15" pageNumber="153">
<emphasis id="EEB5EA8063345F2FFCA4FD38F2EE85A4" bold="true" box="[830,937,700,725]" pageId="15" pageNumber="153">Hal Color</emphasis>
</th>
<th id="21205DAC6334A0DFFB83FD2EF5388596" box="[1049,1151,682,743]" gridcol="5" gridrow="0" pageId="15" pageNumber="153">
<emphasis id="EEB5EA8063345F2FFB83FD38F57985A4" bold="true" box="[1049,1086,700,725]" pageId="15" pageNumber="153">AR</emphasis>
</th>
<th id="21205DAC6334A0DFFB28FD2EF5A38596" box="[1202,1252,682,743]" gridcol="6" gridrow="0" pageId="15" pageNumber="153">
<emphasis id="EEB5EA8063345F2FFB28FD38F59785A4" bold="true" box="[1202,1232,700,725]" pageId="15" pageNumber="153">Ac</emphasis>
</th>
<th id="21205DAC6334A0DFFA99FD2EF4058596" box="[1283,1346,682,743]" gridcol="7" gridrow="0" pageId="15" pageNumber="153">
<emphasis id="EEB5EA8063345F2FFA99FD38F46685A4" bold="true" box="[1283,1313,700,725]" pageId="15" pageNumber="153">Dc</emphasis>
</th>
<th id="21205DAC6334A0DFFAE9FD2EF4E18596" box="[1395,1446,682,743]" gridcol="8" gridrow="0" pageId="15" pageNumber="153">
<emphasis id="EEB5EA8063345F2FFAE9FD38F4C885A4" bold="true" box="[1395,1423,700,725]" pageId="15" pageNumber="153">Sq</emphasis>
</th>
<th id="21205DAC6334A0DFFA54FD2EF7498596" box="[1486,1550,682,743]" gridcol="9" gridrow="0" pageId="15" pageNumber="153">
<emphasis id="EEB5EA8063345F2FFA54FD2EF74B8596" bold="true" box="[1486,1548,682,743]" pageId="15" pageNumber="153">Costa ext. L</emphasis>
</th>
<th id="21205DAC6334A0DFF9D7FD2EF78B8596" box="[1613,1740,682,743]" gridcol="10" gridrow="0" pageId="15" pageNumber="153">
<emphasis id="EEB5EA8063345F2FF9D7FD38F78B85A4" bold="true" box="[1613,1740,700,725]" pageId="15" pageNumber="153">AnP L (μm)</emphasis>
</th>
<th id="21205DAC6334A0DFF89FFD2EF61E8596" box="[1797,1881,682,743]" gridcol="11" gridrow="0" pageId="15" pageNumber="153">
<emphasis id="EEB5EA8063345F2FF89FFD38F61E85A4" bold="true" box="[1797,1881,700,725]" pageId="15" pageNumber="153">AnP LS</emphasis>
</th>
</tr>
<tr id="62F134D06334A0DFFF38FD7CF61E8460" box="[162,1881,760,785]" gridrow="1" pageId="15" pageNumber="153">
<th id="21205DAC6334A0DFFF38FD7CF0288460" box="[162,367,760,785]" gridcol="0" gridrow="1" pageId="15" pageNumber="153">
<taxonomicName id="1BC14D1163345F2FFF38FD7CF04A8460" box="[162,269,760,785]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="153" phylum="Arthropoda" rank="species" species="davoodi">
<emphasis id="EEB5EA8063345F2FFF38FD7CF04A8460" bold="true" box="[162,269,760,785]" italics="true" pageId="15" pageNumber="153">P. davoodi</emphasis>
</taxonomicName>
</th>
<td id="21205DAC6334A0DFFE3EFD7CF3418460" box="[420,518,760,785]" gridcol="1" gridrow="1" pageId="15" pageNumber="153">2.1–2.4</td>
<td id="21205DAC6334A0DFFDBDFD7CF3D68460" box="[551,657,760,785]" gridcol="2" gridrow="1" pageId="15" pageNumber="153">1.3–1.5</td>
<td id="21205DAC6334A0DFFD33FD7CF2458460" box="[681,770,760,785]" gridcol="3" gridrow="1" pageId="15" pageNumber="153">204</td>
<td id="21205DAC6334A0DFFCA4FD7CF2AC8460" box="[830,1003,760,785]" gridcol="4" gridrow="1" pageId="15" pageNumber="153">Whitish</td>
<td id="21205DAC6334A0DFFB83FD7CF5388460" box="[1049,1151,760,785]" gridcol="5" gridrow="1" pageId="15" pageNumber="153">0.9–1.0</td>
<td id="21205DAC6334A0DFFB28FD7CF5A38460" box="[1202,1252,760,785]" gridcol="6" gridrow="1" pageId="15" pageNumber="153">12</td>
<td id="21205DAC6334A0DFFA99FD7CF4058460" box="[1283,1346,760,785]" gridcol="7" gridrow="1" pageId="15" pageNumber="153">9–12</td>
<td id="21205DAC6334A0DFFAE9FD7CF4E18460" box="[1395,1446,760,785]" gridcol="8" gridrow="1" pageId="15" pageNumber="153">4</td>
<td id="21205DAC6334A0DFFA54FD7CF7498460" box="[1486,1550,760,785]" gridcol="9" gridrow="1" pageId="15" pageNumber="153">22</td>
<td id="21205DAC6334A0DFF9D7FD7CF78B8460" box="[1613,1740,760,785]" gridcol="10" gridrow="1" pageId="15" pageNumber="153">22–26</td>
<td id="21205DAC6334A0DFF89FFD7CF61E8460" box="[1797,1881,760,785]" gridcol="11" gridrow="1" pageId="15" pageNumber="153">6</td>
</tr>
<tr id="62F134D06334A0DFFF38FCA7F61E844D" box="[162,1881,803,828]" gridrow="2" pageId="15" pageNumber="153">
<th id="21205DAC6334A0DFFF38FCA7F028844D" box="[162,367,803,828]" gridcol="0" gridrow="2" pageId="15" pageNumber="153">
<taxonomicName id="1BC14D1163345F2FFF38FCA7F1BA844D" authorityName="Saether" authorityYear="1980" box="[162,253,803,828]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="153" phylum="Arthropoda" rank="species" species="glaber">
<emphasis id="EEB5EA8063345F2FFF38FCA7F1BA844D" bold="true" box="[162,253,803,828]" italics="true" pageId="15" pageNumber="153">P. glaber</emphasis>
</taxonomicName>
</th>
<td id="21205DAC6334A0DFFE3EFCA7F341844D" box="[420,518,803,828]" gridcol="1" gridrow="2" pageId="15" pageNumber="153">2.7</td>
<td id="21205DAC6334A0DFFDBDFCA7F3D6844D" box="[551,657,803,828]" gridcol="2" gridrow="2" pageId="15" pageNumber="153">1.6</td>
<td id="21205DAC6334A0DFFD33FCA7F245844D" box="[681,770,803,828]" gridcol="3" gridrow="2" pageId="15" pageNumber="153">281</td>
<td id="21205DAC6334A0DFFCA4FCA7F2AC844D" box="[830,1003,803,828]" gridcol="4" gridrow="2" pageId="15" pageNumber="153">Brown</td>
<td id="21205DAC6334A0DFFB83FCA7F538844D" box="[1049,1151,803,828]" gridcol="5" gridrow="2" pageId="15" pageNumber="153">0.70–0.95</td>
<td id="21205DAC6334A0DFFB28FCA7F5A3844D" box="[1202,1252,803,828]" gridcol="6" gridrow="2" pageId="15" pageNumber="153">8</td>
<td id="21205DAC6334A0DFFA99FCA7F405844D" box="[1283,1346,803,828]" gridcol="7" gridrow="2" pageId="15" pageNumber="153">11</td>
<td id="21205DAC6334A0DFFAE9FCA7F4E1844D" box="[1395,1446,803,828]" gridcol="8" gridrow="2" pageId="15" pageNumber="153">7</td>
<td id="21205DAC6334A0DFFA54FCA7F749844D" box="[1486,1550,803,828]" gridcol="9" gridrow="2" pageId="15" pageNumber="153">54</td>
<td id="21205DAC6334A0DFF9D7FCA7F78B844D" box="[1613,1740,803,828]" gridcol="10" gridrow="2" pageId="15" pageNumber="153">23–31</td>
<td id="21205DAC6334A0DFF89FFCA7F61E844D" box="[1797,1881,803,828]" gridcol="11" gridrow="2" pageId="15" pageNumber="153">9</td>
</tr>
<tr id="62F134D06334A0DFFF38FCC9F61E8417" box="[162,1881,845,870]" gridrow="3" pageId="15" pageNumber="153">
<th id="21205DAC6334A0DFFF38FCC9F0288417" box="[162,367,845,870]" gridcol="0" gridrow="3" pageId="15" pageNumber="153">
<taxonomicName id="1BC14D1163345F2FFF38FCC9F05C8417" baseAuthorityName="Bhattacharyay &amp; Chaudhuri" baseAuthorityYear="1987" box="[162,283,845,870]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="153" phylum="Arthropoda" rank="species" species="insulatus">
<emphasis id="EEB5EA8063345F2FFF38FCC9F05C8417" bold="true" box="[162,283,845,870]" italics="true" pageId="15" pageNumber="153">P. insulatus</emphasis>
</taxonomicName>
</th>
<td id="21205DAC6334A0DFFE3EFCC9F3418417" box="[420,518,845,870]" gridcol="1" gridrow="3" pageId="15" pageNumber="153">2.8–3.0</td>
<td id="21205DAC6334A0DFFDBDFCC9F3D68417" box="[551,657,845,870]" gridcol="2" gridrow="3" pageId="15" pageNumber="153">1.6–1.7</td>
<td id="21205DAC6334A0DFFD33FCC9F2458417" box="[681,770,845,870]" gridcol="3" gridrow="3" pageId="15" pageNumber="153">90</td>
<td id="21205DAC6334A0DFFCA4FCC9F2AC8417" box="[830,1003,845,870]" gridcol="4" gridrow="3" pageId="15" pageNumber="153">Light brown</td>
<td id="21205DAC6334A0DFFB83FCC9F5388417" box="[1049,1151,845,870]" gridcol="5" gridrow="3" pageId="15" pageNumber="153">1</td>
<td id="21205DAC6334A0DFFB28FCC9F5A38417" box="[1202,1252,845,870]" gridcol="6" gridrow="3" pageId="15" pageNumber="153">9</td>
<td id="21205DAC6334A0DFFA99FCC9F4058417" box="[1283,1346,845,870]" gridcol="7" gridrow="3" pageId="15" pageNumber="153">13–14</td>
<td id="21205DAC6334A0DFFAE9FCC9F4E18417" box="[1395,1446,845,870]" gridcol="8" gridrow="3" pageId="15" pageNumber="153">1</td>
<td id="21205DAC6334A0DFFA54FCC9F7498417" box="[1486,1550,845,870]" gridcol="9" gridrow="3" pageId="15" pageNumber="153">~ 66</td>
<td id="21205DAC6334A0DFF9D7FCC9F78B8417" box="[1613,1740,845,870]" gridcol="10" gridrow="3" pageId="15" pageNumber="153">33</td>
<td id="21205DAC6334A0DFF89FFCC9F61E8417" box="[1797,1881,845,870]" gridcol="11" gridrow="3" pageId="15" pageNumber="153">0</td>
</tr>
<tr id="62F134D06334A0DFFF38FCF3F61E84E1" box="[162,1881,887,912]" gridrow="4" pageId="15" pageNumber="153">
<th id="21205DAC6334A0DFFF38FCF3F02884E1" box="[162,367,887,912]" gridcol="0" gridrow="4" pageId="15" pageNumber="153">
<taxonomicName id="1BC14D1163345F2FFF38FCF3F06D84E1" authorityName="Niitsuma" authorityYear="1990" box="[162,298,887,912]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="153" phylum="Arthropoda" rank="species" species="irregularis">
<emphasis id="EEB5EA8063345F2FFF38FCF3F06D84E1" bold="true" box="[162,298,887,912]" italics="true" pageId="15" pageNumber="153">P. irregularis</emphasis>
</taxonomicName>
</th>
<td id="21205DAC6334A0DFFE3EFCF3F34184E1" box="[420,518,887,912]" gridcol="1" gridrow="4" pageId="15" pageNumber="153">1.7–2.3</td>
<td id="21205DAC6334A0DFFDBDFCF3F3D684E1" box="[551,657,887,912]" gridcol="2" gridrow="4" pageId="15" pageNumber="153">1.3–1.6</td>
<td id="21205DAC6334A0DFFD33FCF3F24584E1" box="[681,770,887,912]" gridcol="3" gridrow="4" pageId="15" pageNumber="153">─</td>
<td id="21205DAC6334A0DFFCA4FCF3F2AC84E1" box="[830,1003,887,912]" gridcol="4" gridrow="4" pageId="15" pageNumber="153">Dark brown</td>
<td id="21205DAC6334A0DFFB83FCF3F53884E1" box="[1049,1151,887,912]" gridcol="5" gridrow="4" pageId="15" pageNumber="153">0.24–0.30</td>
<td id="21205DAC6334A0DFFB28FCF3F5A384E1" box="[1202,1252,887,912]" gridcol="6" gridrow="4" pageId="15" pageNumber="153">4–7</td>
<td id="21205DAC6334A0DFFA99FCF3F40584E1" box="[1283,1346,887,912]" gridcol="7" gridrow="4" pageId="15" pageNumber="153">7–10</td>
<td id="21205DAC6334A0DFFAE9FCF3F4E184E1" box="[1395,1446,887,912]" gridcol="8" gridrow="4" pageId="15" pageNumber="153">8–16</td>
<td id="21205DAC6334A0DFFA54FCF3F74984E1" box="[1486,1550,887,912]" gridcol="9" gridrow="4" pageId="15" pageNumber="153">─</td>
<td id="21205DAC6334A0DFF9D7FCF3F78B84E1" box="[1613,1740,887,912]" gridcol="10" gridrow="4" pageId="15" pageNumber="153">40–44</td>
<td id="21205DAC6334A0DFF89FFCF3F61E84E1" box="[1797,1881,887,912]" gridcol="11" gridrow="4" pageId="15" pageNumber="153">11–17</td>
</tr>
<tr id="62F134D06334A0DFFF38FC24F61E8370" box="[162,1881,928,1025]" gridrow="5" pageId="15" pageNumber="153">
<th id="21205DAC6334A0DFFF38FC24F0288370" box="[162,367,928,1025]" gridcol="0" gridrow="5" pageId="15" pageNumber="153">
<taxonomicName id="1BC14D1163345F2FFF38FC40F00184AC" authorityName="Caldwell" authorityYear="1985" box="[162,326,964,989]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="153" phylum="Arthropoda" rank="species" species="millrockensis">
<emphasis id="EEB5EA8063345F2FFF38FC40F00184AC" bold="true" box="[162,326,964,989]" italics="true" pageId="15" pageNumber="153">P. millrockensis</emphasis>
</taxonomicName>
</th>
<td id="21205DAC6334A0DFFE3EFC24F3418370" box="[420,518,928,1025]" gridcol="1" gridrow="5" pageId="15" pageNumber="153">1.8</td>
<td id="21205DAC6334A0DFFDBDFC24F3D68370" box="[551,657,928,1025]" gridcol="2" gridrow="5" pageId="15" pageNumber="153">0.97</td>
<td id="21205DAC6334A0DFFD33FC24F2458370" box="[681,770,928,1025]" gridcol="3" gridrow="5" pageId="15" pageNumber="153">─</td>
<td id="21205DAC6334A0DFFCA4FC24F2AC8370" box="[830,1003,928,1025]" gridcol="4" gridrow="5" pageId="15" pageNumber="153">Basally brown, apically light brown</td>
<td id="21205DAC6334A0DFFB83FC24F5388370" box="[1049,1151,928,1025]" gridcol="5" gridrow="5" pageId="15" pageNumber="153">0.62–0.63</td>
<td id="21205DAC6334A0DFFB28FC24F5A38370" box="[1202,1252,928,1025]" gridcol="6" gridrow="5" pageId="15" pageNumber="153">5</td>
<td id="21205DAC6334A0DFFA99FC24F4058370" box="[1283,1346,928,1025]" gridcol="7" gridrow="5" pageId="15" pageNumber="153">6–9</td>
<td id="21205DAC6334A0DFFAE9FC24F4E18370" box="[1395,1446,928,1025]" gridcol="8" gridrow="5" pageId="15" pageNumber="153">3</td>
<td id="21205DAC6334A0DFFA54FC24F7498370" box="[1486,1550,928,1025]" gridcol="9" gridrow="5" pageId="15" pageNumber="153">─</td>
<td id="21205DAC6334A0DFF9D7FC24F78B8370" box="[1613,1740,928,1025]" gridcol="10" gridrow="5" pageId="15" pageNumber="153">17–26</td>
<td id="21205DAC6334A0DFF89FFC24F61E8370" box="[1797,1881,928,1025]" gridcol="11" gridrow="5" pageId="15" pageNumber="153">2–3</td>
</tr>
<tr id="62F134D06334A0DFFF38FB94F61E8300" box="[162,1881,1040,1137]" gridrow="6" pageId="15" pageNumber="153">
<th id="21205DAC6334A0DFFF38FB94F0288300" box="[162,367,1040,1137]" gridcol="0" gridrow="6" pageId="15" pageNumber="153">
<taxonomicName id="1BC14D1163345F2FFF38FBB0F048833C" authorityName="Chaudhuri &amp; Som" authorityYear="1998" box="[162,271,1076,1101]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="153" phylum="Arthropoda" rank="species" species="missilus">
<emphasis id="EEB5EA8063345F2FFF38FBB0F048833C" bold="true" box="[162,271,1076,1101]" italics="true" pageId="15" pageNumber="153">P. missilus</emphasis>
</taxonomicName>
</th>
<td id="21205DAC6334A0DFFE3EFB94F3418300" box="[420,518,1040,1137]" gridcol="1" gridrow="6" pageId="15" pageNumber="153">2.8–2.85</td>
<td id="21205DAC6334A0DFFDBDFB94F3D68300" box="[551,657,1040,1137]" gridcol="2" gridrow="6" pageId="15" pageNumber="153">1.5</td>
<td id="21205DAC6334A0DFFD33FB94F2458300" box="[681,770,1040,1137]" gridcol="3" gridrow="6" pageId="15" pageNumber="153">─</td>
<td id="21205DAC6334A0DFFCA4FB94F2AC8300" box="[830,1003,1040,1137]" gridcol="4" gridrow="6" pageId="15" pageNumber="153">Basally blackish, apically light brown</td>
<td id="21205DAC6334A0DFFB83FB94F5388300" box="[1049,1151,1040,1137]" gridcol="5" gridrow="6" pageId="15" pageNumber="153">0.27</td>
<td id="21205DAC6334A0DFFB28FB94F5A38300" box="[1202,1252,1040,1137]" gridcol="6" gridrow="6" pageId="15" pageNumber="153">5–6</td>
<td id="21205DAC6334A0DFFA99FB94F4058300" box="[1283,1346,1040,1137]" gridcol="7" gridrow="6" pageId="15" pageNumber="153">8</td>
<td id="21205DAC6334A0DFFAE9FB94F4E18300" box="[1395,1446,1040,1137]" gridcol="8" gridrow="6" pageId="15" pageNumber="153">14</td>
<td id="21205DAC6334A0DFFA54FB94F7498300" box="[1486,1550,1040,1137]" gridcol="9" gridrow="6" pageId="15" pageNumber="153">48–68</td>
<td id="21205DAC6334A0DFF9D7FB94F78B8300" box="[1613,1740,1040,1137]" gridcol="10" gridrow="6" pageId="15" pageNumber="153">36–40</td>
<td id="21205DAC6334A0DFF89FFB94F61E8300" box="[1797,1881,1040,1137]" gridcol="11" gridrow="6" pageId="15" pageNumber="153">2–3</td>
</tr>
<tr id="62F134D06334A0DFFF38FB05F61E83EB" box="[162,1881,1153,1178]" gridrow="7" pageId="15" pageNumber="153">
<th id="21205DAC6334A0DFFF38FB05F02883EB" box="[162,367,1153,1178]" gridcol="0" gridrow="7" pageId="15" pageNumber="153">
<taxonomicName id="1BC14D1163345F2FFF38FB05F04D83EB" authorityName="Steiner" authorityYear="1983" box="[162,266,1153,1178]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="153" phylum="Arthropoda" rank="species" species="mozleyi">
<emphasis id="EEB5EA8063345F2FFF38FB05F04D83EB" bold="true" box="[162,266,1153,1178]" italics="true" pageId="15" pageNumber="153">P. mozleyi</emphasis>
</taxonomicName>
</th>
<td id="21205DAC6334A0DFFE3EFB05F34183EB" box="[420,518,1153,1178]" gridcol="1" gridrow="7" pageId="15" pageNumber="153">3.2</td>
<td id="21205DAC6334A0DFFDBDFB05F3D683EB" box="[551,657,1153,1178]" gridcol="2" gridrow="7" pageId="15" pageNumber="153">─</td>
<td id="21205DAC6334A0DFFD33FB05F24583EB" box="[681,770,1153,1178]" gridcol="3" gridrow="7" pageId="15" pageNumber="153">─</td>
<td id="21205DAC6334A0DFFCA4FB05F2AC83EB" box="[830,1003,1153,1178]" gridcol="4" gridrow="7" pageId="15" pageNumber="153">Dark brown</td>
<td id="21205DAC6334A0DFFB83FB05F53883EB" box="[1049,1151,1153,1178]" gridcol="5" gridrow="7" pageId="15" pageNumber="153">0.84</td>
<td id="21205DAC6334A0DFFB28FB05F5A383EB" box="[1202,1252,1153,1178]" gridcol="6" gridrow="7" pageId="15" pageNumber="153">9</td>
<td id="21205DAC6334A0DFFA99FB05F40583EB" box="[1283,1346,1153,1178]" gridcol="7" gridrow="7" pageId="15" pageNumber="153">6–9</td>
<td id="21205DAC6334A0DFFAE9FB05F4E183EB" box="[1395,1446,1153,1178]" gridcol="8" gridrow="7" pageId="15" pageNumber="153">3–4</td>
<td id="21205DAC6334A0DFFA54FB05F74983EB" box="[1486,1550,1153,1178]" gridcol="9" gridrow="7" pageId="15" pageNumber="153">─</td>
<td id="21205DAC6334A0DFF9D7FB05F78B83EB" box="[1613,1740,1153,1178]" gridcol="10" gridrow="7" pageId="15" pageNumber="153">8–11</td>
<td id="21205DAC6334A0DFF89FFB05F61E83EB" box="[1797,1881,1153,1178]" gridcol="11" gridrow="7" pageId="15" pageNumber="153">0</td>
</tr>
<tr id="62F134D06334A0DFFF38FB2FF61E83B5" box="[162,1881,1195,1220]" gridrow="8" pageId="15" pageNumber="153">
<th id="21205DAC6334A0DFFF38FB2FF02883B5" box="[162,367,1195,1220]" gridcol="0" gridrow="8" pageId="15" pageNumber="153">
<taxonomicName id="1BC14D1163345F2FFF38FB2FF1B583B5" authorityName="sensu Saether" authorityYear="1980" baseAuthorityName="Kieffer" baseAuthorityYear="1913" box="[162,242,1195,1220]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="153" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="EEB5EA8063345F2FFF38FB2FF1B583B5" bold="true" box="[162,242,1195,1220]" italics="true" pageId="15" pageNumber="153">P. niger</emphasis>
</taxonomicName>
</th>
<td id="21205DAC6334A0DFFE3EFB2FF34183B5" box="[420,518,1195,1220]" gridcol="1" gridrow="8" pageId="15" pageNumber="153">1.8–2.4</td>
<td id="21205DAC6334A0DFFDBDFB2FF3D683B5" box="[551,657,1195,1220]" gridcol="2" gridrow="8" pageId="15" pageNumber="153">1.0–1.5</td>
<td id="21205DAC6334A0DFFD33FB2FF24583B5" box="[681,770,1195,1220]" gridcol="3" gridrow="8" pageId="15" pageNumber="153">255–256</td>
<td id="21205DAC6334A0DFFCA4FB2FF2AC83B5" box="[830,1003,1195,1220]" gridcol="4" gridrow="8" pageId="15" pageNumber="153">Dark brown</td>
<td id="21205DAC6334A0DFFB83FB2FF53883B5" box="[1049,1151,1195,1220]" gridcol="5" gridrow="8" pageId="15" pageNumber="153">0.51–0.73</td>
<td id="21205DAC6334A0DFFB28FB2FF5A383B5" box="[1202,1252,1195,1220]" gridcol="6" gridrow="8" pageId="15" pageNumber="153">2–9</td>
<td id="21205DAC6334A0DFFA99FB2FF40583B5" box="[1283,1346,1195,1220]" gridcol="7" gridrow="8" pageId="15" pageNumber="153">8–10</td>
<td id="21205DAC6334A0DFFAE9FB2FF4E183B5" box="[1395,1446,1195,1220]" gridcol="8" gridrow="8" pageId="15" pageNumber="153">6–12</td>
<td id="21205DAC6334A0DFFA54FB2FF74983B5" box="[1486,1550,1195,1220]" gridcol="9" gridrow="8" pageId="15" pageNumber="153">41–54</td>
<td id="21205DAC6334A0DFF9D7FB2FF78B83B5" box="[1613,1740,1195,1220]" gridcol="10" gridrow="8" pageId="15" pageNumber="153">32–55</td>
<td id="21205DAC6334A0DFF89FFB2FF61E83B5" box="[1797,1881,1195,1220]" gridcol="11" gridrow="8" pageId="15" pageNumber="153">6–8</td>
</tr>
<tr id="62F134D06334A0DFFF38FB51F61E839F" box="[162,1881,1237,1262]" gridrow="9" pageId="15" pageNumber="153">
<th id="21205DAC6334A0DFFF38FB51F028839F" box="[162,367,1237,1262]" gridcol="0" gridrow="9" pageId="15" pageNumber="153">
<taxonomicName id="1BC14D1163345F2FFF38FB51F027839F" authorityName="Sasa, Kwai &amp; Ueno" authorityYear="1988" box="[162,352,1237,1262]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="153" phylum="Arthropoda" rank="species" species="oyabeangulatus">
<emphasis id="EEB5EA8063345F2FFF38FB51F027839F" bold="true" box="[162,352,1237,1262]" italics="true" pageId="15" pageNumber="153">P. oyabeangulatus</emphasis>
</taxonomicName>
</th>
<td id="21205DAC6334A0DFFE3EFB51F341839F" box="[420,518,1237,1262]" gridcol="1" gridrow="9" pageId="15" pageNumber="153">1.9</td>
<td id="21205DAC6334A0DFFDBDFB51F3D6839F" box="[551,657,1237,1262]" gridcol="2" gridrow="9" pageId="15" pageNumber="153">1.1</td>
<td id="21205DAC6334A0DFFD33FB51F245839F" box="[681,770,1237,1262]" gridcol="3" gridrow="9" pageId="15" pageNumber="153">─</td>
<td id="21205DAC6334A0DFFCA4FB51F2AC839F" box="[830,1003,1237,1262]" gridcol="4" gridrow="9" pageId="15" pageNumber="153">─</td>
<td id="21205DAC6334A0DFFB83FB51F538839F" box="[1049,1151,1237,1262]" gridcol="5" gridrow="9" pageId="15" pageNumber="153">0.38</td>
<td id="21205DAC6334A0DFFB28FB51F5A3839F" box="[1202,1252,1237,1262]" gridcol="6" gridrow="9" pageId="15" pageNumber="153">10</td>
<td id="21205DAC6334A0DFFA99FB51F405839F" box="[1283,1346,1237,1262]" gridcol="7" gridrow="9" pageId="15" pageNumber="153">8</td>
<td id="21205DAC6334A0DFFAE9FB51F4E1839F" box="[1395,1446,1237,1262]" gridcol="8" gridrow="9" pageId="15" pageNumber="153">5</td>
<td id="21205DAC6334A0DFFA54FB51F749839F" box="[1486,1550,1237,1262]" gridcol="9" gridrow="9" pageId="15" pageNumber="153">~ 86</td>
<td id="21205DAC6334A0DFF9D7FB51F78B839F" box="[1613,1740,1237,1262]" gridcol="10" gridrow="9" pageId="15" pageNumber="153">─</td>
<td id="21205DAC6334A0DFF89FFB51F61E839F" box="[1797,1881,1237,1262]" gridcol="11" gridrow="9" pageId="15" pageNumber="153">─</td>
</tr>
<tr id="62F134D06334A0DFFF38FB7AF61E822E" box="[162,1881,1278,1375]" gridrow="10" pageId="15" pageNumber="153">
<th id="21205DAC6334A0DFFF38FB7AF028822E" box="[162,367,1278,1375]" gridcol="0" gridrow="10" pageId="15" pageNumber="153">
<taxonomicName id="1BC14D1163345F2FFF38FAA6F069824A" authorityName="Hazra &amp; Chaudhuri" authorityYear="2002" box="[162,302,1314,1339]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="153" phylum="Arthropoda" rank="species" species="spinicornis">
<emphasis id="EEB5EA8063345F2FFF38FAA6F069824A" bold="true" box="[162,302,1314,1339]" italics="true" pageId="15" pageNumber="153">P. spinicornis</emphasis>
</taxonomicName>
</th>
<td id="21205DAC6334A0DFFE3EFB7AF341822E" box="[420,518,1278,1375]" gridcol="1" gridrow="10" pageId="15" pageNumber="153">1.6–2.3</td>
<td id="21205DAC6334A0DFFDBDFB7AF3D6822E" box="[551,657,1278,1375]" gridcol="2" gridrow="10" pageId="15" pageNumber="153">0.9–1.1</td>
<td id="21205DAC6334A0DFFD33FB7AF245822E" box="[681,770,1278,1375]" gridcol="3" gridrow="10" pageId="15" pageNumber="153">─</td>
<td id="21205DAC6334A0DFFCA4FB7AF2AC822E" box="[830,1003,1278,1375]" gridcol="4" gridrow="10" pageId="15" pageNumber="153">Basally blackish, apically light brown</td>
<td id="21205DAC6334A0DFFB83FB7AF538822E" box="[1049,1151,1278,1375]" gridcol="5" gridrow="10" pageId="15" pageNumber="153">0.28–0.37</td>
<td id="21205DAC6334A0DFFB28FB7AF5A3822E" box="[1202,1252,1278,1375]" gridcol="6" gridrow="10" pageId="15" pageNumber="153">7–15</td>
<td id="21205DAC6334A0DFFA99FB7AF405822E" box="[1283,1346,1278,1375]" gridcol="7" gridrow="10" pageId="15" pageNumber="153">8–12</td>
<td id="21205DAC6334A0DFFAE9FB7AF4E1822E" box="[1395,1446,1278,1375]" gridcol="8" gridrow="10" pageId="15" pageNumber="153">4–6</td>
<td id="21205DAC6334A0DFFA54FB7AF749822E" box="[1486,1550,1278,1375]" gridcol="9" gridrow="10" pageId="15" pageNumber="153">~ 30</td>
<td id="21205DAC6334A0DFF9D7FB7AF78B822E" box="[1613,1740,1278,1375]" gridcol="10" gridrow="10" pageId="15" pageNumber="153">29–33</td>
<td id="21205DAC6334A0DFF89FFB7AF61E822E" box="[1797,1881,1278,1375]" gridcol="11" gridrow="10" pageId="15" pageNumber="153">2–3</td>
</tr>
<tr id="62F134D06334A0DFFF38FAEBF61E82F8" box="[162,1881,1391,1417]" gridrow="11" pageId="15" pageNumber="153">
<th id="21205DAC6334A0DFFF38FAEBF02882F8" box="[162,367,1391,1417]" gridcol="0" gridrow="11" pageId="15" pageNumber="153">
<emphasis id="EEB5EA8063345F2FFF38FAEBF01582F9" bold="true" box="[162,338,1391,1416]" pageId="15" pageNumber="153">
<taxonomicName id="1BC14D1163345F2FFF38FAEBF1BB82F9" authorityName="Namayandeh &amp; Ghaderi" authorityYear="2025" box="[162,252,1391,1416]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="153" phylum="Arthropoda" rank="species" species="sturae" status="sp. nov.">
<emphasis id="EEB5EA8063345F2FFF38FAEBF1BB82F9" bold="true" box="[162,252,1391,1416]" italics="true" pageId="15" pageNumber="153">P. sturae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F58657FB63345F2FFE98FAEBF01582F9" box="[258,338,1391,1416]" pageId="15" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="21205DAC6334A0DFFE3EFAEBF34182F8" box="[420,518,1391,1417]" gridcol="1" gridrow="11" pageId="15" pageNumber="153">2.1–2.4</td>
<td id="21205DAC6334A0DFFDBDFAEBF3D682F8" box="[551,657,1391,1417]" gridcol="2" gridrow="11" pageId="15" pageNumber="153">1.3–1.04</td>
<td id="21205DAC6334A0DFFD33FAEBF24582F8" box="[681,770,1391,1417]" gridcol="3" gridrow="11" pageId="15" pageNumber="153">242</td>
<td id="21205DAC6334A0DFFCA4FAEBF2AC82F8" box="[830,1003,1391,1417]" gridcol="4" gridrow="11" pageId="15" pageNumber="153">Brown</td>
<td id="21205DAC6334A0DFFB83FAEBF53882F8" box="[1049,1151,1391,1417]" gridcol="5" gridrow="11" pageId="15" pageNumber="153">1.5</td>
<td id="21205DAC6334A0DFFB28FAEBF5A382F8" box="[1202,1252,1391,1417]" gridcol="6" gridrow="11" pageId="15" pageNumber="153">12</td>
<td id="21205DAC6334A0DFFA99FAEBF40582F8" box="[1283,1346,1391,1417]" gridcol="7" gridrow="11" pageId="15" pageNumber="153">7</td>
<td id="21205DAC6334A0DFFAE9FAEBF4E182F8" box="[1395,1446,1391,1417]" gridcol="8" gridrow="11" pageId="15" pageNumber="153">8</td>
<td id="21205DAC6334A0DFFA54FAEBF74982F8" box="[1486,1550,1391,1417]" gridcol="9" gridrow="11" pageId="15" pageNumber="153">61–66</td>
<td id="21205DAC6334A0DFF9D7FAEBF78B82F8" box="[1613,1740,1391,1417]" gridcol="10" gridrow="11" pageId="15" pageNumber="153">21–29</td>
<td id="21205DAC6334A0DFF89FFAEBF61E82F8" box="[1797,1881,1391,1417]" gridcol="11" gridrow="11" pageId="15" pageNumber="153">4–6</td>
</tr>
<tr id="62F134D06334A0DFFF38FA1DF61E82C2" box="[162,1881,1433,1459]" gridrow="12" pageId="15" pageNumber="153">
<th id="21205DAC6334A0DFFF38FA1DF02882C2" box="[162,367,1433,1459]" gridcol="0" gridrow="12" pageId="15" pageNumber="153">
<taxonomicName id="1BC14D1163345F2FFF38FA1DF06A82C3" baseAuthorityName="Sasa" baseAuthorityYear="1983" box="[162,301,1433,1458]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="153" phylum="Arthropoda" rank="species" species="tamabrevis">
<emphasis id="EEB5EA8063345F2FFF38FA1DF06A82C3" bold="true" box="[162,301,1433,1458]" italics="true" pageId="15" pageNumber="153">P. tamabrevis</emphasis>
</taxonomicName>
</th>
<td id="21205DAC6334A0DFFE3EFA1DF34182C2" box="[420,518,1433,1459]" gridcol="1" gridrow="12" pageId="15" pageNumber="153">1.4–1.6</td>
<td id="21205DAC6334A0DFFDBDFA1DF3D682C2" box="[551,657,1433,1459]" gridcol="2" gridrow="12" pageId="15" pageNumber="153">1.0–1.3</td>
<td id="21205DAC6334A0DFFD33FA1DF24582C2" box="[681,770,1433,1459]" gridcol="3" gridrow="12" pageId="15" pageNumber="153">─</td>
<td id="21205DAC6334A0DFFCA4FA1DF2AC82C2" box="[830,1003,1433,1459]" gridcol="4" gridrow="12" pageId="15" pageNumber="153">Dark brown</td>
<td id="21205DAC6334A0DFFB83FA1DF53882C2" box="[1049,1151,1433,1459]" gridcol="5" gridrow="12" pageId="15" pageNumber="153">0.31–0.37</td>
<td id="21205DAC6334A0DFFB28FA1DF5A382C2" box="[1202,1252,1433,1459]" gridcol="6" gridrow="12" pageId="15" pageNumber="153">6–9</td>
<td id="21205DAC6334A0DFFA99FA1DF40582C2" box="[1283,1346,1433,1459]" gridcol="7" gridrow="12" pageId="15" pageNumber="153">8–10</td>
<td id="21205DAC6334A0DFFAE9FA1DF4E182C2" box="[1395,1446,1433,1459]" gridcol="8" gridrow="12" pageId="15" pageNumber="153">2–6</td>
<td id="21205DAC6334A0DFFA54FA1DF74982C2" box="[1486,1550,1433,1459]" gridcol="9" gridrow="12" pageId="15" pageNumber="153">~75</td>
<td id="21205DAC6334A0DFF9D7FA1DF78B82C2" box="[1613,1740,1433,1459]" gridcol="10" gridrow="12" pageId="15" pageNumber="153">21–25</td>
<td id="21205DAC6334A0DFF89FFA1DF61E82C2" box="[1797,1881,1433,1459]" gridcol="11" gridrow="12" pageId="15" pageNumber="153">8–10</td>
</tr>
<tr id="62F134D06334A0DFFF38FA47F61E82AC" box="[162,1881,1475,1501]" gridrow="13" pageId="15" pageNumber="153">
<th id="21205DAC6334A0DFFF38FA47F02882AC" box="[162,367,1475,1501]" gridcol="0" gridrow="13" pageId="15" pageNumber="153">
<taxonomicName id="1BC14D1163345F2FFF38FA47F07082AD" baseAuthorityName="Sasa &amp; Okazawa" baseAuthorityYear="1992" box="[162,311,1475,1500]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="153" phylum="Arthropoda" rank="species" species="togakuroasi">
<emphasis id="EEB5EA8063345F2FFF38FA47F07082AD" bold="true" box="[162,311,1475,1500]" italics="true" pageId="15" pageNumber="153">P. togakuroasi</emphasis>
</taxonomicName>
</th>
<td id="21205DAC6334A0DFFE3EFA47F34182AC" box="[420,518,1475,1501]" gridcol="1" gridrow="13" pageId="15" pageNumber="153">2.6</td>
<td id="21205DAC6334A0DFFDBDFA47F3D682AC" box="[551,657,1475,1501]" gridcol="2" gridrow="13" pageId="15" pageNumber="153">1.6</td>
<td id="21205DAC6334A0DFFD33FA47F24582AC" box="[681,770,1475,1501]" gridcol="3" gridrow="13" pageId="15" pageNumber="153">─</td>
<td id="21205DAC6334A0DFFCA4FA47F2AC82AC" box="[830,1003,1475,1501]" gridcol="4" gridrow="13" pageId="15" pageNumber="153">─</td>
<td id="21205DAC6334A0DFFB83FA47F53882AC" box="[1049,1151,1475,1501]" gridcol="5" gridrow="13" pageId="15" pageNumber="153">0.71</td>
<td id="21205DAC6334A0DFFB28FA47F5A382AC" box="[1202,1252,1475,1501]" gridcol="6" gridrow="13" pageId="15" pageNumber="153">10</td>
<td id="21205DAC6334A0DFFA99FA47F40582AC" box="[1283,1346,1475,1501]" gridcol="7" gridrow="13" pageId="15" pageNumber="153">8</td>
<td id="21205DAC6334A0DFFAE9FA47F4E182AC" box="[1395,1446,1475,1501]" gridcol="8" gridrow="13" pageId="15" pageNumber="153">4–5</td>
<td id="21205DAC6334A0DFFA54FA47F74982AC" box="[1486,1550,1475,1501]" gridcol="9" gridrow="13" pageId="15" pageNumber="153">~ 50</td>
<td id="21205DAC6334A0DFF9D7FA47F78B82AC" box="[1613,1740,1475,1501]" gridcol="10" gridrow="13" pageId="15" pageNumber="153">42</td>
<td id="21205DAC6334A0DFF89FFA47F61E82AC" box="[1797,1881,1475,1501]" gridcol="11" gridrow="13" pageId="15" pageNumber="153">2</td>
</tr>
<tr id="62F134D06334A0DFFF38FA69F61E8176" box="[162,1881,1517,1543]" gridrow="14" pageId="15" pageNumber="153">
<th id="21205DAC6334A0DFFF38FA69F0288176" box="[162,367,1517,1543]" gridcol="0" gridrow="14" pageId="15" pageNumber="153">
<emphasis id="EEB5EA8063345F2FFF38FA69F0288177" bold="true" box="[162,367,1517,1542]" pageId="15" pageNumber="153">
<taxonomicName id="1BC14D1163345F2FFF38FA69F05F8177" authorityName="Namayandeh &amp; Ghaderi" authorityYear="2025" box="[162,280,1517,1542]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="153" phylum="Arthropoda" rank="species" species="torbjorni" status="sp. nov.">
<emphasis id="EEB5EA8063345F2FFF38FA69F05F8177" bold="true" box="[162,280,1517,1542]" italics="true" pageId="15" pageNumber="153">P. torbjorni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F58657FB63345F2FFE84FA69F0288177" box="[286,367,1517,1542]" pageId="15" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="21205DAC6334A0DFFE3EFA69F3418176" box="[420,518,1517,1543]" gridcol="1" gridrow="14" pageId="15" pageNumber="153">2.1</td>
<td id="21205DAC6334A0DFFDBDFA69F3D68176" box="[551,657,1517,1543]" gridcol="2" gridrow="14" pageId="15" pageNumber="153">1.3</td>
<td id="21205DAC6334A0DFFD33FA69F2458176" box="[681,770,1517,1543]" gridcol="3" gridrow="14" pageId="15" pageNumber="153">259</td>
<td id="21205DAC6334A0DFFCA4FA69F2AC8176" box="[830,1003,1517,1543]" gridcol="4" gridrow="14" pageId="15" pageNumber="153">Brown</td>
<td id="21205DAC6334A0DFFB83FA69F5388176" box="[1049,1151,1517,1543]" gridcol="5" gridrow="14" pageId="15" pageNumber="153">─</td>
<td id="21205DAC6334A0DFFB28FA69F5A38176" box="[1202,1252,1517,1543]" gridcol="6" gridrow="14" pageId="15" pageNumber="153">─</td>
<td id="21205DAC6334A0DFFA99FA69F4058176" box="[1283,1346,1517,1543]" gridcol="7" gridrow="14" pageId="15" pageNumber="153">7</td>
<td id="21205DAC6334A0DFFAE9FA69F4E18176" box="[1395,1446,1517,1543]" gridcol="8" gridrow="14" pageId="15" pageNumber="153">6</td>
<td id="21205DAC6334A0DFFA54FA69F7498176" box="[1486,1550,1517,1543]" gridcol="9" gridrow="14" pageId="15" pageNumber="153">40</td>
<td id="21205DAC6334A0DFF9D7FA69F78B8176" box="[1613,1740,1517,1543]" gridcol="10" gridrow="14" pageId="15" pageNumber="153">13</td>
<td id="21205DAC6334A0DFF89FFA69F61E8176" box="[1797,1881,1517,1543]" gridcol="11" gridrow="14" pageId="15" pageNumber="153">0</td>
</tr>
<tr id="62F134D06334A0DFFF38F993F61E8125" box="[162,1881,1559,1620]" gridrow="15" pageId="15" pageNumber="153">
<th id="21205DAC6334A0DFFF38F993F0288125" box="[162,367,1559,1620]" gridcol="0" gridrow="15" pageId="15" pageNumber="153">
<taxonomicName id="1BC14D1163345F2FFF38F9ADF0608133" authorityName="sensu Saether" authorityYear="1980" baseAuthorityName="Brundin" baseAuthorityYear="1947" box="[162,295,1577,1602]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="153" phylum="Arthropoda" rank="species" species="uliginosus">
<emphasis id="EEB5EA8063345F2FFF38F9ADF0608133" bold="true" box="[162,295,1577,1602]" italics="true" pageId="15" pageNumber="153">P. uliginosus</emphasis>
</taxonomicName>
</th>
<td id="21205DAC6334A0DFFE3EF993F3418125" box="[420,518,1559,1620]" gridcol="1" gridrow="15" pageId="15" pageNumber="153">2.0–2.3</td>
<td id="21205DAC6334A0DFFDBDF993F3D68125" box="[551,657,1559,1620]" gridcol="2" gridrow="15" pageId="15" pageNumber="153">1.3–1.4</td>
<td id="21205DAC6334A0DFFD33F993F2458125" box="[681,770,1559,1620]" gridcol="3" gridrow="15" pageId="15" pageNumber="153">224–240</td>
<td id="21205DAC6334A0DFFCA4F993F2AC8125" box="[830,1003,1559,1620]" gridcol="4" gridrow="15" pageId="15" pageNumber="153">Whitish</td>
<td id="21205DAC6334A0DFFB83F993F5388125" box="[1049,1151,1559,1620]" gridcol="5" gridrow="15" pageId="15" pageNumber="153">0.46–0.56</td>
<td id="21205DAC6334A0DFFB28F993F5A38125" box="[1202,1252,1559,1620]" gridcol="6" gridrow="15" pageId="15" pageNumber="153">12– 14</td>
<td id="21205DAC6334A0DFFA99F993F4058125" box="[1283,1346,1559,1620]" gridcol="7" gridrow="15" pageId="15" pageNumber="153">8</td>
<td id="21205DAC6334A0DFFAE9F993F4E18125" box="[1395,1446,1559,1620]" gridcol="8" gridrow="15" pageId="15" pageNumber="153">3–5</td>
<td id="21205DAC6334A0DFFA54F993F7498125" box="[1486,1550,1559,1620]" gridcol="9" gridrow="15" pageId="15" pageNumber="153">82–97</td>
<td id="21205DAC6334A0DFF9D7F993F78B8125" box="[1613,1740,1559,1620]" gridcol="10" gridrow="15" pageId="15" pageNumber="153">19–26</td>
<td id="21205DAC6334A0DFF89FF993F61E8125" box="[1797,1881,1559,1620]" gridcol="11" gridrow="15" pageId="15" pageNumber="153">6</td>
</tr>
</table>
</paragraph>
<paragraph id="DC7E3692632B5F30FF5DFE33F089827C" blockId="16.[151,1437,151,1293]" pageId="16" pageNumber="154">
The examined
<typeStatus id="037A8830632B5F30FEE9FE33F0E486A0" box="[371,419,439,465]" pageId="16" pageNumber="154">type</typeStatus>
specimens of
<taxonomicName id="1BC14D11632B5F30FDD6FE33F3F786A1" authorityName="Namayandeh &amp; Ghaderi" authorityYear="2025" box="[588,688,439,464]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="species" species="sturae" status="sp. nov.">
<emphasis id="EEB5EA80632B5F30FDD6FE33F3F786A1" box="[588,688,439,464]" italics="true" pageId="16" pageNumber="154">P. sturae</emphasis>
</taxonomicName>
<emphasis id="EEB5EA80632B5F30FD23FE33F25186A0" bold="true" box="[697,790,439,465]" pageId="16" pageNumber="154">
<taxonomicNameLabel id="F58657FB632B5F30FD23FE33F25186A0" box="[697,790,439,465]" pageId="16" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="1BC14D11632B5F30FCCBFE33F29386A0" authorityName="Namayandeh &amp; Ghaderi" authorityYear="2025" box="[849,980,439,465]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="species" species="torbjorni" status="sp. nov.">
<emphasis id="EEB5EA80632B5F30FCCBFE33F29386A0" box="[849,980,439,465]" italics="true" pageId="16" pageNumber="154">P. torbjorni</emphasis>
</taxonomicName>
<emphasis id="EEB5EA80632B5F30FC44FE33F57D86A0" bold="true" box="[990,1082,439,465]" pageId="16" pageNumber="154">
<taxonomicNameLabel id="F58657FB632B5F30FC44FE33F57D86A0" box="[990,1082,439,465]" pageId="16" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
lacked antenna, and therefore, for now, we did not include them in the adult male key. However, they are included in the preliminary alternative key based on adult males’ tergite setation, further separating them from other known species of
<taxonomicName id="1BC14D11632B5F30FB7DFE7BF4D28569" authorityName="Brundin" authorityYear="1956" box="[1255,1429,511,536]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="EEB5EA80632B5F30FB7DFE7BF4D28569" box="[1255,1429,511,536]" italics="true" pageId="16" pageNumber="154">Paracricotopus</emphasis>
</taxonomicName>
. Historically, the generic diagnosis of
<taxonomicName id="1BC14D11632B5F30FDA2FDA7F3A1854D" authorityName="Brundin" authorityYear="1956" box="[568,742,547,572]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="EEB5EA80632B5F30FDA2FDA7F3A1854D" box="[568,742,547,572]" italics="true" pageId="16" pageNumber="154">Paracricotopus</emphasis>
</taxonomicName>
only mentioned the abdominal tergites and sternites in having distinct rows of anterior and posterior setae (Thieneman &amp; Harnisch 1931; Cranston e
<emphasis id="EEB5EA80632B5F30FBF7FDC3F5DD8510" box="[1133,1178,583,609]" italics="true" pageId="16" pageNumber="154">t al.</emphasis>
1989). Some previous workers have provided the total number of setae on tergites (
<bibRefCitation id="B8504B63632B5F30FCDFFDEFF2A985F4" author="Caldwell, B. A." box="[837,1006,618,645]" pageId="16" pageNumber="154" pagination="161 - 168" refId="ref14170" refString="Caldwell, B. A. (1985) Paracricotopus millrockensis a new species of Orthocladiinae (Diptera: Chironomidae) from the southeastern United States. Brimleyana, 11, 161 - 168." type="journal article" year="1985">Caldwell 1985</bibRefCitation>
; Chaudhuri &amp; Som 1998;
<bibRefCitation id="B8504B63632B5F30FA83FDEFF19485D8" author="Hazra, N. &amp; Saha, G. K. &amp; Chaudhuri, P. K." pageId="16" pageNumber="154" pagination="41 - 55" refId="ref14575" refString="Hazra, N., Saha, G. K. &amp; Chaudhuri, P. K. (2002) Records of Orthoclad species from the Darjeeling - Sikkim Himalayas of India (Diptera: Chironomidae), with notes on their ecology. Hydrobiologia, 474, 41 - 55. https: // doi. org / 10.1023 / A: 1016511702944" type="journal article" year="2002">
Hazra
<emphasis id="EEB5EA80632B5F30FAF9FDEFF4D285F4" box="[1379,1429,619,645]" italics="true" pageId="16" pageNumber="154">et al</emphasis>
. 2002
</bibRefCitation>
; Nitsuma 1990;
<bibRefCitation id="B8504B63632B5F30FE10FD0BF36E85D8" author="Saether, O. A." box="[394,553,655,681]" pageId="16" pageNumber="154" pagination="129 - 145" refId="ref15286" refString="Saether, O. A. (1980 b) The Females and immatures of Paracricotopus Thienemann and Harnisch, 1932, with the description of a new species (Diptera: Chironomidae). Aquatic Insects, 2, 129 - 145. https: // doi. org / 10.1080 / 01650428009361018" type="journal article" year="1980">Saether 1980b</bibRefCitation>
). However, the pattern of these setae was never investigated. Based on the
<typeStatus id="037A8830632B5F30FAF6FD0BF4DB85D8" box="[1388,1436,655,681]" pageId="16" pageNumber="154">type</typeStatus>
and voucher specimens we have examined and the review of the literature, we found that each
<taxonomicName id="1BC14D11632B5F30FB74FD37F1AF8580" pageId="16" pageNumber="154">
<emphasis id="EEB5EA80632B5F30FB74FD37F4DB85BD" box="[1262,1436,691,716]" italics="true" pageId="16" pageNumber="154">Paracricotopus</emphasis>
species
</taxonomicName>
has a unique tergite setae pattern, which can be further used as a distinguishing characteristic for adults (
<figureCitation id="44FA2A17632B5F30FF05FD7FF1A38464" box="[159,228,763,789]" captionStart="FIGURE 7" captionStartId="13.[151,250,1841,1865]" captionTargetBox="[318,1314,159,1806]" captionTargetId="figure-113@13.[310,1322,151,1814]" captionTargetPageId="13" captionText="FIGURE 7. Abdominal tergites I–V of Paracricotopus Brundin, spp. adult males. A. Paracricotopus davoodi Ghaderi et al., 2023. B. Paracricotopus glaber Saether, 1980. C.Paracricotopus insulatus (Bhattacharyay &amp; Chaudhuri, 1987), tergite I missing. D. Paracricotopus mozleyi Steiner, 1983. E. Paracricotopus niger (Kieffer, 1913) sensu Saether (1980). F. Paracricotopus spinicornis Hazra &amp; Chaudhuri, 2002. G. Paracricotopus sturae sp. nov. H. Paracricotopus tamabrevis (Sasa, 1983). I. Paracricotopus togakuroasi (Sasa &amp; Okazawa, 1992). J. Paracricotopus torbjorni sp. nov. K. Paracricotopus uliginosus (Brundin, 1947) sensu Saether (1980)." figureDoi="http://doi.org/10.5281/zenodo.14704014" httpUri="https://zenodo.org/record/14704014/files/figure.png" pageId="16" pageNumber="154">Fig. 7</figureCitation>
). We designated the most lateral anterior basal and posterior marginal setae as lateral setae, which we found their numbers is a distinguishing characteristic. Further, we found that the anterior basal and posterior marginal setae of tergite I are diagonally parallel and have a unique pattern format that is species-specific. Also, the setation pattern of tergites I to V is much more distinguishing, and as such, we mainly emphasized these tergites. Based on these characteristics, we provided additional descriptions for most
<taxonomicName id="1BC14D11632B5F30FC27FC0FF58184D4" box="[957,1222,907,933]" pageId="16" pageNumber="154">
<emphasis id="EEB5EA80632B5F30FC27FC0FF52C84D5" box="[957,1131,907,932]" italics="true" pageId="16" pageNumber="154">Paracricotopus</emphasis>
species
</taxonomicName>
and an alternative key to the adult male of the genus. Since we only examined a few specimens from each species, in addition to what little was available in the literature, and could not examine the
<typeStatus id="037A8830632B5F30FC01FC57F28C849C" box="[923,971,979,1005]" pageId="16" pageNumber="154">type</typeStatus>
specimens of
<taxonomicName id="1BC14D11632B5F30FBF4FC57F0068360" authority="Nitsuma, 1990" authorityName="Niitsuma" authorityYear="1990" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="species" species="irregularis">
<emphasis id="EEB5EA80632B5F30FBF4FC57F4DB849C" box="[1134,1436,979,1005]" italics="true" pageId="16" pageNumber="154">Paracricotopus irregularis</emphasis>
Nitsuma, 1990
</taxonomicName>
,
<taxonomicName id="1BC14D11632B5F30FED4FC73F2148360" authority="Caldwell, 1985" authorityName="Caldwell" authorityYear="1985" box="[334,851,1014,1041]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="species" species="millrockensis">
<emphasis id="EEB5EA80632B5F30FED4FC73F3DD8360" box="[334,666,1015,1041]" italics="true" pageId="16" pageNumber="154">Paracricotopus millrockensis</emphasis>
<bibRefCitation id="B8504B63632B5F30FD39FC73F2148360" author="Caldwell, B. A." box="[675,851,1014,1041]" pageId="16" pageNumber="154" pagination="161 - 168" refId="ref14170" refString="Caldwell, B. A. (1985) Paracricotopus millrockensis a new species of Orthocladiinae (Diptera: Chironomidae) from the southeastern United States. Brimleyana, 11, 161 - 168." type="journal article" year="1985">Caldwell, 1985</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="1BC14D11632B5F30FCFBFC73F4DF8360" authority="Chaudhuri &amp; Som, 1998" authorityName="Chaudhuri &amp; Som" authorityYear="1998" box="[865,1432,1015,1041]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="species" species="missilus">
<emphasis id="EEB5EA80632B5F30FCFBFC73F5358360" box="[865,1138,1015,1041]" italics="true" pageId="16" pageNumber="154">Paracricotopus missilus</emphasis>
Chaudhuri &amp; Som, 1998
</taxonomicName>
, and
<taxonomicName id="1BC14D11632B5F30FF52FB9FF2218344" authority="Sasa, Kwai &amp; Ueno, 1988" authorityName="Sasa, Kwai &amp; Ueno" authorityYear="1988" box="[200,870,1051,1077]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="species" species="oyabeangulatus">
<emphasis id="EEB5EA80632B5F30FF52FB9FF3778344" box="[200,560,1051,1077]" italics="true" pageId="16" pageNumber="154">Paracricotopus oyabeangulatus</emphasis>
Sasa, Kwai &amp; Ueno, 1988
</taxonomicName>
, our tergite setae key at best can be a preliminary one, until more
<typeStatus id="037A8830632B5F30FED6FBBBF03B8328" box="[332,380,1087,1113]" pageId="16" pageNumber="154">type</typeStatus>
and voucher adult specimens from various locations are examined. Combining the
<specimenCount id="CAC7FD1B632B5F30FAA8FBBBF4DB8328" box="[1330,1436,1087,1113]" count="2" pageId="16" pageNumber="154" type="adult">two adult</specimenCount>
male keys to produce a more comprehensive key would have made sense. For now, two reasons prevented us from doing so. First, the alternative key signifies the importance of tergite setation as a distinguishing characteristic. Further, the absence of antenna and, therefore, its characteristics in
<taxonomicName id="1BC14D11632B5F30FC1AFB2FF2A683B5" authorityName="Namayandeh &amp; Ghaderi" authorityYear="2025" box="[896,993,1195,1220]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="species" species="sturae" status="sp. nov.">
<emphasis id="EEB5EA80632B5F30FC1AFB2FF2A683B5" box="[896,993,1195,1220]" italics="true" pageId="16" pageNumber="154">P. sturae</emphasis>
</taxonomicName>
<emphasis id="EEB5EA80632B5F30FC7DFB2FF57883B4" bold="true" box="[999,1087,1195,1221]" pageId="16" pageNumber="154">
<taxonomicNameLabel id="F58657FB632B5F30FC7DFB2FF57883B4" box="[999,1087,1195,1221]" pageId="16" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="1BC14D11632B5F30FBE9FB2FF5B483B4" authorityName="Namayandeh &amp; Ghaderi" authorityYear="2025" box="[1139,1267,1195,1221]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="species" species="torbjorni" status="sp. nov.">
<emphasis id="EEB5EA80632B5F30FBE9FB2FF5B483B4" box="[1139,1267,1195,1221]" italics="true" pageId="16" pageNumber="154">P. torbjorni</emphasis>
</taxonomicName>
<emphasis id="EEB5EA80632B5F30FB63FB2FF41683B4" bold="true" box="[1273,1361,1195,1221]" pageId="16" pageNumber="154">
<taxonomicNameLabel id="F58657FB632B5F30FB63FB2FF41683B4" box="[1273,1361,1195,1221]" pageId="16" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, along with the absence of the species above, could not provide us with proper options of tergite setae character to separate the
<taxonomicName id="1BC14D11632B5F30FF5AFB77F089827C" box="[192,462,1267,1293]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="154" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="EEB5EA80632B5F30FF5AFB77F029827D" box="[192,366,1267,1292]" italics="true" pageId="16" pageNumber="154">Paracricotopus</emphasis>
species.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>
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