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<document ID-CLB-Dataset="193454" ID-DOI="http://dx.doi.org/10.3897/nl.45.94938" ID-GBIF-Dataset="77c88040-8a55-4fac-85e4-f8393c17ff4e" ID-Pensoft-Pub="2367-5365-45-315" ID-Pensoft-UUID="E5B91E93DB905CFF93FE3F2EDFC544D4" ID-ZooBank="A8BBBDA3D0C048B2B8C469E37531B762" ModsDocID="2367-5365-45-315" checkinTime="1669268527460" checkinUser="pensoft" docAuthor="Mally, Richard, Aarvik, Leif, Karisch, Timm, Lees, David C. &amp; Malm, Tobias" docDate="2022" docId="545CF07C8E0450BB9159B8B528811753" docLanguage="en" docName="NotaLepidopt 45: 315-353" docOrigin="Nota Lepidopterologica 45" docPubDate="2022-11-23" docSource="http://dx.doi.org/10.3897/nl.45.94938" docTitle="Udea hageni Viette 1952" docType="treatment" docVersion="5" id="E5B91E93DB905CFF93FE3F2EDFC544D4" lastPageNumber="315" masterDocId="E5B91E93DB905CFF93FE3F2EDFC544D4" masterDocTitle="Revision of Afrotropical Udea Guenee in Duponchel, 1845, with description of five new species of the U. ferrugalis (Huebner, 1796) group (Lepidoptera, Crambidae, Spilomelinae)" masterLastPageNumber="353" masterPageNumber="315" pageNumber="315" updateTime="1732937943148" updateUser="ExternalLinkService">
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<mods:title id="34D9F11B3590217057974EEE98EE19A9">Revision of Afrotropical Udea Guenee in Duponchel, 1845, with description of five new species of the U. ferrugalis (Huebner, 1796) group (Lepidoptera, Crambidae, Spilomelinae)</mods:title>
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<mods:namePart id="E2581F73C24288789B6B99394E0B3EC1">Mally, Richard</mods:namePart>
<mods:nameIdentifier id="2A5520530712346BE6F226E69C5999F4" type="ORCID">https://orcid.org/0000-0001-5996-9471</mods:nameIdentifier>
<mods:affiliation id="7EFFACC06BA76E3E771560D3E336A3F1">Department of Forest Protection and Entomology, Czech University of Life Sciences, Prague, Kamycka 129, 165 00 Prague 6 - Suchdol, Czech Republic; mally @ fld. czu. cz</mods:affiliation>
<mods:nameIdentifier id="C97DDAFBA8ACE7C9F086CD21EF1A24A7" type="email">mally@fld.czu.cz</mods:nameIdentifier>
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<mods:namePart id="71F236AE16188658EDF87C1009271288">Aarvik, Leif</mods:namePart>
<mods:nameIdentifier id="EEE4D6C233EF778BFF1789AFC63C01FD" type="ORCID">https://orcid.org/0000-0002-0112-8837</mods:nameIdentifier>
<mods:affiliation id="3AF6B25B6F26F1E691B00129F75D0A23">Natural History Museum, University of Oslo, Oslo, Norway; leif. aarvik @ nhm. uio. no</mods:affiliation>
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<mods:namePart id="D6C1282EE7C4B72A56D34B5C45E2021B">Karisch, Timm</mods:namePart>
<mods:affiliation id="2AE08A24F48F94BDE42512AB9C4184E3">Museum fuer Naturkunde und Vorgeschichte, Dessau, Germany; Timm. Karisch @ naturkunde. dessau. de</mods:affiliation>
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<mods:namePart id="6F4523CBFA4EBB4292270DA03DC5C35A">Lees, David C.</mods:namePart>
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<mods:affiliation id="ACE92D646619C28C1564AA0D51FE9890">Insects Division, Natural History Museum, Cromwell Road, SW 7 5 BD, London, UK; david. lees @ nhm. ac. uk</mods:affiliation>
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<mods:namePart id="E82223FAAFD01C3DD124C0605EFA20AD">Malm, Tobias</mods:namePart>
<mods:affiliation id="ADD245B18464523E2C0CAD8327921D5E">Naturhistoriska Riksmuseet, Stockholm, Sweden; tobias. malm @ nrm. se</mods:affiliation>
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<mods:date id="EC05B94E1EC1E8417FE452910D07D069">2022</mods:date>
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<mods:number id="F5D6814C76173389491E6AF100A4EDE0">45</mods:number>
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<treatment id="545CF07C8E0450BB9159B8B528811753" ID-GBIF-Taxon="205134404" LSID="urn:lsid:plazi:treatment:545CF07C8E0450BB9159B8B528811753" httpUri="http://treatment.plazi.org/id/545CF07C8E0450BB9159B8B528811753" lastPageNumber="315" pageId="0" pageNumber="315" scope_family="Crambidae" scope_order="Lepidoptera" scope_subFamily="Spilomelinae">
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<paragraph id="3FB5A4AC9C1899E65EDDF6F7A55C1728" pageId="0" pageNumber="315">
<taxonomicName id="16142B2F0B403A8A0E42787801E724BD" ID-CoL="7DDN6" LSID="545CF07C-8E04-50BB-9159-B8B528811753" authority="Viette, 1952" authorityName="Viette" authorityYear="1952" class="Insecta" family="Crambidae" genus="Udea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udea hageni" order="Lepidoptera" pageId="0" pageNumber="315" phylum="Arthropoda" rank="species" species="hageni">Udea hageni Viette, 1952</taxonomicName>
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<paragraph id="7DDD5961B20C52EC598F1596FDD37D0C" pageId="0" pageNumber="315">
<figureCitation id="2BC65E15F560D5B185691482CD9B7610" captionStart="Figures 15" captionStartId="F1" captionText="Figures 1 - 5. Adults of Afrotropical Udea species. 1. U. delineatalis (Walker in Melliss 1875); a. ♂ (SDEI); b. cotype ♀ (NHMUK); 2. Phlyctaenia epicoena Meyrick, 1937 syn. nov. of U. ferrugalis (Huebner, 1796), paralectotype ♀ (NHMUK); 3. U. hageni Viette, 1952, ♂ (NHMO); 4. U. kirinyaga Mally, sp. nov., holotype ♂ (NHMO); 5. U. momella Mally, sp. nov., holotype ♂, abdomen removed (ZSM). Scale bar represents 5 mm, all specimens to scale." figureDoi="10.3897/nl.45.94938.figures1-5" httpUri="https://binary.pensoft.net/fig/773284" pageId="0" pageNumber="315">Figs 3</figureCitation>
<figureCitation id="5F119AE84768B46F8F69D4E74B3FEC1B" captionStart="Figures 1120" captionStartId="F3" captionText="Figures 11 - 20. Tympanal organs of Afrotropical Udea species and species removed from Udea. 11. U. delineatalis (Walker in Melliss 1875), ♂; 12. U. ferrugalis (Huebner, 1796), ♂; 13. U. hageni Viette, 1952, ♂; 14. U. kirinyaga Mally, sp. nov., ♂; 15. U. momella Mally, sp. nov., ♂; 16. U. nicholsae Mally, sp. nov., ♀; 17. U. meruensis Mally, sp. nov., ♂; 18. U. namaquana Karisch &amp; Mally, sp. nov., ♂; 19. Achyra melanostictalis (Hampson in Poulton 1916), ♀; 20. Lirabotys infuscalis (Zeller, 1852), ♂. Scale bar represents 500 μm, all figures to scale." figureDoi="10.3897/nl.45.94938.figures11-20" httpUri="https://binary.pensoft.net/fig/773286" pageId="0" pageNumber="315">, 13</figureCitation>
<figureCitation id="959F52B8D856D86F2FDBBD6FE0F22974" captionStart="Figures 2123" captionStartId="F4" captionText="Figures 21 - 23. Male genitalia of Udea species, with male genitalia (a), phallus (b), and posterior phallus with cornutus (c). 21. U. delineatalis (Walker in Melliss 1875); 22. U. ferrugalis (Huebner, 1796); 23. U. hageni Viette, 1852. Scale bars: 500 μm (21 a, b- 23 a, b); 100 μm (21 c- 23 c)." figureDoi="10.3897/nl.45.94938.figures21-23" httpUri="https://binary.pensoft.net/fig/773287" pageId="0" pageNumber="315">, 23</figureCitation>
</paragraph>
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<subSubSection id="4A91F7A635F3249EF845A9866E75AD57" pageId="0" pageNumber="315" type="reference_group">
<paragraph id="D487FB5528662DA8AABCEA3E11C2F470" pageId="0" pageNumber="315">
<taxonomicName id="B3799DBA8CA193C660893969A1C1A27D" authorityName="Viette" authorityYear="1952" class="Insecta" family="Crambidae" genus="Udea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udea hageni" order="Lepidoptera" pageId="0" pageNumber="315" phylum="Arthropoda" rank="species" species="hageni">Udea hageni</taxonomicName>
Viette, 1952: 3-5, figs 1, 2. Type locality: Tristan da Cunha.
</paragraph>
</subSubSection>
<subSubSection id="4503C7AA8E17AFC38B4CC316CD03C119" pageId="0" pageNumber="315" type="materials_examined">
<paragraph id="831809ACD991D2C64B39495CAFB625C7" pageId="0" pageNumber="315">Material examined.</paragraph>
<paragraph id="CB692E2D45582B61D73FBADC5DAA8886" pageId="0" pageNumber="315">
<materialsCitation id="38344594B41A11DF8DF5A8C874BBC3EF" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Tristan Da Cunha" location="Tristan Da Cunha" specimenCount="1" specimenCount-male="1">
<collectingCountry id="75CEFA295F3B8C0D7092A1EB935D7615" name="Saint Helena, Ascension and Tristan da Cunha">Tristan Da Cunha</collectingCountry>
<specimenCount id="A5FF542D0E84DBFBBA86351846FA633D" type="male">1 ♂</specimenCount>
;
<normalizedToken id="113346815B94D24126526216A58A4178" originalValue="“St.75”">&quot;St.75&quot;</normalizedToken>
, &quot;Gm 1075 ZM.Oslo&quot;, &quot;
<taxonomicName id="3F39128F32E18E373022E765B63955F8" authorityName="Viette" authorityYear="1952" class="Insecta" family="Crambidae" genus="Udea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udea hageni" order="Lepidoptera" pageId="0" pageNumber="315" phylum="Arthropoda" rank="species" species="hageni">
<emphasis id="273ACA1623F251E206F8426A1FC484B4" italics="true" pageId="0" pageNumber="315">Udea hageni</emphasis>
</taxonomicName>
VH.&quot;; DNA voucher ZMBN Lep520; Mally genitalia dissection no. 1161; NHMO
</materialsCitation>
.
</paragraph>
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<subSubSection id="ECB2C03849759FB091387125392A769F" pageId="0" pageNumber="315" type="diagnosis">
<paragraph id="74AB28AEFF602282645810C64B499BD5" pageId="0" pageNumber="315">Diagnosis.</paragraph>
<paragraph id="FF52EB600085B557F3578778732DBB5D" pageId="0" pageNumber="315">
The species is readily distinguished from other
<taxonomicName id="1AE5836B0AEDF04C79C5FF82AC51D623" class="Insecta" family="Crambidae" genus="Udea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udea" order="Lepidoptera" pageId="0" pageNumber="315" phylum="Arthropoda" rank="genus">
<emphasis id="FB332915EC11694263D01D2A7B51F880" italics="true" pageId="0" pageNumber="315">Udea</emphasis>
</taxonomicName>
species by the narrow, apically pointed fore- and hindwings, more reminiscent of some
<taxonomicName id="F2646032A7D05F3867F2E177A6CDD8C6" class="Insecta" family="Tortricidae" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="315" phylum="Arthropoda" rank="family">Tortricidae</taxonomicName>
than of
<taxonomicName id="BB410C843190050E48ABEC6F82854CD5" lsidName="" pageId="0" pageNumber="315" rank="subFamily" subFamily="Spilomelinae">Spilomelinae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="63DEAF371B4623737AF76CC8F23FD4A2" pageId="0" pageNumber="315" type="description">
<paragraph id="705DB42DA022756EBF7ABF0CFAE7F892" pageId="0" pageNumber="315">Description of adults.</paragraph>
<paragraph id="95EA9C4D76B93D6AE512A7D87DA953B3" pageId="0" pageNumber="315">
<emphasis id="CB3EC2010B48F4A26CE3BF82B8D93F48" bold="true" italics="true" pageId="0" pageNumber="315">Head</emphasis>
: Dorsally greyish-white with interspersed brownish scales, ventral side beige; labial palps porrect, elongate triangular, about twice as long as eye diameter, dorsal and outer sides brown, ventral and inner sides as well as outer
<normalizedToken id="C794E258F8F3BB3B8B875C1CA9ECBEA7" originalValue="sides">side's</normalizedToken>
base of first meron and a narrow ventral area on second meron beige; maxillary palps brown, well developed, long enough to touch each other at apex, terminal meron with spatulate brush of long scales; haustellum fully developed, basally with cream-coloured scales; frons greyish-brown, slightly rounded; compound eyes large, hemispherical; antennae ochre-brown, scapus short, hidden in scales of vertex, pedicellus large, somewhat swollen, flagellum in male anteroventrally with dense ciliation almost half as long as flagellum width; vertex in centre and behind ocelli with tuft of long cream-coloured scales.
<emphasis id="3E918D15DED3AEE8000DE0A22044AFAA" bold="true" italics="true" pageId="0" pageNumber="315">Thorax</emphasis>
: Prothorax, tegulae, forelegs and anterior mesothorax brown, posterior mesothorax, metathorax and mid- and hindlegs light grey. Outer tibial spur 2/3 length of inner spur in midleg, half as long as
<normalizedToken id="7A88CF2A75DBEFD95E8AAC371D88CB96" originalValue="hindlegs">hindlegs'</normalizedToken>
proximal spur pair, and almost as long as inner spur in distal spur pair.
<emphasis id="695EB8B5AC7D9EF50DB6AC828842BC1A" bold="true" italics="true" pageId="0" pageNumber="315">Wings</emphasis>
: (Fig.
<figureCitation id="53CC96292618095BDF8D334348DAE035" captionStart="Figures 15" captionStartId="F1" captionText="Figures 1 - 5. Adults of Afrotropical Udea species. 1. U. delineatalis (Walker in Melliss 1875); a. ♂ (SDEI); b. cotype ♀ (NHMUK); 2. Phlyctaenia epicoena Meyrick, 1937 syn. nov. of U. ferrugalis (Huebner, 1796), paralectotype ♀ (NHMUK); 3. U. hageni Viette, 1952, ♂ (NHMO); 4. U. kirinyaga Mally, sp. nov., holotype ♂ (NHMO); 5. U. momella Mally, sp. nov., holotype ♂, abdomen removed (ZSM). Scale bar represents 5 mm, all specimens to scale." figureDoi="10.3897/nl.45.94938.figures1-5" httpUri="https://binary.pensoft.net/fig/773284" pageId="0" pageNumber="315">3</figureCitation>
) Forewing length of male 8 mm. Wings with reduced area compared to other
<taxonomicName id="775B4319E13BAFEF69325DA423CBFCB3" class="Insecta" family="Crambidae" genus="Udea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udea" order="Lepidoptera" pageId="0" pageNumber="315" phylum="Arthropoda" rank="genus">
<emphasis id="BC2934C1FD61C0891731BA650E938DE6" italics="true" pageId="0" pageNumber="315">Udea</emphasis>
</taxonomicName>
species. Male with single frenulum bristle, condition unknown for female. Forewing slender, with long convex costal margin, acute apex of about 60°, relatively short, straight termen, and short, convex anal margin. Ground colour cream-beige, with broad straight streak of brown colour from forewing base to 2/3 length of forewing in direction of termen-anal angle, second streak from basal of proximal discal stigma over distal stigma, broadening to area between
<normalizedToken id="65FE150264B902C4205AF92C4EB35416" originalValue="forewings">forewing's</normalizedToken>
apex and mid-termen; discoidal stigmata dark brown, proximal discoidal stigma oval, distal stigma a less clearly defined larger spot on anal side of second brown straight forewing streak; no antemedial line present, postmedial line very vague, as an outward-arching line from beyond distal discoidal stigma to distal end of broad straight streak from wing base; distal costa and termen at ends of wing veins with streak-like dark brown dots; fringe cream-coloured. Hindwing ground colour cream-beige, distal area suffused with grey-brown, veins brownish; faint brownish discoidal stigmata, connected by a thin brownish line; termen and anal margin with interrupted brown line; fringe cream-coloured. Underside of forewing more or less uniformly beige-grey, with upper
<normalizedToken id="819311BF38209084DCEA5EA8460AF9EB" originalValue="sides">side's</normalizedToken>
broad streak from wing base as well as distal discoidal stigma as somewhat darker areas; ends of wing veins with streak-like dark brown dots; fringe pale beige; underside of hindwing uniformly pale beige, with small central brown discal spot; vein ends and fringe as in forewing underside.
<emphasis id="908A5F7F9CF0F8EEB8D63C61CBAFAE3C" bold="true" italics="true" pageId="0" pageNumber="315">Abdomen</emphasis>
: Dark to light grey. Tympanal organs (Fig.
<figureCitation id="5AD5682E92665ED114833164A24F4432" captionStart="Figures 1120" captionStartId="F3" captionText="Figures 11 - 20. Tympanal organs of Afrotropical Udea species and species removed from Udea. 11. U. delineatalis (Walker in Melliss 1875), ♂; 12. U. ferrugalis (Huebner, 1796), ♂; 13. U. hageni Viette, 1952, ♂; 14. U. kirinyaga Mally, sp. nov., ♂; 15. U. momella Mally, sp. nov., ♂; 16. U. nicholsae Mally, sp. nov., ♀; 17. U. meruensis Mally, sp. nov., ♂; 18. U. namaquana Karisch &amp; Mally, sp. nov., ♂; 19. Achyra melanostictalis (Hampson in Poulton 1916), ♀; 20. Lirabotys infuscalis (Zeller, 1852), ♂. Scale bar represents 500 μm, all figures to scale." figureDoi="10.3897/nl.45.94938.figures11-20" httpUri="https://binary.pensoft.net/fig/773286" pageId="0" pageNumber="315">13</figureCitation>
) with lobulus absent.
<emphasis id="ED04D91881B4ADFBBB24A828891B3239" bold="true" italics="true" pageId="0" pageNumber="315">Male genitalia</emphasis>
: (Fig.
<figureCitation id="994DD3682D653BF650A5E1547D4AB8F1" captionStart="Figures 2123" captionStartId="F4" captionText="Figures 21 - 23. Male genitalia of Udea species, with male genitalia (a), phallus (b), and posterior phallus with cornutus (c). 21. U. delineatalis (Walker in Melliss 1875); 22. U. ferrugalis (Huebner, 1796); 23. U. hageni Viette, 1852. Scale bars: 500 μm (21 a, b- 23 a, b); 100 μm (21 c- 23 c)." figureDoi="10.3897/nl.45.94938.figures21-23" httpUri="https://binary.pensoft.net/fig/773287" pageId="0" pageNumber="315">23</figureCitation>
) Circular uncus head with stiff bifid, anteriad directed chaetae on dorsal surface, neck slender, elongate, base of uncus broadly attached to tegumen and each side with a small laterad protrusion and notch at attachment to tegumen; tegumen rectangular; transtilla arms short, rounded, far apart from each other; vinculum large, together with saccus forming an oval; saccus broad U-shaped; juxta small, broadly drop-shaped, dorsally with V-shaped longitudinal membranous
<normalizedToken id="54963FB8347462095FC6D003EDABE62B" originalValue="“notch”">&quot;notch&quot;</normalizedToken>
reaching mid-length; valvae elongate, slender, tapering towards apex; costa concave, base broad, slightly inflated, valva apex evenly rounded, ventral valva margin concave in apical half, at sacculus convex; sacculus elongate, ventrodistal tip in close association to central fibula, distal edge straight; fibula emerging near costa base from an elongate base, forming slender posteriad claw-like structure reaching towards distal edge of sacculus, not reaching ventral margin of valva. Phallus slender, thinnest at posterior end, anterior end stout, without coecum; central section with somewhat stronger sclerotisation; posterior tip of phallus apodeme ventrally with short triangular tooth; vesica with short, slender, spine-shaped cornutus.
<emphasis id="049BC93A2B6D542D4F748FF1CFF068FA" bold="true" italics="true" pageId="0" pageNumber="315">Female genitalia</emphasis>
: Unknown.
</paragraph>
</subSubSection>
<subSubSection id="17BD5E6188AB8A0BDE71B3BEC17C249F" pageId="0" pageNumber="315" type="distribution">
<paragraph id="D448D2715A723CB215472BFA9E67B316" pageId="0" pageNumber="315">Distribution.</paragraph>
<paragraph id="B411DE819187D62B5BF1F98C266C3B99" pageId="0" pageNumber="315">Only known from the mid-Atlantic island of Tristan da Cunha.</paragraph>
</subSubSection>
<subSubSection id="253DEA483F9CEB33586F67FA1461E6C3" pageId="0" pageNumber="315" type="genetic data">
<paragraph id="2613F2E5140454385CBC7954D8426D14" pageId="0" pageNumber="315">Genetic data.</paragraph>
<paragraph id="950B64C2BBF7DE3EBC5DA5425DC6B1FD" pageId="0" pageNumber="315">Not available.</paragraph>
</subSubSection>
<subSubSection id="139019019BD8E1D7BE8A49DE566DAAB3" pageId="0" pageNumber="315" type="remarks">
<paragraph id="A4D52128AE840A6C4A852A953558F5AD" pageId="0" pageNumber="315">Remarks.</paragraph>
<paragraph id="3C6DE90C0C3AD763D9C7CAC7C3897B04" pageId="0" pageNumber="315">
Based on the male genitalia illustrated in
<bibRefCitation id="82847C3496BD47E7321FCAE9B348890E" author="Viette, P" journalOrPublisher="Journal of the Entomological Society of South Africa" pageId="0" pageNumber="315" refId="B70" refString="Viette, P, 1952. Lepidoptera. Results of the Norwegian scientific expedition to Tristan da Cunha 1937-1938 23: 1-19. [pls 1-3, map.]" title="Lepidoptera. Results of the Norwegian scientific expedition to Tristan da Cunha 1937 - 1938 23: 1 - 19. [pls 1 - 3, map.]" year="1952">Viette (1952)</bibRefCitation>
,
<bibRefCitation id="2DC769278CB0B168E7F5B053E1D823EB" author="Mally, R" journalOrPublisher="Arthropod Systematics &amp; Phylogeny" pageId="0" pageNumber="315" pagination="55 - 71" publicationUrl="https://www.senckenberg.de/wp-content/uploads/2019/08/asp_69_1_mally_55-71.pdf" refId="B36" refString="Mally, R, Nuss, M, 2011. Molecular and morphological phylogeny of European Udea moths (Insecta: Lepidoptera: Pyraloidea). Arthropod Systematics &amp; Phylogeny 69 (1): 55 - 71, https://www.senckenberg.de/wp-content/uploads/2019/08/asp_69_1_mally_55-71.pdf" title="Molecular and morphological phylogeny of European Udea moths (Insecta: Lepidoptera: Pyraloidea)." url="https://www.senckenberg.de/wp-content/uploads/2019/08/asp_69_1_mally_55-71.pdf" volume="69" year="2011">Mally and Nuss (2011)</bibRefCitation>
placed
<taxonomicName id="487EBA1591F03FC60FBBEE18680681FB" authorityName="Viette" authorityYear="1952" class="Insecta" family="Crambidae" genus="Udea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udea hageni" order="Lepidoptera" pageId="0" pageNumber="315" phylum="Arthropoda" rank="species" species="hageni">
<emphasis id="7585D267524BF811AF39A90806B6A9ED" italics="true" pageId="0" pageNumber="315">Udea hageni</emphasis>
</taxonomicName>
in the
<taxonomicName id="573EFE27009A049580083CF3BB06F043" class="Insecta" family="Amaranthaceae" kingdom="Animalia" lsidName="U. ferrugalis" order="Lepidoptera" pageId="0" pageNumber="315" phylum="Arthropoda" rank="species" species="ferrugalis">
<emphasis id="89436489D7D374A850F290E1B889D70C" italics="true" pageId="0" pageNumber="315">U. ferrugalis</emphasis>
</taxonomicName>
species group. However, examination of a specimen and the results of our phylogenetic analysis suggest that the species is a lineage of the
<taxonomicName id="58D4DF8C1A2E6ECC536B5187B744584F" class="Insecta" family="Amaranthaceae" kingdom="Animalia" lsidName="U. numeralis" order="Lepidoptera" pageId="0" pageNumber="315" phylum="Arthropoda" rank="species" species="numeralis">
<emphasis id="7ADEA8EA42793592E28D0B05C47EC3AB" italics="true" pageId="0" pageNumber="315">U. numeralis</emphasis>
</taxonomicName>
species group. So far,
<taxonomicName id="7D74B2ECF3D84CF662EBA86222B45722" class="Insecta" family="Amaranthaceae" kingdom="Animalia" lsidName="U. hageni" order="Lepidoptera" pageId="0" pageNumber="315" phylum="Arthropoda" rank="species" species="hageni">
<emphasis id="F8A4CEC1FC8E51933216A448537194AA" italics="true" pageId="0" pageNumber="315">U. hageni</emphasis>
</taxonomicName>
is the only island-endemic
<taxonomicName id="AB44348623A953F5E9DF374BA55CF7C0" class="Insecta" family="Crambidae" genus="Udea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udea" order="Lepidoptera" pageId="0" pageNumber="315" phylum="Arthropoda" rank="genus">
<emphasis id="28035B1208C2BA60E4948B08D4019555" italics="true" pageId="0" pageNumber="315">Udea</emphasis>
</taxonomicName>
species with marked wing reduction. To our knowledge, the female sex of this species is still unknown.
</paragraph>
</subSubSection>
</treatment>
</document>