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<mods:title id="029D4D9B3B58FD532991F8420FFFAA70">Taxonomy of some Galeommatoidea (Mollusca, Bivalvia) associated with deep-sea echinoids: A reassessment of the bivalve genera Axinodon Verrill &amp; Bush, 1898 and Kelliola Dall, 1899 with descriptions of new genera Syssitomya gen. nov. and Ptilomyax gen. nov.</mods:title>
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<taxonomicName id="1BC94D617C5EFFE6877320C4FC5DFB56" ID-CoL="3R52T" authority="(Jeffreys, 1876)" baseAuthorityName="Jeffreys" baseAuthorityYear="1876" box="[582,1005,1224,1251]" class="Bivalvia" family="Lasaeidae" genus="Kelliola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="symmetros">
<emphasis id="EEBDEAF07C5EFFE6877320C4FC9DFB56" bold="true" box="[582,813,1224,1251]" italics="true" pageId="6" pageNumber="7">Kelliola symmetros</emphasis>
(
<bibRefCitation id="B8584B137C5EFFE6860920C4FC55FB56" author="Jeffreys" box="[828,997,1224,1251]" firstAuthor="Jeffreys" pageId="6" pageNumber="7" pagination="490 - 499" refId="ref9385" refString="Jeffreys J. G. 1876. New and peculiar Mollusca of the Kellia, Lucina, Cyprina and Corbula families procured in the ' Valorous' Expedition. Annals and Magazine of Natural History ser. 4, 18: 490 - 499." type="journal article" year="1876">Jeffreys, 1876</bibRefCitation>
)
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</paragraph>
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<subSubSection id="94D365697C5EFFE685882106FCFEF937" pageId="6" pageNumber="7" type="reference_group">
<paragraph id="DC7636E27C5EFFE685882106FDC8FA90" blockId="6.[189,632,1290,1317]" box="[189,632,1290,1317]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="FCD911CC7C5EFFE685882106FDC8FA90" box="[189,632,1290,1317]" pageId="6" pageNumber="7">
<treatmentCitation id="5D6810F37C5EFFE685882106FDC5FA90" author="Jeffreys J. G." box="[189,629,1290,1317]" page="491" pageId="6" pageNumber="7" year="1876">
<taxonomicName id="1BC94D617C5EFFE685882106FDC5FA90" ID-CoL="3R4ZK" authority="Jeffreys, 1876: 491" authorityName="Jeffreys" authorityPageNumber="491" authorityYear="1876" box="[189,629,1290,1317]" class="Bivalvia" family="Lasaeidae" genus="Kellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="symmetros">
<emphasis id="EEBDEAF07C5EFFE685882106FE38FA91" box="[189,392,1290,1316]" italics="true" pageId="6" pageNumber="7">Kellia symmetros</emphasis>
<bibRefCitation id="B8584B137C5EFFE684A52106FDC5FA90" author="Jeffreys" box="[400,629,1290,1317]" firstAuthor="Jeffreys" pageId="6" pageNumber="7" pagination="490 - 499" refId="ref9385" refString="Jeffreys J. G. 1876. New and peculiar Mollusca of the Kellia, Lucina, Cyprina and Corbula families procured in the ' Valorous' Expedition. Annals and Magazine of Natural History ser. 4, 18: 490 - 499." type="journal article" year="1876">Jeffreys, 1876: 491</bibRefCitation>
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.
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</paragraph>
<paragraph id="DC7636E27C5EFFE685882141FD67FAD2" blockId="6.[189,733,1357,1456]" box="[189,727,1357,1383]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="FCD911CC7C5EFFE685882141FD67FAD2" box="[189,727,1357,1383]" pageId="6" pageNumber="7">
<taxonomicName id="1BC94D617C5EFFE685882141FDB1FAD2" ID-CoL="3R4ZK" authorityName="Jeffreys" authorityYear="1876" box="[189,513,1357,1383]" class="Bivalvia" family="Lasaeidae" genus="Kellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="symmetros" subGenus="Kelliola">
<emphasis id="EEBDEAF07C5EFFE685882141FEB5FAD2" box="[189,261,1357,1383]" italics="true" pageId="6" pageNumber="7">Kellia</emphasis>
(
<emphasis id="EEBDEAF07C5EFFE684232141FEC5FAD2" box="[278,373,1357,1383]" italics="true" pageId="6" pageNumber="7">Kelliola</emphasis>
) 
<emphasis id="EEBDEAF07C5EFFE684B02142FDB1FAD2" box="[389,513,1358,1383]" italics="true" pageId="6" pageNumber="7">symmetros</emphasis>
</taxonomicName>
– 
<treatmentCitation id="5D6810F37C5EFFE6872A2141FD63FAD2" author="Dall W. H." box="[543,723,1357,1383]" page="890" pageId="6" pageNumber="7" year="1899">
<bibRefCitation id="B8584B137C5EFFE6872A2141FD63FAD2" author="Dall" box="[543,723,1357,1383]" firstAuthor="Dall" pageId="6" pageNumber="7" pagination="873 - 897" refId="ref8857" refString="Dall W. H. 1899. Synopsis of the Recent and Tertiary Leptonacea of North America and the West Indies. Proceedings of the United States National Museum 21 (1177): 873 - 897, 2 pls. http: // dx. doi. org / 10.5479 / si. 00963801.21 - 1177.873" type="book chapter" year="1899">Dall 1899: 890</bibRefCitation>
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.
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</paragraph>
<paragraph id="DC7636E27C5EFFE68588217DFD6DFA3E" blockId="6.[189,733,1357,1456]" box="[189,733,1393,1419]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="FCD911CC7C5EFFE68588217DFD6DFA3E" box="[189,733,1393,1419]" pageId="6" pageNumber="7">
? 
<taxonomicName id="1BC94D617C5EFFE685E4217DFE2CFA3E" ID-CoL="3R4ZK" authorityName="Jeffreys" authorityYear="1876" box="[209,412,1393,1419]" class="Bivalvia" family="Lasaeidae" genus="Kellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="symmetros">
<emphasis id="EEBDEAF07C5EFFE685E4217DFE2CFA3E" box="[209,412,1393,1419]" italics="true" pageId="6" pageNumber="7">Kellia symmetros</emphasis>
</taxonomicName>
– 
<treatmentCitation id="5D6810F37C5EFFE6848F217DFD69FA3E" author="Friele H. &amp; Grieg J. A." box="[442,729,1393,1419]" page="29" pageId="6" pageNumber="7" year="1901">
<bibRefCitation id="B8584B137C5EFFE6848F217DFD69FA3E" author="Friele &amp; Grieg" box="[442,729,1393,1419]" firstAuthor="Friele" pageId="6" pageNumber="7" refId="ref8997" refString="Friele H. &amp; Grieg J. A. 1901. The Norwegian North-Atlantic Expedition 1876 - 1878. Vol. VII: Zoology, Mollusca III. Grondhal &amp; sons, Christiana. http: // dx. doi. org / 10.5962 / bhl. title. 2168" type="book" year="1901">Friele &amp; Grieg 1901: 29</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="DC7636E27C5EFFE685882199FD20FA1A" blockId="6.[189,733,1357,1456]" box="[189,656,1429,1456]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="FCD911CC7C5EFFE685882199FD20FA1A" box="[189,656,1429,1456]" pageId="6" pageNumber="7">
<taxonomicName id="1BC94D617C5EFFE685882199FE01FA1A" authorityName="Verrill &amp; Bush" authorityYear="1898" box="[189,433,1429,1455]" class="Bivalvia" family="Thyasiridae" genus="Axinodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="symmetros">
<emphasis id="EEBDEAF07C5EFFE685882199FE01FA1A" box="[189,433,1429,1455]" italics="true" pageId="6" pageNumber="7">Axinodon symmetros</emphasis>
</taxonomicName>
– 
<treatmentCitation id="5D6810F37C5EFFE684FB2199FD3CFA1A" author="Waren A." box="[462,652,1429,1455]" page="47" pageId="6" pageNumber="7" year="1980">
<bibRefCitation id="B8584B137C5EFFE684FB2199FD3CFA1A" author="Waren" box="[462,652,1429,1455]" firstAuthor="Waren" pageId="6" pageNumber="7" refId="ref10281" refString="Waren A. 1980. Marine Mollusca described by John Gwyn Jeffreys, with the location of the type material. Special Publication 1, Conchological Society of Great Britain and Ireland." type="book" year="1980">Warén 1980: 47</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="DC7636E27C5EFFE6858821D4FC25FA47" blockId="6.[189,1399,1495,1667]" box="[189,917,1495,1522]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="FCD911CC7C5EFFE6858821D4FC25FA47" box="[189,917,1495,1522]" pageId="6" pageNumber="7">
Not 
<taxonomicName id="1BC94D617C5EFFE685C521DBFE0BFA44" ID-CoL="3R4ZK" authorityName="Jeffreys" authorityYear="1876" box="[240,443,1495,1521]" class="Bivalvia" family="Lasaeidae" genus="Kellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="symmetros">
<emphasis id="EEBDEAF07C5EFFE685C521DBFE0BFA44" box="[240,443,1495,1521]" italics="true" pageId="6" pageNumber="7">Kellia symmetros</emphasis>
</taxonomicName>
– 
<treatmentCitation id="5D6810F37C5EFFE684EC21DBFD1DFA47" author="Locard A." box="[473,685,1495,1522]" page="297" pageId="6" pageNumber="7" year="1898">
<bibRefCitation id="B8584B137C5EFFE684EC21DBFD1DFA47" author="Locard" box="[473,685,1495,1522]" firstAuthor="Locard" pageId="6" pageNumber="7" refId="ref9556" refString="Locard A. 1898. Expeditions Scientifiques du Travailleur et du Talisman pendant les annees 1880, 1811, 1882, 1883. Mollusques Testaces 2. Masson, Paris. http: // dx. doi. org / 10.5962 / bhl. title. 10477" type="journal volume" year="1898">Locard 1898: 297</bibRefCitation>
</treatmentCitation>
, pl.XIII, figs 18-20.
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<paragraph id="DC7636E27C5EFFE6858821F0FD1EF98F" blockId="6.[189,1399,1495,1667]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="FCD911CC7C5EFFE6858821F0FBA2F9A3" box="[189,1042,1531,1558]" pageId="6" pageNumber="7">
Not 
<taxonomicName id="1BC94D617C5EFFE685C521F0FE54F9A0" authorityName="Verrill &amp; Bush" authorityYear="1898" box="[240,484,1532,1558]" class="Bivalvia" family="Thyasiridae" genus="Axinodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="symmetros">
<emphasis id="EEBDEAF07C5EFFE685C521F0FE54F9A0" box="[240,484,1532,1558]" italics="true" pageId="6" pageNumber="7">Axinodon symmetros</emphasis>
</taxonomicName>
– 
<treatmentCitation id="5D6810F37C5EFFE6873721F7FC3AF9A0" author="Bouchet P. &amp; Waren A." box="[514,906,1531,1558]" page="216" pageId="6" pageNumber="7" year="1979">
<bibRefCitation id="B8584B137C5EFFE6873721F7FC3AF9A0" author="Bouchet &amp; Waren" box="[514,906,1531,1558]" firstAuthor="Bouchet" pageId="6" pageNumber="7" pagination="211 - 243" refId="ref8392" refString="Bouchet P. &amp; Waren A. 1979. The abyssal molluscan fauna of the Norwegian Sea and its relation to other faunas. Sarsia 64 (3): 211 - 243." type="journal article" year="1979">Bouchet &amp; Warén 1979: 216-217</bibRefCitation>
</treatmentCitation>
, figs 3A-D.
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<treatmentCitationGroup id="FCD911CC7C5EFFE6812F21F0FD1EF98F" pageId="6" pageNumber="7">
— 
<treatmentCitation id="5D6810F37C5EFFE6810821F0FAB9F9A3" author="Aartsen J. J. van" box="[1085,1289,1531,1558]" page="30" pageId="6" pageNumber="7" year="1996">
<bibRefCitation id="B8584B137C5EFFE6810821F0FAB9F9A3" author="Aartsen" box="[1085,1289,1531,1558]" firstAuthor="Aartsen" pageId="6" pageNumber="7" pagination="27 - 53" refId="ref8206" refString="Aartsen J. J. van. 1996. Galeommatacea &amp; Cyamiacea Part II. La Conchiglia 281: 27 - 53." type="journal article" year="1996">Aartsen 1996: 30</bibRefCitation>
</treatmentCitation>
, fig. 5 (is 
<taxonomicName id="1BC94D617C5EFFE68588222CFD11F98F" authority="Verrill &amp; Bush, 1898" authorityName="Verrill &amp; Bush" authorityYear="1898" box="[189,673,1568,1594]" class="Bivalvia" family="Thyasiridae" genus="Axinodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="ellipticus">
<emphasis id="EEBDEAF07C5EFFE68588222CFE11F98F" box="[189,417,1568,1594]" italics="true" pageId="6" pageNumber="7">Axinodon ellipticus</emphasis>
<bibRefCitation id="B8584B137C5EFFE68492222CFD11F98F" author="Verrill &amp; Bush" box="[423,673,1568,1594]" firstAuthor="Verrill" pageId="6" pageNumber="7" pagination="776 - 901" refId="ref10206" refString="Verrill A. E. &amp; Bush K. J. 1898. Revision of the deep-water Mollusca of the Atlantic coast of North America, with descriptions of new genera and species. Part I. - Bivalvia. Proceedings of the United States National Museum 20 (1139): 776 - 901, pls lxxi-xcvii. http: // dx. doi. org / 10.5962 / bhl. title. 1706" type="journal article" year="1898">Verrill &amp; Bush, 1898</bibRefCitation>
</taxonomicName>
).
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</paragraph>
<paragraph id="DC7636E27C5EFFE685882248FBC6F9EB" blockId="6.[189,1399,1495,1667]" box="[189,1142,1604,1631]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="FCD911CC7C5EFFE685882248FBC6F9EB" box="[189,1142,1604,1631]" pageId="6" pageNumber="7">
Not 
<taxonomicName id="1BC94D617C5EFFE685C52249FDCEF9EB" box="[240,638,1604,1630]" class="Bivalvia" family="Lasaeidae" genus="Montacuta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="symmetros" subGenus="Axinodon">
<emphasis id="EEBDEAF07C5EFFE685C52249FEDFF9EB" box="[240,367,1605,1630]" italics="true" pageId="6" pageNumber="7">Montacuta</emphasis>
(
<emphasis id="EEBDEAF07C5EFFE684B52248FE40F9EB" box="[384,496,1604,1630]" italics="true" pageId="6" pageNumber="7">Axinodon</emphasis>
) 
<emphasis id="EEBDEAF07C5EFFE687372249FDCEF9EB" box="[514,638,1605,1630]" italics="true" pageId="6" pageNumber="7">symmetros</emphasis>
</taxonomicName>
– 
<treatmentCitation id="5D6810F37C5EFFE687A92248FBC1F9EB" author="Gage J. D. &amp; Billett D. S. M. &amp; Jensen M. &amp; Tyler P. A." box="[668,1137,1604,1631]" page="189" pageId="6" pageNumber="7" year="1985">
<bibRefCitation id="B8584B137C5EFFE687A92248FBC1F9EB" author="Gage, Billett, Jensen &amp; Tyler" box="[668,1137,1604,1631]" firstAuthor="Gage" pageId="6" pageNumber="7" pagination="173 - 213" refId="ref9212" refString="Gage J. D., Billett D. S. M., Jensen M. &amp; Tyler P. A. 1985. Echinoderms of the Rockall Trough and adjacent areas 2. Echinoidea and Holothurioidea. Bulletin of the British Museum of Natural History (Zoology) 48 (4): 173 - 213." type="journal article" year="1985">Gage, Billett, Jensen &amp; Tyler 1985: 189</bibRefCitation>
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.
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</paragraph>
<paragraph id="DC7636E27C5EFFE685882264FCFEF937" blockId="6.[189,1399,1495,1667]" box="[189,846,1640,1667]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="FCD911CC7C5EFFE685882264FD23F937" box="[189,659,1640,1667]" pageId="6" pageNumber="7">
Not 
<taxonomicName id="1BC94D617C5EFFE685C52264FED0F937" authorityName="Verrill &amp; Bush" authorityYear="1898" box="[240,352,1640,1666]" class="Bivalvia" family="Thyasiridae" genus="Axinodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C5EFFE685C52264FED0F937" box="[240,352,1640,1666]" italics="true" pageId="6" pageNumber="7">Axinodon</emphasis>
</taxonomicName>
sp.1 – 
<treatmentCitation id="5D6810F37C5EFFE684802264FD23F937" author="Olabarria C." box="[437,659,1640,1667]" page="20" pageId="6" pageNumber="7" year="2005">
<bibRefCitation id="B8584B137C5EFFE684802264FD23F937" author="Olabarria" box="[437,659,1640,1667]" firstAuthor="Olabarria" pageId="6" pageNumber="7" pagination="15 - 31" refId="ref9788" refString="Olabarria C. 2005. Patterns of bathymetric zonation of bivalves in the Porcupine Seabight and adjacent Abyssal plain, NE Atlantic. Deep-Sea Research Part I: Oceanographic Research Papers 52 (1): 15 - 31. http: // dx. doi. org / 10.1016 / j. dsr. 2004.09.005" type="journal article" year="2005">Olabarria 2005: 20</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
(is 
<taxonomicName id="1BC94D617C5EFFE6878A2264FCACF937" authorityName="Angas" authorityYear="1877" box="[703,796,1640,1666]" class="Bivalvia" family="Lasaeidae" genus="Mysella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C5EFFE6878A2264FCACF937" box="[703,796,1640,1666]" italics="true" pageId="6" pageNumber="7">Mysella</emphasis>
</taxonomicName>
sp.)
</paragraph>
</subSubSection>
<subSubSection id="94D365697C5EFFE8858822A6FAC6FE6D" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="DC7636E27C5EFFE6858822A6FE1AF971" blockId="6.[189,426,1706,1732]" box="[189,426,1706,1732]" pageId="6" pageNumber="7">
<heading id="873E818E7C5EFFE6858822A6FE1AF971" bold="true" box="[189,426,1706,1732]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="EEBDEAF07C5EFFE6858822A6FE1AF971" bold="true" box="[189,426,1706,1732]" pageId="6" pageNumber="7">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="6CA13CBF7C5EFFE685DF22D6FCC5F889" ID-GBIF-Occurrence="2625650301" collectionCode="USNM" country="United States of America" county="National" elevation="3202" latitude="59.10" location="Davis Strait" longitude="-50.25" municipality="Godthaab" pageId="6" pageNumber="7" specimenCode="USNM170626." specimenCount="1" typeStatus="holotype">
<paragraph id="DC7636E27C5EFFE685DF22D6FEECF941" blockId="6.[189,1399,1754,1852]" box="[234,348,1754,1780]" pageId="6" pageNumber="7">
<emphasis id="EEBDEAF07C5EFFE685DF22D6FEECF941" bold="true" box="[234,348,1754,1780]" pageId="6" pageNumber="7">
<typeStatus id="037288407C5EFFE685DF22D6FEECF941" box="[234,348,1754,1780]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DC7636E27C5EFFE6858822F2FCC5F889" blockId="6.[189,1399,1754,1852]" pageId="6" pageNumber="7">
<specimenCount id="CACFFD6B7C5EFFE6858822F2FF7CF8AD" box="[189,204,1790,1816]" pageId="6" pageNumber="7">1</specimenCount>
shell, 
<locationDeviation id="CDC42FC07C5EFFE6842F22F2FE77F8AD" box="[282,455,1790,1816]" pageId="6" pageNumber="7">North Atlantic</locationDeviation>
, 
<locationDeviation id="CDC42FC07C5EFFE684E622F2FDAFF8AD" box="[467,543,1790,1816]" pageId="6" pageNumber="7">SW of</locationDeviation>
<collectingMunicipality id="3C12AC987C5EFFE6871222F2FD2CF8AD" box="[551,668,1790,1816]" pageId="6" pageNumber="7">Godthaab</collectingMunicipality>
, 
<location id="D91660397C5EFFE6879C22F2FC88F8AD" LSID="urn:lsid:plazi:treatment:546087F47C5EFFEC877320C4FD79F9FF:D91660397C5EFFE6879C22F2FC88F8AD" box="[681,824,1790,1816]" country="United States of America" county="National" municipality="Godthaab" name="Davis Strait" pageId="6" pageNumber="7">Davis Strait</location>
, 
<location id="D91660397C5EFFE6867622F2FC1BF8AD" LSID="urn:lsid:plazi:treatment:546087F47C5EFFEC877320C4FD79F9FF:D91660397C5EFFE6867622F2FC1BF8AD" box="[835,939,1790,1816]" country="United States of America" county="National" municipality="Godthaab" name="Valorous" pageId="6" pageNumber="7">Valorous</location>
stn 9, 
<geoCoordinate id="B9FD50257C5EFFE686C922F1FBEBF8AD" box="[1020,1115,1789,1816]" direction="north" orientation="latitude" pageId="6" pageNumber="7" value="59.10">59˚10'N</geoCoordinate>
<geoCoordinate id="B9FD50257C5EFFE6815622F1FB79F8AD" box="[1123,1225,1789,1816]" direction="west" orientation="longitude" pageId="6" pageNumber="7" value="-50.25">50˚25'W</geoCoordinate>
, 1750 fathoms (
<quantity id="1B319B077C5EFFE685F0232EFE93F889" box="[197,291,1826,1852]" metricMagnitude="3" metricUnit="m" metricValue="3.2019999999999995" pageId="6" pageNumber="7" unit="m" value="3202.0">
<elevation id="57E4D1D17C5EFFE685F0232EFE93F889" box="[197,291,1826,1852]" metricMagnitude="3" metricUnit="m" metricValue="3.2019999999999995" pageId="6" pageNumber="7" unit="m" value="3202.0">3202 m</elevation>
</quantity>
), United States National Museum- 
<specimenCode id="8C6F9E997C5EFFE687F4232EFCC5F889" box="[705,885,1826,1852]" pageId="6" pageNumber="7">USNM170626.</specimenCode>
</paragraph>
</materialsCitation>
<paragraph id="DC7636E27C5EFFE685DF2368FE16F8CB" blockId="6.[189,1399,1892,2027]" box="[234,422,1892,1918]" pageId="6" pageNumber="7">
<emphasis id="EEBDEAF07C5EFFE685DF2368FE16F8CB" bold="true" box="[234,422,1892,1918]" pageId="6" pageNumber="7">Other material</emphasis>
</paragraph>
<paragraph id="DC7636E27C5EFFE885882385FD9DFEFC" blockId="6.[189,1399,1892,2027]" lastBlockId="8.[189,1398,267,329]" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
<materialsCitation id="6CA13CBF7C5EFFE685882385FEF9F85E" ID-GBIF-Occurrence="5004498306" collectingDate="1977-04-30" collectionCode="NMW.Z." collectorName="E. Southward" elevation="4258" latitude="47.29" location="Shackleton" longitude="-9.33" municipality="Bay of Biscay" pageId="6" pageNumber="7" specimenCode="2012.015.1" specimenCount="2">
<specimenCount id="CACFFD6B7C5EFFE685882385FEE0F817" box="[189,336,1928,1955]" count="2" pageId="6" pageNumber="7" type="generic">2 specimens</specimenCount>
, attached to 
<taxonomicName id="1BC94D617C5EFFE684D62385FD5BF816" authority=", Bay" authorityName="Bay" box="[483,747,1929,1955]" class="Echinoidea" family="Aeropsidae" genus="Aeropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="6" pageNumber="7" phylum="Echinodermata" rank="species" species="rostrata">
<emphasis id="EEBDEAF07C5EFFE684D62385FD1DF817" box="[483,685,1929,1955]" italics="true" pageId="6" pageNumber="7">Aeropsis rostrata</emphasis>
, 
<collectingMunicipality id="3C12AC987C5EFFE687892385FCD3F817" box="[700,867,1928,1955]" pageId="6" pageNumber="7">Bay of Biscay</collectingMunicipality>
</taxonomicName>
, 
<location id="D91660397C5EFFE6865A2384FC41F817" LSID="urn:lsid:plazi:treatment:546087F47C5EFFEC877320C4FD79F9FF:D91660397C5EFFE6865A2384FC41F817" box="[879,1009,1928,1954]" municipality="Bay of Biscay" name="Shackleton" pageId="6" pageNumber="7">Shackleton</location>
cruise 1977/5, stn D7, 
<geoCoordinate id="B9FD50257C5EFFE680342384FAC7F816" box="[1281,1399,1928,1955]" direction="north" orientation="latitude" pageId="6" pageNumber="7" value="47.29">47˚29.7'N</geoCoordinate>
<geoCoordinate id="B9FD50257C5EFFE6858823A0FE88F872" box="[189,312,1964,1991]" direction="west" orientation="longitude" pageId="6" pageNumber="7" value="-09.33">09˚33.3'W</geoCoordinate>
, 
<quantity id="1B319B077C5EFFE6847623A0FE52F872" box="[323,482,1964,1991]" metricMagnitude="3" metricUnit="m" metricValue="4.2575" metricValueMax="4.265" metricValueMin="4.25" pageId="6" pageNumber="7" unit="m" value="4257.5" valueMax="4265.0" valueMin="4250.0">
<elevation id="57E4D1D17C5EFFE6847623A0FE52F872" box="[323,482,1964,1991]" metricMagnitude="3" metricUnit="m" metricValue="4.2575" metricValueMax="4.265" metricValueMin="4.25" pageId="6" pageNumber="7" unit="m" value="4257.5" valueMax="4265.0" valueMin="4250.0">4250-4265 m</elevation>
</quantity>
, 
<date id="A87710227C5EFFE684D823A1FD35F872" box="[493,645,1965,1991]" pageId="6" pageNumber="7" value="1977-04-30">
<collectingDate id="B833E9CA7C5EFFE684D823A1FD35F872" box="[493,645,1965,1991]" pageId="6" pageNumber="7" value="1977-04-30">30 Apr. 1977</collectingDate>
</date>
, leg. 
<collectorName id="713C53347C5EFFE687F723A1FCD4F873" box="[706,868,1964,1991]" pageId="6" pageNumber="7">E. Southward</collectorName>
, National Museum Wales, Zoology- 
<collectionCode id="BAD8AE277C5EFFE6803B23A1FAC7F872" box="[1294,1399,1965,1991]" pageId="6" pageNumber="7">NMW.Z.</collectionCode>
<specimenCode id="8C6F9E997C5EFFE6858823DCFEF5F85E" box="[189,325,2000,2027]" pageId="6" pageNumber="7">2012.015.1</specimenCode>
;
</materialsCitation>
<materialsCitation id="6CA13CBF7C5EFFE8846723DDFD9DFEFC" ID-GBIF-Occurrence="5004498301" collectingDate="1977-04-29" collectionCode="NMW.Z." collectorName="E. Southward" elevation="4143" lastPageId="8" lastPageNumber="9" latitude="47.35" location="Shackleton" longitude="-9.44" municipality="Bay of Biscay" pageId="6" pageNumber="7" specimenCode="2012.015.2" specimenCount="4">
<specimenCount id="CACFFD6B7C5EFFE6846723DDFE58F85E" box="[338,488,2001,2027]" count="4" pageId="6" pageNumber="7" type="generic">4 specimens</specimenCount>
, attached to 
<taxonomicName id="1BC94D617C5EFFE687B423DDFC3DF85E" authority=", Bay" authorityName="Bay" box="[641,909,2001,2027]" class="Echinoidea" family="Aeropsidae" genus="Aeropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="6" pageNumber="7" phylum="Echinodermata" rank="species" species="rostrata">
<emphasis id="EEBDEAF07C5EFFE687B423DDFCFDF85E" box="[641,845,2001,2027]" italics="true" pageId="6" pageNumber="7">Aeropsis rostrata</emphasis>
, 
<collectingMunicipality id="3C12AC987C5EFFE6866B23DDFBB9F85E" box="[862,1033,2001,2027]" pageId="6" pageNumber="7">Bay of Biscay</collectingMunicipality>
</taxonomicName>
, 
<location id="D91660397C5EFFE6812223DDFB29F85E" LSID="urn:lsid:plazi:treatment:546087F47C5EFFEC877320C4FD79F9FF:D91660397C5EFFE6812223DDFB29F85E" box="[1047,1177,2001,2027]" municipality="Bay of Biscay" name="Shackleton" pageId="6" pageNumber="7">Shackleton</location>
cruise 1977/5, stn D2, 
<geoCoordinate id="B9FD50257C50FFE885C52507FEC4FE93" box="[240,372,267,294]" direction="north" orientation="latitude" pageId="8" pageNumber="9" value="47.35">47˚35.52'N</geoCoordinate>
<geoCoordinate id="B9FD50257C50FFE884492507FDB7FE93" box="[380,519,267,294]" direction="west" orientation="longitude" pageId="8" pageNumber="9" value="-9.44">09˚44.07'W</geoCoordinate>
, 
<quantity id="1B319B077C50FFE887262507FD03FE93" box="[531,691,267,294]" metricMagnitude="3" metricUnit="m" metricValue="4.1425" metricValueMax="4.165" metricValueMin="4.12" pageId="8" pageNumber="9" unit="m" value="4142.5" valueMax="4165.0" valueMin="4120.0">
<elevation id="57E4D1D17C50FFE887262507FD03FE93" box="[531,691,267,294]" metricMagnitude="3" metricUnit="m" metricValue="4.1425" metricValueMax="4.165" metricValueMin="4.12" pageId="8" pageNumber="9" unit="m" value="4142.5" valueMax="4165.0" valueMin="4120.0">4120-4165 m</elevation>
</quantity>
, 
<date id="A87710227C50FFE887F52507FCECFE90" box="[704,860,267,294]" pageId="8" pageNumber="9" value="1977-04-29">
<collectingDate id="B833E9CA7C50FFE887F52507FCECFE90" box="[704,860,267,294]" pageId="8" pageNumber="9" value="1977-04-29">29 Apr. 1977</collectingDate>
</date>
, leg. 
<collectorName id="713C53347C50FFE886A82500FBF0FE90" box="[925,1088,267,294]" pageId="8" pageNumber="9">E. Southward</collectorName>
, National Museum Wales, Zoology- 
<collectionCode id="BAD8AE277C50FFE884042523FE2BFEFC" box="[305,411,303,329]" pageId="8" pageNumber="9">NMW.Z.</collectionCode>
<specimenCode id="8C6F9E997C50FFE884972522FD99FEFC" box="[418,553,302,329]" pageId="8" pageNumber="9">2012.015.2</specimenCode>
.
</materialsCitation>
</paragraph>
<caption id="88B6666A7C5FFFE785882386FC15F85E" ID-DOI="http://doi.org/10.5281/zenodo.3857755" ID-Zenodo-Dep="3857755" httpUri="https://zenodo.org/record/3857755/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1930,1956]" targetBox="[230,1356,266,1891]" targetPageId="7">
<paragraph id="DC7636E27C5FFFE785882386FC15F85E" blockId="7.[189,1399,1930,2027]" pageId="7" pageNumber="8">
<emphasis id="EEBDEAF07C5FFFE785882386FEBBF811" bold="true" box="[189,267,1930,1956]" pageId="7" pageNumber="8">Fig. 2.</emphasis>
Holotype of 
<taxonomicName id="1BC94D617C5FFFE784922386FC93F811" authority="Jeffreys, 1876" authorityName="Jeffreys" authorityYear="1876" box="[423,803,1930,1956]" class="Bivalvia" family="Lasaeidae" genus="Kellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="symmetros">
<emphasis id="EEBDEAF07C5FFFE784922386FDC3F811" box="[423,627,1930,1956]" italics="true" pageId="7" pageNumber="8">Kellia symmetros</emphasis>
<bibRefCitation id="B8584B137C5FFFE7874E2386FC93F811" author="Jeffreys" box="[635,803,1930,1956]" firstAuthor="Jeffreys" pageId="7" pageNumber="8" pagination="490 - 499" refId="ref9385" refString="Jeffreys J. G. 1876. New and peculiar Mollusca of the Kellia, Lucina, Cyprina and Corbula families procured in the ' Valorous' Expedition. Annals and Magazine of Natural History ser. 4, 18: 490 - 499." type="journal article" year="1876">Jeffreys, 1876</bibRefCitation>
</taxonomicName>
, USNM 170626. 
<emphasis id="EEBDEAF07C5FFFE786C62386FB96F811" bold="true" box="[1011,1062,1930,1956]" pageId="7" pageNumber="8">A-B</emphasis>
. SEM of hinges of right and left valves. 
<emphasis id="EEBDEAF07C5FFFE7847023A1FEC9F872" bold="true" box="[325,377,1965,1991]" pageId="7" pageNumber="8">C-D</emphasis>
. SEM of internal of right and left valves. 
<emphasis id="EEBDEAF07C5FFFE7865023A2FC24F87D" bold="true" box="[869,916,1966,1992]" pageId="7" pageNumber="8">E-F</emphasis>
SEM of external of right and left valves. 
<emphasis id="EEBDEAF07C5FFFE7858823DCFF44F85F" bold="true" box="[189,244,2000,2026]" pageId="7" pageNumber="8">G-H</emphasis>
. photo micrographs of internal and external of right valve.
</paragraph>
</caption>
<paragraph id="DC7636E27C50FFE885882583FEEDFE1D" blockId="8.[189,349,398,425]" box="[189,349,398,425]" pageId="8" pageNumber="9">
<heading id="873E818E7C50FFE885882583FEEDFE1D" bold="true" box="[189,349,398,425]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="EEBDEAF07C50FFE885882583FEEDFE1D" bold="true" box="[189,349,398,425]" pageId="8" pageNumber="9">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="DC7636E27C50FFE8858825B1FAC6FE6D" blockId="8.[189,1398,445,472]" box="[189,1398,445,472]" pageId="8" pageNumber="9">
North Atlantic, Valorous St. 9, SW of Godthaab, Davis Strait, 
<geoCoordinate id="B9FD50257C50FFE886B425B1FC56FE6D" box="[897,998,445,472]" degrees="59" direction="north" minutes="10" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="59.166668">59°10’N</geoCoordinate>
<geoCoordinate id="B9FD50257C50FFE886DE25B1FBE7FE6D" box="[1003,1111,445,472]" degrees="50" direction="west" minutes="25" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="-50.416668">50°25’W</geoCoordinate>
, 1750 fathoms (
<quantity id="1B319B077C50FFE8803825B1FAD9FE6D" box="[1293,1385,445,472]" metricMagnitude="3" metricUnit="m" metricValue="3.2019999999999995" pageId="8" pageNumber="9" unit="m" value="3202.0">3202 m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="94D365697C50FFE885882611FA91F883" pageId="8" pageNumber="9" type="description">
<paragraph id="DC7636E27C50FFE885882611FD3EFD82" blockId="8.[189,654,541,567]" box="[189,654,541,567]" pageId="8" pageNumber="9">
<heading id="873E818E7C50FFE885882611FD3EFD82" bold="true" box="[189,654,541,567]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="EEBDEAF07C50FFE885882611FD3EFD82" bold="true" box="[189,654,541,567]" pageId="8" pageNumber="9">
Redescription of the 
<typeStatus id="037288407C50FFE884F42611FD9DFD82" box="[449,557,541,567]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
(
<figureCitation id="44F22A677C50FFE887092611FD35FD82" box="[572,645,541,567]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[230,1356,266,1891]" captionTargetId="figure@7.[230,1356,266,1891]" captionTargetPageId="7" captionText="Fig. 2. Holotype of Kellia symmetros Jeffreys, 1876, USNM 170626. A-B. SEM of hinges of right and left valves. C-D. SEM of internal of right and left valves. E-F SEM of external of right and left valves. G-H. photo micrographs of internal and external of right valve." figureDoi="http://doi.org/10.5281/zenodo.3857755" httpUri="https://zenodo.org/record/3857755/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="DC7636E27C50FFE885882640FDA0FC12" blockId="8.[189,1399,588,935]" pageId="8" pageNumber="9">
Shell minute, length 1.0 mm, height 
<quantity id="1B319B077C50FFE887532640FD61FDD3" box="[614,721,588,614]" metricMagnitude="-4" metricUnit="m" metricValue="7.7" pageId="8" pageNumber="9" unit="mm" value="0.77">0.77 mm</quantity>
. Thin, translucent. Equivalve. Weakly inequilateral, beaks just behind the midline. Umbos prominent, beaks orthogyrate. Outline subovate, slightly extended anteriorly; anterior dorsal margin sloping into broadly rounded anterior margin; posterior dorsal margin indistinct sloping into broadly rounded posterior margin, this slightly less expanded than anterior margin; ventral margin weakly curved. Sculpture weak, of indistinct commarginal lines most obvious on lateral margins. Prodissoconch II distinct, 373 μm across, with weak commarginal lines. Hinge plate weak, Ligament short, internal, attached to a shallow resilifer situated beneath and posterior of the beaks. Right valve with a single, projecting, cardinal peg, immediately anterior to this tooth a slight depression. Left valve with a short, weak, marginal flange in a posterior lateral position. Adductor scars oval, roughly of equal size; pallial line entire.
</paragraph>
<paragraph id="DC7636E27C50FFE8858827E1FC8EFBB2" blockId="8.[189,830,1005,1031]" box="[189,830,1005,1031]" pageId="8" pageNumber="9">
<heading id="873E818E7C50FFE8858827E1FC8EFBB2" bold="true" box="[189,830,1005,1031]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="EEBDEAF07C50FFE8858827E1FC8EFBB2" bold="true" box="[189,830,1005,1031]" pageId="8" pageNumber="9">Description (based on material from Bay of Biscay)</emphasis>
</heading>
</paragraph>
<paragraph id="DC7636E27C50FFE885882010FAA3FA6B" blockId="8.[189,1399,1052,1503]" pageId="8" pageNumber="9">
SHELL. (
<figureCitation id="44F22A677C50FFE8842C2010FEEEFB83" box="[281,350,1052,1078]" captionStart="Fig" captionStartId="9.[189,232,1895,1921]" captionTargetBox="[244,1343,266,1854]" captionTargetId="figure@9.[244,1343,266,1854]" captionTargetPageId="9" captionText="Fig. 3. Kelliola symmetros (Jeffreys, 1876) from Biscay. A-C. Photo micrographs a external of left valve, internals of both valves. D-E. SEM of hinges of right and left valves. F-H. SEM of internal of both valves and external of left valve. I. SEM of prodissoconch. J. SEM of anterior area showing weak radial sculpture. K. SEM of margin showing transverse grooves." figureDoi="http://doi.org/10.5281/zenodo.3857757" httpUri="https://zenodo.org/record/3857757/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
) Minute, largest of length 
<quantity id="1B319B077C50FFE887A32010FD43FB83" box="[662,755,1052,1078]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="9" unit="mm" value="1.3">1.3 mm</quantity>
, height 1.0 mm. Thin, fragile. Equivalve. Inequilateral, beaks behind midline. Umbos weakly inflated, beaks orthogyrate. Outline subovate, longer than high, length to height ratio 1.3:1, slightly extended anteriorly; anterior dorsal margin sloping, rather straight merging smoothly with rounded anterior; posterior dorsal margin shorter and sloping more steeply than anterior, merging smoothly with rounded posterior margin; anterior slightly more expanded than posterior; ventral margin gently curved. Sculpture weak almost smooth, of fine commarginal lines; radial lines apparent under transmitted light (
<figureCitation id="44F22A677C50FFE887BB20FCFD5CFABF" box="[654,748,1264,1290]" captionStart="Fig" captionStartId="9.[189,232,1895,1921]" captionTargetBox="[244,1343,266,1854]" captionTargetId="figure@9.[244,1343,266,1854]" captionTargetPageId="9" captionText="Fig. 3. Kelliola symmetros (Jeffreys, 1876) from Biscay. A-C. Photo micrographs a external of left valve, internals of both valves. D-E. SEM of hinges of right and left valves. F-H. SEM of internal of both valves and external of left valve. I. SEM of prodissoconch. J. SEM of anterior area showing weak radial sculpture. K. SEM of margin showing transverse grooves." figureDoi="http://doi.org/10.5281/zenodo.3857757" httpUri="https://zenodo.org/record/3857757/files/figure.png" pageId="8" pageNumber="9">Fig. 3A</figureCitation>
) but these very faintly raised (
<figureCitation id="44F22A677C50FFE8816B20FCFB04FABF" box="[1118,1204,1264,1290]" captionStart="Fig" captionStartId="9.[189,232,1895,1921]" captionTargetBox="[244,1343,266,1854]" captionTargetId="figure@9.[244,1343,266,1854]" captionTargetPageId="9" captionText="Fig. 3. Kelliola symmetros (Jeffreys, 1876) from Biscay. A-C. Photo micrographs a external of left valve, internals of both valves. D-E. SEM of hinges of right and left valves. F-H. SEM of internal of both valves and external of left valve. I. SEM of prodissoconch. J. SEM of anterior area showing weak radial sculpture. K. SEM of margin showing transverse grooves." figureDoi="http://doi.org/10.5281/zenodo.3857757" httpUri="https://zenodo.org/record/3857757/files/figure.png" pageId="8" pageNumber="9">Fig. 3J</figureCitation>
). Prodissoconch II distinct, 
<quantity id="1B319B077C50FFE88477211FFE10FA9B" box="[322,416,1299,1326]" metricMagnitude="-1" metricUnit="m" metricValue="3.8" pageId="8" pageNumber="9" unit="mm" value="380.0">380 µm</quantity>
in diameter sculptured with commarginal lines (
<figureCitation id="44F22A677C50FFE886DE211FFBF0FA98" box="[1003,1088,1299,1325]" captionStart="Fig" captionStartId="9.[189,232,1895,1921]" captionTargetBox="[244,1343,266,1854]" captionTargetId="figure@9.[244,1343,266,1854]" captionTargetPageId="9" captionText="Fig. 3. Kelliola symmetros (Jeffreys, 1876) from Biscay. A-C. Photo micrographs a external of left valve, internals of both valves. D-E. SEM of hinges of right and left valves. F-H. SEM of internal of both valves and external of left valve. I. SEM of prodissoconch. J. SEM of anterior area showing weak radial sculpture. K. SEM of margin showing transverse grooves." figureDoi="http://doi.org/10.5281/zenodo.3857757" httpUri="https://zenodo.org/record/3857757/files/figure.png" pageId="8" pageNumber="9">Fig. 3I</figureCitation>
); Prodissoconch I weakly demarcated, 
<quantity id="1B319B077C50FFE88463213BFE04FAE4" box="[342,436,1335,1361]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="9" unit="mm" value="140.0">140 µm</quantity>
in diameter with a punctate micro-sculpture (
<figureCitation id="44F22A677C50FFE886D4213BFB86FAE4" box="[993,1078,1335,1361]" captionStart="Fig" captionStartId="9.[189,232,1895,1921]" captionTargetBox="[244,1343,266,1854]" captionTargetId="figure@9.[244,1343,266,1854]" captionTargetPageId="9" captionText="Fig. 3. Kelliola symmetros (Jeffreys, 1876) from Biscay. A-C. Photo micrographs a external of left valve, internals of both valves. D-E. SEM of hinges of right and left valves. F-H. SEM of internal of both valves and external of left valve. I. SEM of prodissoconch. J. SEM of anterior area showing weak radial sculpture. K. SEM of margin showing transverse grooves." figureDoi="http://doi.org/10.5281/zenodo.3857757" httpUri="https://zenodo.org/record/3857757/files/figure.png" pageId="8" pageNumber="9">Fig. 3I</figureCitation>
). Ligament short, internal, attached to a shallow resilifer situated beneath and posterior of the beaks. Right valve with a single, projecting, cardinal peg, immediately anterior to this tooth a slight depression (
<figureCitation id="44F22A677C50FFE881662171FB01FA2D" box="[1107,1201,1405,1432]" captionStart="Fig" captionStartId="9.[189,232,1895,1921]" captionTargetBox="[244,1343,266,1854]" captionTargetId="figure@9.[244,1343,266,1854]" captionTargetPageId="9" captionText="Fig. 3. Kelliola symmetros (Jeffreys, 1876) from Biscay. A-C. Photo micrographs a external of left valve, internals of both valves. D-E. SEM of hinges of right and left valves. F-H. SEM of internal of both valves and external of left valve. I. SEM of prodissoconch. J. SEM of anterior area showing weak radial sculpture. K. SEM of margin showing transverse grooves." figureDoi="http://doi.org/10.5281/zenodo.3857757" httpUri="https://zenodo.org/record/3857757/files/figure.png" pageId="8" pageNumber="9">Fig. 3D</figureCitation>
). Left valve with a short, weak, marginal flange in a posterior lateral position (
<figureCitation id="44F22A677C50FFE886BA21ADFC5BFA0E" box="[911,1003,1441,1467]" captionStart="Fig" captionStartId="9.[189,232,1895,1921]" captionTargetBox="[244,1343,266,1854]" captionTargetId="figure@9.[244,1343,266,1854]" captionTargetPageId="9" captionText="Fig. 3. Kelliola symmetros (Jeffreys, 1876) from Biscay. A-C. Photo micrographs a external of left valve, internals of both valves. D-E. SEM of hinges of right and left valves. F-H. SEM of internal of both valves and external of left valve. I. SEM of prodissoconch. J. SEM of anterior area showing weak radial sculpture. K. SEM of margin showing transverse grooves." figureDoi="http://doi.org/10.5281/zenodo.3857757" httpUri="https://zenodo.org/record/3857757/files/figure.png" pageId="8" pageNumber="9">Fig. 3E</figureCitation>
). Adductor scars oval, roughly of equal size; pallial line entire. Ventral margin dissected by minute transverse grooves (
<figureCitation id="44F22A677C50FFE8819221C8FAB5FA6B" box="[1191,1285,1476,1502]" captionStart="Fig" captionStartId="9.[189,232,1895,1921]" captionTargetBox="[244,1343,266,1854]" captionTargetId="figure@9.[244,1343,266,1854]" captionTargetPageId="9" captionText="Fig. 3. Kelliola symmetros (Jeffreys, 1876) from Biscay. A-C. Photo micrographs a external of left valve, internals of both valves. D-E. SEM of hinges of right and left valves. F-H. SEM of internal of both valves and external of left valve. I. SEM of prodissoconch. J. SEM of anterior area showing weak radial sculpture. K. SEM of margin showing transverse grooves." figureDoi="http://doi.org/10.5281/zenodo.3857757" httpUri="https://zenodo.org/record/3857757/files/figure.png" pageId="8" pageNumber="9">Fig. 3K</figureCitation>
).
</paragraph>
<paragraph id="DC7636E27C50FFE88588220AFDF7F941" blockId="8.[189,1399,1542,1780]" pageId="8" pageNumber="9">
ANATOMY. (
<figureCitation id="44F22A677C50FFE8840A220AFE2AF995" box="[319,410,1542,1568]" captionStart="Fig" captionStartId="10.[189,232,1895,1921]" captionTargetBox="[189,1399,630,1853]" captionTargetId="figure@10.[189,1399,630,1853]" captionTargetPageId="10" captionText="Fig. 4. Scanning electron micrographs of anatomy. A. K. symmetros (Jeffreys, 1876) from Biscay. B. Syssitomya pourtalesiana sp. nov. from Norwegian Sea. C. Montacuta substriata (Montagu, 1808) from North Sea. D. Excised ctenidium of M. substriata. E. Excised piece of ctenidium from S. pourtalesiana sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3857759" httpUri="https://zenodo.org/record/3857759/files/figure.png" pageId="8" pageNumber="9">Fig. 4A</figureCitation>
) Mantle margin free for most of its length, joined and attached to terminal of gill axis, anterior (pedal) aperture extensive, posterior aperture very small. Adductor muscles of approximately equal size. Foot with a large toe and small heel, byssus functional producing a mass of threads arising from a single stalk. Anterior pedal retractor inserted above the anterior adductor, posterior pedal retractor above the posterior adductor. Ctenidium of a single demibranch, with nine non-reflected filaments in the largest specimen. Filaments rod shaped, lacking abfrontal extension or harbouring symbiotic bacteria. Labial palps small but projecting.
</paragraph>
<paragraph id="DC7636E27C50FFE885882310FA91F883" blockId="8.[189,1313,1819,1846]" box="[189,1313,1819,1846]" pageId="8" pageNumber="9">
ASSOCIATION. Attached by byssus threads to the spines of the echinoid 
<taxonomicName id="1BC94D617C50FFE886C52310FB0AF880" box="[1008,1210,1820,1846]" class="Echinoidea" family="Aeropsidae" genus="Aeropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="rostrata">
<emphasis id="EEBDEAF07C50FFE886C52310FB0AF880" box="[1008,1210,1820,1846]" italics="true" pageId="8" pageNumber="9">Aeropsis rostrata</emphasis>
</taxonomicName>
(
<figureCitation id="44F22A677C50FFE881FF2317FAA2F883" box="[1226,1298,1819,1846]" captionStart="Fig" captionStartId="11.[189,232,881,907]" captionTargetBox="[455,1133,444,839]" captionTargetId="figure@11.[455,1133,444,839]" captionTargetPageId="11" captionText="Fig. 5. Aeropsis rostrata (Wyville Thomson, 1877) with Kelliola symmetros (Jeffreys, 1876) attached, from Biscay." figureDoi="http://doi.org/10.5281/zenodo.3857761" httpUri="https://zenodo.org/record/3857761/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="94D365697C50FFE885882377FB76F85E" pageId="8" pageNumber="9" type="distribution">
<paragraph id="DC7636E27C50FFE885882377FEE6F820" blockId="8.[189,342,1915,1941]" box="[189,342,1915,1941]" pageId="8" pageNumber="9">
<heading id="873E818E7C50FFE885882377FEE6F820" bold="true" box="[189,342,1915,1941]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="EEBDEAF07C50FFE885882377FEE6F820" bold="true" box="[189,342,1915,1941]" pageId="8" pageNumber="9">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DC7636E27C50FFE8858823A6FB76F85E" blockId="8.[189,1399,1962,2027]" pageId="8" pageNumber="9">
<taxonomicName id="1BC94D617C50FFE8858823A6FE2FF871" baseAuthorityName="Jeffreys" baseAuthorityYear="1876" box="[189,415,1962,1988]" class="Bivalvia" family="Lasaeidae" genus="Kelliola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="symmetros">
<emphasis id="EEBDEAF07C50FFE8858823A6FE2FF871" box="[189,415,1962,1988]" italics="true" pageId="8" pageNumber="9">Kelliola symmetros</emphasis>
</taxonomicName>
is known only from the 
<typeStatus id="037288407C50FFE8878A23A7FD42F870" box="[703,754,1963,1989]" pageId="8" pageNumber="9">type</typeStatus>
locality and from the Bay of Biscay, at abyssal depths. The host echinoid is widely distributed in the North Atlantic (
<bibRefCitation id="B8584B137C50FFE886BB23DDFB09F85E" author="Echinoid Directory" box="[910,1209,2001,2027]" firstAuthor="Echinoid Directory" pageId="8" pageNumber="9" refId="ref8948" refString="Echinoid Directory 2012. Smith A. B. &amp; Kroh A. (eds). Available at http: // www. nhm. ac. uk / researchcuration / research / projects / echinoid-directory / index. html [accessed the 21 Mar. 2012]" type="url" year="2012">Echinoid Directory 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="88B6666A7C51FFE98588236BFC02F85E" ID-DOI="http://doi.org/10.5281/zenodo.3857757" ID-Zenodo-Dep="3857757" httpUri="https://zenodo.org/record/3857757/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,232,1895,1921]" targetBox="[244,1343,266,1854]" targetPageId="9">
<paragraph id="DC7636E27C51FFE98588236BFC02F85E" blockId="9.[189,1398,1895,2027]" pageId="9" pageNumber="10">
<emphasis id="EEBDEAF07C51FFE98588236BFEBEF834" bold="true" box="[189,270,1895,1921]" pageId="9" pageNumber="10">Fig. 3.</emphasis>
<taxonomicName id="1BC94D617C51FFE9842C236BFD77F837" authority="(Jeffreys, 1876)" baseAuthorityName="Jeffreys" baseAuthorityYear="1876" box="[281,711,1895,1922]" class="Bivalvia" family="Lasaeidae" genus="Kelliola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="symmetros">
<emphasis id="EEBDEAF07C51FFE9842C236BFE4FF834" box="[281,511,1895,1921]" italics="true" pageId="9" pageNumber="10">Kelliola symmetros</emphasis>
(
<bibRefCitation id="B8584B137C51FFE98727236BFD0FF837" author="Jeffreys" box="[530,703,1895,1922]" firstAuthor="Jeffreys" pageId="9" pageNumber="10" pagination="490 - 499" refId="ref9385" refString="Jeffreys J. G. 1876. New and peculiar Mollusca of the Kellia, Lucina, Cyprina and Corbula families procured in the ' Valorous' Expedition. Annals and Magazine of Natural History ser. 4, 18: 490 - 499." type="journal article" year="1876">Jeffreys, 1876</bibRefCitation>
)
</taxonomicName>
from Biscay. 
<emphasis id="EEBDEAF07C51FFE98643236BFC1AF834" bold="true" box="[886,938,1895,1921]" pageId="9" pageNumber="10">A-C</emphasis>
. Photo micrographs a external of left valve, internals of both valves. 
<emphasis id="EEBDEAF07C51FFE987012386FDD7F811" bold="true" box="[564,615,1930,1956]" pageId="9" pageNumber="10">D-E</emphasis>
. SEM of hinges of right and left valves. 
<emphasis id="EEBDEAF07C51FFE981672386FB35F811" bold="true" box="[1106,1157,1930,1956]" pageId="9" pageNumber="10">F-H</emphasis>
. SEM of internal of both valves and external of left valve. 
<emphasis id="EEBDEAF07C51FFE9874B23A2FD39F87D" bold="true" box="[638,649,1966,1992]" pageId="9" pageNumber="10">I</emphasis>
. SEM of prodissoconch. 
<emphasis id="EEBDEAF07C51FFE9868623A2FC72F87D" bold="true" box="[947,962,1966,1992]" pageId="9" pageNumber="10">J</emphasis>
. SEM of anterior area showing weak radial sculpture. 
<emphasis id="EEBDEAF07C51FFE984B423DDFE28F85E" bold="true" box="[385,408,2001,2027]" pageId="9" pageNumber="10">K</emphasis>
. SEM of margin showing transverse grooves.
</paragraph>
</caption>
<subSubSection id="94D365697C52FFEC85882507FD79F9FF" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="DC7636E27C52FFEA85882507FE79FE90" blockId="10.[189,457,267,293]" box="[189,457,267,293]" pageId="10" pageNumber="11">
<heading id="873E818E7C52FFEA85882507FE79FE90" bold="true" box="[189,457,267,293]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="EEBDEAF07C52FFEA85882507FE79FE90" bold="true" box="[189,457,267,293]" pageId="10" pageNumber="11">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DC7636E27C52FFEA85882535FC4EFDFF" blockId="10.[189,1399,313,586]" pageId="10" pageNumber="11">
The hinges of 
<taxonomicName id="1BC94D617C52FFEA84582536FDBEFEE6" baseAuthorityName="Jeffreys" baseAuthorityYear="1876" box="[365,526,314,339]" class="Bivalvia" family="Lasaeidae" genus="Kelliola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="symmetros">
<emphasis id="EEBDEAF07C52FFEA84582536FDBEFEE6" box="[365,526,314,339]" italics="true" pageId="10" pageNumber="11">K. symmetros</emphasis>
</taxonomicName>
and the 
<taxonomicName id="1BC94D617C52FFEA874F2536FD50FEE1" authorityName="Mortensen" authorityYear="1907" box="[634,736,314,340]" class="Echinoidea" family="Aeropsidae" genus="Aeropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="genus">
<emphasis id="EEBDEAF07C52FFEA874F2536FD50FEE1" box="[634,736,314,340]" italics="true" pageId="10" pageNumber="11">Aeropsis</emphasis>
</taxonomicName>
commensal are almost identical, but 
<taxonomicName id="1BC94D617C52FFEA81912536FAF5FEE6" baseAuthorityName="Jeffreys" baseAuthorityYear="1876" box="[1188,1349,314,339]" class="Bivalvia" family="Lasaeidae" genus="Kelliola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="symmetros">
<emphasis id="EEBDEAF07C52FFEA81912536FAF5FEE6" box="[1188,1349,314,339]" italics="true" pageId="10" pageNumber="11">K. symmetros</emphasis>
</taxonomicName>
has slightly more prominent umbos and lacks the marginal transverse grooves. Given that 
<taxonomicName id="1BC94D617C52FFEA81E02551FAC6FEC3" baseAuthorityName="Jeffreys" baseAuthorityYear="1876" box="[1237,1398,349,374]" class="Bivalvia" family="Lasaeidae" genus="Kelliola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="symmetros">
<emphasis id="EEBDEAF07C52FFEA81E02551FAC6FEC3" box="[1237,1398,349,374]" italics="true" pageId="10" pageNumber="11">K. symmetros</emphasis>
</taxonomicName>
was taken in the same sample along with 
<taxonomicName id="1BC94D617C52FFEA879F258CFCA0FE2F" authorityName="Mortensen" authorityYear="1907" box="[682,784,384,410]" class="Echinoidea" family="Aeropsidae" genus="Aeropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="genus">
<emphasis id="EEBDEAF07C52FFEA879F258CFCA0FE2F" box="[682,784,384,410]" italics="true" pageId="10" pageNumber="11">Aeropsis</emphasis>
</taxonomicName>
, but not attached to it, it is possible that the two are associated. This suggests an ecological affinity with the 
<taxonomicName id="1BC94D617C52FFEA865425AFFC77FE08" authorityName="Mortensen" authorityYear="1907" box="[865,967,419,445]" class="Echinoidea" family="Aeropsidae" genus="Aeropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="genus">
<emphasis id="EEBDEAF07C52FFEA865425AFFC77FE08" box="[865,967,419,445]" italics="true" pageId="10" pageNumber="11">Aeropsis</emphasis>
</taxonomicName>
commensal described here. Despite the wide geographical separation of the samples considered here, 
<taxonomicName id="1BC94D617C52FFEA86FE25CBFB26FE55" box="[971,1174,455,481]" class="Echinoidea" family="Aeropsidae" genus="Aeropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="rostrata">
<emphasis id="EEBDEAF07C52FFEA86FE25CBFB26FE55" box="[971,1174,455,481]" italics="true" pageId="10" pageNumber="11">Aeropsis rostrata</emphasis>
</taxonomicName>
is regarded as pan Atlantic and having an abyssal bathymetric range (WoRMS 2012). With so few specimens at hand and the poor condition of the 
<typeStatus id="037288407C52FFEA84DE2600FDE2FD93" box="[491,594,524,550]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="1BC94D617C52FFEA874F2601FCAAFD93" baseAuthorityName="Jeffreys" baseAuthorityYear="1876" box="[634,794,525,550]" class="Bivalvia" family="Lasaeidae" genus="Kelliola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="symmetros">
<emphasis id="EEBDEAF07C52FFEA874F2601FCAAFD93" box="[634,794,525,550]" italics="true" pageId="10" pageNumber="11">K. symmetros</emphasis>
</taxonomicName>
, we have chosen to be conservative and regard the 
<taxonomicName id="1BC94D617C52FFEA8588263CFE93FDFF" authorityName="Mortensen" authorityYear="1907" box="[189,291,560,586]" class="Echinoidea" family="Aeropsidae" genus="Aeropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="genus">
<emphasis id="EEBDEAF07C52FFEA8588263CFE93FDFF" box="[189,291,560,586]" italics="true" pageId="10" pageNumber="11">Aeropsis</emphasis>
</taxonomicName>
commensal from Biscay conspecific with 
<taxonomicName id="1BC94D617C52FFEA8620263CFC47FDFF" baseAuthorityName="Jeffreys" baseAuthorityYear="1876" box="[789,1015,560,586]" class="Bivalvia" family="Lasaeidae" genus="Kelliola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="symmetros">
<emphasis id="EEBDEAF07C52FFEA8620263CFC47FDFF" box="[789,1015,560,586]" italics="true" pageId="10" pageNumber="11">Kelliola symmetros</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="88B6666A7C52FFEA8588236BFEA8F85E" ID-DOI="http://doi.org/10.5281/zenodo.3857759" ID-Zenodo-Dep="3857759" httpUri="https://zenodo.org/record/3857759/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,232,1895,1921]" targetBox="[189,1399,630,1853]" targetPageId="10">
<paragraph id="DC7636E27C52FFEA8588236BFEA8F85E" blockId="10.[189,1398,1895,2027]" pageId="10" pageNumber="11">
<emphasis id="EEBDEAF07C52FFEA8588236BFEBCF834" bold="true" box="[189,268,1895,1921]" pageId="10" pageNumber="11">Fig. 4.</emphasis>
Scanning electron micrographs of anatomy. 
<emphasis id="EEBDEAF07C52FFEA8613236BFC8BF834" bold="true" box="[806,827,1895,1921]" pageId="10" pageNumber="11">A</emphasis>
. 
<taxonomicName id="1BC94D617C52FFEA86792364FB01F837" authority="(Jeffreys, 1876)" baseAuthorityName="Jeffreys" baseAuthorityYear="1876" box="[844,1201,1895,1922]" class="Bivalvia" family="Lasaeidae" genus="Kelliola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="symmetros">
<emphasis id="EEBDEAF07C52FFEA86792364FC5CF834" box="[844,1004,1896,1921]" italics="true" pageId="10" pageNumber="11">K. symmetros</emphasis>
(
<bibRefCitation id="B8584B137C52FFEA86CB236BFB19F837" author="Jeffreys" box="[1022,1193,1895,1922]" firstAuthor="Jeffreys" pageId="10" pageNumber="11" pagination="490 - 499" refId="ref9385" refString="Jeffreys J. G. 1876. New and peculiar Mollusca of the Kellia, Lucina, Cyprina and Corbula families procured in the ' Valorous' Expedition. Annals and Magazine of Natural History ser. 4, 18: 490 - 499." type="journal article" year="1876">Jeffreys, 1876</bibRefCitation>
)
</taxonomicName>
from Biscay. 
<emphasis id="EEBDEAF07C52FFEA8069236BFAC0F834" bold="true" box="[1372,1392,1895,1921]" pageId="10" pageNumber="11">B</emphasis>
. 
<taxonomicName id="1BC94D617C52FFEA85882387FE59F811" authority="Oliver, 2012" authorityName="Oliver" authorityYear="2012" box="[189,489,1930,1957]" class="Bivalvia" family="Lasaeidae" genus="Syssitomya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="pourtalesiana" status="sp. nov.">
<emphasis id="EEBDEAF07C52FFEA85882387FE59F811" box="[189,489,1930,1957]" italics="true" pageId="10" pageNumber="11">Syssitomya pourtalesiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F58E578B7C52FFEA84DA2387FDF8F810" box="[495,584,1931,1957]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
from Norwegian Sea. 
<emphasis id="EEBDEAF07C52FFEA867B2386FCD3F811" bold="true" box="[846,867,1930,1956]" pageId="10" pageNumber="11">C</emphasis>
. 
<taxonomicName id="1BC94D617C52FFEA86442387FA87F811" authority="(Montagu, 1808)" baseAuthorityName="Montagu" baseAuthorityYear="1808" box="[881,1335,1930,1956]" class="Bivalvia" family="Lasaeidae" genus="Montacuta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="EEBDEAF07C52FFEA86442387FBDBF811" box="[881,1131,1930,1956]" italics="true" pageId="10" pageNumber="11">Montacuta substriata</emphasis>
(Montagu, 1808)
</taxonomicName>
from North Sea. 
<emphasis id="EEBDEAF07C52FFEA847423A2FEE6F87D" bold="true" box="[321,342,1966,1992]" pageId="10" pageNumber="11">D</emphasis>
. Excised ctenidium of 
<taxonomicName id="1BC94D617C52FFEA875023A2FCB1F872" baseAuthorityName="Montagu" baseAuthorityYear="1808" box="[613,769,1965,1991]" class="Bivalvia" family="Lasaeidae" genus="Montacuta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="EEBDEAF07C52FFEA875023A2FCB1F872" box="[613,769,1965,1991]" italics="true" pageId="10" pageNumber="11">M. substriata</emphasis>
</taxonomicName>
. 
<emphasis id="EEBDEAF07C52FFEA863A23A2FC93F87D" bold="true" box="[783,803,1966,1992]" pageId="10" pageNumber="11">E</emphasis>
. Excised piece of ctenidium from 
<taxonomicName id="1BC94D617C52FFEA818323A2FAC6F872" authority="Oliver, 2012" authorityName="Oliver" authorityYear="2012" box="[1206,1398,1965,1992]" class="Bivalvia" family="Lasaeidae" genus="Syssitomya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="pourtalesiana" status="sp. nov.">
<emphasis id="EEBDEAF07C52FFEA818323A2FAC6F872" box="[1206,1398,1965,1992]" italics="true" pageId="10" pageNumber="11">S. pourtalesiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F58E578B7C52FFEA858823DDFEA8F85E" box="[189,280,2001,2027]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="DC7636E27C53FFEC85882507FCC3FE25" blockId="11.[189,1399,267,400]" lastBlockId="12.[189,1399,267,400]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
Consequently, at the family level the hinge and anatomical characters of 
<taxonomicName id="1BC94D617C53FFEB81392507FBDBFE90" authorityName="Dall" authorityYear="1899" box="[1036,1131,267,293]" class="Bivalvia" family="Lasaeidae" genus="Kelliola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C53FFEB81392507FBDBFE90" box="[1036,1131,267,293]" italics="true" pageId="11" pageNumber="12">Kelliola</emphasis>
</taxonomicName>
are entirely in keeping with the 
<taxonomicName id="1BC94D617C53FFEB84172522FE76FEFD" authorityName="Clark" authorityYear="1855" box="[290,454,302,328]" class="Bivalvia" family="Montacutidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="11" pageNumber="12" phylum="Mollusca" rank="family">Montacutidae</taxonomicName>
, consisting of a single cardinal peg in the right valve, a marginal flange in the left valve and an internal ligament. The ligament is attached to an elongate shallow depression extending below the beaks and is therefore most similar to 
<emphasis id="EEBDEAF07C53FFEB87C5257AFBB8FE3A" box="[752,1032,374,399]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="1BC94D617C53FFEB87C5257AFCDFFE3A" authorityName="Turton" authorityYear="1822" box="[752,879,374,399]" class="Bivalvia" family="Lasaeidae" genus="Montacuta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus" sensu="stricto">Montacuta</taxonomicName>
<taxonomicNameLabel id="F58E578B7C53FFEB8640257AFBB8FE3A" box="[885,1032,374,399]" pageId="11" pageNumber="12" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="44F22A677C53FFEB812D2579FBEDFE25" box="[1048,1117,373,400]" captionStart="Fig" captionStartId="11.[189,232,1930,1956]" captionTargetBox="[189,1399,987,1890]" captionTargetId="figure@11.[189,1399,987,1890]" captionTargetPageId="11" captionText="Fig. 6. Montacuta substriata (Montagu, 1808), NMW.Z 2000.101.73. Gulfaks Oil Field, North Sea, 217 m. A-B. SEM of hinges of right and left valves. C. SEM of internal of right valve. D. SEM of external of left valve." figureDoi="http://doi.org/10.5281/zenodo.3857763" httpUri="https://zenodo.org/record/3857763/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
). 
<emphasis id="EEBDEAF07C53FFEB8141257AFAC6FE3A" box="[1140,1398,373,399]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="1BC94D617C53FFEB8141257AFADFFE3A" baseAuthorityName="Montagu" baseAuthorityYear="1808" box="[1140,1391,373,399]" class="Bivalvia" family="Lasaeidae" genus="Montacuta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="substriata">Montacuta substriata</taxonomicName>
,
</emphasis>
the 
<typeStatus id="037288407C54FFEC85D22500FEAAFE93" box="[231,282,268,294]" pageId="12" pageNumber="13">type</typeStatus>
species of 
<emphasis id="EEBDEAF07C54FFEC84AF2500FD90FE90" box="[410,544,268,293]" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="1BC94D617C54FFEC84AF2500FDA9FE90" authorityName="Turton" authorityYear="1822" box="[410,537,268,293]" class="Bivalvia" family="Lasaeidae" genus="Montacuta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">Montacuta</taxonomicName>
,
</emphasis>
has a longer cardinal tooth, has radial ridges and ovate in outline (
<figureCitation id="44F22A677C54FFEC80142507FAD8FE93" box="[1313,1384,267,294]" captionStart="Fig" captionStartId="11.[189,232,1930,1956]" captionTargetBox="[189,1399,987,1890]" captionTargetId="figure@11.[189,1399,987,1890]" captionTargetPageId="11" captionText="Fig. 6. Montacuta substriata (Montagu, 1808), NMW.Z 2000.101.73. Gulfaks Oil Field, North Sea, 217 m. A-B. SEM of hinges of right and left valves. C. SEM of internal of right valve. D. SEM of external of left valve." figureDoi="http://doi.org/10.5281/zenodo.3857763" httpUri="https://zenodo.org/record/3857763/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
). Anatomically 
<taxonomicName id="1BC94D617C54FFEC845E2523FE7CFEFC" authorityName="Dall" authorityYear="1899" box="[363,460,303,329]" class="Bivalvia" family="Lasaeidae" genus="Kelliola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C54FFEC845E2523FE7CFEFC" box="[363,460,303,329]" italics="true" pageId="12" pageNumber="13">Kelliola</emphasis>
</taxonomicName>
is similar to many montacutids, where the ctenidium is reduced to a single demibranch. However, in this genus, the filaments are very few and not reflected (
<figureCitation id="44F22A677C54FFEC81F0255EFA9AFED9" box="[1221,1322,338,364]" captionStart="Fig" captionStartId="10.[189,232,1895,1921]" captionTargetBox="[189,1399,630,1853]" captionTargetId="figure@10.[189,1399,630,1853]" captionTargetPageId="10" captionText="Fig. 4. Scanning electron micrographs of anatomy. A. K. symmetros (Jeffreys, 1876) from Biscay. B. Syssitomya pourtalesiana sp. nov. from Norwegian Sea. C. Montacuta substriata (Montagu, 1808) from North Sea. D. Excised ctenidium of M. substriata. E. Excised piece of ctenidium from S. pourtalesiana sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3857759" httpUri="https://zenodo.org/record/3857759/files/figure.png" pageId="12" pageNumber="13">Fig. 4A</figureCitation>
). This condition could be due to the small size of the specimens.
</paragraph>
<caption id="88B6666A7C53FFEB8588277DFEE3FC1B" ID-DOI="http://doi.org/10.5281/zenodo.3857761" ID-Zenodo-Dep="3857761" httpUri="https://zenodo.org/record/3857761/files/figure.png" pageId="11" pageNumber="12" startId="11.[189,232,881,907]" targetBox="[455,1133,444,839]" targetPageId="11">
<paragraph id="DC7636E27C53FFEB8588277DFEE3FC1B" blockId="11.[189,1398,881,942]" pageId="11" pageNumber="12">
<emphasis id="EEBDEAF07C53FFEB8588277DFEBBFC3E" bold="true" box="[189,267,881,907]" pageId="11" pageNumber="12">Fig. 5.</emphasis>
<taxonomicName id="1BC94D617C53FFEB8426277DFCADFC3E" authority="(Wyville Thomson, 1877)" baseAuthorityName="Wyville Thomson" baseAuthorityYear="1877" box="[275,797,881,907]" class="Echinoidea" family="Aeropsidae" genus="Aeropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="11" pageNumber="12" phylum="Echinodermata" rank="species" species="rostrata">
<emphasis id="EEBDEAF07C53FFEB8426277DFE6DFC3E" box="[275,477,881,907]" italics="true" pageId="11" pageNumber="12">Aeropsis rostrata</emphasis>
(Wyville Thomson, 1877)
</taxonomicName>
with 
<taxonomicName id="1BC94D617C53FFEB8654277DFAB5FC39" authority="(Jeffreys, 1876)" baseAuthorityName="Jeffreys" baseAuthorityYear="1876" box="[865,1285,881,908]" class="Bivalvia" family="Lasaeidae" genus="Kelliola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="symmetros">
<emphasis id="EEBDEAF07C53FFEB8654277DFC70FC3E" box="[865,960,881,907]" italics="true" pageId="11" pageNumber="12">Kelliola</emphasis>
<emphasis id="EEBDEAF07C53FFEB86F2277EFBF3FC3E" box="[967,1091,882,907]" italics="true" pageId="11" pageNumber="12">symmetros</emphasis>
(
<bibRefCitation id="B8584B137C53FFEB8161277DFB4DFC39" author="Jeffreys" box="[1108,1277,881,908]" firstAuthor="Jeffreys" pageId="11" pageNumber="12" pagination="490 - 499" refId="ref9385" refString="Jeffreys J. G. 1876. New and peculiar Mollusca of the Kellia, Lucina, Cyprina and Corbula families procured in the ' Valorous' Expedition. Annals and Magazine of Natural History ser. 4, 18: 490 - 499." type="journal article" year="1876">Jeffreys, 1876</bibRefCitation>
)
</taxonomicName>
attached, from Biscay.
</paragraph>
</caption>
<caption id="88B6666A7C53FFEB85882386FE08F85E" ID-DOI="http://doi.org/10.5281/zenodo.3857763" ID-Zenodo-Dep="3857763" httpUri="https://zenodo.org/record/3857763/files/figure.png" pageId="11" pageNumber="12" startId="11.[189,232,1930,1956]" targetBox="[189,1399,987,1890]" targetPageId="11">
<paragraph id="DC7636E27C53FFEB85882386FE08F85E" blockId="11.[189,1399,1930,2027]" pageId="11" pageNumber="12">
<emphasis id="EEBDEAF07C53FFEB85882386FEBEF811" bold="true" box="[189,270,1930,1956]" pageId="11" pageNumber="12">Fig. 6.</emphasis>
<taxonomicName id="1BC94D617C53FFEB842D2387FD5FF811" authority="(Montagu, 1808)" baseAuthorityName="Montagu" baseAuthorityYear="1808" box="[280,751,1930,1956]" class="Bivalvia" family="Lasaeidae" genus="Montacuta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="substriata">
<emphasis id="EEBDEAF07C53FFEB842D2387FE27F811" box="[280,407,1931,1956]" italics="true" pageId="11" pageNumber="12">Montacuta</emphasis>
<emphasis id="EEBDEAF07C53FFEB84972386FDA7F811" box="[418,535,1930,1956]" italics="true" pageId="11" pageNumber="12">substriata</emphasis>
(Montagu, 1808)
</taxonomicName>
, NMW.Z 2000.101.73. Gulfaks Oil Field, North Sea, 217 m. 
<emphasis id="EEBDEAF07C53FFEB842923A1FEFFF872" bold="true" box="[284,335,1965,1991]" pageId="11" pageNumber="12">A-B</emphasis>
. SEM of hinges of right and left valves. 
<emphasis id="EEBDEAF07C53FFEB867223A1FCD4F872" bold="true" box="[839,868,1965,1991]" pageId="11" pageNumber="12">C.</emphasis>
SEM of internal of right valve. 
<emphasis id="EEBDEAF07C53FFEB81C423A2FAB6F87D" bold="true" box="[1265,1286,1966,1992]" pageId="11" pageNumber="12">D</emphasis>
. SEM of external of left valve.
</paragraph>
</caption>
<paragraph id="DC7636E27C54FFEC858825B3FB51FB80" blockId="12.[189,1399,447,1077]" pageId="12" pageNumber="13">
<bibRefCitation id="B8584B137C54FFEC858825B3FEFAFE6C" author="Dall" box="[189,330,447,473]" firstAuthor="Dall" pageId="12" pageNumber="13" pagination="873 - 897" refId="ref8857" refString="Dall W. H. 1899. Synopsis of the Recent and Tertiary Leptonacea of North America and the West Indies. Proceedings of the United States National Museum 21 (1177): 873 - 897, 2 pls. http: // dx. doi. org / 10.5479 / si. 00963801.21 - 1177.873" type="book chapter" year="1899">Dall (1899)</bibRefCitation>
noted that Jeffreys’ (1876) description of the hinge of 
<taxonomicName id="1BC94D617C54FFEC86DE25CCFB3FFE6C" baseAuthorityName="Jeffreys" baseAuthorityYear="1876" box="[1003,1167,448,473]" class="Bivalvia" family="Lasaeidae" genus="Kelliola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="symmetros">
<emphasis id="EEBDEAF07C54FFEC86DE25CCFB3FFE6C" box="[1003,1167,448,473]" italics="true" pageId="12" pageNumber="13">K. symmetros</emphasis>
</taxonomicName>
was incorrect and, in re-describing it, created the new genus 
<taxonomicName id="1BC94D617C54FFEC878B25EFFCAFFE48" authorityName="Dall" authorityYear="1899" box="[702,799,483,509]" class="Bivalvia" family="Lasaeidae" genus="Kelliola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C54FFEC878B25EFFCAFFE48" box="[702,799,483,509]" italics="true" pageId="12" pageNumber="13">Kelliola</emphasis>
</taxonomicName>
for it. Dall did not make comparisons with other genera but noted that it was similar to 
<taxonomicName id="1BC94D617C54FFEC87A3260AFCCAFD95" authority="Lea, 1846" authorityName="Lea" authorityYear="1846" box="[662,890,518,545]" class="Bivalvia" family="Lasaeidae" genus="Aligena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C54FFEC87A3260AFD43FD95" box="[662,755,518,544]" italics="true" pageId="12" pageNumber="13">Aligena</emphasis>
Lea, 1846
</taxonomicName>
. This is difficult to understand as 
<taxonomicName id="1BC94D617C54FFEC802C260AFAC6FD95" authorityName="Lea" authorityYear="1846" box="[1305,1398,518,544]" class="Bivalvia" family="Lasaeidae" genus="Aligena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C54FFEC802C260AFAC6FD95" box="[1305,1398,518,544]" italics="true" pageId="12" pageNumber="13">Aligena</emphasis>
</taxonomicName>
species have a cardinal peg in each valve (
<bibRefCitation id="B8584B137C54FFEC87992626FC8AFDF0" author="Harry" box="[684,826,554,581]" firstAuthor="Harry" pageId="12" pageNumber="13" pagination="164 - 181" refId="ref9355" refString="Harry H. W. 1969. A review of the living leptonacean bivalves of the genus Aligena. The Veliger 11 (3): 164 - 181." type="journal article" year="1969">Harry 1969</bibRefCitation>
). Comparisons with other montacutid genera are currently complex due to a lack of compatibility in descriptions of characters and the widely varying use of generic names. Such difficulties were amply recognised by 
<bibRefCitation id="B8584B137C54FFEC8135267DFAA0FD3E" author="Gofas &amp; Salas" box="[1024,1296,625,651]" firstAuthor="Gofas" pageId="12" pageNumber="13" pagination="119 - 135" refId="ref9271" refString="Gofas S. &amp; Salas C. 2008. A review of European &quot; Mysella &quot; species (Bivalvia: Montacutidae) with description of Kurtiella new genus. Journal of Molluscan Studies 74 (2): 119 - 135. http: // dx. doi. org / 10.1093 / mollus / eym 053" type="journal article" year="2008">Gofas &amp; Salas (2008)</bibRefCitation>
in their review of 
<taxonomicName id="1BC94D617C54FFEC84022698FD84FD1B" authority="Angas, 1877" authorityName="Angas" authorityYear="1877" box="[311,564,660,686]" class="Bivalvia" family="Lasaeidae" genus="Mysella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C54FFEC84022698FE26FD1B" box="[311,406,660,686]" italics="true" pageId="12" pageNumber="13">Mysella</emphasis>
Angas, 1877
</taxonomicName>
and consequent creation of the genus 
<taxonomicName id="1BC94D617C54FFEC81372698FAC2FD1B" authority="Gofas &amp; Salas, 2008" authorityName="Gofas &amp; Salas" authorityYear="2008" box="[1026,1394,660,686]" class="Bivalvia" family="Lasaeidae" genus="Kurtiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C54FFEC81372698FBDFFD1B" box="[1026,1135,660,686]" italics="true" pageId="12" pageNumber="13">Kurtiella</emphasis>
<bibRefCitation id="B8584B137C54FFEC81432698FAC2FD1B" author="Gofas &amp; Salas" box="[1142,1394,660,686]" firstAuthor="Gofas" pageId="12" pageNumber="13" pagination="119 - 135" refId="ref9271" refString="Gofas S. &amp; Salas C. 2008. A review of European &quot; Mysella &quot; species (Bivalvia: Montacutidae) with description of Kurtiella new genus. Journal of Molluscan Studies 74 (2): 119 - 135. http: // dx. doi. org / 10.1093 / mollus / eym 053" type="journal article" year="2008">Gofas &amp; Salas, 2008</bibRefCitation>
</taxonomicName>
. For the purposes of this paper, comparisons are restricted to genera that have a single cardinal peg in the right valve only and left valve with varying degrees of pseudocardinal development. In hinge characters, 
<taxonomicName id="1BC94D617C54FFEC847D26F3FE19FCAC" authorityName="Dall" authorityYear="1899" box="[328,425,767,793]" class="Bivalvia" family="Lasaeidae" genus="Kelliola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C54FFEC847D26F3FE19FCAC" box="[328,425,767,793]" italics="true" pageId="12" pageNumber="13">Kelliola</emphasis>
</taxonomicName>
is most similar to 
<emphasis id="EEBDEAF07C54FFEC87A5270CFC04FCAC" box="[656,948,768,793]" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="1BC94D617C54FFEC87A5270CFCA2FCAC" authorityName="Turton" authorityYear="1822" box="[656,786,768,793]" class="Bivalvia" family="Lasaeidae" genus="Montacuta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus" sensu="stricto">Montacuta</taxonomicName>
<taxonomicNameLabel id="F58E578B7C54FFEC862E270CFC04FCAC" box="[795,948,768,793]" pageId="12" pageNumber="13" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="44F22A677C54FFEC86F326F3FBA0FCAF" box="[966,1040,767,794]" captionStart="Fig" captionStartId="11.[189,232,1930,1956]" captionTargetBox="[189,1399,987,1890]" captionTargetId="figure@11.[189,1399,987,1890]" captionTargetPageId="11" captionText="Fig. 6. Montacuta substriata (Montagu, 1808), NMW.Z 2000.101.73. Gulfaks Oil Field, North Sea, 217 m. A-B. SEM of hinges of right and left valves. C. SEM of internal of right valve. D. SEM of external of left valve." figureDoi="http://doi.org/10.5281/zenodo.3857763" httpUri="https://zenodo.org/record/3857763/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
) and 
<taxonomicName id="1BC94D617C54FFEC8162270CFAC7FCAC" authority="Gabb, 1873" authorityName="Gabb" authorityYear="1873" box="[1111,1399,767,793]" class="Bivalvia" family="Lasaeidae" genus="Neaeromya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C54FFEC8162270CFB50FCAC" box="[1111,1248,768,793]" italics="true" pageId="12" pageNumber="13">Neaeromya</emphasis>
Gabb, 1873
</taxonomicName>
(
<bibRefCitation id="B8584B137C54FFEC85F0272EFE3FFC89" author="Coan" box="[197,399,802,828]" etAl="et al." firstAuthor="Coan" pageId="12" pageNumber="13" refId="ref8805" refString="Coan E. V., Scott P. V. &amp; Bernard F. R. 2000. Bivalve seashells of western North America. Marine bivalve mollusks from Arctic Alaska to Baja California. Santa Barbara Museum of Natural History Monographs 2. Santa Barbara Museum of Natural History." type="book" year="2000">
Coan 
<emphasis id="EEBDEAF07C54FFEC8438272FFEF9FC89" box="[269,329,802,828]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2000
</bibRefCitation>
) in that the development of the posterior teeth is limited to a marginal flange in the left valve and the ligament is attached to a shallow depression beneath and posterior to the beaks. For 
<taxonomicName id="1BC94D617C54FFEC85882766FD7FFC31" authority="(Carpenter, 1864) Narchi (1969)" authorityName="Narchi" authorityYear="1969" baseAuthorityName="Carpenter" baseAuthorityYear="1864" box="[189,719,873,900]" class="Bivalvia" family="Lasaeidae" genus="Neaeromya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="rugifera">
<emphasis id="EEBDEAF07C54FFEC85882766FEF3FC36" box="[189,323,873,899]" italics="true" pageId="12" pageNumber="13">N. rugifera</emphasis>
(Carpenter, 1864) 
<bibRefCitation id="B8584B137C54FFEC87102765FD7FFC31" author="Narchi" box="[549,719,873,900]" firstAuthor="Narchi" pageId="12" pageNumber="13" pagination="43 - 52" refId="ref9694" refString="Narchi W. 1969. On Pseudopythina rugifera (Carpenter, 1864) Bivalvia. The Veliger 12: 43 - 52" type="journal article" year="1969">Narchi (1969)</bibRefCitation>
</taxonomicName>
states that there is a tooth in each valve, although it may be reduced in the left valve (Paul Valentich-Scott pers comm); a further example of the contradictory descriptions found for the montacutid species. In 
<taxonomicName id="1BC94D617C54FFEC862327BCFBF3FC7F" authority="T. Brown, 1827" authorityName="T. Brown" authorityYear="1827" box="[790,1091,944,971]" class="Bivalvia" family="Lasaeidae" genus="Tellimya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C54FFEC862327BCFCCCFC7F" box="[790,892,944,970]" italics="true" pageId="12" pageNumber="13">Tellimya</emphasis>
T. Brown, 1827
</taxonomicName>
the resilifer is developed and the hinge plate thickened accordingly (see 
<bibRefCitation id="B8584B137C54FFEC863D27D8FC6CFC5A" author="Ockelmann" box="[776,988,980,1007]" firstAuthor="Ockelmann" pageId="12" pageNumber="13" pagination="211 - 221" refId="ref9716" refString="Ockelmann K. W. 1965. Redescription, distribution, biology, and dimorphous sperm of Montacuta tenella Loven (Mollusca, Leptonacea). Ophelia 2 (1): 211 - 221." type="journal article" year="1965">Ockelmann 1965</bibRefCitation>
, 
<figureCitation id="44F22A677C54FFEC86DE27D8FB87FC5B" box="[1003,1079,980,1006]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[230,1356,266,1891]" captionTargetId="figure@7.[230,1356,266,1891]" captionTargetPageId="7" captionText="Fig. 2. Holotype of Kellia symmetros Jeffreys, 1876, USNM 170626. A-B. SEM of hinges of right and left valves. C-D. SEM of internal of right and left valves. E-F SEM of external of right and left valves. G-H. photo micrographs of internal and external of right valve." figureDoi="http://doi.org/10.5281/zenodo.3857755" httpUri="https://zenodo.org/record/3857755/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
). In 
<taxonomicName id="1BC94D617C54FFEC814127D8FAA4FC5B" authorityName="Jespersen, Lutzen &amp; Nielsen" authorityYear="2004" box="[1140,1300,980,1006]" class="Bivalvia" family="Lasaeidae" genus="Montacutella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C54FFEC814127D8FAA4FC5B" box="[1140,1300,980,1006]" italics="true" pageId="12" pageNumber="13">Montacutella</emphasis>
</taxonomicName>
the left valve flange is developed as a small projection (
<bibRefCitation id="B8584B137C54FFEC863127FBFBB5FBA4" author="Jespersen" box="[772,1029,1015,1041]" etAl="et al." firstAuthor="Jespersen" pageId="12" pageNumber="13" pagination="3 - 19" refId="ref9429" refString="Jespersen A., Lutzen J. &amp; Nielsen C. 2004. On three species and two new genera (Montacutella and Brachiomya) of galeommatid bivalves from the irregular Sea Urchin Brissus latecarinatus with emphasis on their reproduction. Zoologischer Anzeiger 243: 3 - 19. http: // dx. doi. org / 10.1016 / j. jcz. 2004.04.001" type="journal article" year="2004">
Jespersen 
<emphasis id="EEBDEAF07C54FFEC86B427F4FC0EFBA4" box="[897,958,1015,1041]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2004
</bibRefCitation>
) and approaches the condition seen in 
<taxonomicName id="1BC94D617C54FFEC842C2017FEC6FB80" authorityName="Lea" authorityYear="1846" box="[281,374,1051,1077]" class="Bivalvia" family="Lasaeidae" genus="Aligena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C54FFEC842C2017FEC6FB80" box="[281,374,1051,1077]" italics="true" pageId="12" pageNumber="13">Aligena</emphasis>
</taxonomicName>
. The shell of 
<taxonomicName id="1BC94D617C54FFEC87292017FD1EFB80" authorityName="Jespersen, Lutzen &amp; Nielsen" authorityYear="2004" box="[540,686,1051,1077]" class="Bivalvia" family="Lasaeidae" genus="Brachiomya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C54FFEC87292017FD1EFB80" box="[540,686,1051,1077]" italics="true" pageId="12" pageNumber="13">Brachiomya</emphasis>
</taxonomicName>
is like that of 
<taxonomicName id="1BC94D617C54FFEC86692017FB6DFB80" authority="(Jespersen et al. 2004)" baseAuthorityName="Jespersen" baseAuthorityYear="2004" box="[860,1245,1051,1077]" class="Bivalvia" family="Lasaeidae" genus="Tellimya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C54FFEC86692017FC72FB80" box="[860,962,1051,1077]" italics="true" pageId="12" pageNumber="13">Tellimya</emphasis>
(
<bibRefCitation id="B8584B137C54FFEC86E72017FB64FB80" author="Jespersen" box="[978,1236,1051,1077]" etAl="et al." firstAuthor="Jespersen" pageId="12" pageNumber="13" pagination="3 - 19" refId="ref9429" refString="Jespersen A., Lutzen J. &amp; Nielsen C. 2004. On three species and two new genera (Montacutella and Brachiomya) of galeommatid bivalves from the irregular Sea Urchin Brissus latecarinatus with emphasis on their reproduction. Zoologischer Anzeiger 243: 3 - 19. http: // dx. doi. org / 10.1016 / j. jcz. 2004.04.001" type="journal article" year="2004">
Jespersen 
<emphasis id="EEBDEAF07C54FFEC817A2010FB3BFB80" box="[1103,1163,1051,1077]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2004
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="DC7636E27C54FFEC85882068FDB2FB70" blockId="12.[189,1399,1124,1222]" pageId="12" pageNumber="13">
<taxonomicName id="1BC94D617C54FFEC85882068FEAEFBCB" authorityName="Dall" authorityYear="1899" box="[189,286,1124,1150]" class="Bivalvia" family="Lasaeidae" genus="Kelliola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C54FFEC85882068FEAEFBCB" box="[189,286,1124,1150]" italics="true" pageId="12" pageNumber="13">Kelliola</emphasis>
</taxonomicName>
is as different from 
<emphasis id="EEBDEAF07C54FFEC87262069FC84FBCB" box="[531,820,1125,1150]" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="1BC94D617C54FFEC87262069FD25FBCB" authorityName="Turton" authorityYear="1822" box="[531,661,1125,1150]" class="Bivalvia" family="Lasaeidae" genus="Montacuta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus" sensu="stricto">Montacuta</taxonomicName>
<taxonomicNameLabel id="F58E578B7C54FFEC87A92069FC84FBCB" box="[668,820,1125,1150]" pageId="12" pageNumber="13" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
as are the other genera and a molecular study is required to evaluate the significance of the morphological characters. 
<taxonomicName id="1BC94D617C54FFEC81242084FBC2FB17" authorityName="Dall" authorityYear="1899" box="[1041,1138,1160,1186]" class="Bivalvia" family="Lasaeidae" genus="Kelliola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C54FFEC81242084FBC2FB17" box="[1041,1138,1160,1186]" italics="true" pageId="12" pageNumber="13">Kelliola</emphasis>
</taxonomicName>
is retained here until such a study is undertaken.
</paragraph>
<paragraph id="DC7636E27C54FFEC858820E1FD79F9FF" blockId="12.[189,1399,1261,1610]" pageId="12" pageNumber="13">
Species level comparisons are restricted to the few abyssal galeommatid species that have been described and none other than that described below under 
<taxonomicName id="1BC94D617C54FFEC86A6211DFB7AFA9F" authority="Oliver, 2012" authorityName="Oliver" authorityYear="2012" box="[915,1226,1296,1323]" class="Bivalvia" family="Lasaeidae" genus="Syssitomya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="pourtalesiana" status="sp. nov.">
<emphasis id="EEBDEAF07C54FFEC86A6211DFB7AFA9F" box="[915,1226,1296,1323]" italics="true" pageId="12" pageNumber="13">Syssitomya pourtalesiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F58E578B7C54FFEC81E1211DFA84FA9E" box="[1236,1332,1297,1323]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
have been found attached to echinoids. The shell of 
<taxonomicName id="1BC94D617C54FFEC87DA2138FC05FAFB" authority="Oliver, 2012" authorityName="Oliver" authorityYear="2012" box="[751,949,1332,1358]" class="Bivalvia" family="Lasaeidae" genus="Syssitomya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="pourtalesiana" status="sp. nov.">
<emphasis id="EEBDEAF07C54FFEC87DA2138FC05FAFB" box="[751,949,1332,1358]" italics="true" pageId="12" pageNumber="13">S. pourtalesiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F58E578B7C54FFEC86892138FBA9FAFB" box="[956,1049,1332,1358]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
is more expanded anteriorly, has a more depressed lunule and lacks marginal notches. The ctenidium is highly modified with laminar filaments whereas that of 
<taxonomicName id="1BC94D617C54FFEC875F2170FCA1FA20" baseAuthorityName="Jeffreys" baseAuthorityYear="1876" box="[618,785,1404,1429]" class="Bivalvia" family="Lasaeidae" genus="Kelliola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="symmetros">
<emphasis id="EEBDEAF07C54FFEC875F2170FCA1FA20" box="[618,785,1404,1429]" italics="true" pageId="12" pageNumber="13">K. symmetros</emphasis>
</taxonomicName>
is not modified in this manner. Other described, Atlantic, deep-sea, galeommatids have been assigned to the genera 
<taxonomicName id="1BC94D617C54FFEC86C32192FBE5FA0D" authorityName="Angas" authorityYear="1877" box="[1014,1109,1438,1464]" class="Bivalvia" family="Lasaeidae" genus="Mysella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C54FFEC86C32192FBE5FA0D" box="[1014,1109,1438,1464]" italics="true" pageId="12" pageNumber="13">Mysella</emphasis>
</taxonomicName>
(now 
<taxonomicName id="1BC94D617C54FFEC81902192FAA2FA0D" authorityName="Gofas &amp; Salas" authorityYear="2008" box="[1189,1298,1438,1464]" class="Bivalvia" family="Lasaeidae" genus="Kurtiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C54FFEC81902192FAA2FA0D" box="[1189,1298,1438,1464]" italics="true" pageId="12" pageNumber="13">Kurtiella</emphasis>
</taxonomicName>
) (
<bibRefCitation id="B8584B137C54FFEC80182192FED5FA69" author="Gofas &amp; Salas" firstAuthor="Gofas" pageId="12" pageNumber="13" pagination="119 - 135" refId="ref9271" refString="Gofas S. &amp; Salas C. 2008. A review of European &quot; Mysella &quot; species (Bivalvia: Montacutidae) with description of Kurtiella new genus. Journal of Molluscan Studies 74 (2): 119 - 135. http: // dx. doi. org / 10.1093 / mollus / eym 053" type="journal article" year="2008">Gofas &amp; Salas 2008</bibRefCitation>
), 
<taxonomicName id="1BC94D617C54FFEC844E21CEFCACFA69" authority="Dall, 1899" authorityName="Dall" authorityYear="1899" box="[379,796,1474,1500]" class="Bivalvia" family="Lasaeidae" genus="Epilepton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C54FFEC844E21CEFE40FA69" box="[379,496,1474,1500]" italics="true" pageId="12" pageNumber="13">Epilepton</emphasis>
<bibRefCitation id="B8584B137C54FFEC84CD21CEFDC8FA69" author="Dall" box="[504,632,1474,1500]" firstAuthor="Dall" pageId="12" pageNumber="13" pagination="873 - 897" refId="ref8857" refString="Dall W. H. 1899. Synopsis of the Recent and Tertiary Leptonacea of North America and the West Indies. Proceedings of the United States National Museum 21 (1177): 873 - 897, 2 pls. http: // dx. doi. org / 10.5479 / si. 00963801.21 - 1177.873" type="book chapter" year="1899">Dall, 1899</bibRefCitation>
(
<bibRefCitation id="B8584B137C54FFEC87BC21CEFCA4FA69" author="Allen" box="[649,788,1474,1500]" firstAuthor="Allen" pageId="12" pageNumber="13" pagination="7 - 14" refId="ref8229" refString="Allen J. A. 2007. A new deep-water species of the genus Epilepton (Bivalvia: Galeommatoidea) from the Atlantic. The Veliger 49 (1): 7 - 14." type="journal article" year="2007">Allen 2007</bibRefCitation>
)
</taxonomicName>
or 
<taxonomicName id="1BC94D617C54FFEC867321CEFB48FA69" authority="Oliver &amp; Lutzen, 2011" authorityName="Oliver &amp; Lutzen" authorityYear="2011" box="[838,1272,1474,1500]" class="Bivalvia" family="Lasaeidae" genus="Draculamya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C54FFEC867321CEFC6CFA69" box="[838,988,1474,1500]" italics="true" pageId="12" pageNumber="13">Draculamya</emphasis>
<bibRefCitation id="B8584B137C54FFEC86D121CEFB48FA69" author="Oliver &amp; Lutzen" box="[996,1272,1474,1500]" firstAuthor="Oliver" pageId="12" pageNumber="13" pagination="365 - 392" refId="ref9894" refString="Oliver P. G. &amp; Lutzen J. 2011. An anatomically bizarre, fluid-feeding, galeommatoidean bivalve, Draculamya porobranchiata gen. et sp. nov. (Mollusca: Bivalvia). Journal of Conchology 40 (4): 365 - 392." type="journal article" year="2011">Oliver &amp; Lützen, 2011</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="B8584B137C54FFEC803D21CEFEE8FA4A" author="Oliver &amp; Lutzen" firstAuthor="Oliver" pageId="12" pageNumber="13" pagination="365 - 392" refId="ref9894" refString="Oliver P. G. &amp; Lutzen J. 2011. An anatomically bizarre, fluid-feeding, galeommatoidean bivalve, Draculamya porobranchiata gen. et sp. nov. (Mollusca: Bivalvia). Journal of Conchology 40 (4): 365 - 392." type="journal article" year="2011">Oliver &amp; Lützen 2011</bibRefCitation>
) none having a dentition identical to 
<taxonomicName id="1BC94D617C54FFEC861A21E9FC20FA4A" authorityName="Dall" authorityYear="1899" box="[815,912,1509,1535]" class="Bivalvia" family="Lasaeidae" genus="Kelliola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C54FFEC861A21E9FC20FA4A" box="[815,912,1509,1535]" italics="true" pageId="12" pageNumber="13">Kelliola</emphasis>
</taxonomicName>
or 
<taxonomicName id="1BC94D617C54FFEC868A21EAFBF1FA4A" authorityName="Turton" authorityYear="1822" box="[959,1089,1510,1535]" class="Bivalvia" family="Lasaeidae" genus="Montacuta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C54FFEC868A21EAFBF1FA4A" box="[959,1089,1510,1535]" italics="true" pageId="12" pageNumber="13">Montacuta</emphasis>
</taxonomicName>
. Among ten undescribed galeommatoids from the deep Atlantic, 
<bibRefCitation id="B8584B137C54FFEC87932205FCF2F996" author="Allen" box="[678,834,1545,1571]" firstAuthor="Allen" pageId="12" pageNumber="13" pagination="57 - 173" refId="ref8264" refString="Allen J. A. 2008. Bivalvia of the Deep Atlantic. Malacologia 50 (1): 57 - 173. http: // dx. doi. org / 10.4002 / 0076 - 2997 - 50.1.57" type="journal article" year="2008">Allen (2008)</bibRefCitation>
lists two undescribed 
<taxonomicName id="1BC94D617C54FFEC81692206FB6EF996" authorityName="Turton" authorityYear="1822" box="[1116,1246,1546,1571]" class="Bivalvia" family="Lasaeidae" genus="Montacuta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="EEBDEAF07C54FFEC81692206FB6EF996" box="[1116,1246,1546,1571]" italics="true" pageId="12" pageNumber="13">Montacuta</emphasis>
</taxonomicName>
species that may or may not be similar to 
<taxonomicName id="1BC94D617C54FFEC8715223CFD72F9FC" baseAuthorityName="Jeffreys" baseAuthorityYear="1876" box="[544,706,1584,1609]" class="Bivalvia" family="Lasaeidae" genus="Kelliola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galeommatida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="symmetros">
<emphasis id="EEBDEAF07C54FFEC8715223CFD72F9FC" box="[544,706,1584,1609]" italics="true" pageId="12" pageNumber="13">K. symmetros</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>