669 lines
89 KiB
XML
669 lines
89 KiB
XML
<document id="38631EB484C2B82078A2E15AC5DF5D4A" ID-DOI="10.5852/ejt.2022.831.1875" ID-GBIF-Dataset="dac72712-c4a7-4a1c-9de0-3ea1c95cecd2" ID-ISSN="2118-9773" ID-Zenodo-Dep="6916932" ID-ZooBank="C82D605B-9BE0-47D1-89E3-8C978CA48B4D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="diego" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658962688836" checkinUser="felipe" docAuthor="Magalhaes, Ivan L. F., Aharon, Shlomi, Ganem, Zeana & Gavish-Regev, Efrat" docDate="2022" docId="54519917FF9CFFC708A9FBBB08A5E88A" docLanguage="en" docName="ejt-831_149-174.pdf" docOrigin="European Journal of Taxonomy 831" docSource="http://dx.doi.org/10.5852/ejt.2022.831.1875" docStyle="DocumentStyle:8DBBCD0342F59C7B93E4D60327A07408.6:EJT.2018-.journal_article.type2" docStyleId="8DBBCD0342F59C7B93E4D60327A07408" docStyleName="EJT.2018-.journal_article.type2" docStyleVersion="6" docTitle="Filistata betarif Magalhaes & Aharon & Ganem & Gavish-Regev 2022, sp. nov." docType="treatment" docUuid="86B4AAD0-D69E-4C79-BF3F-DE5E7DC0EDB1" docUuidSource="ZooBank" docVersion="9" lastPageNumber="167" masterDocId="A868E16FFF90FFD50A18FF9E0A03EC58" masterDocTitle="A new semi-cryptic Filistata from caves in the Levant with comments on the limits of Filistata insidiatrix (Forsskål, 1775) (Arachnida: Araneae: Filistatidae)" masterLastPageNumber="174" masterPageNumber="149" pageNumber="161" updateTime="1701996858087" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="777EAB2A2BD6E5D7413B5A5B60270DA9">A new semi-cryptic Filistata from caves in the Levant with comments on the limits of Filistata insidiatrix (Forsskål, 1775) (Arachnida: Araneae: Filistatidae)</mods:title>
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<mods:roleTerm id="4C09622BC98450964035FA86A233319F">Author</mods:roleTerm>
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<mods:namePart id="DA7636DEFC847889C05021FB22A8A849">Magalhaes, Ivan L. F.</mods:namePart>
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<mods:affiliation id="2D3EC898646AB8D694D09FFC731A2D83">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ”, Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Av. Ángel Gallardo 470, C 1405 DJR, Buenos Aires, Argentina.</mods:affiliation>
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<mods:nameIdentifier id="AD344C2F77A1E027BF07CDDB410D12B6" type="email">magalhaes@macn.gov.ar</mods:nameIdentifier>
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<mods:namePart id="B260E1AEF220D63E89E15EFF5B7A8493">Aharon, Shlomi</mods:namePart>
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<mods:affiliation id="D3A6D4D947F8940881DB0C46F5484E9D">The National Natural History Collections, Edmond J. Safra Campus, Givat Ram, The Hebrew University of Jerusalem, Jerusalem 9190401, Israel. & The Department of Ecology, Evolution and Behavior, Edmond J. Safra Campus, Givat Ram, The Hebrew University of Jerusalem, Jerusalem 9190401, Israel.</mods:affiliation>
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<mods:nameIdentifier id="A525E613D8C13733134480939B754FA4" type="email">shlomi.aharon@gmail.com</mods:nameIdentifier>
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<mods:namePart id="BD1D323FD959B62B8F49640EA3F7A986">Ganem, Zeana</mods:namePart>
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<mods:affiliation id="4F2D509B22508B44A64B73FBC3DE88B3">The National Natural History Collections, Edmond J. Safra Campus, Givat Ram, The Hebrew University of Jerusalem, Jerusalem 9190401, Israel. & The Department of Ecology, Evolution and Behavior, Edmond J. Safra Campus, Givat Ram, The Hebrew University of Jerusalem, Jerusalem 9190401, Israel.</mods:affiliation>
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<mods:nameIdentifier id="F3B7938A3ADA155D98567C1B29E202A0" type="email">zeana.ganem@mail.huji.ac.il</mods:nameIdentifier>
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<mods:namePart id="A47E57F2473F515CE52DB2D9740EDDCD">Gavish-Regev, Efrat</mods:namePart>
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<mods:affiliation id="F4BA483A5FF811E9E0325E147576D5BB">The National Natural History Collections, Edmond J. Safra Campus, Givat Ram, The Hebrew University of Jerusalem, Jerusalem 9190401, Israel.</mods:affiliation>
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<mods:nameIdentifier id="2F40477F27BC2B4F59973981AFFB666D" type="email">efrat.gavish-regev@mail.huji.ac.il</mods:nameIdentifier>
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<mods:title id="F666BB0738594BDD8A1E03DC0BF7FCF2">European Journal of Taxonomy</mods:title>
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<mods:part id="04D747926E8366948B457D594BED76CE">
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<mods:date id="2FB3C3C88A2ABCBFC51EC6745A140E0A">2022</mods:date>
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<mods:number id="5401C4CE980F7A2A44380BAC60E543FA">2022-07-25</mods:number>
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<mods:number id="E674D3D15BD4CA03FA6426B3AEBA6A49">831</mods:number>
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<treatment id="54519917FF9CFFC708A9FBBB08A5E88A" ID-DOI="http://doi.org/10.5281/zenodo.6928202" ID-GBIF-Taxon="197866406" ID-Zenodo-Dep="6928202" LSID="urn:lsid:zoobank.org:act:86B4AAD0-D69E-4C79-BF3F-DE5E7DC0EDB1" httpUri="http://treatment.plazi.org/id/54519917FF9CFFC708A9FBBB08A5E88A" lastPageId="18" lastPageNumber="167" pageId="12" pageNumber="161">
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<subSubSection id="94E27B8AFF9CFFD908A9FBBB09CCE818" box="[689,975,1061,1088]" pageId="12" pageNumber="161" type="nomenclature">
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<paragraph id="DC472801FF9CFFD908A9FBBB09CCE818" blockId="12.[387,1276,1061,1160]" box="[689,975,1061,1088]" pageId="12" pageNumber="161">
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<heading id="870F9F6DFF9CFFD908A9FBBB09CCE818" box="[689,975,1061,1088]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="161" reason="2">
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<taxonomicName id="1BF85382FF9CFFD908A9FBBB096EE818" authority="Magalhaes & Aharon & Ganem & Gavish-Regev, 2022" authorityName="Magalhaes & Aharon & Ganem & Gavish-Regev" authorityYear="2022" box="[689,877,1061,1088]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="161" phylum="Arthropoda" rank="species" species="betarif" status="sp. nov.">
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<emphasis id="EE8CF413FF9CFFD908A9FBBB096EE818" bold="true" box="[689,877,1061,1088]" italics="true" pageId="12" pageNumber="161">Filistata betarif</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F5BF4968FF9CFFD9096CFBB809CCE818" box="[884,975,1062,1088]" pageId="12" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<subSubSection id="94E27B8AFF9CFFD90B9BFBD709F7E8D0" pageId="12" pageNumber="161" type="description">
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<paragraph id="DC472801FF9CFFD90B9BFBD70EFFE83C" blockId="12.[387,1276,1061,1160]" box="[387,1276,1097,1124]" pageId="12" pageNumber="161">
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<uri id="A8692403FF9CFFD90B9BFBD70EFFE83C" box="[387,1276,1097,1124]" pageId="12" pageNumber="161">
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urn:lsid:zoobank.org:act:
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<uuid id="A85E12D4FF9CFFD908BCFBD70EFFE83C" box="[676,1276,1097,1124]" pageId="12" pageNumber="161">86B4AAD0-D69E-4C79-BF3F-DE5E7DC0EDB1</uuid>
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</uri>
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</paragraph>
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<paragraph id="DC472801FF9CFFD90893FBF309F7E8D0" blockId="12.[387,1276,1061,1160]" box="[651,1012,1133,1160]" pageId="12" pageNumber="161">
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<figureCitation id="44C33484FF9CFFD90893FBF308D7E8D0" box="[651,724,1133,1160]" captionStart="Fig" captionStartId="1.[227,270,1965,1991]" captionTargetBox="[227,1437,751,1920]" captionTargetId="figure-216@1.[227,1437,751,1920]" captionTargetPageId="1" captionText="Fig. 1. A–B. Habitat and webs of Filistata betarif sp. nov. in Tinshemet cave. C. Paratype male in life (MACN-Ar 41793). D. Surroundings of Tinshemet cave. Note the entrance to the cave at the right side of the picture. Photos A–B by Shlomi Aharon and C–D by Ivan L.F. Magalhaes." figureDoi="http://doi.org/10.5281/zenodo.6916936" httpUri="https://zenodo.org/record/6916936/files/figure.png" pageId="12" pageNumber="161">
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Figs
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<geoCoordinate id="B9CC4EC6FF9CFFD908DDFBF008D7E8D0" box="[709,724,1134,1160]" degrees="1, 3" direction="east" orientation="longitude" pageId="12" pageNumber="161" precision="5555" value="1.3">1</geoCoordinate>
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</figureCitation>
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,
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<figureCitation id="44C33484FF9CFFD908FBFBF00945E8D0" box="[739,838,1134,1160]" captionStart="Fig" captionStartId="8.[227,270,1754,1780]" captionTargetBox="[273,1385,302,1702]" captionTargetId="figure-137@8.[264,1399,302,1703]" captionTargetPageId="8" captionText="Fig. 3. A–G. Endogynes Filistata Latreille, 1810, ventral view. A. Filistata wunderlichi Zonstein & Marusik, 2019 from Spain, Boca de los Frailes (CRBA IFM-2197). B. Filistata IFMsp191 from Israel, Odem (MACN-Ar 41262). C. Filistata insidiatrix (Forsskål, 1775) from France, Gonfaron (MACN-Ar 39461). D. Filistata insidiatrix from Israel, Beriniki (HUJ-INVAr 21052). E. Filistata betarif sp. nov. from Israel, Oah cave (HUJ-INVAr 21055). F. Filistata betarif sp. nov. from Israel, Tinshemet cave (MACN-Ar 41222). G. Filistata IFMsp99 (subadult ♀?) from Palestine, En Fashkha (HUJ-INVAr 20277). — H–I. ♂♂, left pedipalp, prolateral. H. Filistata betarif sp. nov., holotype from Israel, Tinshemet cave (HUJ -INVAr 21058). I. Filistata insidiatrix from Spain, Mallorca (MACN-Ar 42346). Scale bar = 0.1 mm, all figures to scale. Colours correspond to clades in Fig. 2." figureDoi="http://doi.org/10.5281/zenodo.6916944" httpUri="https://zenodo.org/record/6916944/files/figure.png" pageId="12" pageNumber="161">3E–F, H</figureCitation>
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,
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<figureCitation id="44C33484FF9CFFD9094AFBF309B8E8D0" box="[850,955,1133,1160]" captionStart="Fig" captionStartId="13.[227,270,1928,1954]" captionTargetBox="[227,1437,808,1883]" captionTargetId="figure-274@13.[227,1437,808,1883]" captionTargetPageId="13" captionText="Fig. 5. A, C, E. Filistata betarif sp. nov., paratype, ♂, from Israel, Tinshemet cave, (MACN-Ar 41793), left pedipalp. B, D, F. Filistata insidiatrix (Forsskål, 1775), ♂, from Spain, Mallorca (MACN-Ar 42347), right pedipalp, mirrored. A–B. Prolateral view. C–D. Dorsal view. E–F. Retrolateral view. Abbreviations: k = embolic keel; ps = prolateral setae. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6916952" httpUri="https://zenodo.org/record/6916952/files/figure.png" pageId="12" pageNumber="161">5A, C, E</figureCitation>
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,
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<figureCitation id="44C33484FF9CFFD909D0FBF009F7E8D0" box="[968,1012,1134,1160]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="15.[227,270,1901,1927]" captionStartId-1="16.[227,270,1901,1927]" captionStartId-2="17.[227,270,1965,1991]" captionTargetBox-0="[237,1430,362,1838]" captionTargetBox-1="[233,1434,365,1831]" captionTargetBox-2="[227,1437,1209,1925]" captionTargetId-0="figure-59@15.[227,1437,352,1846]" captionTargetId-1="figure-70@16.[227,1437,352,1846]" captionTargetId-2="figure-320@17.[227,1437,1209,1925]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionText-0="Fig. 7. Filistata betarif sp. nov., habitus. A–C. Holotype, ♂, from Tinshemet cave (HUJ -INVAr 21058). A. Dorsal view. B. Lateral view. C. Ventral view. — D–F. Paratype, ♀, from the same locality (MACN- Ar 41222). D. Lateral view. E. Dorsal view. F. Ventral view. Scale bars = 1 mm." captionText-1="Fig. 8. A–G. Filistata betarif sp. nov., holotype, ♂ (HUJ -INVAr 21058), left pedipalp. A–B, E. Prolateral view. Arrow points to prolateral strong setae in cymbium. C, F–G. Retrolateral view. Arrow points to embolic keel. D. Dorsal view. — H. Filistata insidiatrix (Forsskål, 1775) (MACN-Ar 39461), bulb, retrolateral view. Arrow points to embolic keel. Scale bars: A–F = 0.2 mm; G–H = 0.05 mm." captionText-2="Fig. 9. Filistata betarif sp. nov., female spermathecae, ventral. A. Oah cave (HUJ-INVAr 21055). B. Tinshemet cave (MACN-Ar 41222). C–D. Tinshemet cave (MACN-Ar 41791). Scale bars: A–C = 0.2 mm; D = 0.05 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6916960" figureDoi-1="http://doi.org/10.5281/zenodo.6916964" figureDoi-2="http://doi.org/10.5281/zenodo.6916968" httpUri-0="https://zenodo.org/record/6916960/files/figure.png" httpUri-1="https://zenodo.org/record/6916964/files/figure.png" httpUri-2="https://zenodo.org/record/6916968/files/figure.png" pageId="12" pageNumber="161">7–9</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="94E27B8AFF9CFFD90AFBFB2E09D9E892" box="[227,986,1200,1226]" pageId="12" pageNumber="161" type="reference_group">
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<paragraph id="DC472801FF9CFFD90AFBFB2E09D9E892" blockId="12.[227,986,1200,1226]" box="[227,986,1200,1226]" pageId="12" pageNumber="161">
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||
<taxonomicName id="1BF85382FF9CFFD90AFBFB2E0B45E892" ID-CoL="4K4M" authorityName="Latreille" authorityYear="1810" box="[227,326,1200,1226]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="161" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="EE8CF413FF9CFFD90AFBFB2E0B45E892" box="[227,326,1200,1226]" italics="true" pageId="12" pageNumber="161">Filistata</emphasis>
|
||
</taxonomicName>
|
||
IFMsp142 – Magalhaes & Ramírez in press: 11, 13, 16.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="94E27B8AFF9CFFD90AFBFB6F0F88EA4B" pageId="12" pageNumber="161" type="diagnosis">
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||
<paragraph id="DC472801FF9CFFD90AFBFB6F0B5FE953" blockId="12.[227,348,1265,1291]" box="[227,348,1265,1291]" pageId="12" pageNumber="161">
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||
<heading id="870F9F6DFF9CFFD90AFBFB6F0B5FE953" bold="true" box="[227,348,1265,1291]" fontSize="11" level="3" pageId="12" pageNumber="161" reason="3">
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||
<emphasis id="EE8CF413FF9CFFD90AFBFB6F0B5FE953" bold="true" box="[227,348,1265,1291]" pageId="12" pageNumber="161">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="DC472801FF9CFFD90AFBFABF0F88EA4B" blockId="12.[227,1436,1313,1555]" pageId="12" pageNumber="161">
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Males are similar to those of
|
||
<taxonomicName id="1BF85382FF9CFFD90856FABC08E4E963" baseAuthorityName="Forsskal" baseAuthorityYear="1775" box="[590,743,1313,1339]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="161" phylum="Arthropoda" rank="species" species="insidiatrix">
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||
<emphasis id="EE8CF413FF9CFFD90856FABC08E4E963" box="[590,743,1313,1339]" italics="true" pageId="12" pageNumber="161">F. insidiatrix</emphasis>
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||
</taxonomicName>
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||
and
|
||
<taxonomicName id="1BF85382FF9CFFD9093FFABC0997E963" authorityName="Zonstein & Marusik" authorityYear="2019" box="[807,916,1313,1339]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="161" phylum="Arthropoda" rank="species" species="albens">
|
||
<emphasis id="EE8CF413FF9CFFD9093FFABC0997E963" box="[807,916,1313,1339]" italics="true" pageId="12" pageNumber="161">F. albens</emphasis>
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||
</taxonomicName>
|
||
by having a pedipalp with elongate femur and tibia, and by a subconical copulatory bulb with terminally hooked embolus (
|
||
<figureCitation id="44C33484FF9CFFD90EB5FADB0F33E907" box="[1197,1328,1349,1375]" captionStart="Fig" captionStartId="16.[227,270,1901,1927]" captionTargetBox="[233,1434,365,1831]" captionTargetId="figure-70@16.[227,1437,352,1846]" captionTargetPageId="16" captionText="Fig. 8. A–G. Filistata betarif sp. nov., holotype, ♂ (HUJ -INVAr 21058), left pedipalp. A–B, E. Prolateral view. Arrow points to prolateral strong setae in cymbium. C, F–G. Retrolateral view. Arrow points to embolic keel. D. Dorsal view. — H. Filistata insidiatrix (Forsskål, 1775) (MACN-Ar 39461), bulb, retrolateral view. Arrow points to embolic keel. Scale bars: A–F = 0.2 mm; G–H = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6916964" httpUri="https://zenodo.org/record/6916964/files/figure.png" pageId="12" pageNumber="161">Fig. 8A–B</figureCitation>
|
||
; see also
|
||
<bibRefCitation id="B86955F0FF9CFFD90AFBFAF70821E9DB" author="Zonstein S. & Marusik Y. M." box="[227,546,1385,1411]" pageId="12" pageNumber="161" pagination="53 - 93" refId="ref15464" refString="Zonstein S. & Marusik Y. M. 2019. A revision of the spider genus Filistata (Araneae: Filistatidae). Arachnology 18 (2): 53 - 93. https: // doi. org / 10.13156 / arac. 2018.18.2.53" type="journal article" year="2019">Zonstein & Marusik 2019</bibRefCitation>
|
||
: fig. 27). They differ from both species by the conspicuous embolic keel (
|
||
<figureCitation id="44C33484FF9CFFD90AF3FA120B43E9FF" box="[235,320,1420,1447]" captionStart="Fig" captionStartId="16.[227,270,1901,1927]" captionTargetBox="[233,1434,365,1831]" captionTargetId="figure-70@16.[227,1437,352,1846]" captionTargetPageId="16" captionText="Fig. 8. A–G. Filistata betarif sp. nov., holotype, ♂ (HUJ -INVAr 21058), left pedipalp. A–B, E. Prolateral view. Arrow points to prolateral strong setae in cymbium. C, F–G. Retrolateral view. Arrow points to embolic keel. D. Dorsal view. — H. Filistata insidiatrix (Forsskål, 1775) (MACN-Ar 39461), bulb, retrolateral view. Arrow points to embolic keel. Scale bars: A–F = 0.2 mm; G–H = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6916964" httpUri="https://zenodo.org/record/6916964/files/figure.png" pageId="12" pageNumber="161">Fig. 8F</figureCitation>
|
||
(arrow), 5A, k) (inconspicuous keel in
|
||
<taxonomicName id="1BF85382FF9CFFD90910FA13099EE9FE" baseAuthorityName="Forsskal" baseAuthorityYear="1775" box="[776,925,1420,1446]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="161" phylum="Arthropoda" rank="species" species="insidiatrix">
|
||
<emphasis id="EE8CF413FF9CFFD90910FA13099EE9FE" box="[776,925,1420,1446]" italics="true" pageId="12" pageNumber="161">F. insidiatrix</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="1BF85382FF9CFFD909CCFA130E3FE9FE" authorityName="Zonstein & Marusik" authorityYear="2019" box="[980,1084,1420,1446]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="161" phylum="Arthropoda" rank="species" species="albens">
|
||
<emphasis id="EE8CF413FF9CFFD909CCFA130E3FE9FE" box="[980,1084,1420,1446]" italics="true" pageId="12" pageNumber="161">F. albens</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="44C33484FF9CFFD90E52FA120EA4E9FF" box="[1098,1191,1420,1447]" captionStart="Fig" captionStartId="13.[227,270,1928,1954]" captionTargetBox="[227,1437,808,1883]" captionTargetId="figure-274@13.[227,1437,808,1883]" captionTargetPageId="13" captionText="Fig. 5. A, C, E. Filistata betarif sp. nov., paratype, ♂, from Israel, Tinshemet cave, (MACN-Ar 41793), left pedipalp. B, D, F. Filistata insidiatrix (Forsskål, 1775), ♂, from Spain, Mallorca (MACN-Ar 42347), right pedipalp, mirrored. A–B. Prolateral view. C–D. Dorsal view. E–F. Retrolateral view. Abbreviations: k = embolic keel; ps = prolateral setae. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6916952" httpUri="https://zenodo.org/record/6916952/files/figure.png" pageId="12" pageNumber="161">Figs 5B</figureCitation>
|
||
,
|
||
<figureCitation id="44C33484FF9CFFD90EABFA130EEAE9FF" box="[1203,1257,1421,1447]" captionStart="Fig" captionStartId="19.[227,270,1684,1710]" captionTargetBox="[291,1376,303,1648]" captionTargetId="figure-0@19.[285,1377,302,1651]" captionTargetPageId="19" captionText="Fig. 10. Filistatid morphology. A. Filistata insidiatrix (Forsskål, 1775), ♀, leg tarsus, arrow points to macrosetae. B. Pritha nana (Simon, 1968), arrow points to right calamistrum. C. Sahastata aravaensis Ganem et al., 2022, left calamistrum. D. Zaitunia schmitzi Kulczyński, 1911, left calamistrum, arrow points to gap between setae. E. Filistata insidiatrix, left calamistrum. F. Filistata lubinae Zonstein & Marusik, 2019, ♂, tarsus I. G. Sahastata aravaensis, ♀. H. Pritha IFMsp185, ♀. I. Pritha sp., male abdomen, dorsal view. J. Zaitunia schmitzi, ♂. K. Zaitunia schmitzi, spermathecae ventral view. L. Pritha tenuispina (Strand, 1914), ♂, left pedipalp, prolateral view, arrow points to cymbium. M. Filistata lubinae, left copulatory bulb, prolateral view. N. Zaitunia schmitzi, ♂, left pedipalp, prolateral view, arrow points to cymbium. O. Filistata albens Zonstein & Marusik, 2019, left copulatory bulb, prolateral view. P. Zaitunia schmitzi, male clypeus, lateral view, arrow points to stout setae. Q. Filistata insidiatrix, female clypeus, lateral view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6916970" httpUri="https://zenodo.org/record/6916970/files/figure.png" pageId="12" pageNumber="161">10O</figureCitation>
|
||
). Females have a pair of large spermathecae that appear undivided (
|
||
<figureCitation id="44C33484FF9CFFD90958FA2E0984E993" box="[832,903,1456,1483]" captionStart="Fig" captionStartId="17.[227,270,1965,1991]" captionTargetBox="[227,1437,1209,1925]" captionTargetId="figure-320@17.[227,1437,1209,1925]" captionTargetPageId="17" captionText="Fig. 9. Filistata betarif sp. nov., female spermathecae, ventral. A. Oah cave (HUJ-INVAr 21055). B. Tinshemet cave (MACN-Ar 41222). C–D. Tinshemet cave (MACN-Ar 41791). Scale bars: A–C = 0.2 mm; D = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6916968" httpUri="https://zenodo.org/record/6916968/files/figure.png" pageId="12" pageNumber="161">Fig. 9</figureCitation>
|
||
); they fall within the morphological variation observed for
|
||
<taxonomicName id="1BF85382FF9CFFD90B67FA4B0816E9B6" baseAuthorityName="Forsskal" baseAuthorityYear="1775" box="[383,533,1492,1518]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="161" phylum="Arthropoda" rank="species" species="insidiatrix">
|
||
<emphasis id="EE8CF413FF9CFFD90B67FA4B0816E9B6" box="[383,533,1492,1518]" italics="true" pageId="12" pageNumber="161">F. insidiatrix</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="44C33484FF9CFFD9083EFA4A08A5E9B6" box="[550,678,1492,1518]" captionStart="Fig" captionStartId="8.[227,270,1754,1780]" captionTargetBox="[273,1385,302,1702]" captionTargetId="figure-137@8.[264,1399,302,1703]" captionTargetPageId="8" captionText="Fig. 3. A–G. Endogynes Filistata Latreille, 1810, ventral view. A. Filistata wunderlichi Zonstein & Marusik, 2019 from Spain, Boca de los Frailes (CRBA IFM-2197). B. Filistata IFMsp191 from Israel, Odem (MACN-Ar 41262). C. Filistata insidiatrix (Forsskål, 1775) from France, Gonfaron (MACN-Ar 39461). D. Filistata insidiatrix from Israel, Beriniki (HUJ-INVAr 21052). E. Filistata betarif sp. nov. from Israel, Oah cave (HUJ-INVAr 21055). F. Filistata betarif sp. nov. from Israel, Tinshemet cave (MACN-Ar 41222). G. Filistata IFMsp99 (subadult ♀?) from Palestine, En Fashkha (HUJ-INVAr 20277). — H–I. ♂♂, left pedipalp, prolateral. H. Filistata betarif sp. nov., holotype from Israel, Tinshemet cave (HUJ -INVAr 21058). I. Filistata insidiatrix from Spain, Mallorca (MACN-Ar 42346). Scale bar = 0.1 mm, all figures to scale. Colours correspond to clades in Fig. 2." figureDoi="http://doi.org/10.5281/zenodo.6916944" httpUri="https://zenodo.org/record/6916944/files/figure.png" pageId="12" pageNumber="161">Fig. 3C–D</figureCitation>
|
||
;
|
||
<bibRefCitation id="B86955F0FF9CFFD908ABFA4A09EBE9B6" author="Zonstein S. & Marusik Y. M." box="[691,1000,1492,1518]" pageId="12" pageNumber="161" pagination="53 - 93" refId="ref15464" refString="Zonstein S. & Marusik Y. M. 2019. A revision of the spider genus Filistata (Araneae: Filistatidae). Arachnology 18 (2): 53 - 93. https: // doi. org / 10.13156 / arac. 2018.18.2.53" type="journal article" year="2019">Zonstein & Marusik 2019</bibRefCitation>
|
||
: fig. 17d–h) and thus females are not diagnosable through morphology only; they can be diagnosed using DNA barcoding of the COI gene.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="94E27B8AFF9CFFD90AFBF9A40EF8EAFF" pageId="12" pageNumber="161" type="etymology">
|
||
<paragraph id="DC472801FF9CFFD90AFBF9A40B69EA0C" blockId="12.[227,362,1594,1620]" box="[227,362,1594,1620]" pageId="12" pageNumber="161">
|
||
<heading id="870F9F6DFF9CFFD90AFBF9A40B69EA0C" bold="true" box="[227,362,1594,1620]" fontSize="11" level="3" pageId="12" pageNumber="161" reason="3">
|
||
<emphasis id="EE8CF413FF9CFFD90AFBF9A40B69EA0C" bold="true" box="[227,362,1594,1620]" pageId="12" pageNumber="161">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="DC472801FF9CFFD90AFBF9F70EF8EAFF" blockId="12.[227,1437,1641,1704]" pageId="12" pageNumber="161">
|
||
The specific epithet is a noun in apposition and refers to Bet ‘Arif, the name of the ‘
|
||
<emphasis id="EE8CF413FF9CFFD90EAAF9F70EE8EADB" box="[1202,1259,1641,1667]" italics="true" pageId="12" pageNumber="161">wadi</emphasis>
|
||
’ (dry riverbed) in which the two caves where the species was collected (Tinshemet and Oah) are located.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="94E27B8AFF9CFFDB0AFBF94E0B29ED2F" lastPageId="14" lastPageNumber="163" pageId="12" pageNumber="161" type="materials_examined">
|
||
<paragraph id="DC472801FF9CFFD90AFBF94E0811EAB1" blockId="12.[227,530,1743,1770]" box="[227,530,1743,1770]" pageId="12" pageNumber="161">
|
||
<heading id="870F9F6DFF9CFFD90AFBF94E0811EAB1" bold="true" box="[227,530,1743,1770]" fontSize="11" level="3" pageId="12" pageNumber="161" reason="3">
|
||
<emphasis id="EE8CF413FF9CFFD90AFBF94E0811EAB1" bold="true" box="[227,530,1743,1770]" pageId="12" pageNumber="161">Type material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="DC472801FF9CFFD90B08F9610906EBDD" blockId="12.[227,1436,1791,1925]" pageId="12" pageNumber="161">
|
||
<materialsCitation id="6C90225CFF9CFFD90B08F9610906EBDD" ID-GBIF-Occurrence="3866864336" collectingDate="2020-02-20" collectionCode="HUJ-INVAr" collectorName="I. L. F. Magalhaes & E. Gavish- Regev & S. Aharon" country="Israel" county="Tinshemet cave" elevation="122" latitude="31.99938" location="Judean hills" longLatPrecision="1" longitude="34.96813" municipality="Arif" pageId="12" pageNumber="161" specimenCode="HUJ-INVAr 21058" specimenCount="123" specimenCount-male="1" typeStatus="holotype">
|
||
<emphasis id="EE8CF413FF9CFFD90B08F9610B81EB41" bold="true" box="[272,386,1791,1817]" pageId="12" pageNumber="161">
|
||
<typeStatus id="034396A3FF9CFFD90B08F9610B81EB41" box="[272,386,1791,1817]" pageId="12" pageNumber="161" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="A4EF6891FF9CFFD90AFBF8BD0B49EB65" box="[227,330,1827,1853]" name="Israel" pageId="12" pageNumber="161">ISRAEL</collectingCountry>
|
||
•
|
||
<specimenCount id="CAFEE388FF9CFFD90B46F8BD0B88EB65" box="[350,395,1827,1853]" pageId="12" pageNumber="161" type="male">1 ♂</specimenCount>
|
||
;
|
||
<location id="D9277EDAFF9CFFD90B8EF8BC0821EB64" LSID="urn:lsid:plazi:treatment:54519917FF9CFFC708A9FBBB08A5E88A:D9277EDAFF9CFFD90B8EF8BC0821EB64" box="[406,546,1826,1852]" country="Israel" county="Tinshemet cave" latitude="31.99938" longLatPrecision="1" longitude="34.96813" municipality="Arif" name="Judean hills" pageId="12" pageNumber="161">Judean hills</location>
|
||
,
|
||
<locationDeviation id="CDF53123FF9CFFD90834F8BD08CBEB64" box="[556,712,1826,1853]" pageId="12" pageNumber="161">near Shoham</locationDeviation>
|
||
,
|
||
<location id="D9277EDAFF9CFFD908CAF8BC0E60EB65" LSID="urn:lsid:plazi:treatment:54519917FF9CFFC708A9FBBB08A5E88A:D9277EDAFF9CFFD908CAF8BC0E60EB65" box="[722,1123,1826,1853]" country="Israel" county="Tinshemet cave" latitude="31.99938" longLatPrecision="1" longitude="34.96813" municipality="Arif" name="Hadom Shomeron Nature Reserve" pageId="12" pageNumber="161">Hadom Shomeron Nature Reserve</location>
|
||
;
|
||
<location id="D9277EDAFF9CFFD90E75F8BD0F1BEB64" LSID="urn:lsid:plazi:treatment:54519917FF9CFFC708A9FBBB08A5E88A:D9277EDAFF9CFFD90E75F8BD0F1BEB64" box="[1133,1304,1826,1853]" country="Israel" county="Tinshemet cave" latitude="31.99938" longLatPrecision="1" longitude="34.96813" municipality="Arif" name="Bet ' Arif wadi" pageId="12" pageNumber="161">Bet ‘Arif wadi</location>
|
||
,
|
||
<location id="D9277EDAFF9CFFD90F39F8BC0B1FEB39" LSID="urn:lsid:plazi:treatment:54519917FF9CFFC708A9FBBB08A5E88A:D9277EDAFF9CFFD90F39F8BC0B1FEB39" country="Israel" county="Tinshemet cave" latitude="31.99938" longLatPrecision="1" longitude="34.96813" municipality="Arif" name="Tinshemet cave" pageId="12" pageNumber="161">Tinshemet cave</location>
|
||
,
|
||
<location id="D9277EDAFF9CFFD90B30F8D80BACEB39" LSID="urn:lsid:plazi:treatment:54519917FF9CFFC708A9FBBB08A5E88A:D9277EDAFF9CFFD90B30F8D80BACEB39" box="[296,431,1862,1889]" country="Israel" county="Tinshemet cave" latitude="31.99938" longLatPrecision="1" longitude="34.96813" municipality="Arif" name="inside cave" pageId="12" pageNumber="161">inside cave</location>
|
||
;
|
||
<geoCoordinate id="B9CC4EC6FF9CFFD90BA3F8D90856EB39" box="[443,597,1863,1889]" degrees="31.99938" direction="north" orientation="latitude" pageId="12" pageNumber="161" precision="1" value="31.99938">31.99938° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B9CC4EC6FF9CFFD90879F8D808FBEB39" box="[609,760,1862,1889]" degrees="34.96813" direction="east" orientation="longitude" pageId="12" pageNumber="161" precision="1" value="34.96813">34.96813° E</geoCoordinate>
|
||
;
|
||
<specimenCount id="CAFEE388FF9CFFD9091DF8D9095EEB38" box="[773,861,1862,1889]" pageId="12" pageNumber="161" type="generic">
|
||
<quantity id="1B0085E4FF9CFFD9091DF8D9095BEB38" box="[773,856,1862,1889]" metricMagnitude="2" metricUnit="m" metricValue="1.22" pageId="12" pageNumber="161" unit="m" value="122.0">122 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
a.s.l.;
|
||
<date id="A8460EC1FF9CFFD90985F8D90E3FEB39" box="[925,1084,1862,1889]" pageId="12" pageNumber="161" value="2020-02-20">
|
||
<collectingDate id="B802F729FF9CFFD90985F8D90E3FEB39" box="[925,1084,1862,1889]" pageId="12" pageNumber="161" value="2020-02-20">20 Feb. 2020</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="710D4DD7FF9CFFD90E50F8D90F11EB38" box="[1096,1298,1862,1889]" pageId="12" pageNumber="161">I.L.F. Magalhaes</collectorName>
|
||
,
|
||
<collectorName id="710D4DD7FF9CFFD90F06F8D90B2DEBDD" pageId="12" pageNumber="161">E. Gavish- Regev</collectorName>
|
||
and
|
||
<collectorName id="710D4DD7FF9CFFD90B7FF8F40BDFEBDC" box="[359,476,1898,1924]" pageId="12" pageNumber="161">S. Aharon</collectorName>
|
||
leg.;
|
||
<specimenCode id="8C5E807AFF9CFFD90806F8F40902EBDD" box="[542,769,1898,1925]" collectionCode="HUJ-INVAr" pageId="12" pageNumber="161">HUJ-INVAr 21058</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="DC472801FF9CFFD80B08F83208D6EEB7" blockId="12.[225,1437,1964,2063]" lastBlockId="13.[227,1437,303,752]" lastPageId="13" lastPageNumber="162" pageId="12" pageNumber="161">
|
||
<materialsCitation id="6C90225CFF9CFFD80B08F8320B2FED12" ID-GBIF-Occurrence="3866864369" collectingDate="2014-05-25" collectionCode="HUJ-INVAr" collectorName="S. Aharon & E. Gavish-Regev" country="Israel" county="Oah cave" lastPageId="13" lastPageNumber="162" latitude="32.0053" location="Judean hills" longLatPrecision="6" longitude="34.9722" municipality="Arif" pageId="12" pageNumber="161" specimenCode="HUJ-INVAr 21055" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<emphasis id="EE8CF413FF9CFFD90B08F8320B8EEB9E" bold="true" box="[272,397,1964,1990]" pageId="12" pageNumber="161">
|
||
<typeStatus id="034396A3FF9CFFD90B08F8320B8EEB9E" box="[272,397,1964,1990]" pageId="12" pageNumber="161" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="A4EF6891FF9CFFD90AFBF84E0B49EBB2" box="[227,330,2000,2026]" name="Israel" pageId="12" pageNumber="161">ISRAEL</collectingCountry>
|
||
•
|
||
<specimenCount id="CAFEE388FF9CFFD90B47F84E0B89EBB3" box="[351,394,2000,2027]" pageId="12" pageNumber="161" type="female">1 ♀</specimenCount>
|
||
;
|
||
<location id="D9277EDAFF9CFFD90B80F84E0827EBB2" LSID="urn:lsid:plazi:treatment:54519917FF9CFFC708A9FBBB08A5E88A:D9277EDAFF9CFFD90B80F84E0827EBB2" box="[408,548,2000,2026]" country="Israel" county="Oah cave" latitude="32.0053" longLatPrecision="6" longitude="34.9722" municipality="Arif" name="Judean hills" pageId="12" pageNumber="161">Judean hills</location>
|
||
,
|
||
<locationDeviation id="CDF53123FF9CFFD90837F84F08C9EBB2" box="[559,714,2000,2027]" pageId="12" pageNumber="161">near Shoham</locationDeviation>
|
||
,
|
||
<location id="D9277EDAFF9CFFD908CDF84E0E6BEBB3" LSID="urn:lsid:plazi:treatment:54519917FF9CFFC708A9FBBB08A5E88A:D9277EDAFF9CFFD908CDF84E0E6BEBB3" box="[725,1128,2000,2027]" country="Israel" county="Oah cave" latitude="32.0053" longLatPrecision="6" longitude="34.9722" municipality="Arif" name="Hadom Shomeron Nature Reserve" pageId="12" pageNumber="161">Hadom Shomeron Nature Reserve</location>
|
||
;
|
||
<location id="D9277EDAFF9CFFD90E6AF84F0F1DEBB2" LSID="urn:lsid:plazi:treatment:54519917FF9CFFC708A9FBBB08A5E88A:D9277EDAFF9CFFD90E6AF84F0F1DEBB2" box="[1138,1310,2000,2027]" country="Israel" county="Oah cave" latitude="32.0053" longLatPrecision="6" longitude="34.9722" municipality="Arif" name="Bet ' Arif wadi" pageId="12" pageNumber="161">Bet ‘Arif wadi</location>
|
||
,
|
||
<location id="D9277EDAFF9CFFD90F30F84E0F9BEBB3" LSID="urn:lsid:plazi:treatment:54519917FF9CFFC708A9FBBB08A5E88A:D9277EDAFF9CFFD90F30F84E0F9BEBB3" box="[1320,1432,2000,2027]" country="Israel" county="Oah cave" latitude="32.0053" longLatPrecision="6" longitude="34.9722" municipality="Arif" name="Oah cave" pageId="12" pageNumber="161">Oah cave</location>
|
||
,
|
||
<location id="D9277EDAFF9CFFD90AF9F86A0B6AE457" LSID="urn:lsid:plazi:treatment:54519917FF9CFFC708A9FBBB08A5E88A:D9277EDAFF9CFFD90AF9F86A0B6AE457" box="[225,361,2036,2063]" country="Israel" county="Oah cave" latitude="32.0053" longLatPrecision="6" longitude="34.9722" municipality="Arif" name="inside cave" pageId="12" pageNumber="161">inside cave</location>
|
||
;
|
||
<geoCoordinate id="B9CC4EC6FF9CFFD90B6DF86A0801E457" box="[373,514,2036,2063]" degrees="32.0053" direction="north" orientation="latitude" pageId="12" pageNumber="161" precision="5" value="32.0053">32.0053° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B9CC4EC6FF9CFFD90816F86A0895E457" box="[526,662,2036,2063]" degrees="34.9722" direction="east" orientation="longitude" pageId="12" pageNumber="161" precision="5" value="34.9722">34.9722° E</geoCoordinate>
|
||
;
|
||
<date id="A8460EC1FF9CFFD908BBF86A0947E456" box="[675,836,2036,2063]" pageId="12" pageNumber="161" value="2014-05-25">
|
||
<collectingDate id="B802F729FF9CFFD908BBF86A0947E456" box="[675,836,2036,2063]" pageId="12" pageNumber="161" value="2014-05-25">25 May 2014</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="710D4DD7FF9CFFD90949F86A09C4E456" box="[849,967,2036,2062]" pageId="12" pageNumber="161">S. Aharon</collectorName>
|
||
and
|
||
<collectorName id="710D4DD7FF9CFFD90E19F86B0EC9E456" box="[1025,1226,2036,2063]" pageId="12" pageNumber="161">E. Gavish-Regev</collectorName>
|
||
leg.;
|
||
<specimenCode id="8C5E807AFF9CFFD80F15F86A0B2FED12" collectionCode="HUJ-INVAr" lastPageId="13" lastPageNumber="162" pageId="12" pageNumber="161">HUJ-INVAr 21055</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="6C90225CFF9DFFD80B5DFEB10B83ED35" ID-GBIF-Occurrence="3866864421" collectingDate="2014-03-09" collectionCode="HUJ" collectorName="S. Aharon & E. Gavish-Regev" country="Israel" county="Oah cave" latitude="32.0053" location="Judean" longLatPrecision="6" longitude="34.9722" municipality="Arif" pageId="13" pageNumber="162" specimenCode="HUJ- INVAr 21059" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
|
||
<specimenCount id="CAFEE388FF9DFFD80B5DFEB10B8BED11" box="[325,392,303,329]" pageId="13" pageNumber="162" type="female">2 ♀♀</specimenCount>
|
||
; same locality as for preceding;
|
||
<date id="A8460EC1FF9DFFD8091EFEB1099BED11" box="[774,920,303,329]" pageId="13" pageNumber="162" value="2014-03-09">
|
||
<collectingDate id="B802F729FF9DFFD8091EFEB1099BED11" box="[774,920,303,329]" pageId="13" pageNumber="162" value="2014-03-09">9 Mar. 2014</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="710D4DD7FF9DFFD809BCFEB10E1AED11" box="[932,1049,303,329]" pageId="13" pageNumber="162">S. Aharon</collectorName>
|
||
and
|
||
<collectorName id="710D4DD7FF9DFFD80E4AFEB10F18ED11" box="[1106,1307,303,329]" pageId="13" pageNumber="162">E. Gavish-Regev</collectorName>
|
||
leg.;
|
||
<specimenCode id="8C5E807AFF9DFFD80F45FEB10B83ED35" pageId="13" pageNumber="162">HUJ- INVAr 21059</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="6C90225CFF9DFFD80B8EFECC096DEDEA" ID-GBIF-Occurrence="3866864364" collectingDate="2020-02-20" collectionCode="MACN-Ar" collectorName="I. L. F. Magalhaes & E. Gavish-Regev & S. Aharon" country="Israel" county="Tinshemet cave" elevation="122" latitude="31.99938" location="Judean hills" longLatPrecision="1" longitude="34.96813" municipality="Arif" pageId="13" pageNumber="162" specimenCode="MACN-Ar 41222" specimenCount="129" specimenCount-female="6" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="CAFEE388FF9DFFD80B8EFECC0BC1ED34" box="[406,450,338,364]" pageId="13" pageNumber="162" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="CAFEE388FF9DFFD80BD4FECC080EED34" box="[460,525,338,364]" pageId="13" pageNumber="162" type="female">6 ♀♀</specimenCount>
|
||
;
|
||
<location id="D9277EDAFF9DFFD80802FECC08A6ED34" LSID="urn:lsid:plazi:treatment:54519917FF9CFFC708A9FBBB08A5E88A:D9277EDAFF9DFFD80802FECC08A6ED34" box="[538,677,338,364]" country="Israel" county="Tinshemet cave" latitude="31.99938" longLatPrecision="1" longitude="34.96813" municipality="Arif" name="Judean hills" pageId="13" pageNumber="162">Judean hills</location>
|
||
,
|
||
<locationDeviation id="CDF53123FF9DFFD808A8FECC0949ED34" box="[688,842,338,364]" pageId="13" pageNumber="162">near Shoham</locationDeviation>
|
||
,
|
||
<location id="D9277EDAFF9DFFD8094CFECC0EE7ED34" LSID="urn:lsid:plazi:treatment:54519917FF9CFFC708A9FBBB08A5E88A:D9277EDAFF9DFFD8094CFECC0EE7ED34" box="[852,1252,338,364]" country="Israel" county="Tinshemet cave" latitude="31.99938" longLatPrecision="1" longitude="34.96813" municipality="Arif" name="Hadom Shomeron Nature Reserve" pageId="13" pageNumber="162">Hadom Shomeron Nature Reserve</location>
|
||
,
|
||
<location id="D9277EDAFF9DFFD80EF6FECC0F9BED34" LSID="urn:lsid:plazi:treatment:54519917FF9CFFC708A9FBBB08A5E88A:D9277EDAFF9DFFD80EF6FECC0F9BED34" box="[1262,1432,338,364]" country="Israel" county="Tinshemet cave" latitude="31.99938" longLatPrecision="1" longitude="34.96813" municipality="Arif" name="Bet ' Arif wadi" pageId="13" pageNumber="162">Bet ‘Arif wadi</location>
|
||
,
|
||
<location id="D9277EDAFF9DFFD80AFBFEEB0B9CEDC8" LSID="urn:lsid:plazi:treatment:54519917FF9CFFC708A9FBBB08A5E88A:D9277EDAFF9DFFD80AFBFEEB0B9CEDC8" box="[227,415,373,400]" country="Israel" county="Tinshemet cave" latitude="31.99938" longLatPrecision="1" longitude="34.96813" municipality="Arif" name="Tinshemet cave" pageId="13" pageNumber="162">Tinshemet cave</location>
|
||
,
|
||
<location id="D9277EDAFF9DFFD80BB3FEEB0831EDC8" LSID="urn:lsid:plazi:treatment:54519917FF9CFFC708A9FBBB08A5E88A:D9277EDAFF9DFFD80BB3FEEB0831EDC8" box="[427,562,373,400]" country="Israel" county="Tinshemet cave" latitude="31.99938" longLatPrecision="1" longitude="34.96813" municipality="Arif" name="inside cave" pageId="13" pageNumber="162">inside cave</location>
|
||
;
|
||
<geoCoordinate id="B9CC4EC6FF9DFFD80827FEEB08DAEDC8" box="[575,729,373,400]" degrees="31.99938" direction="north" orientation="latitude" pageId="13" pageNumber="162" precision="1" value="31.99938">31.99938° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B9CC4EC6FF9DFFD808FEFEEB097FEDC8" box="[742,892,373,400]" degrees="34.96813" direction="east" orientation="longitude" pageId="13" pageNumber="162" precision="1" value="34.96813">34.96813° E</geoCoordinate>
|
||
,
|
||
<specimenCount id="CAFEE388FF9DFFD80991FEEB09E2EDD7" box="[905,993,373,399]" pageId="13" pageNumber="162" type="generic">
|
||
<quantity id="1B0085E4FF9DFFD80991FEEB09DFEDD7" box="[905,988,373,399]" metricMagnitude="2" metricUnit="m" metricValue="1.22" pageId="13" pageNumber="162" unit="m" value="122.0">122 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
a.s.l.;
|
||
<date id="A8460EC1FF9DFFD80E3AFEEB0EC2EDD7" box="[1058,1217,373,399]" pageId="13" pageNumber="162" value="2020-02-20">
|
||
<collectingDate id="B802F729FF9DFFD80E3AFEEB0EC2EDD7" box="[1058,1217,373,399]" pageId="13" pageNumber="162" value="2020-02-20">20 Feb. 2020</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="710D4DD7FF9DFFD80ED6FEE80F9BEDD7" box="[1230,1432,373,400]" pageId="13" pageNumber="162">I.L.F. Magalhaes</collectorName>
|
||
,
|
||
<collectorName id="710D4DD7FF9DFFD80AFBFE070BAFEDEA" box="[227,428,408,435]" pageId="13" pageNumber="162">E. Gavish-Regev</collectorName>
|
||
and
|
||
<collectorName id="710D4DD7FF9DFFD80BFDFE060859EDEA" box="[485,602,408,434]" pageId="13" pageNumber="162">S. Aharon</collectorName>
|
||
leg.;
|
||
<specimenCode id="8C5E807AFF9DFFD80885FE06096DEDEA" box="[669,878,408,434]" collectionCode="MACN-Ar" pageId="13" pageNumber="162">MACN-Ar 41222</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="6C90225CFF9DFFD8099FFE060877ED8E" ID-GBIF-Occurrence="3866864418" collectingDate="2020-02-20" collectionCode="HUJ-INVAr" collectorName="I. L. F. Magalhaes & E. Gavish-Regev & S. Aharon" country="Israel" county="Tinshemet cave" elevation="122" latitude="31.99938" location="Judean" longLatPrecision="1" longitude="34.96813" municipality="Arif" pageId="13" pageNumber="162" specimenCode="HUJ-INVAr 20275" specimenCount="7" specimenCount-female="3" specimenCount-male="2" typeStatus="paratype">
|
||
<specimenCount id="CAFEE388FF9DFFD8099FFE0609CFEDEB" box="[903,972,408,435]" pageId="13" pageNumber="162" type="male">2 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="CAFEE388FF9DFFD809C0FE060E1FEDEB" box="[984,1052,408,435]" pageId="13" pageNumber="162" type="female">3 ♀♀</specimenCount>
|
||
,
|
||
<specimenCount id="CAFEE388FF9DFFD80E32FE060E82EDEA" box="[1066,1153,408,434]" pageId="13" pageNumber="162" type="generic">2 imm.</specimenCount>
|
||
; same collection data as for preceding;
|
||
<specimenCode id="8C5E807AFF9DFFD80B8BFE220877ED8E" box="[403,628,443,470]" collectionCode="HUJ-INVAr" pageId="13" pageNumber="162">HUJ-INVAr 20275</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="6C90225CFF9DFFD8088AFE220B2FEDA1" ID-GBIF-Occurrence="3866864329" collectingDate="2020-02-20" collectionCode="MACN-Ar" collectorName="I. L. F. Magalhaes & E. Gavish-Regev & S. Aharon" country="Israel" county="Tinshemet cave" elevation="122" latitude="31.99938" location="Judean" longLatPrecision="1" longitude="34.96813" municipality="Arif" pageId="13" pageNumber="162" specimenCode="MACN-Ar 41791" specimenCount="5" specimenCount-female="4" typeStatus="paratype">
|
||
<specimenCount id="CAFEE388FF9DFFD8088AFE2208D4ED8E" box="[658,727,444,470]" pageId="13" pageNumber="162" type="female">4 ♀♀</specimenCount>
|
||
,
|
||
<specimenCount id="CAFEE388FF9DFFD808FEFE22093CED8D" box="[742,831,443,470]" pageId="13" pageNumber="162" type="generic">1 imm.</specimenCount>
|
||
; same collection data as for preceding;
|
||
<specimenCode id="8C5E807AFF9DFFD80F03FE220B2FEDA1" collectionCode="MACN-Ar" pageId="13" pageNumber="162">MACN-Ar 41791</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="6C90225CFF9DFFD80B51FE410E2FEDA1" ID-GBIF-Occurrence="3866864381" box="[329,1068,479,505]" collectingDate="2020-02-20" collectionCode="MACN-Ar" collectorName="I. L. F. Magalhaes & E. Gavish-Regev & S. Aharon" country="Israel" county="Tinshemet cave" elevation="122" latitude="31.99938" location="Judean" longLatPrecision="1" longitude="34.96813" municipality="Arif" pageId="13" pageNumber="162" specimenCode="MACN-Ar 41793" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="CAFEE388FF9DFFD80B51FE410B79EDA1" box="[329,378,479,505]" pageId="13" pageNumber="162" type="male">1 ♂</specimenCount>
|
||
; same collection data as for preceding;
|
||
<specimenCode id="8C5E807AFF9DFFD80941FE410E2FEDA1" box="[857,1068,479,505]" collectionCode="MACN-Ar" pageId="13" pageNumber="162">MACN-Ar 41793</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="6C90225CFF9DFFD80E52FE410943EE67" ID-GBIF-Occurrence="3866864396" collectedFrom="under stones" collectingDate="2020-02-20" collectionCode="MACN-Ar" collectorName="I. L. F. Magalhaes & E. Gavish-Regev & S. Aharon" country="Israel" county="Tinshemet cave" elevation="122" latitude="31.99938" location="Mediterranean scrubland" longLatPrecision="1" longitude="34.96813" municipality="Arif" pageId="13" pageNumber="162" specimenCode="MACN-Ar 41829" specimenCount="2" typeStatus="paratype">
|
||
<specimenCount id="CAFEE388FF9DFFD80E52FE410EA0EDA1" box="[1098,1187,479,505]" pageId="13" pageNumber="162" type="generic">2 imm.</specimenCount>
|
||
; same locality as for preceding,
|
||
<location id="D9277EDAFF9DFFD80B7DFD9C088FEE44" LSID="urn:lsid:plazi:treatment:54519917FF9CFFC708A9FBBB08A5E88A:D9277EDAFF9DFFD80B7DFD9C088FEE44" box="[357,652,514,540]" country="Israel" county="Tinshemet cave" latitude="31.99938" longLatPrecision="1" longitude="34.96813" municipality="Arif" name="Mediterranean scrubland" pageId="13" pageNumber="162">Mediterranean scrubland</location>
|
||
,
|
||
<locationDeviation id="CDF53123FF9DFFD80880FD9C095BEE44" box="[664,856,514,540]" pageId="13" pageNumber="162">outside the cave</locationDeviation>
|
||
,
|
||
<collectedFrom id="6FE46BFCFF9DFFD8097BFD9C09F9EE44" box="[867,1018,514,540]" pageId="13" pageNumber="162">under stones</collectedFrom>
|
||
;
|
||
<date id="A8460EC1FF9DFFD80E1EFD9C0EA0EE44" box="[1030,1187,514,540]" pageId="13" pageNumber="162" value="2020-02-20">
|
||
<collectingDate id="B802F729FF9DFFD80E1EFD9C0EA0EE44" box="[1030,1187,514,540]" pageId="13" pageNumber="162" value="2020-02-20">20 Feb. 2020</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="710D4DD7FF9DFFD80EB7FD9C0F74EE44" box="[1199,1399,514,540]" pageId="13" pageNumber="162">I.L.F. Magalhaes</collectorName>
|
||
,
|
||
<collectorName id="710D4DD7FF9DFFD80F9BFD9C0B88EE67" pageId="13" pageNumber="162">E. Gavish-Regev</collectorName>
|
||
and
|
||
<collectorName id="710D4DD7FF9DFFD80BA7FDBB0830EE67" box="[447,563,549,575]" pageId="13" pageNumber="162">S. Aharon</collectorName>
|
||
leg.;
|
||
<specimenCode id="8C5E807AFF9DFFD80869FDBB0943EE67" box="[625,832,549,575]" collectionCode="MACN-Ar" pageId="13" pageNumber="162">MACN-Ar 41829</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="6C90225CFF9DFFD8094DFDBB0BF8EEDE" ID-GBIF-Occurrence="3866864395" collectedFrom="outside the cave" collectingDate="2014-05-25" collectionCode="HUJ-INVAr" collectorName="S. Aharon & E. Gavish-Regev" country="Israel" county="Tinshemet cave" elevation="122" latitude="31.9994" location="Mediterranean scrubland" longLatPrecision="6" longitude="34.9681" municipality="Arif" pageId="13" pageNumber="162" specimenCode="HUJ-INVAr 21056" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<specimenCount id="CAFEE388FF9DFFD8094DFDBB0983EE18" box="[853,896,549,576]" pageId="13" pageNumber="162" type="female">1 ♀</specimenCount>
|
||
; same locality as for preceding;
|
||
<location id="D9277EDAFF9DFFD80EE9FDBB0B5BEE3A" LSID="urn:lsid:plazi:treatment:54519917FF9CFFC708A9FBBB08A5E88A:D9277EDAFF9DFFD80EE9FDBB0B5BEE3A" country="Israel" county="Tinshemet cave" latitude="31.9994" longLatPrecision="6" longitude="34.9681" municipality="Arif" name="Mediterranean scrubland" pageId="13" pageNumber="162">Mediterranean scrubland</location>
|
||
,
|
||
<collectedFrom id="6FE46BFCFF9DFFD80B7EFDD60829EE3B" box="[358,554,584,611]" pageId="13" pageNumber="162">outside the cave</collectedFrom>
|
||
;
|
||
<geoCoordinate id="B9CC4EC6FF9DFFD80820FDD608C6EE3B" box="[568,709,584,611]" degrees="31.9994" direction="north" orientation="latitude" pageId="13" pageNumber="162" precision="5" value="31.9994">31.9994° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B9CC4EC6FF9DFFD808CBFDD6095FEE3B" box="[723,860,584,611]" degrees="34.9681" direction="east" orientation="longitude" pageId="13" pageNumber="162" precision="5" value="34.9681">34.9681° E</geoCoordinate>
|
||
;
|
||
<date id="A8460EC1FF9DFFD80973FDD60E0CEE3A" box="[875,1039,584,611]" pageId="13" pageNumber="162" value="2014-05-25">
|
||
<collectingDate id="B802F729FF9DFFD80973FDD60E0CEE3A" box="[875,1039,584,611]" pageId="13" pageNumber="162" value="2014-05-25">25 May 2014</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="710D4DD7FF9DFFD80E05FDD60E96EE3A" box="[1053,1173,584,610]" pageId="13" pageNumber="162">S. Aharon</collectorName>
|
||
and
|
||
<collectorName id="710D4DD7FF9DFFD80ECAFDD70F9FEE3A" box="[1234,1436,584,611]" pageId="13" pageNumber="162">E. Gavish-Regev</collectorName>
|
||
leg.;
|
||
<specimenCode id="8C5E807AFF9DFFD80B05FDF20BF8EEDE" box="[285,507,619,646]" collectionCode="HUJ-INVAr" pageId="13" pageNumber="162">HUJ-INVAr 21056</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="6C90225CFF9DFFD8080BFDF20E32EEF1" ID-GBIF-Occurrence="3866864427" collectedFrom="inside cave" collectingDate="2014-05-25" collectionCode="HUJ-INVAr" collectorName="S. Aharon & E. Gavish-Regev" country="Israel" county="Tinshemet cave" elevation="122" latitude="31.99938" location="Mediterranean" longLatPrecision="1" longitude="34.96813" municipality="Arif" pageId="13" pageNumber="162" specimenCode="HUJ-INVAr 20301" specimenCount="7" specimenCount-female="7" typeStatus="paratype">
|
||
<specimenCount id="CAFEE388FF9DFFD8080BFDF20856EEDE" box="[531,597,620,646]" pageId="13" pageNumber="162" type="female">7 ♀♀</specimenCount>
|
||
; same locality as for preceding;
|
||
<collectedFrom id="6FE46BFCFF9DFFD809D6FDF50E50EEDE" box="[974,1107,619,646]" pageId="13" pageNumber="162">inside cave</collectedFrom>
|
||
;
|
||
<geoCoordinate id="B9CC4EC6FF9DFFD80E47FDF20EF4EEDE" box="[1119,1271,620,646]" degrees="31.99938" direction="north" orientation="latitude" pageId="13" pageNumber="162" precision="1" value="31.99938">31.99938° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B9CC4EC6FF9DFFD80F1AFDF50F94EEDE" box="[1282,1431,619,646]" degrees="34.96813" direction="east" orientation="longitude" pageId="13" pageNumber="162" precision="1" value="34.96813">34.96813° E</geoCoordinate>
|
||
;
|
||
<date id="A8460EC1FF9DFFD80AFBFD110B86EEF1" box="[227,389,655,682]" pageId="13" pageNumber="162" value="2014-05-25">
|
||
<collectingDate id="B802F729FF9DFFD80AFBFD110B86EEF1" box="[227,389,655,682]" pageId="13" pageNumber="162" value="2014-05-25">25 May 2014</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="710D4DD7FF9DFFD80B8AFD11080BEEF1" box="[402,520,655,681]" pageId="13" pageNumber="162">S. Aharon</collectorName>
|
||
and
|
||
<collectorName id="710D4DD7FF9DFFD8085BFD11090EEEF1" box="[579,781,655,681]" pageId="13" pageNumber="162">E. Gavish-Regev</collectorName>
|
||
leg.;
|
||
<specimenCode id="8C5E807AFF9DFFD80949FD110E32EEF1" box="[849,1073,655,681]" collectionCode="HUJ-INVAr" pageId="13" pageNumber="162">HUJ-INVAr 20301</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="6C90225CFF9DFFD80E54FD11086CEE95" ID-GBIF-Occurrence="3866864326" collectedFrom="inside cave" collectingDate="2014-05-25" collectionCode="HUJ-INVAr" collectorName="S. Aharon & E. Gavish-Regev" country="Israel" county="Tinshemet cave" elevation="122" latitude="31.99938" location="Mediterranean" longLatPrecision="1" longitude="34.96813" municipality="Arif" pageId="13" pageNumber="162" specimenCode="HUJ-INVAr 21057" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<specimenCount id="CAFEE388FF9DFFD80E54FD110E79EEF1" box="[1100,1146,655,681]" pageId="13" pageNumber="162" type="female">1 ♀</specimenCount>
|
||
; same collection data as for preceding;
|
||
<specimenCode id="8C5E807AFF9DFFD80B88FD2C086CEE95" box="[400,623,690,717]" collectionCode="HUJ-INVAr" pageId="13" pageNumber="162">HUJ-INVAr 21057</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="6C90225CFF9DFFD80891FD2C08D3EEB7" ID-GBIF-Occurrence="3866864393" collectedFrom="inside cave" collectingDate="2014-03-09" collectionCode="HUJ-INVAr" collectorName="S. Aharon & E. Gavish-Regev" country="Israel" county="Tinshemet cave" elevation="122" latitude="31.99938" location="Mediterranean" longLatPrecision="1" longitude="34.96813" municipality="Arif" pageId="13" pageNumber="162" specimenCode="HUJ-INVAr 21060" specimenCount="1" typeStatus="paratype">
|
||
3 imm; same locality as for preceding;
|
||
<date id="A8460EC1FF9DFFD80E4EFD2C0EEBEE94" box="[1110,1256,690,716]" pageId="13" pageNumber="162" value="2014-03-09">
|
||
<collectingDate id="B802F729FF9DFFD80E4EFD2C0EEBEE94" box="[1110,1256,690,716]" pageId="13" pageNumber="162" value="2014-03-09">9 Mar. 2014</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="710D4DD7FF9DFFD80EEDFD2C0F69EE94" box="[1269,1386,690,716]" pageId="13" pageNumber="162">S. Aharon</collectorName>
|
||
and
|
||
<collectorName id="710D4DD7FF9DFFD80AFBFD480BA8EEB7" box="[227,427,725,752]" pageId="13" pageNumber="162">E. Gavish-Regev</collectorName>
|
||
leg.;
|
||
<specimenCode id="8C5E807AFF9DFFD80BF5FD4B08D3EEB7" box="[493,720,725,751]" collectionCode="HUJ-INVAr" pageId="13" pageNumber="162">HUJ-INVAr 21060</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="88877889FF9DFFD80AFBF8160E5CE455" ID-DOI="http://doi.org/10.5281/zenodo.6916952" ID-Zenodo-Dep="6916952" httpUri="https://zenodo.org/record/6916952/files/figure.png" pageId="13" pageNumber="162" startId="13.[227,270,1928,1954]" targetBox="[227,1437,808,1883]" targetPageId="13">
|
||
<paragraph id="DC472801FF9DFFD80AFBF8160E5CE455" blockId="13.[227,1437,1928,2061]" pageId="13" pageNumber="162">
|
||
<emphasis id="EE8CF413FF9DFFD80AFBF8160B92EBFB" bold="true" box="[227,401,1928,1955]" pageId="13" pageNumber="162">Fig. 5. A, C, E</emphasis>
|
||
.
|
||
<taxonomicName id="1BF85382FF9DFFD80B87F8160854EBFA" authority="Magalhaes & Aharon & Ganem & Gavish-Regev, 2022" authorityName="Magalhaes & Aharon & Ganem & Gavish-Regev" authorityYear="2022" box="[415,599,1928,1954]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="162" phylum="Arthropoda" rank="species" species="betarif" status="sp. nov.">
|
||
<emphasis id="EE8CF413FF9DFFD80B87F8160854EBFA" box="[415,599,1928,1954]" italics="true" pageId="13" pageNumber="162">Filistata betarif</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F5BF4968FF9DFFD80846F81708B9EBFB" box="[606,698,1929,1955]" pageId="13" pageNumber="162" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype, ♂, from Israel, Tinshemet cave, (MACN-Ar 41793), left pedipalp.
|
||
<emphasis id="EE8CF413FF9DFFD80B97F8320BF1EB9E" bold="true" box="[399,498,1964,1990]" pageId="13" pageNumber="162">B, D, F</emphasis>
|
||
.
|
||
<taxonomicName id="1BF85382FF9DFFD8081EF83209C7EB9E" authority="(Forsskal, 1775)" baseAuthorityName="Forsskal" baseAuthorityYear="1775" box="[518,964,1963,1990]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="162" phylum="Arthropoda" rank="species" species="insidiatrix">
|
||
<emphasis id="EE8CF413FF9DFFD8081EF83208ECEB9E" box="[518,751,1964,1990]" italics="true" pageId="13" pageNumber="162">Filistata insidiatrix</emphasis>
|
||
(
|
||
<bibRefCitation id="B86955F0FF9DFFD8091BF83509B9EB9E" author="Forsskal P." box="[771,954,1963,1990]" pageId="13" pageNumber="162" refId="ref13586" refString="Forsskal P. 1775. Descriptiones animalium avium, amphibiorum, piscium, insectorum, vermium; quae in itinere orientali observavit Petrus Forskal [sic]. Moller, Havniae [= Copenhagen]. https: // doi. org / 10.5962 / bhl. title. 2154" type="book" year="1775">Forsskål, 1775</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, ♂, from Spain, Mallorca (MACN-Ar 42347), right pedipalp, mirrored.
|
||
<emphasis id="EE8CF413FF9DFFD80866F85108B6EBB1" bold="true" box="[638,693,1999,2025]" pageId="13" pageNumber="162">A–B</emphasis>
|
||
. Prolateral view.
|
||
<emphasis id="EE8CF413FF9DFFD80988F85109CAEBB1" bold="true" box="[912,969,1999,2025]" pageId="13" pageNumber="162">C–D</emphasis>
|
||
. Dorsal view.
|
||
<emphasis id="EE8CF413FF9DFFD80E98F8510EB7EBB1" bold="true" box="[1152,1204,1999,2025]" pageId="13" pageNumber="162">E–F</emphasis>
|
||
. Retrolateral view. Abbreviations: k = embolic keel; ps = prolateral setae. Scale bars = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DC472801FF9EFFDB0AFBFEB00859ED10" blockId="14.[227,602,302,328]" box="[227,602,302,328]" pageId="14" pageNumber="163">
|
||
<heading id="870F9F6DFF9EFFDB0AFBFEB00859ED10" bold="true" box="[227,602,302,328]" fontSize="11" level="3" pageId="14" pageNumber="163" reason="3">
|
||
<emphasis id="EE8CF413FF9EFFDB0AFBFEB00859ED10" bold="true" box="[227,602,302,328]" pageId="14" pageNumber="163">Additional material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="DC472801FF9EFFDB0AFBFEC30B29ED2F" blockId="14.[227,298,349,375]" box="[227,298,349,375]" pageId="14" pageNumber="163">None.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="94E27B8AFF9EFFC70AFBFE000E2FE866" lastPageId="18" lastPageNumber="167" pageId="14" pageNumber="163" type="description">
|
||
<paragraph id="DC472801FF9EFFDB0AFBFE000B77EDE0" blockId="14.[227,372,414,440]" box="[227,372,414,440]" pageId="14" pageNumber="163">
|
||
<heading id="870F9F6DFF9EFFDB0AFBFE000B77EDE0" bold="true" box="[227,372,414,440]" fontSize="11" level="3" pageId="14" pageNumber="163" reason="3">
|
||
<emphasis id="EE8CF413FF9EFFDB0AFBFE000B77EDE0" bold="true" box="[227,372,414,440]" pageId="14" pageNumber="163">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="DC472801FF9EFFDB0B08FE5308BCEDB0" blockId="14.[227,1437,461,839]" box="[272,703,461,488]" pageId="14" pageNumber="163">
|
||
<emphasis id="EE8CF413FF9EFFDB0B08FE530B53EDBF" bold="true" box="[272,336,461,487]" pageId="14" pageNumber="163">Male</emphasis>
|
||
(
|
||
<typeStatus id="034396A3FF9EFFDB0B47FE530BC9EDBF" box="[351,458,461,487]" pageId="14" pageNumber="163" type="holotype">holotype</typeStatus>
|
||
, HUJ-INVAr 21058)
|
||
</paragraph>
|
||
<paragraph id="DC472801FF9EFFC40AFBFE6E0971EDD7" blockId="14.[227,1437,461,839]" lastBlockId="17.[227,1436,303,399]" lastPageId="17" lastPageNumber="166" pageId="14" pageNumber="163">
|
||
Coloration yellowish cream except where noted. Carapace lined with black pigment, with marbled brown clypeal markings and brown, V-shaped median pattern. Legs I–III with three brown ventral spots. Abdomen brownish gray. Anterior margin of the carapace unmodified, eye apodemes absent. Sternum subrounded, sigilla not visible. Total length 5.25. Carapace length 2.38, width 1.88. Clypeus length 0.28. Eye diameters and interdistances: AME 0.14, PME 0.15, ALE 0.20, PLE 0.18, AME–AME 0.06, PME– PME 0.14. Sternum length 1.26, width 1.05. Pedipalp: femur length 2.34, height 0.30, tibia length 2.10, height 0.34. Leg I: 14.88 (3.82, 1.09, 3.94, 3.91, 2.12). II: 11.29 (3.05, 0.94, 2.78, 3.17, 1.35). III: 10.23 (2.70, 0.84, 2.52, 2.91, 1.26). IV: 12.52 (3.57, 0.90, 3.28, 3.44, 1.33). Abdomen: length 2.97, width 1.72. Pedipalp macrosetae on ventral surface of femur. Leg macrosetae: fe I d1-0-0, dp0-0-2, ti I v0-2-2-2, mt I v2-2-2-4, fe II d1-1-0, dp0-0-2, ti II v0-2-2, mt II v2-2-4, fe III d1-0-0, dp0-0-1, dr0-0-2, ti III d1-2-2, v1-1, mt III dr0-0-1, v1-2-2-4, fe IV d1-1-0, dr0-0-2, ti IV v1-1, mt IV v1-1-2-4, all tarsi with ventral macrosetae. Pedipalp as in diagnosis (
|
||
<figureCitation id="44C33484FF81FFC408B2FECC0956ED34" box="[682,853,338,365]" captionStart="Fig" captionStartId="13.[227,270,1928,1954]" captionTargetBox="[227,1437,808,1883]" captionTargetId="figure-274@13.[227,1437,808,1883]" captionTargetPageId="13" captionText="Fig. 5. A, C, E. Filistata betarif sp. nov., paratype, ♂, from Israel, Tinshemet cave, (MACN-Ar 41793), left pedipalp. B, D, F. Filistata insidiatrix (Forsskål, 1775), ♂, from Spain, Mallorca (MACN-Ar 42347), right pedipalp, mirrored. A–B. Prolateral view. C–D. Dorsal view. E–F. Retrolateral view. Abbreviations: k = embolic keel; ps = prolateral setae. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6916952" httpUri="https://zenodo.org/record/6916952/files/figure.png" pageId="17" pageNumber="166">Figs 5A, C, E</figureCitation>
|
||
,
|
||
<figureCitation id="44C33484FF81FFC4097BFECC09B3ED34" box="[867,944,338,364]" captionStart="Fig" captionStartId="16.[227,270,1901,1927]" captionTargetBox="[233,1434,365,1831]" captionTargetId="figure-70@16.[227,1437,352,1846]" captionTargetPageId="16" captionText="Fig. 8. A–G. Filistata betarif sp. nov., holotype, ♂ (HUJ -INVAr 21058), left pedipalp. A–B, E. Prolateral view. Arrow points to prolateral strong setae in cymbium. C, F–G. Retrolateral view. Arrow points to embolic keel. D. Dorsal view. — H. Filistata insidiatrix (Forsskål, 1775) (MACN-Ar 39461), bulb, retrolateral view. Arrow points to embolic keel. Scale bars: A–F = 0.2 mm; G–H = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6916964" httpUri="https://zenodo.org/record/6916964/files/figure.png" pageId="17" pageNumber="166">8A–G</figureCitation>
|
||
). State of the specimen: good, right leg I removed for molecular analysis, left pedipalp dissected.
|
||
</paragraph>
|
||
<caption id="88877889FF9EFFDB0AFBF817099FE455" ID-DOI="http://doi.org/10.5281/zenodo.6916956" ID-Zenodo-Dep="6916956" httpUri="https://zenodo.org/record/6916956/files/figure.png" pageId="14" pageNumber="163" startId="14.[227,270,1929,1955]" targetBox="[227,1437,898,1883]" targetPageId="14">
|
||
<paragraph id="DC472801FF9EFFDB0AFBF817099FE455" blockId="14.[227,1436,1929,2061]" pageId="14" pageNumber="163">
|
||
<emphasis id="EE8CF413FF9EFFDB0AFBF8170B31EBFB" bold="true" box="[227,306,1929,1955]" pageId="14" pageNumber="163">Fig. 6.</emphasis>
|
||
<taxonomicName id="1BF85382FF9EFFDB0B23F81708EDEBFC" authority="(Forsskal, 1775)" baseAuthorityName="Forsskal" baseAuthorityYear="1775" box="[315,750,1929,1956]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="163" phylum="Arthropoda" rank="species" species="insidiatrix">
|
||
<emphasis id="EE8CF413FF9EFFDB0B23F8170823EBFB" box="[315,544,1929,1955]" italics="true" pageId="14" pageNumber="163">Filistata insidiatrix</emphasis>
|
||
(
|
||
<bibRefCitation id="B86955F0FF9EFFDB0829F81708E6EBFC" author="Forsskal P." box="[561,741,1929,1956]" pageId="14" pageNumber="163" refId="ref13586" refString="Forsskal P. 1775. Descriptiones animalium avium, amphibiorum, piscium, insectorum, vermium; quae in itinere orientali observavit Petrus Forskal [sic]. Moller, Havniae [= Copenhagen]. https: // doi. org / 10.5962 / bhl. title. 2154" type="book" year="1775">Forsskål, 1775</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, left bulb, prolateral (A, C, E, G) and dorsal (B, D, F, H) views.
|
||
<emphasis id="EE8CF413FF9EFFDB0B22F8320B71EB9E" bold="true" box="[314,370,1964,1990]" pageId="14" pageNumber="163">A–B</emphasis>
|
||
. France (MACN-Ar 39461, ♂ collected with ♀ IFM-1702).
|
||
<emphasis id="EE8CF413FF9EFFDB0E78F8320E9AEB9E" bold="true" box="[1120,1177,1964,1990]" pageId="14" pageNumber="163">C–D</emphasis>
|
||
. Spain (CRBA IFM- 2196).
|
||
<emphasis id="EE8CF413FF9EFFDB0B2FF84E0B68EBB2" bold="true" box="[311,363,2000,2026]" pageId="14" pageNumber="163">E–F</emphasis>
|
||
. Israel, Ornit cave (MACN-Ar 41813, IFM-2175).
|
||
<emphasis id="EE8CF413FF9EFFDB09D6F8510E08EBB1" bold="true" box="[974,1035,1999,2025]" pageId="14" pageNumber="163">G–H</emphasis>
|
||
. Israel, Sharakh cave (MACN-Ar 41218, IFM-2144). Scale bar = 0.1 mm, all figures to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="88877889FF9FFFDA0AFBF8F30E92EB96" ID-DOI="http://doi.org/10.5281/zenodo.6916960" ID-Zenodo-Dep="6916960" httpUri="https://zenodo.org/record/6916960/files/figure.png" pageId="15" pageNumber="164" startId="15.[227,270,1901,1927]" targetBox="[237,1430,362,1838]" targetPageId="15">
|
||
<paragraph id="DC472801FF9FFFDA0AFBF8F30E92EB96" blockId="15.[227,1437,1901,1998]" pageId="15" pageNumber="164">
|
||
<emphasis id="EE8CF413FF9FFFDA0AFBF8F30B2CEBDF" bold="true" box="[227,303,1901,1927]" pageId="15" pageNumber="164">Fig. 7.</emphasis>
|
||
<taxonomicName id="1BF85382FF9FFFDA0B2DF8F30BEEEBDF" authority="Magalhaes & Aharon & Ganem & Gavish-Regev, 2022" authorityName="Magalhaes & Aharon & Ganem & Gavish-Regev" authorityYear="2022" box="[309,493,1901,1927]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="164" phylum="Arthropoda" rank="species" species="betarif" status="sp. nov.">
|
||
<emphasis id="EE8CF413FF9FFFDA0B2DF8F30BEEEBDF" box="[309,493,1901,1927]" italics="true" pageId="15" pageNumber="164">Filistata betarif</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F5BF4968FF9FFFDA0BEBF8F3084CEBDF" box="[499,591,1901,1927]" pageId="15" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, habitus.
|
||
<emphasis id="EE8CF413FF9FFFDA08A4F8F308F6EBDF" bold="true" box="[700,757,1901,1927]" pageId="15" pageNumber="164">A–C</emphasis>
|
||
. Holotype, ♂, from Tinshemet cave (HUJ -INVAr 21058).
|
||
<emphasis id="EE8CF413FF9FFFDA0AFBF80E0AFBEBF2" bold="true" box="[227,248,1936,1962]" pageId="15" pageNumber="164">A</emphasis>
|
||
. Dorsal view.
|
||
<emphasis id="EE8CF413FF9FFFDA0BB8F80E0BB7EBF2" bold="true" box="[416,436,1936,1962]" pageId="15" pageNumber="164">B</emphasis>
|
||
. Lateral view.
|
||
<emphasis id="EE8CF413FF9FFFDA0879F80E0875EBF2" bold="true" box="[609,630,1936,1962]" pageId="15" pageNumber="164">C</emphasis>
|
||
. Ventral view. —
|
||
<emphasis id="EE8CF413FF9FFFDA0950F80E097EEBF2" bold="true" box="[840,893,1936,1962]" pageId="15" pageNumber="164">D–F</emphasis>
|
||
. Paratype, ♀, from the same locality (MACN- Ar 41222).
|
||
<emphasis id="EE8CF413FF9FFFDA0B73F82D0B83EB95" bold="true" box="[363,384,1971,1997]" pageId="15" pageNumber="164">D</emphasis>
|
||
. Lateral view.
|
||
<emphasis id="EE8CF413FF9FFFDA0837F82D0840EB95" bold="true" box="[559,579,1971,1997]" pageId="15" pageNumber="164">E</emphasis>
|
||
. Dorsal view.
|
||
<emphasis id="EE8CF413FF9FFFDA08F4F82D08FDEB95" bold="true" box="[748,766,1971,1997]" pageId="15" pageNumber="164">F</emphasis>
|
||
. Ventral view. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="88877889FF80FFC50AFBF8F30F23EBA9" ID-DOI="http://doi.org/10.5281/zenodo.6916964" ID-Zenodo-Dep="6916964" httpUri="https://zenodo.org/record/6916964/files/figure.png" pageId="16" pageNumber="165" startId="16.[227,270,1901,1927]" targetBox="[233,1434,365,1831]" targetPageId="16">
|
||
<paragraph id="DC472801FF80FFC50AFBF8F30F23EBA9" blockId="16.[227,1436,1901,2033]" pageId="16" pageNumber="165">
|
||
<emphasis id="EE8CF413FF80FFC50AFBF8F30B6EEBDF" bold="true" box="[227,365,1901,1927]" pageId="16" pageNumber="165">Fig. 8. A–G</emphasis>
|
||
.
|
||
<taxonomicName id="1BF85382FF80FFC50B62F8F30833EBDF" authority="Magalhaes & Aharon & Ganem & Gavish-Regev, 2022" authorityName="Magalhaes & Aharon & Ganem & Gavish-Regev" authorityYear="2022" box="[378,560,1901,1927]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="165" phylum="Arthropoda" rank="species" species="betarif" status="sp. nov.">
|
||
<emphasis id="EE8CF413FF80FFC50B62F8F30833EBDF" box="[378,560,1901,1927]" italics="true" pageId="16" pageNumber="165">Filistata betarif</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F5BF4968FF80FFC5082DF8F30893EBDF" box="[565,656,1901,1927]" pageId="16" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂ (HUJ -INVAr 21058), left pedipalp.
|
||
<emphasis id="EE8CF413FF80FFC50EDEF8F30F1DEBDF" bold="true" box="[1222,1310,1901,1927]" pageId="16" pageNumber="165">A–B, E</emphasis>
|
||
. Prolateral view. Arrow points to prolateral strong setae in cymbium.
|
||
<emphasis id="EE8CF413FF80FFC50982F80E09F4EBF2" bold="true" box="[922,1015,1936,1962]" pageId="16" pageNumber="165">C, F–G</emphasis>
|
||
. Retrolateral view. Arrow points to embolic keel.
|
||
<emphasis id="EE8CF413FF80FFC50B96F82D0BA0EB95" bold="true" box="[398,419,1971,1997]" pageId="16" pageNumber="165">D</emphasis>
|
||
. Dorsal view. —
|
||
<emphasis id="EE8CF413FF80FFC50867F82D0895EB95" bold="true" box="[639,662,1971,1997]" pageId="16" pageNumber="165">H</emphasis>
|
||
.
|
||
<taxonomicName id="1BF85382FF80FFC508B0F82D0E5FEB96" authority="(Forsskal, 1775)" baseAuthorityName="Forsskal" baseAuthorityYear="1775" box="[680,1116,1971,1998]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="165" phylum="Arthropoda" rank="species" species="insidiatrix">
|
||
<emphasis id="EE8CF413FF80FFC508B0F82D098CEB95" box="[680,911,1971,1997]" italics="true" pageId="16" pageNumber="165">Filistata insidiatrix</emphasis>
|
||
(
|
||
<bibRefCitation id="B86955F0FF80FFC509B9F82D0E57EB96" author="Forsskal P." box="[929,1108,1971,1998]" pageId="16" pageNumber="165" refId="ref13586" refString="Forsskal P. 1775. Descriptiones animalium avium, amphibiorum, piscium, insectorum, vermium; quae in itinere orientali observavit Petrus Forskal [sic]. Moller, Havniae [= Copenhagen]. https: // doi. org / 10.5962 / bhl. title. 2154" type="book" year="1775">Forsskål, 1775</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(MACN-Ar 39461), bulb, retrolateral view. Arrow points to embolic keel. Scale bars: A–F = 0.2 mm; G–H = 0.05 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DC472801FF81FFC40B08FE2908C8ED89" blockId="17.[227,1437,439,852]" box="[272,715,439,465]" pageId="17" pageNumber="166">
|
||
<emphasis id="EE8CF413FF81FFC40B08FE290B68ED89" bold="true" box="[272,363,439,465]" pageId="17" pageNumber="166">Female</emphasis>
|
||
(
|
||
<typeStatus id="034396A3FF81FFC40B63FE290BE6ED89" box="[379,485,439,465]" pageId="17" pageNumber="166" type="paratype">paratype</typeStatus>
|
||
, MACN-Ar 41222)
|
||
</paragraph>
|
||
<paragraph id="DC472801FF81FFC40AFBFE440954EF0C" blockId="17.[227,1437,439,852]" pageId="17" pageNumber="166">
|
||
Coloration as in male. Eye apodemes absent. Sternum subrounded, with one pair of posterior sigilla. Total length 7.76. Carapace length 3.17, width 2.51. Clypeus length 0.40. Eye diameters and interdistances: AME 0.14, PME 0.19, ALE 0.27, PLE 0.23, AME–AME 0.04, PME–PME 0.23. Sternum length 1.64, width 1.50. Pedipalp: femur length 1.70, height 0.67, tibia length 1.06, height 0.65. Leg I: 13.60 (3.66, 1.37, 3.49, 3.23, 1.85). II: 9.66 (2.84, 1.13, 2.26, 2.16, 1.27). III: 7.82 (2.34, 1.03, 1.66, 1.64, 1.15). IV: 10.54 (3.30, 1.25, 2.64, 2.27, 1.08). Abdomen: length 4.67, width 3.08. Palpal macrosetae on ventral surface of tibia and tarsus. Leg macrosetae: fe I d1-0-0, ti I v0-2-0, mt I v2-2-2-4, fe II d1-0-0, ti II v0- 2-0, mt II v2-2-2-4, fe III d1-0-0, ti III d0-2-0, v0-1-2, mt III dr1-1-1, v1-2-2-4, fe IV d1-0-0, ti IV v1-1, mt IV v1-1-1-4, all tarsi with ventral macrosetae. Calamistrum with three staggered rows with <6 setae; calamistrum gap present. Epigastric furrow unmodified. Endogyne of females from the same locality as in the diagnosis (
|
||
<figureCitation id="44C33484FF81FFC40BB3FCA40BF0EF0C" box="[427,499,826,852]" captionStart="Fig" captionStartId="17.[227,270,1965,1991]" captionTargetBox="[227,1437,1209,1925]" captionTargetId="figure-320@17.[227,1437,1209,1925]" captionTargetPageId="17" captionText="Fig. 9. Filistata betarif sp. nov., female spermathecae, ventral. A. Oah cave (HUJ-INVAr 21055). B. Tinshemet cave (MACN-Ar 41222). C–D. Tinshemet cave (MACN-Ar 41791). Scale bars: A–C = 0.2 mm; D = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6916968" httpUri="https://zenodo.org/record/6916968/files/figure.png" pageId="17" pageNumber="166">Fig. 9</figureCitation>
|
||
). State of the specimen: good.
|
||
</paragraph>
|
||
<paragraph id="DC472801FF81FFC40AFBFCE50B5AEFCD" blockId="17.[227,345,891,917]" box="[227,345,891,917]" pageId="17" pageNumber="166">
|
||
<heading id="870F9F6DFF81FFC40AFBFCE50B5AEFCD" bold="true" box="[227,345,891,917]" fontSize="11" level="3" pageId="17" pageNumber="166" reason="3">
|
||
<emphasis id="EE8CF413FF81FFC40AFBFCE50B5AEFCD" bold="true" box="[227,345,891,917]" pageId="17" pageNumber="166">Variation</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="DC472801FF81FFC40AFBFC340E5CE875" blockId="17.[227,1437,938,1070]" pageId="17" pageNumber="166">Males (N = 3): total length 4.89–5.67 (5.27), carapace length 2.25–2.78 (2.47), femur I length 3.62–4.04 (3.83), femur/carapace ratio 1.45–1.61 (1.56). Females (N = 5): total length 6.47–8.95 (7.79), carapace length 3.17–3.98 (3.57), femur I length 3.5–4.18 (3.86), femur/carapace ratio 1.01–1.15 (1.08). The shape of the spermathecae varies only slightly among the dissected females.</paragraph>
|
||
<caption id="88877889FF81FFC40AFBF8330BF3E456" ID-DOI="http://doi.org/10.5281/zenodo.6916968" ID-Zenodo-Dep="6916968" httpUri="https://zenodo.org/record/6916968/files/figure.png" pageId="17" pageNumber="166" startId="17.[227,270,1965,1991]" targetBox="[227,1437,1209,1925]" targetPageId="17">
|
||
<paragraph id="DC472801FF81FFC40AFBF8330BF3E456" blockId="17.[227,1437,1965,2062]" pageId="17" pageNumber="166">
|
||
<emphasis id="EE8CF413FF81FFC40AFBF8330B36EB9F" bold="true" box="[227,309,1965,1991]" pageId="17" pageNumber="166">Fig. 9.</emphasis>
|
||
<taxonomicName id="1BF85382FF81FFC40B5AF8330803EB9F" authority="Magalhaes & Aharon & Ganem & Gavish-Regev, 2022" authorityName="Magalhaes & Aharon & Ganem & Gavish-Regev" authorityYear="2022" box="[322,512,1965,1991]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="166" phylum="Arthropoda" rank="species" species="betarif" status="sp. nov.">
|
||
<emphasis id="EE8CF413FF81FFC40B5AF8330803EB9F" box="[322,512,1965,1991]" italics="true" pageId="17" pageNumber="166">Filistata betarif</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F5BF4968FF81FFC40814F833086DEB9F" box="[524,622,1965,1991]" pageId="17" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, female spermathecae, ventral.
|
||
<emphasis id="EE8CF413FF81FFC409ECF8330E0AEB9F" bold="true" box="[1012,1033,1965,1991]" pageId="17" pageNumber="166">A</emphasis>
|
||
. Oah cave (HUJ-INVAr 21055).
|
||
<emphasis id="EE8CF413FF81FFC40AFBF84E0AF4EBB2" bold="true" box="[227,247,2000,2026]" pageId="17" pageNumber="166">B</emphasis>
|
||
. Tinshemet cave (MACN-Ar 41222).
|
||
<emphasis id="EE8CF413FF81FFC408D8F84E08FAEBB2" bold="true" box="[704,761,2000,2026]" pageId="17" pageNumber="166">C–D</emphasis>
|
||
. Tinshemet cave (MACN-Ar 41791). Scale bars: A–C = 0.2 mm; D = 0.05 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DC472801FF82FFC70AFBFEB00BA7ED10" blockId="18.[227,420,302,328]" box="[227,420,302,328]" pageId="18" pageNumber="167">
|
||
<heading id="870F9F6DFF82FFC70AFBFEB00BA7ED10" bold="true" box="[227,420,302,328]" fontSize="11" level="3" pageId="18" pageNumber="167" reason="3">
|
||
<emphasis id="EE8CF413FF82FFC70AFBFEB00BA7ED10" bold="true" box="[227,420,302,328]" pageId="18" pageNumber="167">Natural history</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="DC472801FF82FFC70AFBFEC30EE6EF11" blockId="18.[227,1437,349,841]" pageId="18" pageNumber="167">
|
||
<taxonomicName id="1BF85382FF82FFC70AFBFEC30B99ED2F" authority="Magalhaes & Aharon & Ganem & Gavish-Regev, 2022" authorityName="Magalhaes & Aharon & Ganem & Gavish-Regev" authorityYear="2022" box="[227,410,349,375]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="167" phylum="Arthropoda" rank="species" species="betarif" status="sp. nov.">
|
||
<emphasis id="EE8CF413FF82FFC70AFBFEC30B99ED2F" box="[227,410,349,375]" italics="true" pageId="18" pageNumber="167">Filistata betarif</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F5BF4968FF82FFC70BB9FEC00BF9ED20" box="[417,506,350,376]" pageId="18" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is a troglophilic species, inhabiting both epigean and hypogean (caves) habitats. It is known from the mesic Mediterranean region of
|
||
<collectingCountry id="A4EF6891FF82FFC70953FE1F098CEDC3" box="[843,911,385,411]" name="Israel" pageId="18" pageNumber="167">Israel</collectingCountry>
|
||
, in scrubland habitats, and usually hides in a refugium made of cribellate silk in crevices, holes or under stones and on rocky walls (
|
||
<figureCitation id="44C33484FF82FFC70EFAFE3B0F67EDE7" box="[1250,1380,421,447]" captionStart="Fig" captionStartId="1.[227,270,1965,1991]" captionTargetBox="[227,1437,751,1920]" captionTargetId="figure-216@1.[227,1437,751,1920]" captionTargetPageId="1" captionText="Fig. 1. A–B. Habitat and webs of Filistata betarif sp. nov. in Tinshemet cave. C. Paratype male in life (MACN-Ar 41793). D. Surroundings of Tinshemet cave. Note the entrance to the cave at the right side of the picture. Photos A–B by Shlomi Aharon and C–D by Ivan L.F. Magalhaes." figureDoi="http://doi.org/10.5281/zenodo.6916936" httpUri="https://zenodo.org/record/6916936/files/figure.png" pageId="18" pageNumber="167">Fig. 1A–B</figureCitation>
|
||
). All specimens were collected from Bet ‘Arif wadi near Shoham, Hadom Shomeron Nature Reserve, central
|
||
<collectingCountry id="A4EF6891FF82FFC70AFBFE720B2BEE5E" box="[227,296,492,518]" name="Israel" pageId="18" pageNumber="167">Israel</collectingCountry>
|
||
. The species was found in low abundances outside of caves (
|
||
<figureCitation id="44C33484FF82FFC709FAFE720E38EE5F" box="[994,1083,492,519]" captionStart="Fig" captionStartId="1.[227,270,1965,1991]" captionTargetBox="[227,1437,751,1920]" captionTargetId="figure-216@1.[227,1437,751,1920]" captionTargetPageId="1" captionText="Fig. 1. A–B. Habitat and webs of Filistata betarif sp. nov. in Tinshemet cave. C. Paratype male in life (MACN-Ar 41793). D. Surroundings of Tinshemet cave. Note the entrance to the cave at the right side of the picture. Photos A–B by Shlomi Aharon and C–D by Ivan L.F. Magalhaes." figureDoi="http://doi.org/10.5281/zenodo.6916936" httpUri="https://zenodo.org/record/6916936/files/figure.png" pageId="18" pageNumber="167">Fig. 1D</figureCitation>
|
||
) and in very high abundances, year-around, in Tinshemet cave that is characterized by a high guano amount of the Egyptian fruit bat [
|
||
<taxonomicName id="1BF85382FF82FFC70AF5FDAB08C5EE16" authority="(Geoffroy, 1810)" baseAuthorityName="Geoffroy" baseAuthorityYear="1810" box="[237,710,563,590]" class="Mammalia" family="Pteropodidae" genus="Rousettus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="167" phylum="Chordata" rank="species" species="aegyptiacus">
|
||
<emphasis id="EE8CF413FF82FFC70AF5FDAB0BF1EE16" box="[237,498,565,590]" italics="true" pageId="18" pageNumber="167">Rousettus aegyptiacus</emphasis>
|
||
(Geoffroy, 1810)
|
||
</taxonomicName>
|
||
]. Tinshemet (Arabic name: Mugharet al-Watwat) is a small to medium-sized Paleolithic cave, formed in a limestone rock of the Turonian Bina formation. It is located on the east bank of Bet ‘Arif wadi (Arabic name: Wadi Adasiyeh), 95 meters a.s.l. and 15 meters above the wadi (
|
||
<bibRefCitation id="B86955F0FF82FFC70B4DFD010847EEE2" author="Frumkin A. & Langford B. & Marder O. & Ullman M." box="[341,580,671,698]" pageId="18" pageNumber="167" pagination="246 - 258" refId="ref13636" refString="Frumkin A., Langford B., Marder O. & Ullman M. 2016. Paleolithic caves and hillslope processes in south-western Samaria, Israel: Environmental and archaeological implications. Quaternary International 398: 246 - 258. https: // doi. org / 10.1016 / j. quaint. 2015.05.064" type="journal article" year="2016">
|
||
Frumkin
|
||
<emphasis id="EE8CF413FF82FFC70BDBFD3E0BF5EEE1" box="[451,502,671,697]" italics="true" pageId="18" pageNumber="167">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
). It has a main opening on the side of the cliff, and an additional chimney opening, which the frugivorous bats frequently use (personal observation).
|
||
<taxonomicName id="1BF85382FF82FFC70E57FD5D0F04EE85" authority="Magalhaes & Aharon & Ganem & Gavish-Regev, 2022" authorityName="Magalhaes & Aharon & Ganem & Gavish-Regev" authorityYear="2022" box="[1103,1287,707,733]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="167" phylum="Arthropoda" rank="species" species="betarif" status="sp. nov.">
|
||
<emphasis id="EE8CF413FF82FFC70E57FD5D0F04EE85" box="[1103,1287,707,733]" italics="true" pageId="18" pageNumber="167">Filistata betarif</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F5BF4968FF82FFC70F16FD5A0F6BEE86" box="[1294,1384,708,734]" pageId="18" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
|
||
was also found in lower abundances in the nearby Oah cave that houses no bats (
|
||
<bibRefCitation id="B86955F0FF82FFC70E68FD790F31EF59" author="Cuff J. P. & Aharon S. & Armiach Steinpress I. & Seifan M. & Lubin Y. & Gavish-Regev E." box="[1136,1330,743,769]" pageId="18" pageNumber="167" pagination="576" refId="ref13474" refString="Cuff J. P., Aharon S., Armiach Steinpress I., Seifan M., Lubin Y. & Gavish-Regev E. 2021. It's all about the zone: spider assemblages in different ecological zones of Levantine caves. Diversity 13 (576). https: // doi. org / 10.3390 / d 13110576" type="journal article" year="2021">
|
||
Cuff
|
||
<emphasis id="EE8CF413FF82FFC70EA8FD760EE6EF59" box="[1200,1253,743,769]" italics="true" pageId="18" pageNumber="167">et al</emphasis>
|
||
. 2021
|
||
</bibRefCitation>
|
||
; Gavish- Regev
|
||
<emphasis id="EE8CF413FF82FFC70B20FC920B6CEF7D" box="[312,367,779,805]" italics="true" pageId="18" pageNumber="167">et al</emphasis>
|
||
. 2021). Oah is a small cave, situated on the same east bank of Bet ‘Arif wadi, north to Tinshemet cave, and is formed in a similar rock and age (Boaz Langford, pers. comm.).
|
||
</paragraph>
|
||
<paragraph id="DC472801FF82FFC70AFBFCEF0E2FE866" blockId="18.[227,1437,880,1086]" pageId="18" pageNumber="167">
|
||
In both caves,
|
||
<taxonomicName id="1BF85382FF82FFC70B82FCEE0854EFD2" authority="Magalhaes & Aharon & Ganem & Gavish-Regev, 2022" authorityName="Magalhaes & Aharon & Ganem & Gavish-Regev" authorityYear="2022" box="[410,599,880,906]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="167" phylum="Arthropoda" rank="species" species="betarif" status="sp. nov.">
|
||
<emphasis id="EE8CF413FF82FFC70B82FCEE0854EFD2" box="[410,599,880,906]" italics="true" pageId="18" pageNumber="167">Filistata betarif</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F5BF4968FF82FFC7087AFCEF08C2EFD3" box="[610,705,881,907]" pageId="18" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
|
||
was found mainly at the entrance zones, on the cave walls (
|
||
<figureCitation id="44C33484FF82FFC70AF3FC0A0B6CEFF6" box="[235,367,916,942]" captionStart="Fig" captionStartId="1.[227,270,1965,1991]" captionTargetBox="[227,1437,751,1920]" captionTargetId="figure-216@1.[227,1437,751,1920]" captionTargetPageId="1" captionText="Fig. 1. A–B. Habitat and webs of Filistata betarif sp. nov. in Tinshemet cave. C. Paratype male in life (MACN-Ar 41793). D. Surroundings of Tinshemet cave. Note the entrance to the cave at the right side of the picture. Photos A–B by Shlomi Aharon and C–D by Ivan L.F. Magalhaes." figureDoi="http://doi.org/10.5281/zenodo.6916936" httpUri="https://zenodo.org/record/6916936/files/figure.png" pageId="18" pageNumber="167">Fig. 1A–B</figureCitation>
|
||
). Other caves surveyed by us, at the mesic Mediterranean, semi-arid, and arid regions of
|
||
<collectingCountry id="A4EF6891FF82FFC70AFBFC260B27EF8A" box="[227,292,952,978]" name="Israel" pageId="18" pageNumber="167">Israel</collectingCountry>
|
||
and
|
||
<collectingCountry id="A4EF6891FF82FFC70B7CFC260BD2EF8A" box="[356,465,952,978]" name="Palestine" pageId="18" pageNumber="167">Palestine</collectingCountry>
|
||
, were not found to be occupied by
|
||
<taxonomicName id="1BF85382FF82FFC7099FFC2709F6EF8A" authority="Magalhaes & Aharon & Ganem & Gavish-Regev, 2022" authorityName="Magalhaes & Aharon & Ganem & Gavish-Regev" authorityYear="2022" box="[903,1013,952,978]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="167" phylum="Arthropoda" rank="species" species="betarif" status="sp. nov.">
|
||
<emphasis id="EE8CF413FF82FFC7099FFC2709F6EF8A" box="[903,1013,952,978]" italics="true" pageId="18" pageNumber="167">F. betarif</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F5BF4968FF82FFC70E18FC270E62EF8B" box="[1024,1121,953,979]" pageId="18" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, but instead by its sibling troglophilic species
|
||
<taxonomicName id="1BF85382FF82FFC70BC4FC43087AEFAE" baseAuthorityName="Forsskal" baseAuthorityYear="1775" box="[476,633,988,1014]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="167" phylum="Arthropoda" rank="species" species="insidiatrix">
|
||
<emphasis id="EE8CF413FF82FFC70BC4FC43087AEFAE" box="[476,633,988,1014]" italics="true" pageId="18" pageNumber="167">F. insidiatrix</emphasis>
|
||
</taxonomicName>
|
||
[Berniki, Yir’on, Yonim (Galilee); Ezba’, Oren (Karmel); Sal’it (northern Samaria); Andartat HaBiqa’ (central
|
||
<collectingCountry id="A4EF6891FF82FFC7091EFB9E0957E842" box="[774,852,1024,1050]" name="Jordan" pageId="18" pageNumber="167">Jordan</collectingCountry>
|
||
Valley); Perat (northern Judean desert); Qumeran (northern Dead-Sea area)] (
|
||
<bibRefCitation id="B86955F0FF82FFC7083CFBBD08E0E866" author="Cuff J. P. & Aharon S. & Armiach Steinpress I. & Seifan M. & Lubin Y. & Gavish-Regev E." box="[548,739,1059,1086]" pageId="18" pageNumber="167" pagination="576" refId="ref13474" refString="Cuff J. P., Aharon S., Armiach Steinpress I., Seifan M., Lubin Y. & Gavish-Regev E. 2021. It's all about the zone: spider assemblages in different ecological zones of Levantine caves. Diversity 13 (576). https: // doi. org / 10.3390 / d 13110576" type="journal article" year="2021">
|
||
Cuff
|
||
<emphasis id="EE8CF413FF82FFC7087BFBBA0895E865" box="[611,662,1059,1085]" italics="true" pageId="18" pageNumber="167">et al</emphasis>
|
||
. 2021
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B86955F0FF82FFC708F7FBBD0E1CE866" author="Gavish-Regev E. & Aharon S. & Steinpress I. A. & Seifan M. & Lubin Y." box="[751,1055,1059,1086]" pageId="18" pageNumber="167" pagination="5" refId="ref13691" refString="Gavish-Regev E., Aharon S., Steinpress I. A., Seifan M. & Lubin Y. 2021. A primer on spider assemblages in Levantine caves: the neglected subterranean habitats of the levant - a biodiversity mine. Diversity 13 (5). https: // doi. org / 10.3390 / d 13050179" type="journal article" year="2021">
|
||
Gavish-Regev
|
||
<emphasis id="EE8CF413FF82FFC70986FBBA09D2E865" box="[926,977,1059,1085]" italics="true" pageId="18" pageNumber="167">et al</emphasis>
|
||
. 2021
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="94E27B8AFF82FFC70AFBFBFB08A5E88A" pageId="18" pageNumber="167" type="distribution">
|
||
<paragraph id="DC472801FF82FFC70AFBFBFB0B7FE827" blockId="18.[227,380,1125,1151]" box="[227,380,1125,1151]" pageId="18" pageNumber="167">
|
||
<heading id="870F9F6DFF82FFC70AFBFBFB0B7FE827" bold="true" box="[227,380,1125,1151]" fontSize="11" level="3" pageId="18" pageNumber="167" reason="3">
|
||
<emphasis id="EE8CF413FF82FFC70AFBFBFB0B7FE827" bold="true" box="[227,380,1125,1151]" pageId="18" pageNumber="167">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="DC472801FF82FFC70AFBFB0B08A5E88A" blockId="18.[227,1437,1172,1235]" pageId="18" pageNumber="167">
|
||
Known only from two caves and under stones in their surroundings in Bet ‘Arif wadi, Hadom Shomeron Nature Reserve, central
|
||
<collectingCountry id="A4EF6891FF82FFC70BE6FB26083CE88A" box="[510,575,1208,1234]" name="Israel" pageId="18" pageNumber="167">Israel</collectingCountry>
|
||
(
|
||
<figureCitation id="44C33484FF82FFC70857FB26089BE88A" box="[591,664,1208,1234]" captionStart="Fig" captionStartId="11.[227,270,1792,1818]" captionTargetBox="[227,1430,352,1735]" captionTargetId="figure-3270@11.[282,837,852,1279]" captionTargetPageId="11" captionText="Fig. 4. Distribution maps. A. Filistata insidiatrix (Forsskål, 1775) in the Mediterranean and Middle East (note the species has been recorded from Angola, Venezuela and the islands of Socotra, Azores and Cabo Verde, but these records seem to be introduced); literature records have been taken from Benoit (1968), Brignoli (1982), Marusik & Zonstein (2014), Marusik & Zamani (2015a) and Zonstein & Marusik (2019). B. Filistata insidiatrix (circles) and F. betarif sp. nov. (star) in Israel and Palestine; only specimens examined by us are included in the map. Darker shades of orange are records that include at least one male specimen. C. Other species of Filistata Latreille, 1810 in Israel (except F. insidiatrix)." figureDoi="http://doi.org/10.5281/zenodo.6916946" httpUri="https://zenodo.org/record/6916946/files/figure.png" pageId="18" pageNumber="167">Fig. 4</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |