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<document id="F353192F8303781A33220553A9BFB4BE" ID-CLB-Dataset="5618" ID-DOI="10.11646/zootaxa.275.1.1" ID-GBIF-Dataset="9322f2b2-a384-40bf-9d57-d789c76ce5c4" ID-ISSN="1175­5334" ID-Zenodo-Dep="5019235" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624466010041" checkinUser="felipe" docAuthor="PRATHER, AYSHA L." docDate="2003" docId="546087A8FFC82A6EFEA9FD08FE22E392" docLanguage="en" docName="zt00275.pdf" docOrigin="Zootaxa 275 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.275.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Phylloicus spectabilis Martynov Segment IX 1912" docType="treatment" docVersion="7" lastPageNumber="95" masterDocId="A859FFD0FF952A30FFA1FF8CFC30E154" masterDocTitle="Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae)" masterLastPageNumber="214" masterPageNumber="1" pageNumber="94" updateTime="1699020181560" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B190EF861719AE5F3FAA10CCC7812347">Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae)</mods:title>
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<treatment id="546087A8FFC82A6EFEA9FD08FE22E392" ID-DOI="http://doi.org/10.5281/zenodo.5019245" ID-GBIF-Taxon="183402345" ID-Zenodo-Dep="5019245" LSID="urn:lsid:plazi:treatment:546087A8FFC82A6EFEA9FD08FE22E392" httpUri="http://treatment.plazi.org/id/546087A8FFC82A6EFEA9FD08FE22E392" lastPageId="94" lastPageNumber="95" pageId="93" pageNumber="94">
<subSubSection id="94D36535FFC82A6DFEA9FD08FEA6E3CA" box="[264,662,644,670]" pageId="93" pageNumber="94" type="nomenclature">
<paragraph id="DC7636BEFFC82A6DFEA9FD08FEA6E3CA" blockId="93.[264,662,644,705]" box="[264,662,644,670]" pageId="93" pageNumber="94">
<heading id="873E81D2FFC82A6DFEA9FD08FEA6E3CA" bold="true" box="[264,662,644,670]" fontSize="11" level="1" pageId="93" pageNumber="94" reason="1">
<taxonomicName id="1BC94D3DFFC82A6DFEA9FD08FEA6E3CA" ID-CoL="4H7KJ" authority="Martynov" authorityName="Martynov Segment IX" authorityYear="1912" box="[264,662,644,670]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="spectabilis">
<emphasis id="EEBDEAACFFC82A6DFEA9FD08FEA6E3CA" bold="true" box="[264,662,644,670]" pageId="93" pageNumber="94">Phylloicus spectabilis Martynov</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="94D36535FFC82A6DFEA9FD2BFD6BE395" box="[264,347,679,705]" pageId="93" pageNumber="94" type="description">
<paragraph id="DC7636BEFFC82A6DFEA9FD2BFD6BE395" blockId="93.[264,662,644,705]" box="[264,347,679,705]" pageId="93" pageNumber="94">
<figureCitation id="44F22A3BFFC82A6DFEA9FD2BFD6BE395" box="[264,347,679,705]" captionStart="FIGURE 99" captionStartId="202.[264,368,1808,1832]" captionTargetBox="[279,1314,329,1790]" captionTargetId="figure-48@202.[277,1315,326,1795]" captionTargetPageId="202" captionText="FIGURE 99. Phylloicus spectabilis. Male (holotype): A—lateral view; B—dorsal view; C— sterna VIII, IX, and inferior appendages, ventral view; D—phallus, lateral view; E—phallus, ventral view; F—tergum IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019474" httpUri="https://zenodo.org/record/5019474/files/figure.png" pageId="93" pageNumber="94">Fig. 99</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="94D36535FFC82A6DFEA9FD7EFF6FE54A" pageId="93" pageNumber="94" type="reference_group">
<paragraph id="DC7636BEFFC82A6DFEA9FD7EF8BFE25C" blockId="93.[264,1167,753,777]" box="[264,1167,753,777]" pageId="93" pageNumber="94">
<treatmentCitationGroup id="FCD91190FFC82A6DFEA9FD7EF8BFE25C" box="[264,1167,753,777]" pageId="93" pageNumber="94">
<treatmentCitation id="5D6810AFFFC82A6DFEA9FD7EFE87E25C" author="Martynov, A. B." box="[264,695,753,777]" page="9" pageId="93" pageNumber="94" year="1912">
<taxonomicName id="1BC94D3DFFC82A6DFEA9FD7EFE87E25C" ID-CoL="4H7KJ" authority="Martynov, 1912: 9" authorityName="Martynov Segment IX" authorityPageNumber="9" authorityYear="1912" box="[264,695,753,777]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="spectabilis">
<emphasis id="EEBDEAACFFC82A6DFEA9FD7EFDC1E25D" box="[264,497,754,777]" italics="true" pageId="93" pageNumber="94">Phylloicus spectabilis</emphasis>
<materialsCitation id="6CA13CE3FFC82A6DFE59FD7DFE87E25C" ID-GBIF-Occurrence="3319300764" box="[504,695,753,776]" country="Peru" location="Callanga" pageId="93" pageNumber="94" specimenCount="1">
<bibRefCitation id="B8584B4FFFC82A6DFE59FD7DFE87E25C" author="Martynov, A. B." box="[504,695,753,776]" pageId="93" pageNumber="94" pagination="1 - 40" refId="ref75186" refString="Martynov, A. B. (1912) On two collections of Trichoptera from Peru. Annuaire de Musee Zoologique de l'Academie Imperial des Sciences de St. Petersbourg, 17, 1 - 40." type="journal article" year="1912">Martynov, 1912:9</bibRefCitation>
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[
<typeStatus id="0372881CFFC82A6DFD65FD7DFECCE25C" box="[708,764,753,776]" pageId="93" pageNumber="94">Type</typeStatus>
locality:
<collectingCountry id="A4DE762EFFC82A6DFCC1FD7DFFA4E25C" box="[864,916,753,776]" name="Peru" pageId="93" pageNumber="94">Peru</collectingCountry>
,
<location id="D9166065FFC82A6DFC3FFD7DF830E25C" LSID="urn:lsid:plazi:treatment:546087A8FFC82A6EFEA9FD08FE22E392:D9166065FFC82A6DFC3FFD7DF830E25C" box="[926,1024,753,776]" country="Peru" name="Callanga" pageId="93" pageNumber="94">Callanga</location>
; ASL; male].
</treatmentCitationGroup>
</paragraph>
<paragraph id="DC7636BEFFC82A6DFEA9FCB0FF6FE54A" blockId="93.[264,1323,828,1894]" pageId="93" pageNumber="94">
This species is known only from the badly rubbed
<typeStatus id="0372881CFFC82A6DFCCCFCB0FFE8E202" box="[877,984,828,854]" pageId="93" pageNumber="94" type="lectotype">lectotype</typeStatus>
male and
<specimenCount id="CACFFD37FFC82A6DFBF8FCB0FD74E22A" pageId="93" pageNumber="94" type="male" typeStatus="paralectotype">one paralectotype male</specimenCount>
, which is missing an abdomen. The modifications of the male abdomen are extremely elaborate in this species. It is similar to a number of other species, but is distinguished by the shape of the lateral sclerite of tergum IV, the arrangement and number of lobes of the coremata (
<figureCitation id="44F22A3BFFC82A6DFDB8FC50FEB3E2A2" box="[537,643,988,1014]" captionStart="FIGURE 99" captionStartId="202.[264,368,1808,1832]" captionTargetBox="[279,1314,329,1790]" captionTargetId="figure-48@202.[277,1315,326,1795]" captionTargetPageId="202" captionText="FIGURE 99. Phylloicus spectabilis. Male (holotype): A—lateral view; B—dorsal view; C— sterna VIII, IX, and inferior appendages, ventral view; D—phallus, lateral view; E—phallus, ventral view; F—tergum IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019474" httpUri="https://zenodo.org/record/5019474/files/figure.png" pageId="93" pageNumber="94">Fig. 99F</figureCitation>
), and the shape of sternum VIII (
<figureCitation id="44F22A3BFFC82A6DFBB3FC50F8B0E2A2" box="[1042,1152,988,1014]" captionStart="FIGURE 99" captionStartId="202.[264,368,1808,1832]" captionTargetBox="[279,1314,329,1790]" captionTargetId="figure-48@202.[277,1315,326,1795]" captionTargetPageId="202" captionText="FIGURE 99. Phylloicus spectabilis. Male (holotype): A—lateral view; B—dorsal view; C— sterna VIII, IX, and inferior appendages, ventral view; D—phallus, lateral view; E—phallus, ventral view; F—tergum IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019474" httpUri="https://zenodo.org/record/5019474/files/figure.png" pageId="93" pageNumber="94">Fig. 99C</figureCitation>
). The description of color features is based on the
<typeStatus id="0372881CFFC82A6DFD18FB88FF6BE54A" box="[697,859,1028,1054]" pageId="93" pageNumber="94" type="paralectotype">paralectotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="94D36535FFC82A6EFE99FBA0FE22E392" lastPageId="94" lastPageNumber="95" pageId="93" pageNumber="94" type="description">
<paragraph id="DC7636BEFFC82A6DFE99FBA0FF3AE512" blockId="93.[264,1323,828,1894]" box="[312,778,1068,1094]" pageId="93" pageNumber="94">Adult. Forewing length 11.6 mm, n = 2.</paragraph>
<paragraph id="DC7636BEFFC82A6DFE99FBD8FE90E4D2" blockId="93.[264,1323,828,1894]" pageId="93" pageNumber="94">Head chestnut brown. Maxillary palps chestnut brown. Antenna chestnut brown, with narrow patches of pale sensilla on anteromesal surface of each flagellomere. Dorsal pterothorax chestnut brown; ventrolateral thorax chestnut brown. Legs chestnut brown. Metathoracic leg of male with posterior fringe of long setae. Tibial spur formula 2,4,4. Forewing flat; chestnut brown; with two transverse bands; proximal band golden, reaching posterior wing margin, at least 1/2 width of wing; distal band golden, not reaching either wing margin, 1/2 width of wing or less; with two basal stripes, golden. Hind wing basal brush present in male, dark brown.</paragraph>
<paragraph id="DC7636BEFFC82A6EFE99FA18FE73E372" blockId="93.[264,1323,828,1894]" lastBlockId="94.[264,1322,284,710]" lastPageId="94" lastPageNumber="95" pageId="93" pageNumber="94">
Male. Preterminalic abdominal terga with anteromesal notch. Corematic structures present. Tergum III with sclerotized setose processes of posterior pleural membrane. Tergum IV with paired posterior processes and paired lateral sclerites, mesal coremata and lateral coremata; posterior process truncate; lateral sclerite spatulate, directed laterally; lateral coremata with basal globose lobes and long tubular posterior lobe; mesal coremata bilobed, setose, posterior lobe three times length of anterior lobe. Tergum V without sclerotized modifications (
<figureCitation id="44F22A3BFFC82A6DFDB2F908FE4CE7CA" box="[531,636,1668,1694]" captionStart="FIGURE 99" captionStartId="202.[264,368,1808,1832]" captionTargetBox="[279,1314,329,1790]" captionTargetId="figure-48@202.[277,1315,326,1795]" captionTargetPageId="202" captionText="FIGURE 99. Phylloicus spectabilis. Male (holotype): A—lateral view; B—dorsal view; C— sterna VIII, IX, and inferior appendages, ventral view; D—phallus, lateral view; E—phallus, ventral view; F—tergum IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019474" httpUri="https://zenodo.org/record/5019474/files/figure.png" pageId="93" pageNumber="94">Fig. 99F</figureCitation>
). Sternum VII with short, acute anteromesal process. Sternum VIII enclosing base of elongate sternum IX; posteromesal process absent and posterior of segment semimembranous (
<figureCitation id="44F22A3BFFC82A6DFD4DF958FF6CE7BA" box="[748,860,1748,1774]" captionStart="FIGURE 99" captionStartId="202.[264,368,1808,1832]" captionTargetBox="[279,1314,329,1790]" captionTargetId="figure-48@202.[277,1315,326,1795]" captionTargetPageId="202" captionText="FIGURE 99. Phylloicus spectabilis. Male (holotype): A—lateral view; B—dorsal view; C— sterna VIII, IX, and inferior appendages, ventral view; D—phallus, lateral view; E—phallus, ventral view; F—tergum IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019474" httpUri="https://zenodo.org/record/5019474/files/figure.png" pageId="93" pageNumber="94">Fig. 99C</figureCitation>
). Tergum IX with paired mesal ridges extending length of segment; posterior margin continuous with basodorsal process of tergum X (
<figureCitation id="44F22A3BFFC82A6DFEC8F8A8FDE5E66A" box="[361,469,1828,1854]" captionStart="FIGURE 99" captionStartId="202.[264,368,1808,1832]" captionTargetBox="[279,1314,329,1790]" captionTargetId="figure-48@202.[277,1315,326,1795]" captionTargetPageId="202" captionText="FIGURE 99. Phylloicus spectabilis. Male (holotype): A—lateral view; B—dorsal view; C— sterna VIII, IX, and inferior appendages, ventral view; D—phallus, lateral view; E—phallus, ventral view; F—tergum IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019474" httpUri="https://zenodo.org/record/5019474/files/figure.png" pageId="93" pageNumber="94">Fig. 99B</figureCitation>
); lateral ridge absent; dorsal pleural setae approximately 5, ventral pleural setae approximately 3 (
<figureCitation id="44F22A3BFFC82A6DFDF5F8C0FEF6E632" box="[596,710,1868,1894]" captionStart="FIGURE 99" captionStartId="202.[264,368,1808,1832]" captionTargetBox="[279,1314,329,1790]" captionTargetId="figure-48@202.[277,1315,326,1795]" captionTargetPageId="202" captionText="FIGURE 99. Phylloicus spectabilis. Male (holotype): A—lateral view; B—dorsal view; C— sterna VIII, IX, and inferior appendages, ventral view; D—phallus, lateral view; E—phallus, ventral view; F—tergum IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019474" httpUri="https://zenodo.org/record/5019474/files/figure.png" pageId="93" pageNumber="94">Fig. 99A</figureCitation>
). Preanal appendage at least 11/2 times length of tergum X, widest apically, setae filamentous, longer than appendage (
<figureCitation id="44F22A3BFFCB2A6EFBEFFE90F8D1E062" box="[1102,1249,284,310]" captionStart="FIGURE 99" captionStartId="202.[264,368,1808,1832]" captionTargetBox="[279,1314,329,1790]" captionTargetId="figure-48@202.[277,1315,326,1795]" captionTargetPageId="202" captionText="FIGURE 99. Phylloicus spectabilis. Male (holotype): A—lateral view; B—dorsal view; C— sterna VIII, IX, and inferior appendages, ventral view; D—phallus, lateral view; E—phallus, ventral view; F—tergum IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019474" httpUri="https://zenodo.org/record/5019474/files/figure.png" pageId="94" pageNumber="95">Fig. 99A, B</figureCitation>
). Tergum X without basal lobes; basodorsal process long and digitate; basolateral processes absent; apex, in lateral view, rounded, in dorsal view, entire (
<figureCitation id="44F22A3BFFCB2A6EFBACFEE0F897E0D2" box="[1037,1191,364,390]" captionStart="FIGURE 99" captionStartId="202.[264,368,1808,1832]" captionTargetBox="[279,1314,329,1790]" captionTargetId="figure-48@202.[277,1315,326,1795]" captionTargetPageId="202" captionText="FIGURE 99. Phylloicus spectabilis. Male (holotype): A—lateral view; B—dorsal view; C— sterna VIII, IX, and inferior appendages, ventral view; D—phallus, lateral view; E—phallus, ventral view; F—tergum IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019474" httpUri="https://zenodo.org/record/5019474/files/figure.png" pageId="94" pageNumber="95">Fig. 99A, B</figureCitation>
). Harpago rounded; peglike setae many, apical (
<figureCitation id="44F22A3BFFCB2A6EFD1EFE18FF60E0FA" box="[703,848,404,430]" captionStart="FIGURE 99" captionStartId="202.[264,368,1808,1832]" captionTargetBox="[279,1314,329,1790]" captionTargetId="figure-48@202.[277,1315,326,1795]" captionTargetPageId="202" captionText="FIGURE 99. Phylloicus spectabilis. Male (holotype): A—lateral view; B—dorsal view; C— sterna VIII, IX, and inferior appendages, ventral view; D—phallus, lateral view; E—phallus, ventral view; F—tergum IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019474" httpUri="https://zenodo.org/record/5019474/files/figure.png" pageId="94" pageNumber="95">Fig. 99A, C</figureCitation>
). Phallic endotheca with paired basolateral lobes, basolateral lobes large and round; phallotremal sclerites average size, longest dimension less than diameter of phallobase; dorsal sclerite ovoid, in dorsal view horseshoe­shaped (
<figureCitation id="44F22A3BFFCB2A6EFE09FD80FE05E372" box="[424,565,524,550]" captionStart="FIGURE 99" captionStartId="202.[264,368,1808,1832]" captionTargetBox="[279,1314,329,1790]" captionTargetId="figure-48@202.[277,1315,326,1795]" captionTargetPageId="202" captionText="FIGURE 99. Phylloicus spectabilis. Male (holotype): A—lateral view; B—dorsal view; C— sterna VIII, IX, and inferior appendages, ventral view; D—phallus, lateral view; E—phallus, ventral view; F—tergum IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019474" httpUri="https://zenodo.org/record/5019474/files/figure.png" pageId="94" pageNumber="95">Fig. 99D, E</figureCitation>
).
</paragraph>
<paragraph id="DC7636BEFFCB2A6EFE99FDB8FE28E31A" blockId="94.[264,1322,284,710]" box="[312,536,564,590]" pageId="94" pageNumber="95">Female. Unknown.</paragraph>
<paragraph id="DC7636BEFFCB2A6EFE99FDD0FEAEE3CA" blockId="94.[264,1322,284,710]" pageId="94" pageNumber="95">
<materialsCitation id="6CA13CE3FFCB2A6EFE99FDD0F91AE322" ID-GBIF-Occurrence="3319300840" box="[312,1322,603,630]" collectorName="Staudinger &amp; Bang-Haas" country="Peru" location="Cuzco" pageId="94" pageNumber="95" specimenCount="1" stateProvince="Cusco" typeStatus="lectotype">
Material examined.
<emphasis id="EEBDEAACFFCB2A6EFD89FDD7FED3E321" bold="true" box="[552,739,603,629]" pageId="94" pageNumber="95">
<collectingCountry id="A4DE762EFFCB2A6EFD89FDD7FE4CE321" box="[552,636,603,629]" name="Peru" pageId="94" pageNumber="95">PERU</collectingCountry>
:
<collectingRegion id="1E0DF85CFFCB2A6EFD2DFDD7FEEEE321" box="[652,734,603,629]" country="Peru" name="Cusco" pageId="94" pageNumber="95">Cuzco</collectingRegion>
:
</emphasis>
Callanga,
<collectorName id="713C5368FFCB2A6EFCC6FDD0FFD5E322" box="[871,997,604,630]" pageId="94" pageNumber="95">Staudinger</collectorName>
&amp;
<collectorName id="713C5368FFCB2A6EFBB1FDD0F8A2E322" box="[1040,1170,604,630]" pageId="94" pageNumber="95">Bang­Haas</collectorName>
—
<typeStatus id="0372881CFFCB2A6EFB1EFDD0F91AE322" box="[1215,1322,604,630]" pageId="94" pageNumber="95" type="lectotype">lectotype</typeStatus>
</materialsCitation>
<materialsCitation id="6CA13CE3FFCB2A6EFEA9FD08FEAAE3CA" ID-GBIF-Occurrence="3319300803" box="[264,666,644,670]" collectionCode="ASL" country="Peru" pageId="94" pageNumber="95" specimenCount="1" specimenCount-male="1" stateProvince="Cusco" typeStatus="paralectotype">
male,
<specimenCount id="CACFFD37FFCB2A6EFEEEFD08FDAFE3CA" box="[335,415,644,670]" pageId="94" pageNumber="95" type="male">1 male</specimenCount>
<typeStatus id="0372881CFFCB2A6EFE07FD08FE74E3CA" box="[422,580,644,670]" pageId="94" pageNumber="95" type="paralectotype">paralectotype</typeStatus>
(
<collectionCode id="BAD8AE7BFFCB2A6EFDF5FD08FEA1E3CA" box="[596,657,644,670]" pageId="94" pageNumber="95">ASL</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="DC7636BEFFCB2A6EFE99FD20FE22E392" blockId="94.[264,1322,284,710]" box="[312,530,684,710]" pageId="94" pageNumber="95">
Distribution.
<collectingCountry id="A4DE762EFFCB2A6EFE75FD20FE3EE392" box="[468,526,684,710]" name="Peru" pageId="94" pageNumber="95">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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