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<document id="484E8DA8BE74841BE84D84D5A4C13F24" ID-CLB-Dataset="5618" ID-DOI="10.11646/zootaxa.275.1.1" ID-GBIF-Dataset="9322f2b2-a384-40bf-9d57-d789c76ce5c4" ID-ISSN="1175­5334" ID-Zenodo-Dep="5019235" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624466010041" checkinUser="felipe" docAuthor="PRATHER, AYSHA L." docDate="2003" docId="546087A8FFF42A52FEA9F88AFD5DE762" docLanguage="en" docName="zt00275.pdf" docOrigin="Zootaxa 275 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.275.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Phylloicus yolandae Prather 2003, new species" docType="treatment" docVersion="8" lastPageNumber="99" masterDocId="A859FFD0FF952A30FFA1FF8CFC30E154" masterDocTitle="Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae)" masterLastPageNumber="214" masterPageNumber="1" pageNumber="98" updateTime="1699020181560" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8B5A1DB2BC3B75ADD8624805ABDF7769">Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae)</mods:title>
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<mods:namePart id="F1F167BAF19A7A78B5D97EE70DBBDA97">PRATHER, AYSHA L.</mods:namePart>
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<mods:title id="2845E7869FBC240D41E99F89F5484181">Zootaxa</mods:title>
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<mods:date id="BE124A3CD486022BEB529C56B3758E25">2003</mods:date>
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<mods:classification id="0DF257EADC853137BB5C18406BA5576B">journal article</mods:classification>
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<mods:identifier id="6A2789F721D70FE21A38ADAE2908B23E" type="DOI">10.11646/zootaxa.275.1.1</mods:identifier>
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<treatment id="546087A8FFF42A52FEA9F88AFD5DE762" ID-DOI="http://doi.org/10.5281/zenodo.5019255" ID-GBIF-Taxon="183402330" ID-Zenodo-Dep="5019255" LSID="urn:lsid:plazi:treatment:546087A8FFF42A52FEA9F88AFD5DE762" httpUri="http://treatment.plazi.org/id/546087A8FFF42A52FEA9F88AFD5DE762" lastPageId="98" lastPageNumber="99" pageId="97" pageNumber="98">
<subSubSection id="94D36535FFF42A51FEA9F88AFEAFE674" box="[264,671,1798,1824]" pageId="97" pageNumber="98" type="nomenclature">
<paragraph id="DC7636BEFFF42A51FEA9F88AFEAFE674" blockId="97.[264,671,1798,1859]" box="[264,671,1798,1824]" pageId="97" pageNumber="98">
<heading id="873E81D2FFF42A51FEA9F88AFEAFE674" bold="true" box="[264,671,1798,1824]" fontSize="11" level="1" pageId="97" pageNumber="98" reason="1">
<emphasis id="EEBDEAACFFF42A51FEA9F88AFEAFE674" bold="true" box="[264,671,1798,1824]" pageId="97" pageNumber="98">
<taxonomicName id="1BC94D3DFFF42A51FEA9F88AFE32E674" authority="Prather, 2003" authorityName="Prather " authorityYear="2003" box="[264,514,1798,1824]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="yolandae" status="sp. nov.">Phylloicus yolandae</taxonomicName>
,
<taxonomicNameLabel id="F58E57D7FFF42A51FDAEF88AFEAFE674" box="[527,671,1798,1824]" pageId="97" pageNumber="98" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="94D36535FFF42A52FEA9F8A5FE28E43A" lastPageId="98" lastPageNumber="99" pageId="97" pageNumber="98" type="description">
<paragraph id="DC7636BEFFF42A51FEA9F8A5FD9FE617" blockId="97.[264,671,1798,1859]" box="[264,431,1833,1859]" pageId="97" pageNumber="98">
<figureCitation id="44F22A3BFFF42A51FEA9F8A5FD47E617" box="[264,375,1833,1859]" captionStart="FIGURE 104" captionStartId="207.[264,368,1642,1666]" captionTargetBox="[266,1320,328,1617]" captionTargetId="figure-41@207.[266,1321,326,1617]" captionTargetPageId="207" captionText="FIGURE 104. Phylloicus yolandae. Male (holotype): A—lateral view; B—dorsal view; C—sternum IX and inferior appendages, ventral view; D—phallus, lateral view; E—phallus, caudoventral view." figureDoi="http://doi.org/10.5281/zenodo.5019494" httpUri="https://zenodo.org/record/5019494/files/figure.png" pageId="97" pageNumber="98">Figs. 104</figureCitation>
,
<figureCitation id="44F22A3BFFF42A51FE25F8A5FD9FE617" box="[388,431,1833,1859]" captionStart="FIGURES 113-116. — 113" captionStartId="210.[264,382,1773,1797]" captionTargetBox="[269,1319,314,1749]" captionTargetId="figure-149@210.[490,1318,675,1042]" captionTargetPageId="210" captionText="FIGURES 113-116. — 113. Phylloicus paucartambo (composite of UMSP000010031-8). Forewing. — 114. Phylloicus pulchrus (composite of UMSP000009766 and UMSP000068395). Forewing. — 115. Phylloicus superbus (UMSP000009816 and UMSP000009824). Forewing. — 116. Phylloicus yolandae (holotype). Forewing." figureDoi="http://doi.org/10.5281/zenodo.5019500" httpUri="https://zenodo.org/record/5019500/files/figure.png" pageId="97" pageNumber="98">116</figureCitation>
</paragraph>
<paragraph id="DC7636BEFFF72A52FEA9FE90FF2DE0FA" blockId="98.[264,1323,284,1590]" pageId="98" pageNumber="99">
<taxonomicName id="1BC94D3DFFF72A52FEA9FE90FDC5E061" authorityName="Prather Table Of Contents" authorityYear="2003" box="[264,501,284,309]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="yolandae">
<emphasis id="EEBDEAACFFF72A52FEA9FE90FDC5E061" box="[264,501,284,309]" italics="true" pageId="98" pageNumber="99">Phylloicus yolandae</emphasis>
</taxonomicName>
is one of the prettiest and most unusual species of
<taxonomicName id="1BC94D3DFFF72A52FBE4FE90F8F1E061" box="[1093,1217,284,309]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis id="EEBDEAACFFF72A52FBE4FE90F8F1E061" box="[1093,1217,284,309]" italics="true" pageId="98" pageNumber="99">Phylloicus</emphasis>
</taxonomicName>
. With its partially clear forewings and dramatic coloration (
<figureCitation id="44F22A3BFFF72A52FCFCFEC8FFF4E00A" box="[861,964,324,350]" captionStart="FIGURES 113-116. — 113" captionStartId="210.[264,382,1773,1797]" captionTargetBox="[269,1319,314,1749]" captionTargetId="figure-149@210.[490,1318,675,1042]" captionTargetPageId="210" captionText="FIGURES 113-116. — 113. Phylloicus paucartambo (composite of UMSP000010031-8). Forewing. — 114. Phylloicus pulchrus (composite of UMSP000009766 and UMSP000068395). Forewing. — 115. Phylloicus superbus (UMSP000009816 and UMSP000009824). Forewing. — 116. Phylloicus yolandae (holotype). Forewing." figureDoi="http://doi.org/10.5281/zenodo.5019500" httpUri="https://zenodo.org/record/5019500/files/figure.png" pageId="98" pageNumber="99">Fig. 116</figureCitation>
), this species resembles clearwinged moth species found in the region. The male genitalia are recognized by the cordate tergum X and the digitate preanal appendage.
</paragraph>
<paragraph id="DC7636BEFFF72A52FE99FE30FECDE082" blockId="98.[264,1323,284,1590]" box="[312,765,444,470]" pageId="98" pageNumber="99">Adult. Forewing length 8.1 mm, n = 1.</paragraph>
<paragraph id="DC7636BEFFF72A52FE99FE68F88FE242" blockId="98.[264,1323,284,1590]" pageId="98" pageNumber="99">
Head dark brown or black. Maxillary palps dark brown or black. Antenna twice forewing length; dark brown, with narrow patches of pale sensilla on anteromesal surface of each flagellomere. Dorsal pterothorax dark brown or black; ventrolateral thorax dark brown. Legs dark brown. Metathoracic leg of male without posterior fringe. Tibial spur formula 2,4,4. Forewing flat; dark brown, or black; with two transverse bands; proximal band ivory, reaching posterior wing margin, at least 1/2 width of wing, interrupted by clear patch (
<figureCitation id="44F22A3BFFF72A52FEF8FD58FD8FE3BA" box="[345,447,724,750]" captionStart="FIGURES 113-116. — 113" captionStartId="210.[264,382,1773,1797]" captionTargetBox="[269,1319,314,1749]" captionTargetId="figure-149@210.[490,1318,675,1042]" captionTargetPageId="210" captionText="FIGURES 113-116. — 113. Phylloicus paucartambo (composite of UMSP000010031-8). Forewing. — 114. Phylloicus pulchrus (composite of UMSP000009766 and UMSP000068395). Forewing. — 115. Phylloicus superbus (UMSP000009816 and UMSP000009824). Forewing. — 116. Phylloicus yolandae (holotype). Forewing." figureDoi="http://doi.org/10.5281/zenodo.5019500" httpUri="https://zenodo.org/record/5019500/files/figure.png" pageId="98" pageNumber="99">Fig. 116</figureCitation>
); distal band ivory, not reaching either wing margin, 1/2 width of wing or less; with single basal stripe, ivory; large clear patch in center of wing (
<figureCitation id="44F22A3BFFF72A52FBEDFD70F882E242" box="[1100,1202,764,790]" captionStart="FIGURES 113-116. — 113" captionStartId="210.[264,382,1773,1797]" captionTargetBox="[269,1319,314,1749]" captionTargetId="figure-149@210.[490,1318,675,1042]" captionTargetPageId="210" captionText="FIGURES 113-116. — 113. Phylloicus paucartambo (composite of UMSP000010031-8). Forewing. — 114. Phylloicus pulchrus (composite of UMSP000009766 and UMSP000068395). Forewing. — 115. Phylloicus superbus (UMSP000009816 and UMSP000009824). Forewing. — 116. Phylloicus yolandae (holotype). Forewing." figureDoi="http://doi.org/10.5281/zenodo.5019500" httpUri="https://zenodo.org/record/5019500/files/figure.png" pageId="98" pageNumber="99">Fig. 116</figureCitation>
).
</paragraph>
<paragraph id="DC7636BEFFF72A52FE99FCA8FEA2E412" blockId="98.[264,1323,284,1590]" pageId="98" pageNumber="99">
Male. Preterminalic abdominal terga without anteromesal notches. Corematic structures absent, terga III­V unmodified, without membranous lobes or sclerotized processes. Sternum VIII similar to anterior sterna, sternum IX not elongate. Tergum IX without mesal ridge; posterior margin notched (
<figureCitation id="44F22A3BFFF72A52FD3FFC10FF2DE2E2" box="[670,797,924,950]" captionStart="FIGURE 104" captionStartId="207.[264,368,1642,1666]" captionTargetBox="[266,1320,328,1617]" captionTargetId="figure-41@207.[266,1321,326,1617]" captionTargetPageId="207" captionText="FIGURE 104. Phylloicus yolandae. Male (holotype): A—lateral view; B—dorsal view; C—sternum IX and inferior appendages, ventral view; D—phallus, lateral view; E—phallus, caudoventral view." figureDoi="http://doi.org/10.5281/zenodo.5019494" httpUri="https://zenodo.org/record/5019494/files/figure.png" pageId="98" pageNumber="99">Fig. 104B</figureCitation>
); lateral ridge present; dorsal pleural setae approximately 8, ventral pleural setae approximately 2 (
<figureCitation id="44F22A3BFFF72A52FC0BFC48F818E28A" box="[938,1064,964,990]" captionStart="FIGURE 104" captionStartId="207.[264,368,1642,1666]" captionTargetBox="[266,1320,328,1617]" captionTargetId="figure-41@207.[266,1321,326,1617]" captionTargetPageId="207" captionText="FIGURE 104. Phylloicus yolandae. Male (holotype): A—lateral view; B—dorsal view; C—sternum IX and inferior appendages, ventral view; D—phallus, lateral view; E—phallus, caudoventral view." figureDoi="http://doi.org/10.5281/zenodo.5019494" httpUri="https://zenodo.org/record/5019494/files/figure.png" pageId="98" pageNumber="99">Fig. 104A</figureCitation>
); sternum IX without mesolateral ridges; sternum IX (
<figureCitation id="44F22A3BFFF72A52FD23FC60FECDE552" box="[642,765,1004,1030]" captionStart="FIGURE 104" captionStartId="207.[264,368,1642,1666]" captionTargetBox="[266,1320,328,1617]" captionTargetId="figure-41@207.[266,1321,326,1617]" captionTargetPageId="207" captionText="FIGURE 104. Phylloicus yolandae. Male (holotype): A—lateral view; B—dorsal view; C—sternum IX and inferior appendages, ventral view; D—phallus, lateral view; E—phallus, caudoventral view." figureDoi="http://doi.org/10.5281/zenodo.5019494" httpUri="https://zenodo.org/record/5019494/files/figure.png" pageId="98" pageNumber="99">Fig. 104C</figureCitation>
). Preanal appendage shorter than tergum X, but greater than 2/3 length, narrowly elliptic, setae long, but not filamentous or longer than appendage (
<figureCitation id="44F22A3BFFF72A52FE3AFBB0FE73E502" box="[411,579,1084,1110]" captionStart="FIGURE 104" captionStartId="207.[264,368,1642,1666]" captionTargetBox="[266,1320,328,1617]" captionTargetId="figure-41@207.[266,1321,326,1617]" captionTargetPageId="207" captionText="FIGURE 104. Phylloicus yolandae. Male (holotype): A—lateral view; B—dorsal view; C—sternum IX and inferior appendages, ventral view; D—phallus, lateral view; E—phallus, caudoventral view." figureDoi="http://doi.org/10.5281/zenodo.5019494" httpUri="https://zenodo.org/record/5019494/files/figure.png" pageId="98" pageNumber="99">Fig. 104A, B</figureCitation>
). Tergum X without basal lobes; basodorsal process absent; basolateral processes absent; apex, in lateral view, acute, in dorsal view, notched, notch deep and round; setose basodorsally; broad basally, constricted at mid­length (
<figureCitation id="44F22A3BFFF72A52FB0BFB00FD2EE59A" captionStart="FIGURE 104" captionStartId="207.[264,368,1642,1666]" captionTargetBox="[266,1320,328,1617]" captionTargetId="figure-41@207.[266,1321,326,1617]" captionTargetPageId="207" captionText="FIGURE 104. Phylloicus yolandae. Male (holotype): A—lateral view; B—dorsal view; C—sternum IX and inferior appendages, ventral view; D—phallus, lateral view; E—phallus, caudoventral view." figureDoi="http://doi.org/10.5281/zenodo.5019494" httpUri="https://zenodo.org/record/5019494/files/figure.png" pageId="98" pageNumber="99">Fig. 104A, B</figureCitation>
). Harpago sharply tapered; peglike setae few, apical (
<figureCitation id="44F22A3BFFF72A52FC00FB38F872E59A" box="[929,1090,1204,1230]" captionStart="FIGURE 104" captionStartId="207.[264,368,1642,1666]" captionTargetBox="[266,1320,328,1617]" captionTargetId="figure-41@207.[266,1321,326,1617]" captionTargetPageId="207" captionText="FIGURE 104. Phylloicus yolandae. Male (holotype): A—lateral view; B—dorsal view; C—sternum IX and inferior appendages, ventral view; D—phallus, lateral view; E—phallus, caudoventral view." figureDoi="http://doi.org/10.5281/zenodo.5019494" httpUri="https://zenodo.org/record/5019494/files/figure.png" pageId="98" pageNumber="99">Fig. 104A, C</figureCitation>
). Phallic endotheca with paired apicolateral lobes, apicolateral lobes large and rounded; phallotremal sclerites small, longest dimension 1/2 diameter of phallobase; dorsal sclerite ovoid, in dorsal view horseshoe­shaped (
<figureCitation id="44F22A3BFFF72A52FE49FAA0FEB4E412" box="[488,644,1324,1350]" captionStart="FIGURE 104" captionStartId="207.[264,368,1642,1666]" captionTargetBox="[266,1320,328,1617]" captionTargetId="figure-41@207.[266,1321,326,1617]" captionTargetPageId="207" captionText="FIGURE 104. Phylloicus yolandae. Male (holotype): A—lateral view; B—dorsal view; C—sternum IX and inferior appendages, ventral view; D—phallus, lateral view; E—phallus, caudoventral view." figureDoi="http://doi.org/10.5281/zenodo.5019494" httpUri="https://zenodo.org/record/5019494/files/figure.png" pageId="98" pageNumber="99">Fig. 104D, E</figureCitation>
).
</paragraph>
<paragraph id="DC7636BEFFF72A52FE99FAD8FE28E43A" blockId="98.[264,1323,284,1590]" box="[312,536,1364,1390]" pageId="98" pageNumber="99">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="94D36535FFF72A52FE99FAF0FD5DE762" pageId="98" pageNumber="99" type="materials_examined">
<paragraph id="DC7636BEFFF72A52FE99FAF0F88AE4EA" blockId="98.[264,1323,284,1590]" pageId="98" pageNumber="99">
<materialsCitation id="6CA13CE3FFF72A52FE99FAF0F88AE4EA" ID-GBIF-Occurrence="3319301097" collectingDate="1998-04-04" collectingDateMax="1998-04-07" collectingDateMin="1998-04-04" collectionCode="MZUSP" collectorName="Holzenthal &amp; Huisman" country="Brazil" elevation="700" latitude="-24.898056" location="Rio Novo" longLatPrecision="21" longitude="-53.248333" municipality="Municipio Corbelia" pageId="98" pageNumber="99" specimenCount="1" stateProvince="Parana" typeStatus="holotype">
<emphasis id="EEBDEAACFFF72A52FE99FAF0FECBE4C2" bold="true" box="[312,763,1404,1430]" pageId="98" pageNumber="99">
<typeStatus id="0372881CFFF72A52FE99FAF0FD9AE4C2" box="[312,426,1404,1430]" pageId="98" pageNumber="99" type="holotype">Holotype</typeStatus>
male:
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:
<collectingRegion id="1E0DF85CFFF72A52FD37FAF0FEC5E4C2" box="[662,757,1404,1430]" country="Brazil" name="Parana" pageId="98" pageNumber="99">Paraná</collectingRegion>
:
</emphasis>
<collectingMunicipality id="3C12ACC4FFF72A52FCABFAF0FFCBE4C2" box="[778,1019,1404,1430]" pageId="98" pageNumber="99">Municipio Corbélia</collectingMunicipality>
,
<location id="D9166065FFF72A52FBAFFAF0F8BCE4C2" LSID="urn:lsid:plazi:treatment:546087A8FFF42A52FEA9F88AFD5DE762:D9166065FFF72A52FBAFFAF0F8BCE4C2" box="[1038,1164,1404,1430]" country="Brazil" latitude="-24.898056" longLatPrecision="21" longitude="-53.248333" municipality="Municipio Corbelia" name="Rio Novo" pageId="98" pageNumber="99" stateProvince="Parana">Rio Novo</location>
, headwaters,
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,
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,
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</quantity>
,
<date id="A877107EFFF72A52FD33FA28FF29E4EA" box="[658,793,1444,1470]" pageId="98" pageNumber="99" value="1998-04-04" valueMax="1998-04-07" valueMin="1998-04-04">
<collectingDate id="B833E996FFF72A52FD33FA28FF29E4EA" box="[658,793,1444,1470]" pageId="98" pageNumber="99" value="1998-04-04" valueMax="1998-04-07" valueMin="1998-04-04">4­7.iv.1998</collectingDate>
</date>
,
<collectorName id="713C5368FFF72A52FC87FA28FF96E4EA" box="[806,934,1444,1470]" pageId="98" pageNumber="99">Holzenthal</collectorName>
&amp;
<collectorName id="713C5368FFF72A52FC6DFA28F805E4EA" box="[972,1077,1444,1470]" pageId="98" pageNumber="99">Huisman</collectorName>
(
<collectionCode id="BAD8AE7BFFF72A52FBE4FA28F89DE4EA" box="[1093,1197,1444,1470]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="98" pageNumber="99">MZUSP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="DC7636BEFFF72A52FE99FA40FE13E4B2" blockId="98.[264,1323,284,1590]" box="[312,547,1484,1510]" pageId="98" pageNumber="99">
Distribution.
<collectingCountry id="A4DE762EFFF72A52FE75FA40FE2FE4B2" box="[468,543,1484,1510]" name="Brazil" pageId="98" pageNumber="99">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="DC7636BEFFF72A52FE99FA78FD5DE762" blockId="98.[264,1323,284,1590]" pageId="98" pageNumber="99">Etymology. I am very pleased to name this species for Jolanda Huisman, who collected it.</paragraph>
</subSubSection>
</treatment>
</document>
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