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<document id="53A4BC87C54F6409A8F9416857934E9E" ID-DOI="10.11646/phytotaxa.184.4.4" ID-ISSN="1179-3163" ID-Zenodo-Dep="5146779" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1627600570188" checkinUser="felipe" docAuthor="Valdespino, Iván A., López, Christian &amp; De Araújo Góes-Neto, Luiz A." docDate="2014" docId="545B2A589757FFBAA2FFFB37FBF3FD53" docLanguage="en" docName="Phytotaxa.184.4.235-244.pdf" docOrigin="Phytotaxa 184 (4)" docSource="http://dx.doi.org/10.11646/phytotaxa.184.4.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Selaginella myriostachya Valdespino, C. Lopez &amp; L. Goes 2014, sp. nov." docType="treatment" docVersion="3" lastPageNumber="239" masterDocId="A86252209756FFBEA277FFA3FFBEFFDF" masterDocTitle="Additions to Cuban Selaginella (Selaginellaceae)" masterLastPageNumber="244" masterPageNumber="235" pageNumber="236" updateTime="1699030341196" updateUser="plazi" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="77B0029FEBB311882D97AFA51533F9D7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="01E68E2AD47533F9F01D497D47110BA2">
<mods:title id="012170D0ED19B7F2B82CAB10FEE08651">Additions to Cuban Selaginella (Selaginellaceae)</mods:title>
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<mods:name id="0B08BD2582BA11BEC6B1F5951672B19D" type="personal">
<mods:role id="EA123C5E6ECDCB1FD35AE1EA755D56F4">
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<mods:namePart id="B5BA3811ABF8A624F9EA02FB52060503">Valdespino, Iván A.</mods:namePart>
<mods:affiliation id="DE0DE0A58E44A55E3046C3618B2AAC72">Departamento de Botánica, Facultad de Ciencias Naturales, Exactas y Tecnología, Universidad de Panamá, Apartado Postal 0824 - 00073, Panamá, Panamá. E-mail: iavaldespino @ gmail. com</mods:affiliation>
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<mods:namePart id="A2BE22C5FC5DBD5C3A59CA3D7DE7C6C3">López, Christian</mods:namePart>
<mods:affiliation id="63EA25BC4AEDE4A6E0C76662E59EE4BF">Departamento de Botánica, Facultad de Ciencias Naturales, Exactas y Tecnología, Universidad de Panamá, Apartado Postal 0824 - 00073, Panamá, Panamá. E-mail: iavaldespino @ gmail. com &amp; Apartado Postal 0819 - 12099, El Dorado. Panamá, Panamá. E-mail: adolfolopezs @ hotmail. com</mods:affiliation>
</mods:name>
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<mods:role id="3898ED88CEF23301E4A5FC67E925884C">
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</mods:role>
<mods:namePart id="47158EF5D36F9330D0BF890ED5DDE359">De Araújo Góes-Neto, Luiz A.</mods:namePart>
<mods:affiliation id="FF31A12B62FF975D31FE73843A564B8A">Departamento de Botânica, Instituto de Ciências Biológicas, Universidade Federal de Minas Gerais, CP 486, 31270 - 901, Belo Horizonte, MG, Brazil. E-mail: lgoes-neto @ hotmail. com</mods:affiliation>
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<mods:typeOfResource id="4A8D9C0C659D76939CC2608AF57EB977">text</mods:typeOfResource>
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<mods:title id="74209F490303A27D3DA43713F92BCD6D">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="DF26F4CDEC2C5AFE28B2884E35657B8B">
<mods:date id="3DFB2A22E83FED4FE4698D9256E148C1">2014</mods:date>
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<mods:number id="A16880F55ED8729DBD3339A58AE51407">2014-11-07</mods:number>
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<mods:detail id="92C6975770E8624AC4A563BB1D8A4D22" type="volume">
<mods:number id="C3931C0053FFF00E5421289695552F41">184</mods:number>
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<mods:number id="A819453E3A97DCE1500AAC329521CF24">4</mods:number>
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<mods:start id="00CD1526C430AA315BE9079D1E475E4B">235</mods:start>
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<mods:url id="7F29E657D2038C7E7A3527EC3095BCCB">http://dx.doi.org/10.11646/phytotaxa.184.4.4</mods:url>
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<mods:classification id="85862158BFE1A10879A7935265E07F0B">journal article</mods:classification>
<mods:identifier id="065925009B9434CA562660E46F1768A9" type="DOI">10.11646/phytotaxa.184.4.4</mods:identifier>
<mods:identifier id="31D6C0D657249BA638BAB27886DBCD4B" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="A4CD708920C1A709E29011A67CD059B6" type="Zenodo-Dep">5146779</mods:identifier>
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<treatment id="545B2A589757FFBAA2FFFB37FBF3FD53" ID-DOI="http://doi.org/10.5281/zenodo.5150162" ID-Zenodo-Dep="5150162" LSID="urn:lsid:plazi:treatment:545B2A589757FFBAA2FFFB37FBF3FD53" httpUri="http://treatment.plazi.org/id/545B2A589757FFBAA2FFFB37FBF3FD53" lastPageId="4" lastPageNumber="239" pageId="1" pageNumber="236">
<subSubSection id="94E8C8C59757FFBFA2FFFB37FBA7FB71" box="[136,1049,1172,1199]" pageId="1" pageNumber="236" type="nomenclature">
<paragraph id="DC4D9B4E9757FFBFA2FFFB37FBA7FB71" blockId="1.[136,1049,1172,1199]" box="[136,1049,1172,1199]" pageId="1" pageNumber="236">
<heading id="87052C229757FFBFA2FFFB37FBA7FB71" box="[136,1049,1172,1199]" fontSize="11" level="2" pageId="1" pageNumber="236" reason="3">
<taxonomicName id="1BF2E0CD9757FFBFA2FFFB37FC9FFB71" ID-CoL="4WC5R" authority="Valdespino, C. Lopez &amp; L. Goes" authorityName="Valdespino, C. Lopez &amp; L. Goes" authorityYear="2014" box="[136,801,1172,1199]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="1" pageNumber="236" phylum="Tracheophyta" rank="species" species="myriostachya" status="sp. nov.">
<emphasis id="EE86475C9757FFBFA2FFFB37FE14FB70" bold="true" box="[136,426,1172,1199]" italics="true" pageId="1" pageNumber="236">Selaginella myriostachya</emphasis>
Valdespino, C. López &amp; L. Góes
</taxonomicName>
,
<emphasis id="EE86475C9757FFBFA15AFB36FC3CFB71" box="[813,898,1173,1198]" italics="true" pageId="1" pageNumber="236">
<taxonomicNameLabel id="F5B5FA279757FFBFA15AFB36FC3CFB71" box="[813,898,1173,1198]" pageId="1" pageNumber="236" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="44C987CB9757FFBFA1E7FB37FC63FB71" box="[912,989,1172,1198]" captionStart="FIGURE 1" captionStartId="2.[136,229,1953,1975]" captionTargetBox="[173,1415,183,1922]" captionTargetId="figure-21@2.[173,1415,183,1922]" captionTargetPageId="2" captionText="FIGURE 1. SEM micrographs of stem section and leaves below first branch of Selaginella myriostachya (Valdespino et al. 2013-1, PMA). A. Stem section showing appressed, slightly dimorphic median (m) and lateral (l) leaves. B. Close up of A. C. Upper surface of median leaf. D. Close up of apical region of median leaf upper surface. E. Close up of basal region of median leaf upper surface. F. Stem section showing appressed lateral leaves. G. Lower surface of lateral leaf. H. Close up of mid region of lateral leaf lower surface." figureDoi="http://doi.org/10.5281/zenodo.5146781" httpUri="https://zenodo.org/record/5146781/files/figure.png" pageId="1" pageNumber="236">Figs. 1</figureCitation>
&amp;
<figureCitation id="44C987CB9757FFBFA676FB37FBAEFB71" box="[1025,1040,1172,1198]" captionStart="FIGURE 2" captionStartId="3.[136,229,1133,1155]" captionTargetBox="[151,1435,209,1109]" captionTargetId="figure-18@3.[151,1435,209,1109]" captionTargetPageId="3" captionText="FIGURE 2. SEM micrographs of stem and leaves of Selaginella myriostachya (Shafer 13794, NY). A. Upper surface of stem. B. Upper surface of median leaf. C. Lower surface of stem. D. Lower surface of lateral leaf." figureDoi="http://doi.org/10.5281/zenodo.5146785" httpUri="https://zenodo.org/record/5146785/files/figure.png" pageId="1" pageNumber="236">2</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="94E8C8C59757FFBFA2FFFB43FDFEFA5A" pageId="1" pageNumber="236" type="reference_group">
<paragraph id="DC4D9B4E9757FFBFA2FFFB43FDFEFA5A" blockId="1.[136,1452,1248,1486]" pageId="1" pageNumber="236">
<taxonomicName id="1BF2E0CD9757FFBFA2FFFB43FEC0FB2A" ID-CoL="4WC5R" authorityName="Valdespino, C. Lopez &amp; L. Goes" authorityYear="2014" box="[136,382,1248,1269]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="1" pageNumber="236" phylum="Tracheophyta" rank="species" species="myriostachya">
<emphasis id="EE86475C9757FFBFA2FFFB43FEC0FB2A" box="[136,382,1248,1269]" italics="true" pageId="1" pageNumber="236">Selaginella myriostachya</emphasis>
</taxonomicName>
differs from
<taxonomicName id="1BF2E0CD9757FFBFA071FB43FBC5FB29" ID-CoL="4WBS8" authority="(Desvaux ex Poiret 1814: 558) Spring (1843: 234)" authorityName="Spring" authorityPageNumber="234" authorityYear="1843" baseAuthorityName="Desvaux ex Poiret" baseAuthorityPageNumber="558" baseAuthorityYear="1814" box="[518,1147,1248,1270]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="1" pageNumber="236" phylum="Tracheophyta" rank="species" species="flabellum">
<emphasis id="EE86475C9757FFBFA071FB43FDC5FB2A" box="[518,635,1248,1269]" italics="true" pageId="1" pageNumber="236">S. flabellum</emphasis>
(Desvaux ex Poiret 1814: 558)
<bibRefCitation id="B863E6BF9757FFBFA1C8FB43FBC5FB29" author="Spring, A. F." box="[959,1147,1248,1270]" pageId="1" pageNumber="236" pagination="225 - 235" refId="ref6778" refString="Spring, A. F. (1843) Enumeratio Lycopodinearum. Bulletins de L'Academie Royale des Sciences de Bruxelles 10: 135 - 146, 225 - 235." type="journal article" year="1843">Spring (1843: 234)</bibRefCitation>
</taxonomicName>
by its very slightly dimorphic (vs. obviously dimorphic) leaves below the first branch of the stem, ovate-oblong or lanceolate-oblong lateral leaves (vs. narrowly ovate) that are short-ciliate (
<quantity id="1B0A36AB9757FFBFA3A6FA8BFDEBFAE2" box="[465,597,1320,1342]" metricMagnitude="-5" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="1" pageNumber="236" unit="mm" value="0.075" valueMax="0.1" valueMin="0.05">0.05–0.1 mm</quantity>
) along basal ¼ of acroscopic margin (vs. long-ciliate,
<quantity id="1B0A36AB9757FFBFA61DFA8BFB5CFAE2" box="[1130,1250,1320,1342]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="1" pageNumber="236" unit="mm" value="0.35" valueMax="0.5" valueMin="0.2">0.2–0.5 mm</quantity>
, along basal ½), and lanceolate or ovate-elliptic median leaves (vs. narrowly lanceolate) with a weekly developed outer auricle attached to the stem (vs. well-developed and free from the stem).
</paragraph>
</subSubSection>
<subSubSection id="94E8C8C59757FFBDA2FFFA37FECFF986" lastPageId="3" lastPageNumber="238" pageId="1" pageNumber="236" type="materials_examined">
<paragraph id="DC4D9B4E9757FFBFA2FFFA37FDE0FA11" blockId="1.[136,1452,1248,1486]" pageId="1" pageNumber="236">
<emphasis id="EE86475C9757FFBFA2FFFA37FF04FA76" bold="true" box="[136,186,1428,1449]" pageId="1" pageNumber="236">Type</emphasis>
:—
<materialsCitation id="6C9A91139757FFBFA2AFFA37FDE5FA11" collectingDate="2013-07-06" collectionCode="I, PMA, B, HAJB, K, NY, UC" collectorName="I. Valdespino" country="Cuba" county="Vinales" location="Cueva de Santo Tomas" municipality="Parque Nacional Vinales" pageId="1" pageNumber="236" specimenCount="1" stateProvince="Pinar del Rio" typeStatus="holotype">
<collectingCountry id="A4E5DBDE9757FFBFA2AFFA37FEA1FA76" box="[216,287,1428,1449]" name="Cuba" pageId="1" pageNumber="236">CUBA</collectingCountry>
.
<collectingRegion id="1E3655AC9757FFBFA351FA37FE18FA75" box="[294,422,1428,1450]" country="Cuba" name="Pinar del Rio" pageId="1" pageNumber="236">Pinar del Río</collectingRegion>
:
<collectingCounty id="352CE3C29757FFBFA3D9FA37FE46FA75" box="[430,504,1428,1450]" pageId="1" pageNumber="236">Viñales</collectingCounty>
,
<collectingMunicipality id="3C2901349757FFBFA076FA37FD53FA75" box="[513,749,1428,1450]" pageId="1" pageNumber="236">Parque Nacional Viñales</collectingMunicipality>
,
<location id="D92DCD959757FFBFA081FA37FC6FFA75" LSID="urn:lsid:plazi:treatment:545B2A589757FFBAA2FFFB37FBF3FD53:D92DCD959757FFBFA081FA37FC6FFA75" box="[758,977,1428,1450]" country="Cuba" county="Vinales" municipality="Parque Nacional Vinales" name="Cueva de Santo Tomas" pageId="1" pageNumber="236" stateProvince="Pinar del Rio">Cueva de Santo Tomás</location>
,
<date id="A84CBD8E9757FFBFA1AEFA37FBF7FA76" box="[985,1097,1428,1450]" pageId="1" pageNumber="236" value="2013-07-06">
<collectingDate id="B80844669757FFBFA1AEFA37FBF7FA76" box="[985,1097,1428,1450]" pageId="1" pageNumber="236" value="2013-07-06">6 July 2013</collectingDate>
</date>
,
<emphasis id="EE86475C9757FFBFA625FA37FAF5FA76" box="[1106,1355,1428,1449]" italics="true" pageId="1" pageNumber="236">
<collectorName id="7107FE989757FFBFA625FA37FB70FA76" box="[1106,1230,1428,1449]" pageId="1" pageNumber="236">
<collectionCode id="BAE3038B9757FFBFA625FA37FBE3FA76" box="[1106,1117,1428,1449]" country="Romania" lsid="urn:lsid:biocol.org:col:14415" name="&amp;quot;Alexandru Ioan Cuza&amp;quot; University" pageId="1" pageNumber="236" type="Herbarium">I</collectionCode>
. Valdespino
</collectorName>
et al. 2013-1
</emphasis>
(
<typeStatus id="034925EC9757FFBFA721FA37FA12FA75" box="[1366,1452,1428,1450]" pageId="1" pageNumber="236" type="holotype">holotype</typeStatus>
<collectionCode id="BAE3038B9757FFBFA2CAFA1BFF46FA12" box="[189,248,1464,1485]" country="Canada" httpUri="http://grbio.org/cool/a5s1-0agm" name="Provincial Museum of Alberta" pageId="1" pageNumber="236">PMA</collectionCode>
!, isotypes
<collectionCode id="BAE3038B9757FFBFA32DFA1BFED1FA12" box="[346,367,1464,1485]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="1" pageNumber="236" type="Herbarium">B</collectionCode>
!,
<collectionCode id="BAE3038B9757FFBFA309FA1BFE7EFA12" box="[382,448,1464,1485]" country="Cuba" httpUri="http://grbio.org/cool/7xtp-2qmk" name="Jardín Botánico Nacional" pageId="1" pageNumber="236" type="Herbarium">HAJB</collectionCode>
!,
<collectionCode id="BAE3038B9757FFBFA3BAFA1BFE5AFA12" box="[461,484,1464,1485]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="1" pageNumber="236" type="Herbarium">K</collectionCode>
!,
<collectionCode id="BAE3038B9757FFBFA386FA1BFDA4FA12" box="[497,538,1464,1485]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="1" pageNumber="236" type="Herbarium">NY</collectionCode>
!,
<collectionCode id="BAE3038B9757FFBFA050FA1BFDF1FA11" box="[551,591,1464,1486]" country="USA" name="Upjohn Culture Collection" pageId="1" pageNumber="236">UC</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<paragraph id="DC4D9B4E9757FFBDA2FFFA5FFB5EF9CF" blockId="1.[136,1452,1532,2063]" lastBlockId="3.[136,1452,1238,2093]" lastPageId="3" lastPageNumber="238" pageId="1" pageNumber="236">
<emphasis id="EE86475C9757FFBFA2FFFA5FFF6EF9C9" box="[136,208,1532,1558]" italics="true" pageId="1" pageNumber="236">Plants</emphasis>
epipetric.
<emphasis id="EE86475C9757FFBFA33FFA5FFE37F9C9" box="[328,393,1532,1558]" italics="true" pageId="1" pageNumber="236">Stems</emphasis>
suberect to erect, pale-green,
<quantity id="1B0A36AB9757FFBFA0AAFA5FFC81F9C8" box="[733,831,1532,1559]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="1" pageNumber="236" unit="cm" value="12.5" valueMax="20.0" valueMin="5.0">5–20 cm</quantity>
long,
<quantity id="1B0A36AB9757FFBFA1F0FA5FFC44F9C8" box="[903,1018,1532,1559]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="1" pageNumber="236" unit="mm" value="0.9" valueMax="1.0" valueMin="0.8">0.8–1 mm</quantity>
diam., inarticulate, not flagelliform or stoloniferous, 2 or 3-branched.
<emphasis id="EE86475C9757FFBFA39FF983FDCDF9E5" box="[488,627,1568,1594]" italics="true" pageId="1" pageNumber="236">Rhizophores</emphasis>
ventral, restricted to the base, filiform,
<quantity id="1B0A36AB9757FFBFA645F983FB07F9E4" box="[1074,1209,1568,1595]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="1" pageNumber="236" unit="mm" value="0.2" valueMax="0.3" valueMin="0.1">0.1–0.3 mm</quantity>
diam.
<emphasis id="EE86475C9757FFBFA771F982FAEAF9E5" box="[1286,1364,1569,1594]" italics="true" pageId="1" pageNumber="236">Leaves</emphasis>
at stem base very slightly dimorphic (seemingly monomorphic), obviously dimorphic shortly below first branch and throughout the rest of stem, membranaceous.
<emphasis id="EE86475C9757FFBFA07BF9CBFD12F95D" box="[524,684,1640,1666]" italics="true" pageId="1" pageNumber="236">Lateral leaves</emphasis>
distant, spreading to ascending, ovate-oblong or lanceolate-oblong, 1.8–2.2 × 0.8–1.0 mm; base rounded to slightly truncate, acroscopic base overlapping the stem, basiscopic base free from the stem; margins hyaline, 2 or 3 cells wide, the cells elongate and papillate parallel to margin, papillae in a single row over cell lumen, short-ciliate (
<quantity id="1B0A36AB9757FFBFA071F977FD14F930" box="[518,682,1748,1775]" metricMagnitude="-5" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="1" pageNumber="236" unit="mm" value="0.075" valueMax="0.1" valueMin="0.05">0.05–0.10 mm</quantity>
) along basal ¼, otherwise serrate apically; apex cuspidate (
<quantity id="1B0A36AB9757FFBFA74DF977FF0BF8CC" metricMagnitude="-5" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="1" pageNumber="236" unit="mm" value="0.075" valueMax="0.1" valueMin="0.05">0.05–0.10 mm</quantity>
) or acute if tips broken off; both surfaces glabrous, upper surface made up of rounded to quadrangular, sinuatewalled cells, without stomata or idioblasts, lower surface made up of elongated, sinuate-walled cells, with stomata along central portion of midrib and with obscure, papillate idioblasts.
<emphasis id="EE86475C9757FFBFA1CBF8E3FBDCF885" box="[956,1122,1856,1882]" italics="true" pageId="1" pageNumber="236">Median leaves</emphasis>
distant to slightly imbricate apically, ascending, lanceolate or ovate-elliptic, 0.8–1.3 ×
<quantity id="1B0A36AB9757FFBFA169F8C7FC17F8A0" box="[798,937,1892,1919]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="11.0" metricValueMin="6.0" pageId="1" pageNumber="236" unit="mm" value="0.85" valueMax="1.1" valueMin="0.6">0.6–1.1 mm</quantity>
; base oblique, inner base truncate, outer base slightly auriculate, auricle attached to stem; margins narrowly hyaline made up of a band 1–3 cells wide, the cells elongate and papillate parallel to margin, papillae in a single row over cell lumen, short-ciliate (
<quantity id="1B0A36AB9757FFBFA6A6F80FFAE2F818" box="[1233,1372,1964,1991]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="1" pageNumber="236" unit="mm" value="0.15" valueMax="0.2" valueMin="0.1">0.1–0.2 mm</quantity>
) along basal ¼, otherwise apically serrate; apex acuminate to short-aristate (
<quantity id="1B0A36AB9757FFBFA1D2F873FB8AF834" box="[933,1076,2000,2027]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="1" pageNumber="236" unit="mm" value="0.25" valueMax="0.4" valueMin="0.1">0.1–0.4 mm</quantity>
); both surfaces glabrous without idioblasts, upper surface made up of rounded to quadrangular, sinuate-walled cells, some papillate, papillae 3–7 over cell lumen, with stomata along midrib, lower surface made up of elongated, sinuate-walled cells, without stomata.
<emphasis id="EE86475C9755FFBDA2FFFB59FE8EFACB" box="[136,304,1274,1300]" italics="true" pageId="3" pageNumber="238">Axillary leaves</emphasis>
similar to lateral leaves.
<emphasis id="EE86475C9755FFBDA03AFB59FD1EFACB" box="[589,672,1274,1300]" italics="true" pageId="3" pageNumber="238">Strobili</emphasis>
terminal on branch tips, one per branch, loosely quadrangular,
<quantity id="1B0A36AB9755FFBDA71BFB59FF0AFAE6" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="5.0" metricValueMin="1.5" pageId="3" pageNumber="238" unit="mm" value="3.25" valueMax="5.0" valueMin="1.5">1.5–5 mm</quantity>
long.
<emphasis id="EE86475C9755FFBDA28BFABDFE3FFAE7" box="[252,385,1310,1336]" italics="true" pageId="3" pageNumber="238">Sporophylls</emphasis>
monomorphic, without a laminar flap, ovate, 1.0–1.2 ×
<quantity id="1B0A36AB9755FFBDA18DFABDFB3DFAE6" box="[1018,1155,1310,1337]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="3" pageNumber="238" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.5–0.6 mm</quantity>
, with a slightly developed keel apically along midrib on upper surface; base rounded; margins hyaline, dentate; apex acuminate (
<quantity id="1B0A36AB9755FFBDA681FAE1FA21FA82" box="[1270,1439,1346,1373]" metricMagnitude="-5" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="3" pageNumber="238" unit="mm" value="0.075" valueMax="0.1" valueMin="0.05">0.05–0.10 mm</quantity>
); both surfaces with idioblasts;
<emphasis id="EE86475C9755FFBDA393FAC5FD06FA5F" box="[484,696,1382,1408]" italics="true" pageId="3" pageNumber="238">dorsal sporophylls</emphasis>
with upper surface green, except for the half that is overlapping the ventral sporophylls where it is hyaline, lower surface greenish-hyaline;
<emphasis id="EE86475C9755FFBDA184FA29FB70FA7B" box="[1011,1230,1418,1444]" italics="true" pageId="3" pageNumber="238">ventral sporophylls</emphasis>
with both surfaces greenish-hyaline.
<emphasis id="EE86475C9755FFBDA327FA0CFDBFFA17" box="[336,513,1455,1480]" italics="true" pageId="3" pageNumber="238">Megasporangia</emphasis>
along basal half of ventral rows;
<emphasis id="EE86475C9755FFBDA102FA0CFC44FA17" box="[885,1018,1455,1480]" italics="true" pageId="3" pageNumber="238">megaspores</emphasis>
deep orange, proximal and distal faces rugose-reticulate with echinate microstructure, 340–357 µm diam.
<emphasis id="EE86475C9755FFBDA1F8FA70FBFAFA33" box="[911,1092,1491,1516]" italics="true" pageId="3" pageNumber="238">Microsporangia</emphasis>
in dorsal rows and apically in ventral rows;
<emphasis id="EE86475C9755FFBDA355FA54FE14F9CF" box="[290,426,1527,1552]" italics="true" pageId="3" pageNumber="238">microspores</emphasis>
deep orange, proximal face rugose, distal face baculate, 30–40 µm diam.
</paragraph>
<caption id="888DCBC69754FFBCA2FFF802FAA7F7FC" ID-DOI="http://doi.org/10.5281/zenodo.5146781" ID-Zenodo-Dep="5146781" httpUri="https://zenodo.org/record/5146781/files/figure.png" pageId="2" pageNumber="237" startId="2.[136,229,1953,1975]" targetBox="[173,1415,183,1922]" targetPageId="2">
<paragraph id="DC4D9B4E9754FFBCA2FFF802FAA7F7FC" blockId="2.[136,1452,1953,2083]" pageId="2" pageNumber="237">
<emphasis id="EE86475C9754FFBCA2FFF802FF49F869" bold="true" box="[136,247,1953,1975]" pageId="2" pageNumber="237">FIGURE 1</emphasis>
. SEM micrographs of stem section and leaves below first branch of
<taxonomicName id="1BF2E0CD9754FFBCA103F802FBD9F869" authorityName="Valdespino, C. Lopez &amp; L. Goes" authorityYear="2014" box="[884,1127,1953,1974]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="2" pageNumber="237" phylum="Tracheophyta" rank="species" species="myriostachya">
<emphasis id="EE86475C9754FFBCA103F802FBD9F869" box="[884,1127,1953,1974]" italics="true" pageId="2" pageNumber="237">Selaginella myriostachya</emphasis>
</taxonomicName>
(
<emphasis id="EE86475C9754FFBCA602F802FAE0F869" box="[1141,1374,1953,1975]" italics="true" pageId="2" pageNumber="237">Valdespino et al. 2013-1</emphasis>
, PMA).
<emphasis id="EE86475C9754FFBCA2FFF866FF27F805" bold="true" box="[136,153,1989,2010]" pageId="2" pageNumber="237">A</emphasis>
. Stem section showing appressed, slightly dimorphic median (m) and lateral (l) leaves.
<emphasis id="EE86475C9754FFBCA19AF866FC43F805" bold="true" box="[1005,1021,1989,2010]" pageId="2" pageNumber="237">B</emphasis>
. Close up of A.
<emphasis id="EE86475C9754FFBCA6EAF866FB10F804" bold="true" box="[1181,1198,1989,2011]" pageId="2" pageNumber="237">C</emphasis>
. Upper surface of median leaf.
<emphasis id="EE86475C9754FFBCA2CFF84AFF77F821" bold="true" box="[184,201,2025,2046]" pageId="2" pageNumber="237">D</emphasis>
. Close up of apical region of median leaf upper surface.
<emphasis id="EE86475C9754FFBCA09CF84AFD45F821" bold="true" box="[747,763,2025,2046]" pageId="2" pageNumber="237">E</emphasis>
. Close up of basal region of median leaf upper surface.
<emphasis id="EE86475C9754FFBCA762F84AFA9AF821" bold="true" box="[1301,1316,2025,2046]" pageId="2" pageNumber="237">F</emphasis>
. Stem section showing appressed lateral leaves.
<emphasis id="EE86475C9754FFBCA3B9F7AEFE5FF7FC" bold="true" box="[462,481,2061,2083]" pageId="2" pageNumber="237">G</emphasis>
. Lower surface of lateral leaf.
<emphasis id="EE86475C9754FFBCA17FF7AEFC9CF7FD" bold="true" box="[776,802,2061,2082]" pageId="2" pageNumber="237">H.</emphasis>
Close up of mid region of lateral leaf lower surface.
</paragraph>
</caption>
<caption id="888DCBC69755FFBDA2FFFBCEFC21FB77" ID-DOI="http://doi.org/10.5281/zenodo.5146785" ID-Zenodo-Dep="5146785" httpUri="https://zenodo.org/record/5146785/files/figure.png" pageId="3" pageNumber="238" startId="3.[136,229,1133,1155]" targetBox="[151,1435,209,1109]" targetPageId="3">
<paragraph id="DC4D9B4E9755FFBDA2FFFBCEFC21FB77" blockId="3.[136,1451,1133,1192]" pageId="3" pageNumber="238">
<emphasis id="EE86475C9755FFBDA2FFFBCEFF46FB5D" bold="true" box="[136,248,1133,1155]" pageId="3" pageNumber="238">FIGURE 2</emphasis>
. SEM micrographs of stem and leaves of
<taxonomicName id="1BF2E0CD9755FFBDA0FFFBCDFCC5FB5C" authorityName="Valdespino, C. Lopez &amp; L. Goes" authorityYear="2014" box="[648,891,1134,1155]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="3" pageNumber="238" phylum="Tracheophyta" rank="species" species="myriostachya">
<emphasis id="EE86475C9755FFBDA0FFFBCDFCC5FB5C" box="[648,891,1134,1155]" italics="true" pageId="3" pageNumber="238">Selaginella myriostachya</emphasis>
</taxonomicName>
(
<emphasis id="EE86475C9755FFBDA1FCFBCDFBB2FB5C" box="[907,1036,1134,1155]" italics="true" pageId="3" pageNumber="238">Shafer 13794</emphasis>
, NY).
<emphasis id="EE86475C9755FFBDA627FBCEFBDFFB5D" bold="true" box="[1104,1121,1133,1154]" pageId="3" pageNumber="238">A</emphasis>
. Upper surface of stem.
<emphasis id="EE86475C9755FFBDA724FBCDFADDFB5C" bold="true" box="[1363,1379,1134,1155]" pageId="3" pageNumber="238">B</emphasis>
. Upper surface of median leaf.
<emphasis id="EE86475C9755FFBDA31DFB32FEC5FB78" bold="true" box="[362,379,1169,1191]" pageId="3" pageNumber="238">C</emphasis>
. Lower surface of stem.
<emphasis id="EE86475C9755FFBDA01CFB31FDC2FB78" bold="true" box="[619,636,1170,1191]" pageId="3" pageNumber="238">D</emphasis>
. Lower surface of lateral leaf.
</paragraph>
</caption>
<paragraph id="DC4D9B4E9755FFBDA2CAF9B9FECFF986" blockId="3.[136,1452,1238,2093]" pageId="3" pageNumber="238">
<emphasis id="EE86475C9755FFBDA2CAF9B9FE57F9EB" bold="true" box="[189,489,1562,1588]" pageId="3" pageNumber="238">Habitat and Distribution</emphasis>
:—
<collectingCountry id="A4E5DBDE9755FFBDA07BF9B9FDF2F9EB" box="[524,588,1562,1588]" name="Cuba" pageId="3" pageNumber="238">Cuba</collectingCountry>
; growing in patches on limestone rocks in caves with open ceiling and high humidity; ca.
<specimenCount id="CAF450C79755FFBDA354F99DFECFF986" box="[291,369,1598,1625]" pageId="3" pageNumber="238" type="generic">
<quantity id="1B0A36AB9755FFBDA354F99DFED3F986" box="[291,365,1598,1625]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="238" unit="m" value="400.0">400 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="94E8C8C59755FFBDA2CAF9C1FAF4F81F" pageId="3" pageNumber="238" type="etymology">
<paragraph id="DC4D9B4E9755FFBDA2CAF9C1FD34F97F" blockId="3.[136,1452,1238,2093]" pageId="3" pageNumber="238">
<emphasis id="EE86475C9755FFBDA2CAF9C1FE80F9A3" bold="true" box="[189,318,1634,1660]" pageId="3" pageNumber="238">Etymology</emphasis>
:
<emphasis id="EE86475C9755FFBDA331F9C1FEDCF9A3" bold="true" box="[326,354,1634,1660]" pageId="3" pageNumber="238">—</emphasis>
The specific epithet derives from the Greek
<emphasis id="EE86475C9755FFBDA113F9C0FC1BF9A3" box="[868,933,1635,1660]" italics="true" pageId="3" pageNumber="238">myrio</emphasis>
, countless, and
<emphasis id="EE86475C9755FFBDA615F9C1FB77F9A3" box="[1122,1225,1634,1660]" italics="true" pageId="3" pageNumber="238">stachyus,</emphasis>
relating to a spike; referring to the many strobili on fertile plants.
</paragraph>
<paragraph id="DC4D9B4E9755FFBDA2CAF909FBE9F8EF" blockId="3.[136,1452,1238,2093]" pageId="3" pageNumber="238">
<emphasis id="EE86475C9755FFBDA2CAF909FE10F91B" bold="true" box="[189,430,1706,1732]" pageId="3" pageNumber="238">Conservation Status</emphasis>
:
<emphasis id="EE86475C9755FFBDA3C1F909FE6CF91B" bold="true" box="[438,466,1706,1732]" pageId="3" pageNumber="238">—</emphasis>
<materialsCitation id="6C9A91139755FFBDA3A5F909FBECF8EF" collectionCode="VU" country="Cuba" county="According" location="Although" municipality="It" pageId="3" pageNumber="238" specimenCount="1" stateProvince="Pinar del Rio">
<location id="D92DCD959755FFBDA3A5F909FD82F91B" LSID="urn:lsid:plazi:treatment:545B2A589757FFBAA2FFFB37FBF3FD53:D92DCD959755FFBDA3A5F909FD82F91B" box="[466,572,1706,1732]" country="Cuba" county="According" municipality="It" name="Although" pageId="3" pageNumber="238" stateProvince="Pinar del Rio">Although</location>
this species is to be expected on or around
<location id="D92DCD959755FFBDA664F908FBCBF91A" LSID="urn:lsid:plazi:treatment:545B2A589757FFBAA2FFFB37FBF3FD53:D92DCD959755FFBDA664F908FBCBF91A" box="[1043,1141,1707,1733]" country="Cuba" county="According" municipality="It" name="Mogotes" pageId="3" pageNumber="238" stateProvince="Pinar del Rio">Mogotes</location>
along
<location id="D92DCD959755FFBDA6C8F909FABCF91B" LSID="urn:lsid:plazi:treatment:545B2A589757FFBAA2FFFB37FBF3FD53:D92DCD959755FFBDA6C8F909FABCF91B" box="[1215,1282,1706,1732]" country="Cuba" county="According" municipality="It" name="Sierra" pageId="3" pageNumber="238" stateProvince="Pinar del Rio">Sierra</location>
de los
<location id="D92DCD959755FFBDA73AF908FA15F919" LSID="urn:lsid:plazi:treatment:545B2A589757FFBAA2FFFB37FBF3FD53:D92DCD959755FFBDA73AF908FA15F919" box="[1357,1451,1707,1734]" country="Cuba" county="According" municipality="It" name="Organos" pageId="3" pageNumber="238" stateProvince="Pinar del Rio">Órganos</location>
in
<collectingRegion id="1E3655AC9755FFBDA2DEF96DFE0CF937" box="[169,434,1742,1768]" country="Cuba" name="Pinar del Rio" pageId="3" pageNumber="238">Pinar del Río province</collectingRegion>
in western
<collectingCountry id="A4E5DBDE9755FFBDA048F96DFDC3F937" box="[575,637,1742,1768]" name="Cuba" pageId="3" pageNumber="238">Cuba</collectingCountry>
, it has a limited range of known distribution.
<collectingMunicipality id="3C2901349755FFBDA6E8F96CFB0EF936" box="[1183,1200,1743,1769]" pageId="3" pageNumber="238">It</collectingMunicipality>
is found in localized populations that may be threatened by agriculture expansion and ecotourism activities such as caving.
<collectingCounty id="352CE3C29755FFBDA760F951FA33F8D3" box="[1303,1421,1778,1804]" pageId="3" pageNumber="238">According</collectingCounty>
to
<bibRefCitation id="B863E6BF9755FFBDA2FFF8B5FEA0F8EF" author="IUCN &amp; Cambridge, U. K." box="[136,286,1814,1840]" pageId="3" pageNumber="238" refId="ref6456" refString="IUCN. (2012) IUCN Red List Categories: Version 3.1. Second edition. Gland, Switzerland and Cambridge, U. K.: IUCN. iv + 32 pp." type="book" year="2012">IUCN (2012)</bibRefCitation>
categories and criteria, it may be tentatively considered vulnerable (
<collectionCode id="BAE3038B9755FFBDA66AF8B5FBF5F8EF" box="[1053,1099,1814,1840]" country="Russia" lsid="urn:lsid:biocol.org:col:15248" name="Voronezh State University" pageId="3" pageNumber="238" type="Herbarium">VU</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="DC4D9B4E9755FFBDA2CAF899FAF4F81F" blockId="3.[136,1452,1238,2093]" pageId="3" pageNumber="238">
<materialsCitation id="6C9A91139755FFBDA2CAF899FC73F88B" box="[189,973,1850,1877]" country="Cuba" location="Pinar del Rio" pageId="3" pageNumber="238" specimenCount="2" stateProvince="Pinar del Rio" typeStatus="paratype">
<emphasis id="EE86475C9755FFBDA2CAF899FD7FF88B" bold="true" box="[189,705,1850,1876]" pageId="3" pageNumber="238">
Additional Specimens Examined (
<typeStatus id="034925EC9755FFBDA035F899FD04F88B" box="[578,698,1850,1876]" pageId="3" pageNumber="238" type="paratype">Paratypes</typeStatus>
)
</emphasis>
:
<emphasis id="EE86475C9755FFBDA0BEF899FD5BF88B" bold="true" box="[713,741,1850,1876]" pageId="3" pageNumber="238">—</emphasis>
<collectingCountry id="A4E5DBDE9755FFBDA092F899FC8BF88B" box="[741,821,1850,1876]" name="Cuba" pageId="3" pageNumber="238">CUBA</collectingCountry>
.
<collectingRegion id="1E3655AC9755FFBDA14AF899FC73F88B" box="[829,973,1850,1876]" country="Cuba" name="Pinar del Rio" pageId="3" pageNumber="238">Pinar del Río</collectingRegion>
</materialsCitation>
:
<materialsCitation id="6C9A91139755FFBDA1A2F898FAF8F81F" collectingDate="1910-09-12" collectingDateMax="1912-08-26" collectingDateMin="1910-09-12" collectionCode="S, GH, NY, UC, N, BM, B, MO, P, W" collectorName="La Guira" country="United States of America" county="Sumidero" location="Banos de San Vicente" municipality="Sumidero" pageId="3" pageNumber="238" specimenCount="1">
Loma Harenales [Arenales],
<locationDeviation id="CDFF826C9755FFBDA77CF899FA16F88B" box="[1291,1448,1850,1876]" location="Sumidero" pageId="3" pageNumber="238">
<collectionCode id="BAE3038B9755FFBDA77CF899FAA4F88B" box="[1291,1306,1850,1876]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="3" pageNumber="238" type="Herbarium">S</collectionCode>
of
<collectingCounty id="352CE3C29755FFBDA74DF899FA16F88B" box="[1338,1448,1850,1876]" pageId="3" pageNumber="238">Sumidero</collectingCounty>
</locationDeviation>
,
<date id="A84CBD8E9755FFBDA2FFF8FDFE84F8A7" box="[136,314,1886,1912]" pageId="3" pageNumber="238" value="1912-08-20">
<collectingDate id="B80844669755FFBDA2FFF8FDFE84F8A7" box="[136,314,1886,1912]" pageId="3" pageNumber="238" value="1912-08-20">20 August 1912</collectingDate>
</date>
,
<emphasis id="EE86475C9755FFBDA331F8FDFE64F8A7" box="[326,474,1886,1912]" italics="true" pageId="3" pageNumber="238">Shafer 13794</emphasis>
(F-2 sheets,
<collectionCode id="BAE3038B9755FFBDA01DF8FDFD2AF8A7" box="[618,660,1886,1912]" country="USA" lsid="urn:lsid:biocol.org:col:15631" name="Harvard University - Gray Herbarium" pageId="3" pageNumber="238" type="Herbarium">GH</collectionCode>
,
<collectionCode id="BAE3038B9755FFBDA0D7F8FCFD79F8A6" box="[672,711,1887,1913]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="3" pageNumber="238" type="Herbarium">NY</collectionCode>
,
<collectionCode id="BAE3038B9755FFBDA0A4F8FDFD42F8A7" box="[723,764,1886,1912]" country="USA" name="Upjohn Culture Collection" pageId="3" pageNumber="238">UC</collectionCode>
,
<collectingCountry id="A4E5DBDE9755FFBDA17FF8FDFC91F8A7" box="[776,815,1886,1912]" name="United States of America" pageId="3" pageNumber="238">US</collectingCountry>
); Lavadero to
<collectorName id="7107FE989755FFBDA1A7F8FCFB8BF8A7" box="[976,1077,1886,1913]" pageId="3" pageNumber="238">La Guira</collectorName>
,
<locationDeviation id="CDFF826C9755FFBDA636F8FCFB57F8A7" box="[1089,1257,1886,1913]" location="Sumidero" pageId="3" pageNumber="238">
<collectionCode id="BAE3038B9755FFBDA636F8FCFBEBF8A6" box="[1089,1109,1887,1913]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="3" pageNumber="238" type="Herbarium">N</collectionCode>
of
<collectingMunicipality id="3C2901349755FFBDA60BF8FDFB57F8A7" box="[1148,1257,1886,1912]" pageId="3" pageNumber="238">Sumidero</collectingMunicipality>
</locationDeviation>
,
<date id="A84CBD8E9755FFBDA682F8FDFA19F8A7" box="[1269,1447,1886,1912]" pageId="3" pageNumber="238" value="1912-08-26">
<collectingDate id="B80844669755FFBDA682F8FDFA19F8A7" box="[1269,1447,1886,1912]" pageId="3" pageNumber="238" value="1912-08-26">26 August 1912</collectingDate>
</date>
,
<emphasis id="EE86475C9755FFBDA2FFF821FEA4F843" box="[136,282,1922,1948]" italics="true" pageId="3" pageNumber="238">Shafer 13845</emphasis>
(
<collectionCode id="BAE3038B9755FFBDA35FF821FEE8F843" box="[296,342,1922,1948]" country="United Kingdom" name="Bristol Museum" pageId="3" pageNumber="238">BM</collectionCode>
,
<collectionCode id="BAE3038B9755FFBDA316F820FE33F842" box="[353,397,1923,1949]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="3" pageNumber="238" type="Herbarium">NY</collectionCode>
);
<collectingCounty id="352CE3C29755FFBDA3E8F820FD30F843" box="[415,654,1922,1949]" pageId="3" pageNumber="238">
<location id="D92DCD959755FFBDA3E8F820FD30F843" LSID="urn:lsid:plazi:treatment:545B2A589757FFBAA2FFFB37FBF3FD53:D92DCD959755FFBDA3E8F820FD30F843" box="[415,654,1922,1949]" country="United States of America" county="Sumidero" municipality="Sumidero" name="Banos de San Vicente" pageId="3" pageNumber="238">Baños de San Vicente</location>
</collectingCounty>
,
<date id="A84CBD8E9755FFBDA0EFF821FC28F843" box="[664,918,1922,1948]" pageId="3" pageNumber="238" value="1910-09-12" valueMax="1910-09-16" valueMin="1910-09-12">
<collectingDate id="B80844669755FFBDA0EFF821FC28F843" box="[664,918,1922,1948]" pageId="3" pageNumber="238" value="1910-09-12" valueMax="1910-09-16" valueMin="1910-09-12">12–16 September 1910</collectingDate>
</date>
,
<emphasis id="EE86475C9755FFBDA1D6F820FBCBF843" box="[929,1141,1922,1948]" italics="true" pageId="3" pageNumber="238">
<collectingMunicipality id="3C2901349755FFBDA1D6F820FC50F843" box="[929,1006,1923,1948]" pageId="3" pageNumber="238">
<location id="D92DCD959755FFBDA1D6F820FC50F843" LSID="urn:lsid:plazi:treatment:545B2A589757FFBAA2FFFB37FBF3FD53:D92DCD959755FFBDA1D6F820FC50F843" box="[929,1006,1923,1948]" country="United States of America" county="Sumidero" municipality="Sumidero" name="Britton" pageId="3" pageNumber="238">Britton</location>
</collectingMunicipality>
et al. 7341a
</emphasis>
(
<collectionCode id="BAE3038B9755FFBDA6F5F821FB11F843" box="[1154,1199,1922,1948]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="3" pageNumber="238" type="Herbarium">NY</collectionCode>
);
<location id="D92DCD959755FFBDA6B5F821FADCF843" LSID="urn:lsid:plazi:treatment:545B2A589757FFBAA2FFFB37FBF3FD53:D92DCD959755FFBDA6B5F821FADCF843" box="[1218,1378,1922,1948]" country="United States of America" county="Sumidero" municipality="Sumidero" name="Cuba Orientali" pageId="3" pageNumber="238">Cuba Orientali</location>
[Pinar del Río],
<location id="D92DCD959755FFBDA299F805FEFDF81F" LSID="urn:lsid:plazi:treatment:545B2A589757FFBAA2FFFB37FBF3FD53:D92DCD959755FFBDA299F805FEFDF81F" box="[238,323,1958,1984]" country="United States of America" county="Sumidero" municipality="Sumidero" name="Vinales" pageId="3" pageNumber="238">Viñales</location>
, 1859–1860,
<emphasis id="EE86475C9755FFBDA3A1F805FDE0F81F" box="[470,606,1958,1984]" italics="true" pageId="3" pageNumber="238">
<location id="D92DCD959755FFBDA3A1F805FD9FF81F" LSID="urn:lsid:plazi:treatment:545B2A589757FFBAA2FFFB37FBF3FD53:D92DCD959755FFBDA3A1F805FD9FF81F" box="[470,545,1958,1984]" country="United States of America" county="Sumidero" municipality="Sumidero" name="Wright" pageId="3" pageNumber="238">Wright</location>
1823
</emphasis>
(
<collectionCode id="BAE3038B9755FFBDA019F805FD3CF81F" box="[622,642,1958,1984]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="3" pageNumber="238" type="Herbarium">B</collectionCode>
,
<collectionCode id="BAE3038B9755FFBDA0F9F804FD05F81E" box="[654,699,1959,1985]" country="United Kingdom" name="Bristol Museum" pageId="3" pageNumber="238">BM</collectionCode>
,
<collectionCode id="BAE3038B9755FFBDA0B1F805FD4EF81F" box="[710,752,1958,1984]" country="USA" lsid="urn:lsid:biocol.org:col:15631" name="Harvard University - Gray Herbarium" pageId="3" pageNumber="238" type="Herbarium">GH</collectionCode>
, K-2 sheets,
<collectionCode id="BAE3038B9755FFBDA10AF805FC12F81F" box="[893,940,1958,1984]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="3" pageNumber="238" type="Herbarium">MO</collectionCode>
,
<collectionCode id="BAE3038B9755FFBDA1C0F804FC78F81E" box="[951,966,1959,1985]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="238" type="Herbarium">P</collectionCode>
,
<collectingCountry id="A4E5DBDE9755FFBDA1A6F805FC49F81F" box="[977,1015,1958,1984]" name="United States of America" pageId="3" pageNumber="238">US</collectingCountry>
,
<collectionCode id="BAE3038B9755FFBDA676F804FBA1F81E" box="[1025,1055,1959,1985]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="3" pageNumber="238" type="Herbarium">W</collectionCode>
); 1924,
<emphasis id="EE86475C9755FFBDA600F805FAB9F81F" box="[1143,1287,1958,1984]" italics="true" pageId="3" pageNumber="238">
<location id="D92DCD959755FFBDA600F805FB69F81F" LSID="urn:lsid:plazi:treatment:545B2A589757FFBAA2FFFB37FBF3FD53:D92DCD959755FFBDA600F805FB69F81F" box="[1143,1239,1958,1984]" country="United States of America" county="Sumidero" municipality="Sumidero" name="Chrysler" pageId="3" pageNumber="238">Chrysler</location>
15a
</emphasis>
(
<collectingCountry id="A4E5DBDE9755FFBDA761F805FA80F81F" box="[1302,1342,1958,1984]" name="United States of America" pageId="3" pageNumber="238">US</collectingCountry>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="94E8C8C59755FFBAA2CAF869FBF3FD53" lastPageId="4" lastPageNumber="239" pageId="3" pageNumber="238" type="discussion">
<paragraph id="DC4D9B4E9755FFBAA2CAF869FD57FEB3" blockId="3.[136,1452,1238,2093]" lastBlockId="4.[136,1452,158,653]" lastPageId="4" lastPageNumber="239" pageId="3" pageNumber="238">
<emphasis id="EE86475C9755FFBDA2CAF869FE85F83B" bold="true" box="[189,315,1994,2020]" pageId="3" pageNumber="238">Discussion</emphasis>
:
<emphasis id="EE86475C9755FFBDA334F869FEE1F83B" bold="true" box="[323,351,1994,2020]" pageId="3" pageNumber="238">—</emphasis>
<taxonomicName id="1BF2E0CD9755FFBDA328F869FDC3F83B" authorityName="Valdespino, C. Lopez &amp; L. Goes" authorityYear="2014" box="[351,637,1994,2020]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="3" pageNumber="238" phylum="Tracheophyta" rank="species" species="myriostachya">
<emphasis id="EE86475C9755FFBDA328F869FDC3F83B" box="[351,637,1994,2020]" italics="true" pageId="3" pageNumber="238">Selaginella myriostachya</emphasis>
</taxonomicName>
is characterized by its suberect to erect stems covered by very slightly dimorphic, appressed leaves below the first branch (
<figureCitation id="44C987CB9755FFBDA0B7F84DFD40F7D7" box="[704,766,2030,2056]" captionStart="FIGURE 1" captionStartId="2.[136,229,1953,1975]" captionTargetBox="[173,1415,183,1922]" captionTargetId="figure-21@2.[173,1415,183,1922]" captionTargetPageId="2" captionText="FIGURE 1. SEM micrographs of stem section and leaves below first branch of Selaginella myriostachya (Valdespino et al. 2013-1, PMA). A. Stem section showing appressed, slightly dimorphic median (m) and lateral (l) leaves. B. Close up of A. C. Upper surface of median leaf. D. Close up of apical region of median leaf upper surface. E. Close up of basal region of median leaf upper surface. F. Stem section showing appressed lateral leaves. G. Lower surface of lateral leaf. H. Close up of mid region of lateral leaf lower surface." figureDoi="http://doi.org/10.5281/zenodo.5146781" httpUri="https://zenodo.org/record/5146781/files/figure.png" pageId="3" pageNumber="238">Fig. 1</figureCitation>
) and oblique bases of median leaves, with a weekly developed outer auricle attached to the stem (
<figureCitation id="44C987CB9755FFBDA063F7B1FDD0F7F3" box="[532,622,2066,2092]" captionStart="FIGURE 2" captionStartId="3.[136,229,1133,1155]" captionTargetBox="[151,1435,209,1109]" captionTargetId="figure-18@3.[151,1435,209,1109]" captionTargetPageId="3" captionText="FIGURE 2. SEM micrographs of stem and leaves of Selaginella myriostachya (Shafer 13794, NY). A. Upper surface of stem. B. Upper surface of median leaf. C. Lower surface of stem. D. Lower surface of lateral leaf." figureDoi="http://doi.org/10.5281/zenodo.5146785" httpUri="https://zenodo.org/record/5146785/files/figure.png" pageId="3" pageNumber="238">Fig. 2B</figureCitation>
). Interestingly, below the first branch of the stem, the cell pattern on the upper surface of median and the lower surface of lateral leaves varies slightly than that on leaves above this section. In this region, the upper surface of median leaves is made up of more conspicuously quadrangular cells, most of which are covered by 4–11 papillae (
<figureCitation id="44C987CB9752FFBAA3AFFF45FDF1FEDF" box="[472,591,230,256]" captionStart="FIGURE 1" captionStartId="2.[136,229,1953,1975]" captionTargetBox="[173,1415,183,1922]" captionTargetId="figure-21@2.[173,1415,183,1922]" captionTargetPageId="2" captionText="FIGURE 1. SEM micrographs of stem section and leaves below first branch of Selaginella myriostachya (Valdespino et al. 2013-1, PMA). A. Stem section showing appressed, slightly dimorphic median (m) and lateral (l) leaves. B. Close up of A. C. Upper surface of median leaf. D. Close up of apical region of median leaf upper surface. E. Close up of basal region of median leaf upper surface. F. Stem section showing appressed lateral leaves. G. Lower surface of lateral leaf. H. Close up of mid region of lateral leaf lower surface." figureDoi="http://doi.org/10.5281/zenodo.5146781" httpUri="https://zenodo.org/record/5146781/files/figure.png" pageId="4" pageNumber="239">Fig. 1C–E</figureCitation>
), whereas the lower surface of lateral leaves is made up of elongate, straightwalled cells, many of which are covered by 7–18 papillae (
<figureCitation id="44C987CB9752FFBAA164FEA9FC1CFEFA" box="[787,930,266,293]" captionStart="FIGURE 1" captionStartId="2.[136,229,1953,1975]" captionTargetBox="[173,1415,183,1922]" captionTargetId="figure-21@2.[173,1415,183,1922]" captionTargetPageId="2" captionText="FIGURE 1. SEM micrographs of stem section and leaves below first branch of Selaginella myriostachya (Valdespino et al. 2013-1, PMA). A. Stem section showing appressed, slightly dimorphic median (m) and lateral (l) leaves. B. Close up of A. C. Upper surface of median leaf. D. Close up of apical region of median leaf upper surface. E. Close up of basal region of median leaf upper surface. F. Stem section showing appressed lateral leaves. G. Lower surface of lateral leaf. H. Close up of mid region of lateral leaf lower surface." figureDoi="http://doi.org/10.5281/zenodo.5146781" httpUri="https://zenodo.org/record/5146781/files/figure.png" pageId="4" pageNumber="239">Fig. 1G &amp; H</figureCitation>
). In addition, above the first branch of the stem the lower surface of lateral leaves have obscure idioblasts when observed with a stereoscope, which are conspicuous and obviously papillate as observed on SEM (
<figureCitation id="44C987CB9752FFBAA0F5FEF1FD62FEB3" box="[642,732,338,364]" captionStart="FIGURE 2" captionStartId="3.[136,229,1133,1155]" captionTargetBox="[151,1435,209,1109]" captionTargetId="figure-18@3.[151,1435,209,1109]" captionTargetPageId="3" captionText="FIGURE 2. SEM micrographs of stem and leaves of Selaginella myriostachya (Shafer 13794, NY). A. Upper surface of stem. B. Upper surface of median leaf. C. Lower surface of stem. D. Lower surface of lateral leaf." figureDoi="http://doi.org/10.5281/zenodo.5146785" httpUri="https://zenodo.org/record/5146785/files/figure.png" pageId="4" pageNumber="239">Fig. 2D</figureCitation>
).
</paragraph>
<paragraph id="DC4D9B4E9752FFBAA2CAFED5FBF3FD53" blockId="4.[136,1452,158,653]" pageId="4" pageNumber="239">
<taxonomicName id="1BF2E0CD9752FFBAA2CAFED5FE65FE4F" authorityName="Valdespino, C. Lopez &amp; L. Goes" authorityYear="2014" box="[189,475,374,400]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="4" pageNumber="239" phylum="Tracheophyta" rank="species" species="myriostachya">
<emphasis id="EE86475C9752FFBAA2CAFED5FE65FE4F" box="[189,475,374,400]" italics="true" pageId="4" pageNumber="239">Selaginella myriostachya</emphasis>
</taxonomicName>
may be confused with the morphologically similar
<taxonomicName id="1BF2E0CD9752FFBAA65DFED4FB0CFE4F" authorityName="Spring" authorityYear="1843" baseAuthorityName="Desvaux ex Poiret" baseAuthorityYear="1814" box="[1066,1202,374,401]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="4" pageNumber="239" phylum="Tracheophyta" rank="species" species="flabellum">
<emphasis id="EE86475C9752FFBAA65DFED4FB86FE4E" box="[1066,1080,375,401]" italics="true" pageId="4" pageNumber="239">S</emphasis>
.
<emphasis id="EE86475C9752FFBAA63FFED5FB0CFE4F" box="[1096,1202,374,400]" italics="true" pageId="4" pageNumber="239">flabellum</emphasis>
</taxonomicName>
but differs from it by characters provided in the diagnosis and by having lateral leaves on branches that are mostly ascending (vs. spreading, giving a pectinate appearance to branches in
<taxonomicName id="1BF2E0CD9752FFBAA0E4FE1CFCA2FE07" authorityName="Spring" authorityYear="1843" baseAuthorityName="Desvaux ex Poiret" baseAuthorityYear="1814" box="[659,796,446,473]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="4" pageNumber="239" phylum="Tracheophyta" rank="species" species="flabellum">
<emphasis id="EE86475C9752FFBAA0E4FE1CFD1FFE06" box="[659,673,447,473]" italics="true" pageId="4" pageNumber="239">S</emphasis>
.
<emphasis id="EE86475C9752FFBAA0C5FE1DFCA2FE07" box="[690,796,446,472]" italics="true" pageId="4" pageNumber="239">flabellum</emphasis>
</taxonomicName>
). In addition,
<taxonomicName id="1BF2E0CD9752FFBAA1B3FE1CFBC4FE07" authorityName="Valdespino, C. Lopez &amp; L. Goes" authorityYear="2014" box="[964,1146,446,473]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="4" pageNumber="239" phylum="Tracheophyta" rank="species" species="myriostachya">
<emphasis id="EE86475C9752FFBAA1B3FE1CFC6CFE06" box="[964,978,447,473]" italics="true" pageId="4" pageNumber="239">S</emphasis>
.
<emphasis id="EE86475C9752FFBAA194FE1DFBC4FE07" box="[995,1146,446,472]" italics="true" pageId="4" pageNumber="239">myriostachya</emphasis>
</taxonomicName>
grows on limestone rocks associated with caves in Mogotes, which are habitats of high endemism in
<collectingCountry id="A4E5DBDE9752FFBAA1B5FE41FC43FE23" box="[962,1021,482,508]" name="Cuba" pageId="4" pageNumber="239">Cuba</collectingCountry>
(
<bibRefCitation id="B863E6BF9752FFBAA67CFE41FB69FE23" author="Rivera, C. &amp; Urquiola, A. &amp; Valdes, N. &amp; Paneque, I. &amp; Nieto, M." box="[1035,1239,482,509]" pageId="4" pageNumber="239" pagination="1 - 7" refId="ref6706" refString="Rivera, C., Urquiola, A., Valdes, N., Paneque, I. &amp; Nieto, M. (2008) Flora y vegetacion de Mogotes. Avances 10: 1 - 7." type="journal article" year="2008">
Rivera
<emphasis id="EE86475C9752FFBAA629FE40FB30FE23" box="[1118,1166,482,508]" italics="true" pageId="4" pageNumber="239">et al</emphasis>
. 2008
</bibRefCitation>
), particularly those around the Viñales Valley of
<collectingRegion id="1E3655AC9752FFBAA3BCFDA5FD78FDFF" box="[459,710,518,544]" country="Cuba" name="Pinar del Rio" pageId="4" pageNumber="239">Pinar del Río province</collectingRegion>
in the western part of the island.
<taxonomicName id="1BF2E0CD9752FFBAA64EFDA5FA96FDFF" authorityName="Spring" authorityYear="1843" baseAuthorityName="Desvaux ex Poiret" baseAuthorityYear="1814" box="[1081,1320,518,544]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="4" pageNumber="239" phylum="Tracheophyta" rank="species" species="flabellum">
<emphasis id="EE86475C9752FFBAA64EFDA5FA96FDFF" box="[1081,1320,518,544]" italics="true" pageId="4" pageNumber="239">Selaginella flabellum</emphasis>
</taxonomicName>
, previously known only from
<collectingCountry id="A4E5DBDE9752FFBAA326FD89FE4BFD9B" box="[337,501,554,580]" name="Dominican Republic" pageId="4" pageNumber="239">the Dominican</collectingCountry>
Republic and
<collectingCountry id="A4E5DBDE9752FFBAA0EFFD89FD71FD9B" box="[664,719,554,580]" name="Haiti" pageId="4" pageNumber="239">Haiti</collectingCountry>
(
<bibRefCitation id="B863E6BF9752FFBAA0A9FD89FCD2FD9A" author="Alston, A. H. G." box="[734,876,554,581]" pageId="4" pageNumber="239" pagination="27 - 47" refId="ref6169" refString="Alston, A. H. G. (1952) A revision of the West Indian species of Selaginella. Bulletin of The British Museum (Natural History), Botany 1: 27 - 47." type="journal article" year="1952">Alston 1952</bibRefCitation>
), is recorded in eastern
<collectingCountry id="A4E5DBDE9752FFBAA601FD89FB0FFD9B" box="[1142,1201,554,580]" name="Cuba" pageId="4" pageNumber="239">Cuba</collectingCountry>
from the provinces of
<collectingRegion id="1E3655AC9752FFBAA2FFFDEDFF5AFDB7" box="[136,228,590,616]" country="Cuba" name="Holguin" pageId="4" pageNumber="239">Holguín</collectingRegion>
(e.g.,
<emphasis id="EE86475C9752FFBAA35BFDECFD97FDB7" box="[300,553,590,617]" italics="true" pageId="4" pageNumber="239">
Morton &amp;
<taxonomicName id="1BF2E0CD9752FFBAA3D4FDECFE54FDB6" box="[419,490,591,617]" class="Magnoliopsida" family="Ericaceae" genus="Acuna" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="239" phylum="Tracheophyta" rank="genus">Acuña</taxonomicName>
3300
</emphasis>
,
<collectingCountry id="A4E5DBDE9752FFBAA040FDEDFDDCFDB7" box="[567,610,590,616]" name="United States of America" pageId="4" pageNumber="239">US</collectingCountry>
!;
<emphasis id="EE86475C9752FFBAA004FDEDFD41FDB7" box="[627,767,590,616]" italics="true" pageId="4" pageNumber="239">Ekman 2229</emphasis>
, B!) and Santiago de Cuba (e.g.,
<emphasis id="EE86475C9752FFBAA60FFDEDFAAAFDB7" box="[1144,1300,590,616]" italics="true" pageId="4" pageNumber="239">Clement 6671</emphasis>
,
<collectingCountry id="A4E5DBDE9752FFBAA755FDEDFAF3FDB7" box="[1314,1357,590,616]" name="United States of America" pageId="4" pageNumber="239">US</collectingCountry>
!;
<emphasis id="EE86475C9752FFBAA729FDECFF73FD53" italics="true" pageId="4" pageNumber="239">
<taxonomicName id="1BF2E0CD9752FFBAA729FDECFA12FDB6" box="[1374,1452,591,617]" class="Magnoliopsida" family="Euphorbiaceae" genus="Garcia" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="239" phylum="Tracheophyta" rank="genus">Garcia</taxonomicName>
51652
</emphasis>
, HAJB!;
<emphasis id="EE86475C9752FFBAA34EFDD0FDB5FD53" box="[313,523,626,653]" italics="true" pageId="4" pageNumber="239">
Roy &amp;
<taxonomicName id="1BF2E0CD9752FFBAA3F0FDD0FE70FD52" box="[391,462,627,653]" class="Magnoliopsida" family="Ericaceae" genus="Acuna" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="239" phylum="Tracheophyta" rank="genus">Acuña</taxonomicName>
8172
</emphasis>
,
<collectingCountry id="A4E5DBDE9752FFBAA06EFDD1FDFAFD53" box="[537,580,626,652]" name="United States of America" pageId="4" pageNumber="239">US</collectingCountry>
!). These plants grow on wet soil stream banks.
</paragraph>
</subSubSection>
</treatment>
</document>
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