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<document id="2CAB550FDD1F5484879558913EB0326F" ID-CLB-Dataset="5618" ID-DOI="10.11646/zootaxa.275.1.1" ID-GBIF-Dataset="9322f2b2-a384-40bf-9d57-d789c76ce5c4" ID-ISSN="1175­5334" ID-Zenodo-Dep="5019235" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624466010041" checkinUser="felipe" docAuthor="PRATHER, AYSHA L." docDate="2003" docId="546087A8FFF42A51FEA9FEE7FF7EE7FD" docLanguage="en" docName="zt00275.pdf" docOrigin="Zootaxa 275 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.275.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Phylloicus trichothylax Prather 2003, new species" docType="treatment" docVersion="9" lastPageNumber="98" masterDocId="A859FFD0FF952A30FFA1FF8CFC30E154" masterDocTitle="Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae)" masterLastPageNumber="214" masterPageNumber="1" pageNumber="98" updateTime="1699020181560" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="7F432E6FC8B822BB7E25B84E8C069E6A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5E876A46F3162AE9696DF5A5A31092E5">
<mods:title id="0715F40945CBA55AE6C7B0C0D3627E51">Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="B0739F326E49F282AA602326DB23D822">PRATHER, AYSHA L.</mods:namePart>
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<mods:title id="5C8E0CA72B8D34580B33074F5B63492E">Zootaxa</mods:title>
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<mods:date id="7849254D8716E2EFE0AEBBA1AE0213E6">2003</mods:date>
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<mods:number id="50467495C96301E1E2F60D8FEF5856DE">2003-08-29</mods:number>
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<mods:number id="BD53457268FB561B3A1E8DF6066CC86F">275</mods:number>
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</mods:location>
<mods:classification id="F06A9A540089AFECFC8F8B34D43966D3">journal article</mods:classification>
<mods:identifier id="006E2CB18CFCE33AA4F23A1C2B7F792C" type="CLB-Dataset">5618</mods:identifier>
<mods:identifier id="82AF2E5C054CF722C3357B8A3A13985D" type="DOI">10.11646/zootaxa.275.1.1</mods:identifier>
<mods:identifier id="B52894E86A09535A690DD40CF7FCD2C4" type="GBIF-Dataset">9322f2b2-a384-40bf-9d57-d789c76ce5c4</mods:identifier>
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<treatment id="546087A8FFF42A51FEA9FEE7FF7EE7FD" ID-DOI="http://doi.org/10.5281/zenodo.5081061" ID-GBIF-Taxon="183402329" ID-Zenodo-Dep="5081061" LSID="urn:lsid:plazi:treatment:546087A8FFF42A51FEA9FEE7FF7EE7FD" httpUri="http://treatment.plazi.org/id/546087A8FFF42A51FEA9FEE7FF7EE7FD" lastPageNumber="98" pageId="97" pageNumber="98">
<subSubSection id="94D36535FFF42A51FEA9FEE7FEF7E0D1" box="[264,711,363,389]" pageId="97" pageNumber="98" type="nomenclature">
<paragraph id="DC7636BEFFF42A51FEA9FEE7FEF7E0D1" blockId="97.[264,711,363,425]" box="[264,711,363,389]" pageId="97" pageNumber="98">
<heading id="873E81D2FFF42A51FEA9FEE7FEF7E0D1" bold="true" box="[264,711,363,389]" fontSize="11" level="1" pageId="97" pageNumber="98" reason="1">
<emphasis id="EEBDEAACFFF42A51FEA9FEE7FEF7E0D1" bold="true" box="[264,711,363,389]" pageId="97" pageNumber="98">
<taxonomicName id="1BC94D3DFFF42A51FEA9FEE7FE1BE0D1" authority="Prather, 2003" authorityName="Prather" authorityYear="2003" box="[264,555,363,389]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="trichothylax" status="sp. nov.">Phylloicus trichothylax</taxonomicName>
,
<taxonomicNameLabel id="F58E57D7FFF42A51FD96FEE7FEF7E0D1" box="[567,711,363,389]" pageId="97" pageNumber="98" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="94D36535FFF42A51FEA9FE03FE28E4B5" pageId="97" pageNumber="98" type="description">
<paragraph id="DC7636BEFFF42A51FEA9FE03FD5AE0FD" blockId="97.[264,711,363,425]" box="[264,362,399,425]" pageId="97" pageNumber="98">
<figureCitation id="44F22A3BFFF42A51FEA9FE03FD5AE0FD" box="[264,362,399,425]" captionStart="FIGURE 103" captionStartId="206.[264,368,1495,1519]" captionTargetBox="[264,1322,333,1471]" captionTargetId="figure-40@206.[262,1324,326,1471]" captionTargetPageId="206" captionText="FIGURE 103. Phylloicus trichothylax. Male (holotype): A—lateral view; B—dorsal view; C— ventral view; D—phallus, lateral view; F—tergum IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019492" httpUri="https://zenodo.org/record/5019492/files/figure.png" pageId="97" pageNumber="98">Fig. 103</figureCitation>
</paragraph>
<paragraph id="DC7636BEFFF42A51FEA9FE53F8D5E3CD" blockId="97.[264,1323,479,1705]" pageId="97" pageNumber="98">
<taxonomicName id="1BC94D3DFFF42A51FEA9FE53FE28E0AC" authorityName="Prather Table Of Contents" authorityYear="2003" box="[264,536,479,504]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="trichothylax">
<emphasis id="EEBDEAACFFF42A51FEA9FE53FE28E0AC" box="[264,536,479,504]" italics="true" pageId="97" pageNumber="98">Phylloicus trichothylax</emphasis>
</taxonomicName>
is known only from a single teneral male, and thus details of coloration are difficult to identify and the forewings are too twisted for accurate measurement. It is distinguished by the abdominal coremata, which consist of a pair of invaginated membranous pouches containing long setae and a single, eversible, digitate membranous lobe (
<figureCitation id="44F22A3BFFF42A51FEB0FDF3FDB7E3CD" box="[273,391,639,665]" captionStart="FIGURE 103" captionStartId="206.[264,368,1495,1519]" captionTargetBox="[264,1322,333,1471]" captionTargetId="figure-40@206.[262,1324,326,1471]" captionTargetPageId="206" captionText="FIGURE 103. Phylloicus trichothylax. Male (holotype): A—lateral view; B—dorsal view; C— ventral view; D—phallus, lateral view; F—tergum IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019492" httpUri="https://zenodo.org/record/5019492/files/figure.png" pageId="97" pageNumber="98">Fig. 103F</figureCitation>
), and by the long digitate basodorsal process of tergum X (
<figureCitation id="44F22A3BFFF42A51FB9AFDF3F8E7E3CD" box="[1083,1239,639,665]" captionStart="FIGURE 103" captionStartId="206.[264,368,1495,1519]" captionTargetBox="[264,1322,333,1471]" captionTargetId="figure-40@206.[262,1324,326,1471]" captionTargetPageId="206" captionText="FIGURE 103. Phylloicus trichothylax. Male (holotype): A—lateral view; B—dorsal view; C— ventral view; D—phallus, lateral view; F—tergum IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019492" httpUri="https://zenodo.org/record/5019492/files/figure.png" pageId="97" pageNumber="98">Fig. 103A, B</figureCitation>
).
</paragraph>
<paragraph id="DC7636BEFFF42A51FE99FD2BFD79E235" blockId="97.[264,1323,479,1705]" pageId="97" pageNumber="98">Adult. Head golden brown, with dorsomesal crest of golden brown setae. Maxillary palps dark brown. Antenna golden brown. Dorsal pterothorax golden brown; ventrolateral thorax golden brown. Legs golden brown. Metathoracic leg of male with posterior fringe of long setae. Tibial spur formula 2,4,4. Forewing flat. Hind wing basal brush present in male.</paragraph>
<paragraph id="DC7636BEFFF42A51FE99FCE3FEE3E4ED" blockId="97.[264,1323,479,1705]" pageId="97" pageNumber="98">
Male. Preterminalic abdominal terga with anteromesal notch. Corematic structures present. Tergum IV with expanded lateral flanges, mesal coremata; mesal coremata an invaginated setose pouch, enclosing small, digitate eversible lobe. Tergum V without sclerotized modifications (
<figureCitation id="44F22A3BFFF42A51FDB0FC6BFEB8E555" box="[529,648,999,1025]" captionStart="FIGURE 103" captionStartId="206.[264,368,1495,1519]" captionTargetBox="[264,1322,333,1471]" captionTargetId="figure-40@206.[262,1324,326,1471]" captionTargetPageId="206" captionText="FIGURE 103. Phylloicus trichothylax. Male (holotype): A—lateral view; B—dorsal view; C— ventral view; D—phallus, lateral view; F—tergum IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019492" httpUri="https://zenodo.org/record/5019492/files/figure.png" pageId="97" pageNumber="98">Fig. 103F</figureCitation>
). Sternum VII without anteromesal process. Sternum VIII similar to anterior sterna, sternum IX not elongate. Tergum IX with paired mesal ridges extending length of segment; posterior margin smoothly rounded; thinly sclerotized anteromesally (
<figureCitation id="44F22A3BFFF42A51FE25FBD3FE30E52D" box="[388,512,1119,1145]" captionStart="FIGURE 103" captionStartId="206.[264,368,1495,1519]" captionTargetBox="[264,1322,333,1471]" captionTargetId="figure-40@206.[262,1324,326,1471]" captionTargetPageId="206" captionText="FIGURE 103. Phylloicus trichothylax. Male (holotype): A—lateral view; B—dorsal view; C— ventral view; D—phallus, lateral view; F—tergum IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019492" httpUri="https://zenodo.org/record/5019492/files/figure.png" pageId="97" pageNumber="98">Fig. 103B</figureCitation>
); lateral ridge present; dorsal pleural setae approximately 8, ventral pleural setae approximately 6 (
<figureCitation id="44F22A3BFFF42A51FDDFFB0BFECDE5F5" box="[638,765,1159,1185]" captionStart="FIGURE 103" captionStartId="206.[264,368,1495,1519]" captionTargetBox="[264,1322,333,1471]" captionTargetId="figure-40@206.[262,1324,326,1471]" captionTargetPageId="206" captionText="FIGURE 103. Phylloicus trichothylax. Male (holotype): A—lateral view; B—dorsal view; C— ventral view; D—phallus, lateral view; F—tergum IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019492" httpUri="https://zenodo.org/record/5019492/files/figure.png" pageId="97" pageNumber="98">Fig. 103A</figureCitation>
); sternum IX without mesolateral ridges; sternum IX (
<figureCitation id="44F22A3BFFF42A51FED0FB23FDDBE59D" box="[369,491,1199,1225]" captionStart="FIGURE 103" captionStartId="206.[264,368,1495,1519]" captionTargetBox="[264,1322,333,1471]" captionTargetId="figure-40@206.[262,1324,326,1471]" captionTargetPageId="206" captionText="FIGURE 103. Phylloicus trichothylax. Male (holotype): A—lateral view; B—dorsal view; C— ventral view; D—phallus, lateral view; F—tergum IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019492" httpUri="https://zenodo.org/record/5019492/files/figure.png" pageId="97" pageNumber="98">Fig. 103C</figureCitation>
). Preanal appendage shorter than tergum X, but greater than 2/3 length, narrowly elliptic, setae long, but not filamentous or longer than appendage (
<figureCitation id="44F22A3BFFF42A51FB21FB5BF92CE5A5" box="[1152,1308,1239,1265]" captionStart="FIGURE 103" captionStartId="206.[264,368,1495,1519]" captionTargetBox="[264,1322,333,1471]" captionTargetId="figure-40@206.[262,1324,326,1471]" captionTargetPageId="206" captionText="FIGURE 103. Phylloicus trichothylax. Male (holotype): A—lateral view; B—dorsal view; C— ventral view; D—phallus, lateral view; F—tergum IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019492" httpUri="https://zenodo.org/record/5019492/files/figure.png" pageId="97" pageNumber="98">Fig. 103A, B</figureCitation>
). Tergum X without basal lobes; basodorsal process long and digitate; basolateral processes absent; apex, in lateral view, truncate, in dorsal view, entire (
<figureCitation id="44F22A3BFFF42A51FBA2FAABF89AE415" box="[1027,1194,1319,1345]" captionStart="FIGURE 103" captionStartId="206.[264,368,1495,1519]" captionTargetBox="[264,1322,333,1471]" captionTargetId="figure-40@206.[262,1324,326,1471]" captionTargetPageId="206" captionText="FIGURE 103. Phylloicus trichothylax. Male (holotype): A—lateral view; B—dorsal view; C— ventral view; D—phallus, lateral view; F—tergum IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019492" httpUri="https://zenodo.org/record/5019492/files/figure.png" pageId="97" pageNumber="98">Fig. 103A, B</figureCitation>
). Harpago slightly tapered; peglike setae few, apical (
<figureCitation id="44F22A3BFFF42A51FCAAFAC3FF9EE43D" box="[779,942,1359,1385]" captionStart="FIGURE 103" captionStartId="206.[264,368,1495,1519]" captionTargetBox="[264,1322,333,1471]" captionTargetId="figure-40@206.[262,1324,326,1471]" captionTargetPageId="206" captionText="FIGURE 103. Phylloicus trichothylax. Male (holotype): A—lateral view; B—dorsal view; C— ventral view; D—phallus, lateral view; F—tergum IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019492" httpUri="https://zenodo.org/record/5019492/files/figure.png" pageId="97" pageNumber="98">Fig. 103A, C</figureCitation>
). Phallotremal sclerites average size, longest dimension less than diameter of phallobase; dorsal sclerite ovoid, in dorsal view horseshoe­shaped (
<figureCitation id="44F22A3BFFF42A51FD88FA13FEF5E4ED" box="[553,709,1439,1465]" captionStart="FIGURE 103" captionStartId="206.[264,368,1495,1519]" captionTargetBox="[264,1322,333,1471]" captionTargetId="figure-40@206.[262,1324,326,1471]" captionTargetPageId="206" captionText="FIGURE 103. Phylloicus trichothylax. Male (holotype): A—lateral view; B—dorsal view; C— ventral view; D—phallus, lateral view; F—tergum IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019492" httpUri="https://zenodo.org/record/5019492/files/figure.png" pageId="97" pageNumber="98">Fig. 103D, E</figureCitation>
).
</paragraph>
<paragraph id="DC7636BEFFF42A51FE99FA4BFE28E4B5" blockId="97.[264,1323,479,1705]" box="[312,536,1479,1505]" pageId="97" pageNumber="98">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="94D36535FFF42A51FE99FA62FF7EE7FD" pageId="97" pageNumber="98" type="materials_examined">
<paragraph id="DC7636BEFFF42A51FE99FA62FE65E765" blockId="97.[264,1323,479,1705]" pageId="97" pageNumber="98">
<materialsCitation id="6CA13CE3FFF42A51FE99FA62FE65E765" ID-GBIF-Occurrence="3319300453" collectingDate="1975-07-01" collectionCode="NMNH" collectorName="Langley &amp; Cohen" country="Ecuador" elevation="1372" location="Latacunga" pageId="97" pageNumber="98" specimenCount="1" stateProvince="Cotopaxi" typeStatus="holotype">
<emphasis id="EEBDEAACFFF42A51FE99FA62FF17E75C" bold="true" box="[312,807,1518,1544]" pageId="97" pageNumber="98">
<typeStatus id="0372881CFFF42A51FE99FA62FD9AE75C" box="[312,426,1518,1544]" pageId="97" pageNumber="98" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="A4DE762EFFF42A51FDA5FA62FEADE75C" box="[516,669,1518,1544]" name="Ecuador" pageId="97" pageNumber="98">ECUADOR</collectingCountry>
:
<collectingRegion id="1E0DF85CFFF42A51FD0DFA62FF12E75C" box="[684,802,1518,1544]" country="Ecuador" name="Cotopaxi" pageId="97" pageNumber="98">Cotopaxi</collectingRegion>
:
</emphasis>
<location id="D9166065FFF42A51FC93FA63FF9FE75D" LSID="urn:lsid:plazi:treatment:546087A8FFF42A51FEA9FEE7FF7EE7FD:D9166065FFF42A51FC93FA63FF9FE75D" box="[818,943,1519,1545]" country="Ecuador" name="Latacunga" pageId="97" pageNumber="98" stateProvince="Cotopaxi">Latacunga</location>
,
<quantity id="1B319B5BFFF42A51FC1CFA63F83AE75D" box="[957,1034,1519,1545]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="97" pageNumber="98" unit="km" value="13.0">13 km</quantity>
W,
<quantity id="1B319B5BFFF42A51FB9EFA63F8AEE75D" box="[1087,1182,1519,1545]" metricMagnitude="3" metricUnit="m" metricValue="1.3719999999999999" pageId="97" pageNumber="98" unit="m" value="1372.0">
<elevation id="57E4D18DFFF42A51FB9EFA63F8AEE75D" box="[1087,1182,1519,1545]" metricMagnitude="3" metricUnit="m" metricValue="1.3719999999999999" pageId="97" pageNumber="98" unit="m" value="1372.0">1372 m</elevation>
</quantity>
,
<date id="A877107EFFF42A51FB0DFA63F915E75D" box="[1196,1317,1519,1545]" pageId="97" pageNumber="98" value="1975-07-01">
<collectingDate id="B833E996FFF42A51FB0DFA63F915E75D" box="[1196,1317,1519,1545]" pageId="97" pageNumber="98" value="1975-07-01">1.vii.1975</collectingDate>
</date>
,
<collectorName id="713C5368FFF42A51FEA9F99BFD58E765" box="[264,360,1559,1585]" pageId="97" pageNumber="98">Langley</collectorName>
&amp;
<collectorName id="713C5368FFF42A51FE2CF99BFDE9E765" box="[397,473,1559,1585]" pageId="97" pageNumber="98">Cohen</collectorName>
(
<collectionCode id="BAD8AE7BFFF42A51FE48F99BFE79E765" box="[489,585,1559,1585]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="97" pageNumber="98" type="Museum">NMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="DC7636BEFFF42A51FE99F9B3FE0BE70D" blockId="97.[264,1323,479,1705]" box="[312,571,1599,1625]" pageId="97" pageNumber="98">
Distribution.
<collectingCountry id="A4DE762EFFF42A51FE75F9B3FE08E70D" box="[468,568,1599,1625]" name="Ecuador" pageId="97" pageNumber="98">Ecuador</collectingCountry>
.
</paragraph>
<paragraph id="DC7636BEFFF42A51FE99F9EBFF7EE7FD" blockId="97.[264,1323,479,1705]" pageId="97" pageNumber="98">
Etymology.
<emphasis id="EEBDEAACFFF42A51FE69F9EBFE6CE7D4" box="[456,604,1639,1664]" italics="true" pageId="97" pageNumber="98">Trichothylax</emphasis>
, from the Greek
<emphasis id="EEBDEAACFFF42A51FC8DF9EBFF4DE7D4" box="[812,893,1639,1664]" italics="true" pageId="97" pageNumber="98">trichos</emphasis>
, “hair” and
<emphasis id="EEBDEAACFFF42A51FBB3F9EBF86AE7D4" box="[1042,1114,1639,1664]" italics="true" pageId="97" pageNumber="98">thylax</emphasis>
, “pouch,” for the setae­enclosing membranous pouch of tergum IV.
</paragraph>
</subSubSection>
</treatment>
</document>
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