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<document id="EB820294FD7D0A089729B7C2B4CA3BD9" ID-DOI="10.11646/zootaxa.5569.1.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="14703998" ID-ZooBank="EA597E31-AF95-4D31-A14E-9455D6B335B1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737371943224" checkinUser="plazi" docAuthor="Namayandeh, Armin &amp; Ghaderi, Edris" docDate="2025" docId="5468878463305F2CFF0DFB6CF57481E5" docLanguage="en" docName="zootaxa.5569.1.5.pdf" docOrigin="Zootaxa 5569 (1)" docSource="https://doi.org/10.11646/zootaxa.5569.1.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paracricotopus glaber Saether 1980" docType="treatment" docVersion="4" lastPageNumber="150" masterDocId="A851FFFC633B5F20FF9AFF84F1478771" masterDocTitle="A review of Paracricotopus (Brundin, 1947), with the description of two new species and redescription of the adult males of Paracricotopus niger (Kieffer, 1913) and Paracricotopus uliginosus (Brundin, 1947) sensu Saether (1980) (Chironomidae: Diptera)" masterLastPageNumber="158" masterPageNumber="138" pageNumber="149" updateTime="1737392568075" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CCB08BB2C3BFCC5006723F3C1CA7E544">A review of Paracricotopus (Brundin, 1947), with the description of two new species and redescription of the adult males of Paracricotopus niger (Kieffer, 1913) and Paracricotopus uliginosus (Brundin, 1947) sensu Saether (1980) (Chironomidae: Diptera)</mods:title>
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<mods:namePart id="2FA927FD28CC596118FD5BB2D182E158">Namayandeh, Armin</mods:namePart>
<mods:affiliation id="0C1A15E9B1A736F76250AB133500C4CA">Department of Environmental and Life Sciences, Trent University, 1600 West Bank Drive, Peterborough, Ontario, Canada.</mods:affiliation>
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<mods:namePart id="1C33FF333235C84FF4309304D3AC026F">Ghaderi, Edris</mods:namePart>
<mods:affiliation id="39699C649E0ECFD48B04751F90CF6FF6">Department of Fisheries Sciences, Faculty of Natural Resources, University of Kurdistan, Sanandaj, Iran. &amp; Department of Fisheries and Aquatic Ecology, Faculty of Fisheries and Environmental Sciences, Gorgan University of Agricultural Sciences and Natural Resources, Gorgan, Iran.</mods:affiliation>
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<mods:date id="266D2939AB127D14CD88F806D2BDD1FD">2025</mods:date>
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<treatment id="5468878463305F2CFF0DFB6CF57481E5" ID-DOI="http://doi.org/10.5281/zenodo.14705208" ID-Zenodo-Dep="14705208" LSID="urn:lsid:plazi:treatment:5468878463305F2CFF0DFB6CF57481E5" httpUri="http://treatment.plazi.org/id/5468878463305F2CFF0DFB6CF57481E5" lastPageId="12" lastPageNumber="150" pageId="11" pageNumber="149">
<subSubSection id="94DB651963305F2BFF0DFB6CF3708273" box="[151,567,1255,1282]" pageId="11" pageNumber="149" type="nomenclature">
<paragraph id="DC7E369263305F2BFF0DFB6CF3708273" blockId="11.[151,567,1255,1319]" box="[151,567,1255,1282]" pageId="11" pageNumber="149">
<heading id="873681FE63305F2BFF0DFB6CF3708273" bold="true" box="[151,567,1255,1282]" fontSize="11" level="1" pageId="11" pageNumber="149" reason="1">
<taxonomicName id="1BC14D1163305F2BFF0DFB6CF3708273" ID-CoL="4CRJQ" authority="Saether, 1980" authorityName="Saether" authorityYear="1980" box="[151,567,1255,1282]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="149" phylum="Arthropoda" rank="species" species="glaber">
<emphasis id="EEB5EA8063305F2BFF0DFB6CF3708273" bold="true" box="[151,567,1255,1282]" pageId="11" pageNumber="149">
<emphasis id="EEB5EA8063305F2BFF0DFB6CF0D08273" bold="true" box="[151,407,1255,1282]" italics="true" pageId="11" pageNumber="149">Paracricotopus glaber</emphasis>
Saether, 1980
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="94DB651963305F2BFF0DFA88F06F8256" box="[151,296,1292,1319]" pageId="11" pageNumber="149" type="description">
<paragraph id="DC7E369263305F2BFF0DFA88F06F8256" blockId="11.[151,567,1255,1319]" box="[151,296,1292,1319]" pageId="11" pageNumber="149">
<figureCitation id="44FA2A1763305F2BFF0DFA88F1A48257" box="[151,227,1292,1318]" captionStart="FIGURE 6" captionStartId="12.[151,250,1124,1148]" captionTargetBox="[161,1426,207,1087]" captionTargetId="figure-247@12.[151,1436,197,1096]" captionTargetPageId="12" captionText="FIGURE 6. Paracricotopus glaber Saether, 1980, male (A–E), pupa (F).A. antenna.B. head. C. thorax.D. wing.E. hypopygium. F. thoracic horn. Scale bars are 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14704012" httpUri="https://zenodo.org/record/14704012/files/figure.png" pageId="11" pageNumber="149">Figs. 6</figureCitation>
&amp;
<figureCitation id="44FA2A1763305F2BFE9DFA89F06F8256" box="[263,296,1293,1319]" captionStart="FIGURE 7" captionStartId="13.[151,250,1841,1865]" captionTargetBox="[318,1314,159,1806]" captionTargetId="figure-113@13.[310,1322,151,1814]" captionTargetPageId="13" captionText="FIGURE 7. Abdominal tergites I–V of Paracricotopus Brundin, spp. adult males. A. Paracricotopus davoodi Ghaderi et al., 2023. B. Paracricotopus glaber Saether, 1980. C.Paracricotopus insulatus (Bhattacharyay &amp; Chaudhuri, 1987), tergite I missing. D. Paracricotopus mozleyi Steiner, 1983. E. Paracricotopus niger (Kieffer, 1913) sensu Saether (1980). F. Paracricotopus spinicornis Hazra &amp; Chaudhuri, 2002. G. Paracricotopus sturae sp. nov. H. Paracricotopus tamabrevis (Sasa, 1983). I. Paracricotopus togakuroasi (Sasa &amp; Okazawa, 1992). J. Paracricotopus torbjorni sp. nov. K. Paracricotopus uliginosus (Brundin, 1947) sensu Saether (1980)." figureDoi="http://doi.org/10.5281/zenodo.14704014" httpUri="https://zenodo.org/record/14704014/files/figure.png" pageId="11" pageNumber="149">7B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="94DB651963305F2BFF0DFAD0F55082E2" pageId="11" pageNumber="149" type="materials_examined">
<paragraph id="DC7E369263305F2BFF0DFAD0F55082E2" blockId="11.[151,1436,1364,2038]" pageId="11" pageNumber="149">
<emphasis id="EEB5EA8063305F2BFF0DFAD0F0C4821F" bold="true" box="[151,387,1364,1390]" pageId="11" pageNumber="149">Material examined.</emphasis>
<specimenCount id="CAC7FD1B63305F2BFE16FAD0F09B821F" box="[396,476,1364,1390]" count="1" pageId="11" pageNumber="149" type="male">1 male</specimenCount>
, 3 pharate males,
<materialsCitation id="6CA93CCF63305F2BFD34FAD0F55482E2" collectingDate="1977-04-17" collectorName="P. L. Hudson" country="United States of America" county="Oconee Co" location="Hwy" municipality="Jocassee Overlook" pageId="11" pageNumber="149" specimenCount="2" specimenCount-pupa="2" stateProvince="South Carolina">
<specimenCount id="CAC7FD1B63305F2BFD34FAD0F24B821E" box="[686,780,1364,1391]" count="2" pageId="11" pageNumber="149" type="pupa">2 pupae</specimenCount>
;
<collectingCountry id="A4D6760263305F2BFC80FAD0F212821F" box="[794,853,1364,1390]" name="United States of America" pageId="11" pageNumber="149">USA</collectingCountry>
,
<collectingRegion id="1E05F87063305F2BFCF8FAD0F557821F" box="[866,1040,1364,1390]" country="United States of America" name="South Carolina" pageId="11" pageNumber="149">South Carolina</collectingRegion>
,
<collectingCounty id="351F4E1E63305F2BFB87FAD0F5E7821F" box="[1053,1184,1364,1390]" pageId="11" pageNumber="149">Oconee Co</collectingCounty>
; seepage area, granite outcrop,
<collectingMunicipality id="3C1AACE863305F2BFF61FAFDF08882E3" box="[251,463,1400,1427]" pageId="11" pageNumber="149">Jocassee Overlook</collectingMunicipality>
,
<location id="D91E604963305F2BFE40FAFDF35782E2" LSID="urn:lsid:plazi:treatment:5468878463305F2CFF0DFB6CF57481E5:D91E604963305F2BFE40FAFDF35782E2" box="[474,528,1401,1427]" country="United States of America" county="Oconee Co" municipality="Jocassee Overlook" name="Hwy" pageId="11" pageNumber="149" stateProvince="South Carolina">Hwy</location>
413,
<date id="A87F105263305F2BFDD5FAFCF38F82E3" box="[591,712,1400,1426]" pageId="11" pageNumber="149" value="1977-04-17">
<collectingDate id="B83BE9BA63305F2BFDD5FAFCF38F82E3" box="[591,712,1400,1426]" pageId="11" pageNumber="149" value="1977-04-17">17.iv.1977</collectingDate>
</date>
; leg.
<collectorName id="7134534463305F2BFC9EFAFDF2DE82E3" box="[772,921,1400,1427]" pageId="11" pageNumber="149">P. L. Hudson</collectorName>
; dep. PLH
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="94DB651963305F2CFF5DFA18F3C28101" lastPageId="12" lastPageNumber="150" pageId="11" pageNumber="149" type="description">
<paragraph id="DC7E369263305F2BFF5DFA18F42982C7" blockId="11.[151,1436,1364,2038]" box="[199,1390,1436,1463]" pageId="11" pageNumber="149">
<emphasis id="EEB5EA8063305F2BFF5DFA18F01E82C7" bold="true" box="[199,345,1436,1462]" pageId="11" pageNumber="149">Description.</emphasis>
The adult male as described by
<bibRefCitation id="B8504B6363305F2BFD58FA18F22882C7" author="Saether, O. A." box="[706,879,1436,1462]" pageId="11" pageNumber="149" pagination="129 - 145" refId="ref15286" refString="Saether, O. A. (1980 b) The Females and immatures of Paracricotopus Thienemann and Harnisch, 1932, with the description of a new species (Diptera: Chironomidae). Aquatic Insects, 2, 129 - 145. https: // doi. org / 10.1080 / 01650428009361018" type="journal article" year="1980">Saether (1980b)</bibRefCitation>
with the following additions and corrections:
</paragraph>
<paragraph id="DC7E369263305F2BFF5DFA44F32B82AB" blockId="11.[151,1436,1364,2038]" box="[199,620,1472,1499]" pageId="11" pageNumber="149">Male (n = 4, unless otherwise stated).</paragraph>
<paragraph id="DC7E369263305F2BFF5DFA60F41B828F" blockId="11.[151,1436,1364,2038]" box="[199,1372,1508,1535]" pageId="11" pageNumber="149">
Total length
<quantity id="1B399B7763305F2BFECEFA60F099828E" box="[340,478,1508,1535]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.8" metricValueMin="2.7" pageId="11" pageNumber="149" unit="mm" value="2.75" valueMax="2.8" valueMin="2.7">2.7–2.8 mm</quantity>
. Wing length
<quantity id="1B399B7763305F2BFDE3FA60F396828E" box="[633,721,1508,1535]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="149" unit="mm" value="1.6">1.6 mm</quantity>
, wing width
<quantity id="1B399B7763305F2BFCF9FA60F2FE828E" box="[867,953,1508,1535]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="149" unit="mm" value="0.5">0.5 mm</quantity>
(n = 1). Haltere 281 μm long (n = 1).
</paragraph>
<paragraph id="DC7E369263305F2BFF5DF98CF0BE8153" blockId="11.[151,1436,1364,2038]" box="[199,505,1544,1571]" pageId="11" pageNumber="149">
<emphasis id="EEB5EA8063305F2BFF5DF98CF0058153" box="[199,322,1544,1570]" italics="true" pageId="11" pageNumber="149">Coloration</emphasis>
. Haltere brown.
</paragraph>
<paragraph id="DC7E369263305F2BFF5DF9A8F44981FF" blockId="11.[151,1436,1364,2038]" pageId="11" pageNumber="149">
<emphasis id="EEB5EA8063305F2BFF5DF9A8F0438137" box="[199,260,1580,1606]" italics="true" pageId="11" pageNumber="149">Head</emphasis>
(
<figureCitation id="44FA2A1763305F2BFE89F9A8F0138137" box="[275,340,1580,1606]" captionStart="FIGURE 6" captionStartId="12.[151,250,1124,1148]" captionTargetBox="[161,1426,207,1087]" captionTargetId="figure-247@12.[151,1436,197,1096]" captionTargetPageId="12" captionText="FIGURE 6. Paracricotopus glaber Saether, 1980, male (A–E), pupa (F).A. antenna.B. head. C. thorax.D. wing.E. hypopygium. F. thoracic horn. Scale bars are 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14704012" httpUri="https://zenodo.org/record/14704012/files/figure.png" pageId="11" pageNumber="149">Fig. 6</figureCitation>
A-B). AR 0.70–0.95; last flagellomere with 8 sensilla chaetica (n = 1). Temporal setae 3, including 2 inner verticals and 1 frontal (n = 1). Tentorium 207 μm long (n = 1). Clypeus 72 μm long and 135 μm wide bearing 12 setae, setae 78–86, 82 μm long (n = 1). Palpal segments length (in μm): 69, 44, 74, 97, 147 (n = 1).
</paragraph>
<paragraph id="DC7E369263305F2BFF5DF91CF3AD81C3" blockId="11.[151,1436,1364,2038]" box="[199,746,1688,1715]" pageId="11" pageNumber="149">
<emphasis id="EEB5EA8063305F2BFF5DF91CF05F81C3" box="[199,280,1688,1714]" italics="true" pageId="11" pageNumber="149">Thorax</emphasis>
(n = 1;
<figureCitation id="44FA2A1763305F2BFEEBF91CF09481C3" box="[369,467,1688,1714]" captionStart="FIGURE 6" captionStartId="12.[151,250,1124,1148]" captionTargetBox="[161,1426,207,1087]" captionTargetId="figure-247@12.[151,1436,197,1096]" captionTargetPageId="12" captionText="FIGURE 6. Paracricotopus glaber Saether, 1980, male (A–E), pupa (F).A. antenna.B. head. C. thorax.D. wing.E. hypopygium. F. thoracic horn. Scale bars are 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14704012" httpUri="https://zenodo.org/record/14704012/files/figure.png" pageId="11" pageNumber="149">Figs. 6C</figureCitation>
). Prealars 3, scutellars 6.
</paragraph>
<paragraph id="DC7E369263305F2BFF5DF939F28781A7" blockId="11.[151,1436,1364,2038]" box="[199,960,1724,1751]" pageId="11" pageNumber="149">
<emphasis id="EEB5EA8063305F2BFF5DF939F04681A7" box="[199,257,1725,1750]" italics="true" pageId="11" pageNumber="149">Wing</emphasis>
(n = 1;
<figureCitation id="44FA2A1763305F2BFEC0F938F0F981A7" box="[346,446,1724,1750]" captionStart="FIGURE 6" captionStartId="12.[151,250,1124,1148]" captionTargetBox="[161,1426,207,1087]" captionTargetId="figure-247@12.[151,1436,197,1096]" captionTargetPageId="12" captionText="FIGURE 6. Paracricotopus glaber Saether, 1980, male (A–E), pupa (F).A. antenna.B. head. C. thorax.D. wing.E. hypopygium. F. thoracic horn. Scale bars are 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14704012" httpUri="https://zenodo.org/record/14704012/files/figure.png" pageId="11" pageNumber="149">Figs. 6D</figureCitation>
). R with 6 setae. Costa extension 54 μm long.
</paragraph>
<paragraph id="DC7E369263305F2BFF5DF965F4C9806F" blockId="11.[151,1436,1364,2038]" pageId="11" pageNumber="149">
<emphasis id="EEB5EA8063305F2BFF5DF965F1BC818B" box="[199,251,1761,1786]" italics="true" pageId="11" pageNumber="149">Legs</emphasis>
(n = 1). Hind and mid femurs with keel. Fore tibia spur 43 μm long, mid tibia spurs 20 and 16 μm long, hind tibia spurs 50 and 17 μm long, hind tibia comb with around 14 spines. Lengths and proportions of legs as in
<tableCitation id="9143032963305F2BFAACF880F4CD806F" box="[1334,1418,1796,1822]" captionStart="TABLE 6" captionStartId="12.[152,244,1268,1292]" captionText="TABLE 6. Lengths (in μm) and proportions of leg segments of Paracricotopus glaber Saether, 1980 male." pageId="11" pageNumber="149">Table 6</tableCitation>
.
</paragraph>
<paragraph id="DC7E369263305F2BFF5DF8ACF21C80FA" blockId="11.[151,1436,1364,2038]" pageId="11" pageNumber="149">
<emphasis id="EEB5EA8063305F2BFF5DF8ACF0B68033" box="[199,497,1832,1858]" italics="true" pageId="11" pageNumber="149">Abdominal tergite setation</emphasis>
(
<figureCitation id="44FA2A1763305F2BFD9AF8ACF31F8033" box="[512,600,1832,1858]" captionStart="FIGURE 7" captionStartId="13.[151,250,1841,1865]" captionTargetBox="[318,1314,159,1806]" captionTargetId="figure-113@13.[310,1322,151,1814]" captionTargetPageId="13" captionText="FIGURE 7. Abdominal tergites I–V of Paracricotopus Brundin, spp. adult males. A. Paracricotopus davoodi Ghaderi et al., 2023. B. Paracricotopus glaber Saether, 1980. C.Paracricotopus insulatus (Bhattacharyay &amp; Chaudhuri, 1987), tergite I missing. D. Paracricotopus mozleyi Steiner, 1983. E. Paracricotopus niger (Kieffer, 1913) sensu Saether (1980). F. Paracricotopus spinicornis Hazra &amp; Chaudhuri, 2002. G. Paracricotopus sturae sp. nov. H. Paracricotopus tamabrevis (Sasa, 1983). I. Paracricotopus togakuroasi (Sasa &amp; Okazawa, 1992). J. Paracricotopus torbjorni sp. nov. K. Paracricotopus uliginosus (Brundin, 1947) sensu Saether (1980)." figureDoi="http://doi.org/10.5281/zenodo.14704014" httpUri="https://zenodo.org/record/14704014/files/figure.png" pageId="11" pageNumber="149">Fig. 7B</figureCitation>
). Tergites I with 2 lateral setae, anterior basal setae, and posterior marginal setae arranged in 4 diagonal rows in 1/1/3/1 format. Tergites II–VIII with 4 lateral setae, anterior basal setae, and posterior marginal setae in two parallel, slightly diagonal rows.
</paragraph>
<paragraph id="DC7E369263305F2BFF5DF811F3BF80A3" blockId="11.[151,1436,1364,2038]" pageId="11" pageNumber="149">
<emphasis id="EEB5EA8063305F2BFF5DF811F01180DF" box="[199,342,1941,1966]" italics="true" pageId="11" pageNumber="149">Hypopygium</emphasis>
(
<figureCitation id="44FA2A1763305F2BFEF2F810F08580DF" box="[360,450,1940,1966]" captionStart="FIGURE 6" captionStartId="12.[151,250,1124,1148]" captionTargetBox="[161,1426,207,1087]" captionTargetId="figure-247@12.[151,1436,197,1096]" captionTargetPageId="12" captionText="FIGURE 6. Paracricotopus glaber Saether, 1980, male (A–E), pupa (F).A. antenna.B. head. C. thorax.D. wing.E. hypopygium. F. thoracic horn. Scale bars are 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14704012" httpUri="https://zenodo.org/record/14704012/files/figure.png" pageId="11" pageNumber="149">Fig. 6E</figureCitation>
). Anal point 23–31, 27 μm long and 15–24, 19 μm wide at the base. HV 3.6 (n = 1). Gonostylus with conspicuous preapical crista dorsalis.
</paragraph>
<paragraph id="DC7E369263305F2BFF5DF858F33B8087" blockId="11.[151,1436,1364,2038]" box="[199,636,2012,2038]" pageId="11" pageNumber="149">Female. As described by Saether, 1980.</paragraph>
<caption id="88BE661A63375F2CFF0DFBE0F36B83D0" ID-DOI="http://doi.org/10.5281/zenodo.14704012" ID-Zenodo-Dep="14704012" httpUri="https://zenodo.org/record/14704012/files/figure.png" pageId="12" pageNumber="150" startId="12.[151,250,1124,1148]" targetBox="[161,1426,207,1087]" targetPageId="12" targetType="figure">
<paragraph id="DC7E369263375F2CFF0DFBE0F36B83D0" blockId="12.[151,1437,1124,1185]" pageId="12" pageNumber="150">
<emphasis id="EEB5EA8063375F2CFF0DFBE0F056830D" bold="true" box="[151,273,1124,1148]" pageId="12" pageNumber="150">FIGURE 6.</emphasis>
<taxonomicName id="1BC14D1163375F2CFE8CFBE1F3C3830C" authority="Saether, 1980" authorityName="Saether" authorityYear="1980" box="[278,644,1124,1149]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="150" phylum="Arthropoda" rank="species" species="glaber">
<emphasis id="EEB5EA8063375F2CFE8CFBE1F0BD830D" box="[278,506,1124,1148]" italics="true" pageId="12" pageNumber="150">Paracricotopus glaber</emphasis>
Saether, 1980
</taxonomicName>
, male (A–E), pupa (F).A. antenna. B. head. C. thorax. D. wing. E. hypopygium. F. thoracic horn. Scale bars are 100 µm.
</paragraph>
</caption>
<caption id="88BE661A63375F2CFF02FB70F44E827D" box="[152,1289,1268,1293]" pageId="12" pageNumber="150" startId="12.[152,244,1268,1292]" targetType="table">
<paragraph id="DC7E369263375F2CFF02FB70F44E827D" blockId="12.[152,1289,1268,1293]" box="[152,1289,1268,1293]" pageId="12" pageNumber="150">
<emphasis id="EEB5EA8063375F2CFF02FB70F049827D" bold="true" box="[152,270,1268,1292]" pageId="12" pageNumber="150">TABLE 6.</emphasis>
Lengths (in μm) and proportions of leg segments of
<taxonomicName id="1BC14D1163375F2CFCD9FB71F580827C" authority="Saether, 1980" authorityName="Saether" authorityYear="1980" box="[835,1223,1268,1293]" class="Insecta" family="Chironomidae" genus="Paracricotopus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="150" phylum="Arthropoda" rank="species" species="glaber">
<emphasis id="EEB5EA8063375F2CFCD9FB71F572827D" box="[835,1077,1268,1292]" italics="true" pageId="12" pageNumber="150">Paracricotopus glaber</emphasis>
Saether, 1980
</taxonomicName>
male.
</paragraph>
</caption>
<paragraph id="DC7E369263375F2CFE8DFAA4F40882CB" pageId="12" pageNumber="150">
<table id="AEC1C4326337A0DFFF39FAA4F40882CB" box="[163,1359,1312,1466]" gridcols="11" gridrows="4" pageId="12" pageNumber="150">
<tr id="62F134D06337A0DFFF39FAA4F408824F" box="[163,1359,1312,1342]" gridrow="0" pageId="12" pageNumber="150" rowspan-0="1">
<th id="21205DAC6337A0DFFE8DFAA4F07A824F" box="[279,317,1312,1342]" gridcol="1" gridrow="0" pageId="12" pageNumber="150">
<emphasis id="EEB5EA8063375F2CFE8DFAA4F06C8249" bold="true" box="[279,299,1312,1336]" pageId="12" pageNumber="150">fe</emphasis>
</th>
<th id="21205DAC6337A0DFFE11FAA4F0F6824F" box="[395,433,1312,1342]" gridcol="2" gridrow="0" pageId="12" pageNumber="150">
<emphasis id="EEB5EA8063375F2CFE11FAA4F0DD8249" bold="true" box="[395,410,1312,1336]" pageId="12" pageNumber="150">ti</emphasis>
</th>
<th id="21205DAC6337A0DFFD9AFAA4F361824F" box="[512,550,1312,1342]" gridcol="3" gridrow="0" pageId="12" pageNumber="150">
<emphasis id="EEB5EA8063375F2CFD9AFAA4F3528249" bold="true" box="[512,533,1312,1336]" pageId="12" pageNumber="150">ta</emphasis>
<emphasis id="EEB5EA8063375F2CFD8FFAB4F35B824F" bold="true" box="[533,540,1328,1342]" pageId="12" pageNumber="150">1</emphasis>
</th>
<th id="21205DAC6337A0DFFDEEFAA4F3DD824F" box="[628,666,1312,1342]" gridcol="4" gridrow="0" pageId="12" pageNumber="150">
<emphasis id="EEB5EA8063375F2CFDEEFAA4F3CE8249" bold="true" box="[628,649,1312,1336]" pageId="12" pageNumber="150">ta</emphasis>
<emphasis id="EEB5EA8063375F2CFD13FAB4F3D7824F" bold="true" box="[649,656,1328,1342]" pageId="12" pageNumber="150">2</emphasis>
</th>
<th id="21205DAC6337A0DFFD73FAA4F248824F" box="[745,783,1312,1342]" gridcol="5" gridrow="0" pageId="12" pageNumber="150">
<emphasis id="EEB5EA8063375F2CFD73FAA4F3B98249" bold="true" box="[745,766,1312,1336]" pageId="12" pageNumber="150">ta</emphasis>
<emphasis id="EEB5EA8063375F2CFD64FAB4F242824F" bold="true" box="[766,773,1328,1342]" pageId="12" pageNumber="150">3</emphasis>
</th>
<th id="21205DAC6337A0DFFCC7FAA4F2C4824F" box="[861,899,1312,1342]" gridcol="6" gridrow="0" pageId="12" pageNumber="150">
<emphasis id="EEB5EA8063375F2CFCC7FAA4F2358249" bold="true" box="[861,882,1312,1336]" pageId="12" pageNumber="150">ta</emphasis>
<emphasis id="EEB5EA8063375F2CFCE8FAB4F23E824F" bold="true" box="[882,889,1328,1342]" pageId="12" pageNumber="150">4</emphasis>
</th>
<th id="21205DAC6337A0DFFC4BFAA4F2B0824F" box="[977,1015,1312,1342]" gridcol="7" gridrow="0" pageId="12" pageNumber="150">
<emphasis id="EEB5EA8063375F2CFC4BFAA4F2A18249" bold="true" box="[977,998,1312,1336]" pageId="12" pageNumber="150">ta</emphasis>
<emphasis id="EEB5EA8063375F2CFC7DFAB4F2A9824F" bold="true" box="[999,1006,1328,1342]" pageId="12" pageNumber="150">5</emphasis>
</th>
<th id="21205DAC6337A0DFFBDCFAA4F52E824F" box="[1094,1129,1312,1342]" gridcol="8" gridrow="0" pageId="12" pageNumber="150">
<emphasis id="EEB5EA8063375F2CFBDCFAA4F52E8249" bold="true" box="[1094,1129,1312,1336]" pageId="12" pageNumber="150">LR</emphasis>
</th>
<th id="21205DAC6337A0DFFB20FAA4F59A824F" box="[1210,1245,1312,1342]" gridcol="9" gridrow="0" pageId="12" pageNumber="150">
<emphasis id="EEB5EA8063375F2CFB20FAA4F59A8249" bold="true" box="[1210,1245,1312,1336]" pageId="12" pageNumber="150">BV</emphasis>
</th>
<th id="21205DAC6337A0DFFAB5FAA4F408824F" box="[1327,1359,1312,1342]" gridcol="10" gridrow="0" pageId="12" pageNumber="150">
<emphasis id="EEB5EA8063375F2CFAB5FAA4F4088249" bold="true" box="[1327,1359,1312,1336]" pageId="12" pageNumber="150">SV</emphasis>
</th>
</tr>
<tr id="62F134D06337A0DFFF39FAC9F408821A" box="[163,1359,1357,1387]" gridrow="1" pageId="12" pageNumber="150">
<th id="21205DAC6337A0DFFF39FAC9F1FA821A" box="[163,189,1357,1387]" gridcol="0" gridrow="1" pageId="12" pageNumber="150">
<emphasis id="EEB5EA8063375F2CFF39FAC9F1F58214" bold="true" box="[163,178,1357,1381]" pageId="12" pageNumber="150">P</emphasis>
<emphasis id="EEB5EA8063375F2CFF28FAD9F1FE821A" bold="true" box="[178,185,1373,1387]" pageId="12" pageNumber="150">1</emphasis>
</th>
<td id="21205DAC6337A0DFFE8DFAC9F07A821A" box="[279,317,1357,1387]" gridcol="1" gridrow="1" pageId="12" pageNumber="150">536</td>
<td id="21205DAC6337A0DFFE11FAC9F0F6821A" box="[395,433,1357,1387]" gridcol="2" gridrow="1" pageId="12" pageNumber="150">652</td>
<td id="21205DAC6337A0DFFD9AFAC9F361821A" box="[512,550,1357,1387]" gridcol="3" gridrow="1" pageId="12" pageNumber="150">400</td>
<td id="21205DAC6337A0DFFDEEFAC9F3DD821A" box="[628,666,1357,1387]" gridcol="4" gridrow="1" pageId="12" pageNumber="150">293</td>
<td id="21205DAC6337A0DFFD73FAC9F248821A" box="[745,783,1357,1387]" gridcol="5" gridrow="1" pageId="12" pageNumber="150">213</td>
<td id="21205DAC6337A0DFFCC7FAC9F2C4821A" box="[861,899,1357,1387]" gridcol="6" gridrow="1" pageId="12" pageNumber="150">148</td>
<td id="21205DAC6337A0DFFC4BFAC9F2B0821A" box="[977,1015,1357,1387]" gridcol="7" gridrow="1" pageId="12" pageNumber="150">104</td>
<td id="21205DAC6337A0DFFBDCFAC9F52E821A" box="[1094,1129,1357,1387]" gridcol="8" gridrow="1" pageId="12" pageNumber="150">0.6</td>
<td id="21205DAC6337A0DFFB20FAC9F59A821A" box="[1210,1245,1357,1387]" gridcol="9" gridrow="1" pageId="12" pageNumber="150">2.1</td>
<td id="21205DAC6337A0DFFAB5FAC9F408821A" box="[1327,1359,1357,1387]" gridcol="10" gridrow="1" pageId="12" pageNumber="150">3.0</td>
</tr>
<tr id="62F134D06337A0DFFF39FAF3F40882E4" box="[163,1359,1399,1429]" gridrow="2" pageId="12" pageNumber="150">
<th id="21205DAC6337A0DFFF39FAF3F1FA82E4" box="[163,189,1399,1429]" gridcol="0" gridrow="2" pageId="12" pageNumber="150">
<emphasis id="EEB5EA8063375F2CFF39FAF3F1F582FE" bold="true" box="[163,178,1399,1423]" pageId="12" pageNumber="150">P</emphasis>
<emphasis id="EEB5EA8063375F2CFF28FA03F1FE82E4" bold="true" box="[178,185,1415,1429]" pageId="12" pageNumber="150">2</emphasis>
</th>
<td id="21205DAC6337A0DFFE8DFAF3F07A82E4" box="[279,317,1399,1429]" gridcol="1" gridrow="2" pageId="12" pageNumber="150">558</td>
<td id="21205DAC6337A0DFFE11FAF3F0F682E4" box="[395,433,1399,1429]" gridcol="2" gridrow="2" pageId="12" pageNumber="150">592</td>
<td id="21205DAC6337A0DFFD9AFAF3F36182E4" box="[512,550,1399,1429]" gridcol="3" gridrow="2" pageId="12" pageNumber="150">290</td>
<td id="21205DAC6337A0DFFDEEFAF3F3DD82E4" box="[628,666,1399,1429]" gridcol="4" gridrow="2" pageId="12" pageNumber="150">159</td>
<td id="21205DAC6337A0DFFD73FAF3F24882E4" box="[745,783,1399,1429]" gridcol="5" gridrow="2" pageId="12" pageNumber="150">116</td>
<td id="21205DAC6337A0DFFCC7FAF3F2C482E4" box="[861,899,1399,1429]" gridcol="6" gridrow="2" pageId="12" pageNumber="150">69</td>
<td id="21205DAC6337A0DFFC4BFAF3F2B082E4" box="[977,1015,1399,1429]" gridcol="7" gridrow="2" pageId="12" pageNumber="150">82</td>
<td id="21205DAC6337A0DFFBDCFAF3F52E82E4" box="[1094,1129,1399,1429]" gridcol="8" gridrow="2" pageId="12" pageNumber="150">0.5</td>
<td id="21205DAC6337A0DFFB20FAF3F59A82E4" box="[1210,1245,1399,1429]" gridcol="9" gridrow="2" pageId="12" pageNumber="150">3.4</td>
<td id="21205DAC6337A0DFFAB5FAF3F40882E4" box="[1327,1359,1399,1429]" gridcol="10" gridrow="2" pageId="12" pageNumber="150">4.0</td>
</tr>
<tr id="62F134D06337A0DFFF39FA25F40882CB" box="[163,1359,1441,1466]" gridrow="3" pageId="12" pageNumber="150">
<th id="21205DAC6337A0DFFF39FA25F1FA82CB" box="[163,189,1441,1466]" gridcol="0" gridrow="3" pageId="12" pageNumber="150">P3</th>
<td id="21205DAC6337A0DFFE8DFA25F07A82CB" box="[279,317,1441,1466]" gridcol="1" gridrow="3" pageId="12" pageNumber="150">566</td>
<td id="21205DAC6337A0DFFE11FA25F0F682CB" box="[395,433,1441,1466]" gridcol="2" gridrow="3" pageId="12" pageNumber="150">679</td>
<td id="21205DAC6337A0DFFD9AFA25F36182CB" box="[512,550,1441,1466]" gridcol="3" gridrow="3" pageId="12" pageNumber="150">395</td>
<td id="21205DAC6337A0DFFDEEFA25F3DD82CB" box="[628,666,1441,1466]" gridcol="4" gridrow="3" pageId="12" pageNumber="150">226</td>
<td id="21205DAC6337A0DFFD73FA25F24882CB" box="[745,783,1441,1466]" gridcol="5" gridrow="3" pageId="12" pageNumber="150">183</td>
<td id="21205DAC6337A0DFFCC7FA25F2C482CB" box="[861,899,1441,1466]" gridcol="6" gridrow="3" pageId="12" pageNumber="150">88</td>
<td id="21205DAC6337A0DFFC4BFA25F2B082CB" box="[977,1015,1441,1466]" gridcol="7" gridrow="3" pageId="12" pageNumber="150">91</td>
<td id="21205DAC6337A0DFFBDCFA25F52E82CB" box="[1094,1129,1441,1466]" gridcol="8" gridrow="3" pageId="12" pageNumber="150">0.6</td>
<td id="21205DAC6337A0DFFB20FA25F59A82CB" box="[1210,1245,1441,1466]" gridcol="9" gridrow="3" pageId="12" pageNumber="150">2.8</td>
<td id="21205DAC6337A0DFFAB5FA25F40882CB" box="[1327,1359,1441,1466]" gridcol="10" gridrow="3" pageId="12" pageNumber="150">3.2</td>
</tr>
</table>
</paragraph>
<paragraph id="DC7E369263375F2CFF5DF98BF0FF813D" blockId="12.[151,1436,1550,1684]" pageId="12" pageNumber="150">
Pupa (n = 5). As described by
<bibRefCitation id="B8504B6363375F2CFD8FF98AF3858159" author="Saether, O. A." box="[533,706,1550,1576]" pageId="12" pageNumber="150" pagination="129 - 145" refId="ref15286" refString="Saether, O. A. (1980 b) The Females and immatures of Paracricotopus Thienemann and Harnisch, 1932, with the description of a new species (Diptera: Chironomidae). Aquatic Insects, 2, 129 - 145. https: // doi. org / 10.1080 / 01650428009361018" type="journal article" year="1980">Saether (1980b)</bibRefCitation>
with the following addition: Thoracic horn 100–135 μm long and 25–42 μm wide (
<figureCitation id="44FA2A1763375F2CFECCF9B6F0EC813D" box="[342,427,1586,1612]" captionStart="FIGURE 6" captionStartId="12.[151,250,1124,1148]" captionTargetBox="[161,1426,207,1087]" captionTargetId="figure-247@12.[151,1436,197,1096]" captionTargetPageId="12" captionText="FIGURE 6. Paracricotopus glaber Saether, 1980, male (A–E), pupa (F).A. antenna.B. head. C. thorax.D. wing.E. hypopygium. F. thoracic horn. Scale bars are 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14704012" httpUri="https://zenodo.org/record/14704012/files/figure.png" pageId="12" pageNumber="150">Fig. 6F</figureCitation>
).
</paragraph>
<paragraph id="DC7E369263375F2CFF5DF9D3F3C28101" blockId="12.[151,1436,1550,1684]" box="[199,645,1622,1649]" pageId="12" pageNumber="150">
Larva. As described by
<bibRefCitation id="B8504B6363375F2CFE4AF9D2F3C68101" author="Saether, O. A." box="[464,641,1622,1648]" pageId="12" pageNumber="150" pagination="129 - 145" refId="ref15286" refString="Saether, O. A. (1980 b) The Females and immatures of Paracricotopus Thienemann and Harnisch, 1932, with the description of a new species (Diptera: Chironomidae). Aquatic Insects, 2, 129 - 145. https: // doi. org / 10.1080 / 01650428009361018" type="journal article" year="1980">Saether (1980b)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="94DB651963375F2CFF5DF9FEF57481E5" box="[199,1075,1658,1684]" pageId="12" pageNumber="150" type="distribution">
<paragraph id="DC7E369263375F2CFF5DF9FEF57481E5" blockId="12.[151,1436,1550,1684]" box="[199,1075,1658,1684]" pageId="12" pageNumber="150">
<emphasis id="EEB5EA8063375F2CFF5DF9FEF02681E5" bold="true" box="[199,353,1658,1684]" pageId="12" pageNumber="150">Distribution.</emphasis>
Nearctic:
<collectingCountry id="A4D6760263375F2CFE4FF9FEF34C81E5" box="[469,523,1658,1684]" name="United States of America" pageId="12" pageNumber="150">USA</collectingCountry>
(
<collectingRegion id="1E05F87063375F2CFD80F9FEF3C381E5" box="[538,644,1658,1684]" country="United States of America" name="Alabama" pageId="12" pageNumber="150">Alabama</collectingRegion>
,
<collectingCountry id="A4D6760263375F2CFD15F9FEF3AB81E5" box="[655,748,1658,1684]" name="Georgia" pageId="12" pageNumber="150">Georgia</collectingCountry>
, North and South Carolinas).
</paragraph>
</subSubSection>
</treatment>
</document>
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