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<document id="C3F69B07B0F80DCFE58F5D2E328EF411" ID-CLB-Dataset="5618" ID-DOI="10.11646/zootaxa.275.1.1" ID-GBIF-Dataset="9322f2b2-a384-40bf-9d57-d789c76ce5c4" ID-ISSN="1175­5334" ID-Zenodo-Dep="5019235" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624466010041" checkinUser="felipe" docAuthor="PRATHER, AYSHA L." docDate="2003" docId="546087A8FFB72A13FEA9FE37FD72E59A" docLanguage="en" docName="zt00275.pdf" docOrigin="Zootaxa 275 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.275.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Phylloicus blahniki Prather 2003, new species" docType="treatment" docVersion="9" lastPageNumber="36" masterDocId="A859FFD0FF952A30FFA1FF8CFC30E154" masterDocTitle="Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae)" masterLastPageNumber="214" masterPageNumber="1" pageNumber="35" updateTime="1699020181560" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="534AE4EFC8AAE64A9D4D2A8E655353C3">Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae)</mods:title>
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<treatment id="546087A8FFB72A13FEA9FE37FD72E59A" ID-DOI="http://doi.org/10.5281/zenodo.5019233" ID-GBIF-Taxon="183402361" ID-Zenodo-Dep="5019233" LSID="urn:lsid:plazi:treatment:546087A8FFB72A13FEA9FE37FD72E59A" httpUri="http://treatment.plazi.org/id/546087A8FFB72A13FEA9FE37FD72E59A" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
<subSubSection id="94D36535FFB72A12FEA9FE37FEA6E081" box="[264,662,443,469]" pageId="34" pageNumber="35" type="nomenclature">
<paragraph id="DC7636BEFFB72A12FEA9FE37FEA6E081" blockId="34.[264,662,443,505]" box="[264,662,443,469]" pageId="34" pageNumber="35">
<heading id="873E81D2FFB72A12FEA9FE37FEA6E081" bold="true" box="[264,662,443,469]" fontSize="11" level="1" pageId="34" pageNumber="35" reason="1">
<emphasis id="EEBDEAACFFB72A12FEA9FE37FEA6E081" bold="true" box="[264,662,443,469]" pageId="34" pageNumber="35">
<taxonomicName id="1BC94D3DFFB72A12FEA9FE37FDCAE081" authority="Prather, 2003" authorityName="Prather" authorityYear="2003" box="[264,506,443,469]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="blahniki" status="sp. nov.">Phylloicus blahniki</taxonomicName>
,
<taxonomicNameLabel id="F58E57D7FFB72A12FDA6FE37FEA6E081" box="[519,662,443,469]" pageId="34" pageNumber="35" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="94D36535FFB72A13FEA9FE53FDBBE392" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="description">
<paragraph id="DC7636BEFFB72A12FEA9FE53FDA2E0AD" blockId="34.[264,662,443,505]" box="[264,402,479,505]" pageId="34" pageNumber="35">
<figureCitation id="44F22A3BFFB72A12FEA9FE53FD59E0AD" box="[264,361,479,505]" captionStart="FIGURE 28" captionStartId="131.[264,367,1815,1839]" captionTargetBox="[275,1311,315,1803]" captionTargetId="figure-41@131.[273,1313,314,1803]" captionTargetPageId="131" captionText="FIGURE 28. Phylloicus blahniki. Male (UMSP000048732): A—lateral view; B—dorsal view; C— ventral view; D—phallus, lateral view; E—phallotremal sclerites, dorsal view; F—terga III­IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019285" httpUri="https://zenodo.org/record/5019285/files/figure.png" pageId="34" pageNumber="35">Figs. 28</figureCitation>
,
<figureCitation id="44F22A3BFFB72A12FED4FE53FDA2E0AD" box="[373,402,479,505]" captionStart="FIGURE 29" captionStartId="132.[264,368,1118,1142]" captionTargetBox="[264,1322,327,1094]" captionTargetId="figure-44@132.[263,1323,326,1094]" captionTargetPageId="132" captionText="FIGURE 29. Phylloicus blahniki. Female (UMSP000067615): A—sterna IX, X and vaginal apparatus, ventral view; B—terga IX and X, dorsal view; C—sternum VIII, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5019287" httpUri="https://zenodo.org/record/5019287/files/figure.png" pageId="34" pageNumber="35">29</figureCitation>
</paragraph>
<paragraph id="DC7636BEFFB72A12FEA9FDA3FDBCE235" blockId="34.[264,1323,559,1865]" pageId="34" pageNumber="35">
<taxonomicName id="1BC94D3DFFB72A12FEA9FDA3FDDDE31C" authorityName="Prather Table Of Contents" authorityYear="2003" box="[264,493,559,584]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="blahniki">
<emphasis id="EEBDEAACFFB72A12FEA9FDA3FDDDE31C" box="[264,493,559,584]" italics="true" pageId="34" pageNumber="35">Phylloicus blahniki</emphasis>
</taxonomicName>
is distinguished by the modifications of abdominal segments III and IV of the male. Tergite III is flanged anterolaterally; tergite IV bears a truncate posterior process and the lateral sclerite is short and rounded apically. Only lateral coremata are present, and they consist of four lobes, three of them short and rounded, the posterior lobe slightly elongate (
<figureCitation id="44F22A3BFFB72A12FE78FD43FE71E3BD" box="[473,577,719,745]" captionStart="FIGURE 28" captionStartId="131.[264,367,1815,1839]" captionTargetBox="[275,1311,315,1803]" captionTargetId="figure-41@131.[273,1313,314,1803]" captionTargetPageId="131" captionText="FIGURE 28. Phylloicus blahniki. Male (UMSP000048732): A—lateral view; B—dorsal view; C— ventral view; D—phallus, lateral view; E—phallotremal sclerites, dorsal view; F—terga III­IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019285" httpUri="https://zenodo.org/record/5019285/files/figure.png" pageId="34" pageNumber="35">Fig. 28F</figureCitation>
). The few specimens on which this description is based were all teneral or rubbed, so the description of the wing pattern is incomplete. None of the male specimens had the phallic endotheca fully everted, so I am unable to describe the membranous lobes.
</paragraph>
<paragraph id="DC7636BEFFB72A12FE99FCE3FF3BE2DD" blockId="34.[264,1323,559,1865]" box="[312,779,879,905]" pageId="34" pageNumber="35">Adult. Forewing length 11.7 mm, n = 8.</paragraph>
<paragraph id="DC7636BEFFB72A12FE99FC1BFE7CE5F5" blockId="34.[264,1323,559,1865]" pageId="34" pageNumber="35">Head golden brown. Maxillary palps golden brown. Antenna twice forewing length; dark brown, with narrow patches of pale sensilla on anteromesal surface of each flagellomere. Dorsal pterothorax golden brown, anterolateral margins dark brown; ventrolateral thorax golden brown. Femora golden brown; tibiae golden brown; tarsi dark brown. Metathoracic leg of male without posterior fringe. Tibial spur formula 2,4,4. Forewing flat; chestnut brown; with longitudinal stripes; stripes pale tan. Hind wing basal brush present in male and female.</paragraph>
<paragraph id="DC7636BEFFB72A12FE99FB23FE34E61D" blockId="34.[264,1323,559,1865]" pageId="34" pageNumber="35">
Male. Preterminalic abdominal terga with anteromesal notch. Corematic structures present. Tergum III with anterolateral flanges and short posterolateral projections. Tergum IV with paired posterior processes and paired lateral sclerites, lateral coremata; posterior process truncate; lateral sclerite narrowed apically; lateral coremata with basal globose lobes. Tergum V without sclerotized modifications (
<figureCitation id="44F22A3BFFB72A12FC23FAC3FFDFE43D" box="[898,1007,1359,1385]" captionStart="FIGURE 28" captionStartId="131.[264,367,1815,1839]" captionTargetBox="[275,1311,315,1803]" captionTargetId="figure-41@131.[273,1313,314,1803]" captionTargetPageId="131" captionText="FIGURE 28. Phylloicus blahniki. Male (UMSP000048732): A—lateral view; B—dorsal view; C— ventral view; D—phallus, lateral view; E—phallotremal sclerites, dorsal view; F—terga III­IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019285" httpUri="https://zenodo.org/record/5019285/files/figure.png" pageId="34" pageNumber="35">Fig. 28F</figureCitation>
). Sternum VII with short, acute anteromesal process. Sternum VIII similar to anterior sterna, sternum IX not elongate. Tergum IX without mesal ridge; posterior margin smoothly rounded; thinly sclerotized anteromesally (
<figureCitation id="44F22A3BFFB72A12FDA3FA4BFE41E4B5" box="[514,625,1479,1505]" captionStart="FIGURE 28" captionStartId="131.[264,367,1815,1839]" captionTargetBox="[275,1311,315,1803]" captionTargetId="figure-41@131.[273,1313,314,1803]" captionTargetPageId="131" captionText="FIGURE 28. Phylloicus blahniki. Male (UMSP000048732): A—lateral view; B—dorsal view; C— ventral view; D—phallus, lateral view; E—phallotremal sclerites, dorsal view; F—terga III­IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019285" httpUri="https://zenodo.org/record/5019285/files/figure.png" pageId="34" pageNumber="35">Fig. 28B</figureCitation>
); lateral ridge present; dorsal pleural setae absent, ventral pleural setae approximately 10 (
<figureCitation id="44F22A3BFFB72A12FD25FA63FEC3E75D" box="[644,755,1519,1545]" captionStart="FIGURE 28" captionStartId="131.[264,367,1815,1839]" captionTargetBox="[275,1311,315,1803]" captionTargetId="figure-41@131.[273,1313,314,1803]" captionTargetPageId="131" captionText="FIGURE 28. Phylloicus blahniki. Male (UMSP000048732): A—lateral view; B—dorsal view; C— ventral view; D—phallus, lateral view; E—phallotremal sclerites, dorsal view; F—terga III­IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019285" httpUri="https://zenodo.org/record/5019285/files/figure.png" pageId="34" pageNumber="35">Fig. 28A</figureCitation>
); sternum IX with faint mesolateral ridges; sternum IX (
<figureCitation id="44F22A3BFFB72A12FEDBF99BFDDAE765" box="[378,490,1559,1585]" captionStart="FIGURE 28" captionStartId="131.[264,367,1815,1839]" captionTargetBox="[275,1311,315,1803]" captionTargetId="figure-41@131.[273,1313,314,1803]" captionTargetPageId="131" captionText="FIGURE 28. Phylloicus blahniki. Male (UMSP000048732): A—lateral view; B—dorsal view; C— ventral view; D—phallus, lateral view; E—phallotremal sclerites, dorsal view; F—terga III­IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019285" httpUri="https://zenodo.org/record/5019285/files/figure.png" pageId="34" pageNumber="35">Fig. 28C</figureCitation>
). Preanal appendage longer than tergum X, but less than 11/2 times length, of uniform diameter throughout length, setae long, but not filamentous or longer than appendage (
<figureCitation id="44F22A3BFFB72A12FE73F9EBFE53E7D5" box="[466,611,1639,1665]" captionStart="FIGURE 28" captionStartId="131.[264,367,1815,1839]" captionTargetBox="[275,1311,315,1803]" captionTargetId="figure-41@131.[273,1313,314,1803]" captionTargetPageId="131" captionText="FIGURE 28. Phylloicus blahniki. Male (UMSP000048732): A—lateral view; B—dorsal view; C— ventral view; D—phallus, lateral view; E—phallotremal sclerites, dorsal view; F—terga III­IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019285" httpUri="https://zenodo.org/record/5019285/files/figure.png" pageId="34" pageNumber="35">Fig. 28A, B</figureCitation>
). Tergum X without basal lobes; basodorsal process absent; basolateral processes absent; apex, in lateral view, acute, in dorsal view, notched, notch triangular; base of tergum X setose; tapered apically (
<figureCitation id="44F22A3BFFB72A12FCFEF93BFFDCE785" box="[863,1004,1719,1745]" captionStart="FIGURE 28" captionStartId="131.[264,367,1815,1839]" captionTargetBox="[275,1311,315,1803]" captionTargetId="figure-41@131.[273,1313,314,1803]" captionTargetPageId="131" captionText="FIGURE 28. Phylloicus blahniki. Male (UMSP000048732): A—lateral view; B—dorsal view; C— ventral view; D—phallus, lateral view; E—phallotremal sclerites, dorsal view; F—terga III­IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019285" httpUri="https://zenodo.org/record/5019285/files/figure.png" pageId="34" pageNumber="35">Fig. 28A, B</figureCitation>
). Harpago slightly tapered; peglike setae few, apical (
<figureCitation id="44F22A3BFFB72A12FD97F953FEF4E7AD" box="[566,708,1759,1785]" captionStart="FIGURE 28" captionStartId="131.[264,367,1815,1839]" captionTargetBox="[275,1311,315,1803]" captionTargetId="figure-41@131.[273,1313,314,1803]" captionTargetPageId="131" captionText="FIGURE 28. Phylloicus blahniki. Male (UMSP000048732): A—lateral view; B—dorsal view; C— ventral view; D—phallus, lateral view; E—phallotremal sclerites, dorsal view; F—terga III­IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019285" httpUri="https://zenodo.org/record/5019285/files/figure.png" pageId="34" pageNumber="35">Fig. 28A, C</figureCitation>
). Phallotremal sclerites average size, longest dimension less than diameter of phallobase; dorsal sclerite ovoid, in dorsal view horseshoeshaped (
<figureCitation id="44F22A3BFFB72A12FEC8F8A3FDC7E61D" box="[361,503,1839,1865]" captionStart="FIGURE 28" captionStartId="131.[264,367,1815,1839]" captionTargetBox="[275,1311,315,1803]" captionTargetId="figure-41@131.[273,1313,314,1803]" captionTargetPageId="131" captionText="FIGURE 28. Phylloicus blahniki. Male (UMSP000048732): A—lateral view; B—dorsal view; C— ventral view; D—phallus, lateral view; E—phallotremal sclerites, dorsal view; F—terga III­IV, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5019285" httpUri="https://zenodo.org/record/5019285/files/figure.png" pageId="34" pageNumber="35">Fig. 28D, E</figureCitation>
).
</paragraph>
<paragraph id="DC7636BEFFB62A13FE99FE90FDBBE392" blockId="35.[264,1324,284,1230]" pageId="35" pageNumber="36">
Female. Preterminal abdominal terga with anteromesal notch. Sternum VII with short pointed anteromesal process. Tergum VIII without posterolateral brush; sternum VIII cleft posteromesally to anterior ridge; sternum VIII (
<figureCitation id="44F22A3BFFB62A13FC99FEE0FF94E0D2" box="[824,932,364,390]" captionStart="FIGURE 29" captionStartId="132.[264,368,1118,1142]" captionTargetBox="[264,1322,327,1094]" captionTargetId="figure-44@132.[263,1323,326,1094]" captionTargetPageId="132" captionText="FIGURE 29. Phylloicus blahniki. Female (UMSP000067615): A—sterna IX, X and vaginal apparatus, ventral view; B—terga IX and X, dorsal view; C—sternum VIII, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5019287" httpUri="https://zenodo.org/record/5019287/files/figure.png" pageId="35" pageNumber="36">Fig. 29C</figureCitation>
). Tergum IX without mesal ridge (
<figureCitation id="44F22A3BFFB62A13FEB0FE18FDB0E0FA" box="[273,384,404,430]" captionStart="FIGURE 29" captionStartId="132.[264,368,1118,1142]" captionTargetBox="[264,1322,327,1094]" captionTargetId="figure-44@132.[263,1323,326,1094]" captionTargetPageId="132" captionText="FIGURE 29. Phylloicus blahniki. Female (UMSP000067615): A—sterna IX, X and vaginal apparatus, ventral view; B—terga IX and X, dorsal view; C—sternum VIII, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5019287" httpUri="https://zenodo.org/record/5019287/files/figure.png" pageId="35" pageNumber="36">Fig. 29B</figureCitation>
). Sternum IX anterior and posterior lobes darkly sclerotized and striate, with patch of lightly sclerotized cuticle lateral to vaginal opening (
<figureCitation id="44F22A3BFFB62A13FBBDFE30F8A2E082" box="[1052,1170,444,470]" captionStart="FIGURE 29" captionStartId="132.[264,368,1118,1142]" captionTargetBox="[264,1322,327,1094]" captionTargetId="figure-44@132.[263,1323,326,1094]" captionTargetPageId="132" captionText="FIGURE 29. Phylloicus blahniki. Female (UMSP000067615): A—sterna IX, X and vaginal apparatus, ventral view; B—terga IX and X, dorsal view; C—sternum VIII, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5019287" httpUri="https://zenodo.org/record/5019287/files/figure.png" pageId="35" pageNumber="36">Fig. 29A</figureCitation>
). Tergum X appendage longer than mesal lobe, base indistinct, apex rounded; mesal lobe lightly sclerotized; digitate lateral processes long, at least twice diameter and often asymmetrical (
<figureCitation id="44F22A3BFFB62A13FEB0FDB8FDB0E31A" box="[273,384,564,590]" captionStart="FIGURE 29" captionStartId="132.[264,368,1118,1142]" captionTargetBox="[264,1322,327,1094]" captionTargetId="figure-44@132.[263,1323,326,1094]" captionTargetPageId="132" captionText="FIGURE 29. Phylloicus blahniki. Female (UMSP000067615): A—sterna IX, X and vaginal apparatus, ventral view; B—terga IX and X, dorsal view; C—sternum VIII, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5019287" httpUri="https://zenodo.org/record/5019287/files/figure.png" pageId="35" pageNumber="36">Fig. 29B</figureCitation>
). Sternum X with patches of short fine setae posterolaterally to anal opening (
<figureCitation id="44F22A3BFFB62A13FEB0FDD0FDB3E322" box="[273,387,604,630]" captionStart="FIGURE 29" captionStartId="132.[264,368,1118,1142]" captionTargetBox="[264,1322,327,1094]" captionTargetId="figure-44@132.[263,1323,326,1094]" captionTargetPageId="132" captionText="FIGURE 29. Phylloicus blahniki. Female (UMSP000067615): A—sterna IX, X and vaginal apparatus, ventral view; B—terga IX and X, dorsal view; C—sternum VIII, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5019287" httpUri="https://zenodo.org/record/5019287/files/figure.png" pageId="35" pageNumber="36">Fig. 29A</figureCitation>
). Vaginal apparatus anterior and posterior sclerites equal in length; anterior sclerite rounded anteriorly, posterolateral projections acute; posterior sclerite triangular (
<figureCitation id="44F22A3BFFB62A13FEB0FD20FD4EE392" box="[273,382,684,710]" captionStart="FIGURE 29" captionStartId="132.[264,368,1118,1142]" captionTargetBox="[264,1322,327,1094]" captionTargetId="figure-44@132.[263,1323,326,1094]" captionTargetPageId="132" captionText="FIGURE 29. Phylloicus blahniki. Female (UMSP000067615): A—sterna IX, X and vaginal apparatus, ventral view; B—terga IX and X, dorsal view; C—sternum VIII, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5019287" httpUri="https://zenodo.org/record/5019287/files/figure.png" pageId="35" pageNumber="36">Fig. 29A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="94D36535FFB62A13FE99FD58FD72E59A" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="DC7636BEFFB62A13FE99FD58FD43E26A" blockId="35.[264,1324,284,1230]" pageId="35" pageNumber="36">
<materialsCitation id="6CA13CE3FFB62A13FE99FD58FD43E26A" ID-GBIF-Occurrence="3319300538" collectingDate="1989-04-10" collectionCode="UMSP" collectorName="Holzenthal &amp; Blahnik" country="Costa Rica" elevation="20" latitude="8.581944" location="Parque Nacional Corcovado" longLatPrecision="21" longitude="-83.708885" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
<emphasis id="EEBDEAACFFB62A13FE99FD58FF6EE3BA" bold="true" box="[312,862,724,750]" pageId="35" pageNumber="36">
<typeStatus id="0372881CFFB62A13FE99FD58FD9AE3BA" box="[312,426,724,750]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="A4DE762EFFB62A13FE5EFD58FE88E3BA" box="[511,696,724,750]" name="Costa Rica" pageId="35" pageNumber="36">COSTA RICA</collectingCountry>
:
<collectingRegion id="1E0DF85CFFB62A13FD64FD58FF68E3BA" box="[709,856,724,750]" country="Costa Rica" name="Puntarenas" pageId="35" pageNumber="36">Puntarenas</collectingRegion>
:
</emphasis>
<location id="D9166065FFB62A13FCC4FD58F882E3BA" LSID="urn:lsid:plazi:treatment:546087A8FFB72A13FEA9FE37FD72E59A:D9166065FFB62A13FCC4FD58F882E3BA" box="[869,1202,724,750]" country="Costa Rica" latitude="8.581944" longLatPrecision="21" longitude="-83.708885" name="Parque Nacional Corcovado" pageId="35" pageNumber="36" stateProvince="Puntarenas">Parque Nacional Corcovado</location>
, unnamed stream,
<collectorName id="713C5368FFB62A13FEC4FD70FE21E242" box="[357,529,764,790]" pageId="35" pageNumber="36">Piedra el Arco</collectorName>
,
<geoCoordinate id="B9FD5079FFB62A13FDBCFD70FE9AE242" box="[541,682,764,790]" degrees="08" direction="north" minutes="34" orientation="latitude" pageId="35" pageNumber="36" precision="15" seconds="55" value="8.581944">08°34'55”N</geoCoordinate>
,
<geoCoordinate id="B9FD5079FFB62A13FD17FD70FF77E242" box="[694,839,764,790]" degrees="83" direction="west" minutes="42" orientation="longitude" pageId="35" pageNumber="36" precision="15" seconds="32" value="-83.708885">83°42'32&quot;W</geoCoordinate>
,
<quantity id="1B319B5BFFB62A13FCF2FD70FFA0E242" box="[851,912,764,790]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="36" unit="m" value="20.0">
<elevation id="57E4D18DFFB62A13FCF2FD70FFA0E242" box="[851,912,764,790]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="36" unit="m" value="20.0">20 m</elevation>
</quantity>
,
<date id="A877107EFFB62A13FC3CFD70F82BE242" box="[925,1051,764,790]" pageId="35" pageNumber="36" value="1989-04-10">
<collectingDate id="B833E996FFB62A13FC3CFD70F82BE242" box="[925,1051,764,790]" pageId="35" pageNumber="36" value="1989-04-10">10.iv.1989</collectingDate>
</date>
,
<collectorName id="713C5368FFB62A13FB86FD70F897E242" box="[1063,1191,764,790]" pageId="35" pageNumber="36">Holzenthal</collectorName>
&amp;
<collectorName id="713C5368FFB62A13FB6CFD70F91AE242" box="[1229,1322,764,790]" pageId="35" pageNumber="36">Blahnik</collectorName>
(
<collectionCode id="BAD8AE7BFFB62A13FEB1FCA8FD53E26A" box="[272,355,804,830]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="35" pageNumber="36" type="University or college">UMSP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="DC7636BEFFB62A13FE99FCC0FDC5E502" blockId="35.[264,1324,284,1230]" pageId="35" pageNumber="36">
<materialsCitation id="6CA13CE3FFB62A13FE99FCC0FD5EE2E2" ID-GBIF-Occurrence="3319300624" collectingDate="1986-02-15" collectionCode="UMSP" collectorName="Holzenthal &amp; Morse &amp; Fasth" country="Costa Rica" elevation="15" latitude="8.641944" location="Golfito" longLatPrecision="21" longitude="-83.193054" municipality="Quebrada Pita" pageId="35" pageNumber="36" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
<emphasis id="EEBDEAACFFB62A13FE99FCC0FF1FE232" bold="true" box="[312,815,844,870]" pageId="35" pageNumber="36">
<typeStatus id="0372881CFFB62A13FE99FCC0FD8AE232" box="[312,442,844,870]" pageId="35" pageNumber="36" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="A4DE762EFFB62A13FE68FCC0FEB6E232" box="[457,646,844,870]" name="Costa Rica" pageId="35" pageNumber="36">COSTA RICA</collectingCountry>
:
<collectingRegion id="1E0DF85CFFB62A13FD37FCC0FF19E232" box="[662,809,844,870]" country="Costa Rica" name="Puntarenas" pageId="35" pageNumber="36">Puntarenas</collectingRegion>
:
</emphasis>
<collectingMunicipality id="3C12ACC4FFB62A13FC98FCC0FFD7E232" box="[825,999,844,870]" pageId="35" pageNumber="36">Quebrada Pita</collectingMunicipality>
, ca.
<quantity id="1B319B5BFFB62A13FB83FCC0F850E232" box="[1058,1120,844,870]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="35" pageNumber="36" unit="km" value="3.0">3 km</quantity>
(air) W
<location id="D9166065FFB62A13FB6EFCC0F917E232" LSID="urn:lsid:plazi:treatment:546087A8FFB72A13FEA9FE37FD72E59A:D9166065FFB62A13FB6EFCC0F917E232" box="[1231,1319,844,870]" country="Costa Rica" latitude="8.641944" longLatPrecision="21" longitude="-83.193054" municipality="Quebrada Pita" name="Golfito" pageId="35" pageNumber="36" stateProvince="Puntarenas">Golfito</location>
,
<geoCoordinate id="B9FD5079FFB62A13FEA9FCF8FDA5E2DA" box="[264,405,884,910]" degrees="08" direction="north" minutes="38" orientation="latitude" pageId="35" pageNumber="36" precision="15" seconds="31" value="8.641944">08°38'31”N</geoCoordinate>
,
<geoCoordinate id="B9FD5079FFB62A13FE0AFCF8FE09E2DA" box="[427,569,884,910]" degrees="83" direction="west" minutes="11" orientation="longitude" pageId="35" pageNumber="36" precision="15" seconds="35" value="-83.193054">83°11'35&quot;W</geoCoordinate>
,
<quantity id="1B319B5BFFB62A13FDEEFCF8FEA6E2DA" box="[591,662,884,910]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="35" pageNumber="36" unit="m" value="15.0">
<elevation id="57E4D18DFFB62A13FDEEFCF8FEA6E2DA" box="[591,662,884,910]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="35" pageNumber="36" unit="m" value="15.0">15 m</elevation>
</quantity>
,
<date id="A877107EFFB62A13FD0AFCF8FF14E2DA" box="[683,804,884,910]" pageId="35" pageNumber="36" value="1986-02-15">
<collectingDate id="B833E996FFB62A13FD0AFCF8FF14E2DA" box="[683,804,884,910]" pageId="35" pageNumber="36" value="1986-02-15">15.ii.1986</collectingDate>
</date>
,
<collectorName id="713C5368FFB62A13FC98FCF8FF8DE2DA" box="[825,957,884,910]" pageId="35" pageNumber="36">Holzenthal</collectorName>
,
<collectorName id="713C5368FFB62A13FC73FCF8F810E2DA" box="[978,1056,884,910]" pageId="35" pageNumber="36">Morse</collectorName>
, &amp;
<collectorName id="713C5368FFB62A13FBFAFCF8F8AAE2DA" box="[1115,1178,884,910]" pageId="35" pageNumber="36">Fasth</collectorName>
<specimenCount id="CACFFD37FFB62A13FB72FCF8F91BE2DA" box="[1235,1323,884,910]" pageId="35" pageNumber="36" type="male">1 male</specimenCount>
(
<collectionCode id="BAD8AE7BFFB62A13FEB1FC10FD56E2E2" box="[272,358,924,950]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="35" pageNumber="36" type="University or college">UMSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6CA13CE3FFB62A13FE20FC10FD40E28A" ID-GBIF-Occurrence="3319300926" collectingDate="1979-03-15" collectingDateMax="1979-03-22" collectingDateMin="1979-03-15" collectionCode="INBIO" collectorName="Janzen" country="Costa Rica" location="Osa Peninsula" municipality="Corcovado National Park" pageId="35" pageNumber="36" specimenCount="2" specimenCount-male="2" stateProvince="Puntarenas" typeStatus="paratype">
<collectingMunicipality id="3C12ACC4FFB62A13FE20FC10FE89E2E2" box="[385,697,924,950]" pageId="35" pageNumber="36">Corcovado National Park</collectingMunicipality>
,
<location id="D9166065FFB62A13FD6AFC10FF4BE2E2" LSID="urn:lsid:plazi:treatment:546087A8FFB72A13FEA9FE37FD72E59A:D9166065FFB62A13FD6AFC10FF4BE2E2" box="[715,891,924,950]" country="Costa Rica" municipality="Corcovado National Park" name="Osa Peninsula" pageId="35" pageNumber="36" stateProvince="Puntarenas">Osa Peninsula</location>
,
<date id="A877107EFFB62A13FC2DFC10F805E2E2" box="[908,1077,924,950]" pageId="35" pageNumber="36" value="1979-03-15" valueMax="1979-03-22" valueMin="1979-03-15">
<collectingDate id="B833E996FFB62A13FC2DFC10F805E2E2" box="[908,1077,924,950]" pageId="35" pageNumber="36" value="1979-03-15" valueMax="1979-03-22" valueMin="1979-03-15">15­22.iii.1979</collectingDate>
</date>
,
<collectorName id="713C5368FFB62A13FBE6FC10F8A6E2E2" box="[1095,1174,924,950]" pageId="35" pageNumber="36">Janzen</collectorName>
<specimenCount id="CACFFD37FFB62A13FB6BFC10F91AE2E2" box="[1226,1322,924,950]" pageId="35" pageNumber="36" type="male">2 males</specimenCount>
(
<collectionCode id="BAD8AE7BFFB62A13FEB1FC48FD58E28A" box="[272,360,964,990]" pageId="35" pageNumber="36">INBIO</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6CA13CE3FFB62A13FE21FC48F827E552" ID-GBIF-Occurrence="3319300754" collectingDate="1989-04-13" collectionCode="UMSP" collectorName="Holzenthal &amp; Blahnik" country="Costa Rica" elevation="30" latitude="8.481944" location="Rio Camaronal" longLatPrecision="21" longitude="83.58889" municipality="Parque Nacional Corcovado" pageId="35" pageNumber="36" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Puntarenas" typeStatus="paratype">
<collectingMunicipality id="3C12ACC4FFB62A13FE21FC48FEE6E28A" box="[384,726,964,990]" pageId="35" pageNumber="36">Parque Nacional Corcovado</collectingMunicipality>
,
<location id="D9166065FFB62A13FD44FC48FFAEE28A" LSID="urn:lsid:plazi:treatment:546087A8FFB72A13FEA9FE37FD72E59A:D9166065FFB62A13FD44FC48FFAEE28A" box="[741,926,964,990]" country="Costa Rica" latitude="8.481944" longLatPrecision="21" longitude="83.58889" municipality="Parque Nacional Corcovado" name="Rio Camaronal" pageId="35" pageNumber="36" stateProvince="Puntarenas">Rio Camaronal</location>
,
<geoCoordinate id="B9FD5079FFB62A13FC0CFC48F80BE28A" box="[941,1083,964,990]" degrees="08" direction="north" minutes="28" orientation="latitude" pageId="35" pageNumber="36" precision="15" seconds="55" value="8.481944">08°28'55”N</geoCoordinate>
,
<geoCoordinate id="B9FD5079FFB62A13FBEAFC48F8E4E28A" box="[1099,1236,964,990]" degrees="83" direction="east" minutes="35" orientation="longitude" pageId="35" pageNumber="36" precision="15" seconds="20" value="83.58889">83°35'20&quot;E</geoCoordinate>
,
<quantity id="1B319B5BFFB62A13FB45FC48F916E28A" box="[1252,1318,964,990]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="35" pageNumber="36" unit="m" value="30.0">
<elevation id="57E4D18DFFB62A13FB45FC48F916E28A" box="[1252,1318,964,990]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="35" pageNumber="36" unit="m" value="30.0">30 m</elevation>
</quantity>
,
<date id="A877107EFFB62A13FEA9FC60FDB6E552" box="[264,390,1004,1030]" pageId="35" pageNumber="36" value="1989-04-13">
<collectingDate id="B833E996FFB62A13FEA9FC60FDB6E552" box="[264,390,1004,1030]" pageId="35" pageNumber="36" value="1989-04-13">13.iv.1989</collectingDate>
</date>
,
<collectorName id="713C5368FFB62A13FE34FC60FE26E552" box="[405,534,1004,1030]" pageId="35" pageNumber="36">Holzenthal</collectorName>
&amp;
<collectorName id="713C5368FFB62A13FDE3FC60FEAFE552" box="[578,671,1004,1030]" pageId="35" pageNumber="36">Blahnik</collectorName>
<specimenCount id="CACFFD37FFB62A13FD6CFC60FF09E552" box="[717,825,1004,1030]" pageId="35" pageNumber="36" type="female">1 female</specimenCount>
,
<specimenCount id="CACFFD37FFB62A13FCE9FC60FF95E552" box="[840,933,1004,1030]" pageId="35" pageNumber="36" type="male">2 males</specimenCount>
(
<collectionCode id="BAD8AE7BFFB62A13FC19FC60F83EE552" box="[952,1038,1004,1030]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="35" pageNumber="36" type="University or college">UMSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="6CA13CE3FFB62A13FB87FC60FDC0E502" ID-GBIF-Occurrence="3319300911" collectingDate="1988-09-28" collectingDateMax="1988-10-04" collectingDateMin="1988-09-28" collectionCode="NMNH" collectorName="Wolda" country="Panama" county="Canal Zone" location="Snyder-Molino" municipality="Barro Colorado Island" pageId="35" pageNumber="36" specimenCount="1" specimenCount-male="1" stateProvince="Panama" typeStatus="paratype">
<emphasis id="EEBDEAACFFB62A13FB87FC60F91AE552" bold="true" box="[1062,1322,1004,1030]" pageId="35" pageNumber="36">
<collectingCountry id="A4DE762EFFB62A13FB87FC60F89BE552" box="[1062,1195,1004,1030]" name="Panama" pageId="35" pageNumber="36">PANAMA</collectingCountry>
:
<collectingRegion id="1E0DF85CFFB62A13FB1BFC60F915E552" box="[1210,1317,1004,1030]" country="Panama" name="Panama" pageId="35" pageNumber="36">Panama</collectingRegion>
:
</emphasis>
<collectingCounty id="35174E32FFB62A13FEA9FB98FDA2E57A" box="[264,402,1044,1070]" pageId="35" pageNumber="36">Canal Zone</collectingCounty>
,
<collectingMunicipality id="3C12ACC4FFB62A13FE3CFB98FE95E57A" box="[413,677,1044,1070]" pageId="35" pageNumber="36">Barro Colorado Island</collectingMunicipality>
,
<location id="D9166065FFB62A13FD10FB98FF54E57A" LSID="urn:lsid:plazi:treatment:546087A8FFB72A13FEA9FE37FD72E59A:D9166065FFB62A13FD10FB98FF54E57A" box="[689,868,1044,1070]" country="Panama" county="Canal Zone" municipality="Barro Colorado Island" name="Snyder-Molino" pageId="35" pageNumber="36" stateProvince="Panama">Snyder­Molino</location>
trail, marker 3,
<date id="A877107EFFB62A13FBBCFB98F8E3E57A" box="[1053,1235,1044,1070]" pageId="35" pageNumber="36" value="1988-09-28" valueMax="1988-10-04" valueMin="1988-09-28">
<collectingDate id="B833E996FFB62A13FBBCFB98F8E3E57A" box="[1053,1235,1044,1070]" pageId="35" pageNumber="36" value="1988-09-28" valueMax="1988-10-04" valueMin="1988-09-28">28.ix.­4.x.1988</collectingDate>
</date>
,
<collectorName id="713C5368FFB62A13FB7EFB98F91BE57A" box="[1247,1323,1044,1070]" pageId="35" pageNumber="36">Wolda</collectorName>
<specimenCount id="CACFFD37FFB62A13FE8BFBB0FD48E502" box="[298,376,1084,1110]" pageId="35" pageNumber="36" type="male">1 male</specimenCount>
(
<collectionCode id="BAD8AE7BFFB62A13FE29FBB0FDD8E502" box="[392,488,1084,1110]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="35" pageNumber="36" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="DC7636BEFFB62A13FE99FBE8FEF7E52A" blockId="35.[264,1324,284,1230]" box="[312,711,1124,1150]" pageId="35" pageNumber="36">
Distribution.
<collectingCountry id="A4DE762EFFB62A13FE75FBE8FE66E52A" box="[468,598,1124,1150]" name="Costa Rica" pageId="35" pageNumber="36">Costa Rica</collectingCountry>
,
<collectingCountry id="A4DE762EFFB62A13FDC3FBE8FEF3E52A" box="[610,707,1124,1150]" name="Panama" pageId="35" pageNumber="36">Panama</collectingCountry>
.
</paragraph>
<paragraph id="DC7636BEFFB62A13FE99FB00FD72E59A" blockId="35.[264,1324,284,1230]" pageId="35" pageNumber="36">
Etymology. This species is named for Roger J. Blahnik, who collected the
<typeStatus id="0372881CFFB62A13FB0DFB00F8EEE5F2" box="[1196,1246,1164,1190]" pageId="35" pageNumber="36">type</typeStatus>
specimen.
</paragraph>
</subSubSection>
</treatment>
</document>