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<mods:title id="DD338F82090D1DD46D4D56D93B19C0D7">A new semi-cryptic Filistata from caves in the Levant with comments on the limits of Filistata insidiatrix (Forsskål, 1775) (Arachnida: Araneae: Filistatidae)</mods:title>
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<mods:namePart id="F14178C5F3CF8F8E1D5B329ACE619767">Magalhaes, Ivan L. F.</mods:namePart>
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<mods:affiliation id="246447D732737F8C0CCEDC6E589EF0FB">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ”, Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Av. Ángel Gallardo 470, C 1405 DJR, Buenos Aires, Argentina.</mods:affiliation>
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<mods:namePart id="30B0C87742D2B0FC74107801CE1A84CB">Aharon, Shlomi</mods:namePart>
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<mods:affiliation id="8E3579541DF5F3D51A190613429857A8">The National Natural History Collections, Edmond J. Safra Campus, Givat Ram, The Hebrew University of Jerusalem, Jerusalem 9190401, Israel. &amp; The Department of Ecology, Evolution and Behavior, Edmond J. Safra Campus, Givat Ram, The Hebrew University of Jerusalem, Jerusalem 9190401, Israel.</mods:affiliation>
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<mods:namePart id="D596040C5DB3BA416EDE5D518DDAC0DD">Ganem, Zeana</mods:namePart>
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<mods:affiliation id="EDB6CA399A3489EE51A05D73AA711233">The National Natural History Collections, Edmond J. Safra Campus, Givat Ram, The Hebrew University of Jerusalem, Jerusalem 9190401, Israel. &amp; The Department of Ecology, Evolution and Behavior, Edmond J. Safra Campus, Givat Ram, The Hebrew University of Jerusalem, Jerusalem 9190401, Israel.</mods:affiliation>
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<mods:affiliation id="AECED81F1F85B6F67000807603625456">The National Natural History Collections, Edmond J. Safra Campus, Givat Ram, The Hebrew University of Jerusalem, Jerusalem 9190401, Israel.</mods:affiliation>
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Key to the 
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of 
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and 
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The key below can be used to identify known species of crevice weavers from 
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and 
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. For more information, readers are referred to recent taxonomic revisions of 
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<emphasis id="EE8CF413FF82FFC70E62FAC60EDEE929" box="[1146,1245,1368,1393]" italics="true" pageId="18" pageNumber="167">Zaitunia</emphasis>
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(
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), 
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<emphasis id="EE8CF413FF82FFC70857FAE508B1E9CD" box="[591,690,1403,1429]" italics="true" pageId="18" pageNumber="167">Filistata</emphasis>
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(
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; 
<bibRefCitation id="B86955F0FF82FFC70E3CFAE50F5FE9CD" author="Zamani A. &amp; Marusik Y. M." box="[1060,1372,1403,1429]" pageId="18" pageNumber="167" pagination="121 - 126" refId="ref15367" refString="Zamani A. &amp; Marusik Y. M. 2020. New species of Filistatidae, Palpimanidae and Scytodidae (Arachnida: Araneae) from southern Iran. Acta Arachnologica 69 (2): 121 - 126. https: // doi. org / 10.2476 / asjaa. 69.121" type="journal article" year="2020">Zamani &amp; Marusik 2020</bibRefCitation>
) and 
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<emphasis id="EE8CF413FF82FFC70AFBFA010B54E9E1" box="[227,343,1439,1465]" italics="true" pageId="18" pageNumber="167">Sahastata</emphasis>
<bibRefCitation id="B86955F0FF82FFC70B46FA010BF4E9E1" author="Benoit P. L. G." box="[350,503,1439,1465]" pageId="18" pageNumber="167" pagination="92 - 102" refId="ref13241" refString="Benoit P. L. G. 1968. Synopsis des Filistatidae africains. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot; 77: 92 - 102." type="journal article" year="1968">Benoit, 1968</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="B86955F0FF82FFC7081FFA010912E9E1" author="Magalhaes I. L. F. &amp; Stockmann M. &amp; Marusik Y. M. &amp; Zonstein S. L." box="[519,785,1439,1466]" pageId="18" pageNumber="167" pagination="215 - 246" refId="ref14527" refString="Magalhaes I. L. F., Stockmann M., Marusik Y. M. &amp; Zonstein S. L. 2020. On Sahastata (Araneae: Filistatidae): complementary description of the generotype and two new species from Oman and Morocco. Zootaxa 4899 (1): 215 - 246. https: // doi. org / 10.11646 / zootaxa. 4899.1.12" type="journal article" year="2020">
Magalhaes 
<emphasis id="EE8CF413FF82FFC70888FA3E08C7E9E1" box="[656,708,1439,1465]" italics="true" pageId="18" pageNumber="167">et al</emphasis>
. 2020
</bibRefCitation>
; 
<bibRefCitation id="B86955F0FF82FFC70906FA010E52E9E1" author="Gavish-Regev E. &amp; Steinpress I. A. &amp; Salman I. N. A. &amp; Segev N. &amp; Uzan A. &amp; Byun Y. &amp; Levy T. &amp; Aharon S. &amp; Zvik Y. &amp; Shtuhin R. &amp; Shapira Y. &amp; Majer M. &amp; Ganem Z. &amp; Zonstein S. &amp; Magalhaes I. L. F. &amp; Lubin Y." box="[798,1105,1439,1466]" pageId="18" pageNumber="167" pagination="101" refId="ref13752" refString="Gavish-Regev E., Steinpress I. A., Salman I. N. A., Segev N., Uzan A., Byun Y., Levy T., Aharon S., Zvik Y., Shtuhin R., Shapira Y., Majer M., Ganem Z., Zonstein S., Magalhaes I. L. F. &amp; Lubin Y. 2022. Five-year monitoring of a desert burrow-dwelling spider following an environmental disaster indicates long-term impacts. Insects 13: 101. https: // doi. org / 10.3390 / insects 13010101" type="journal article" year="2022">
Gavish-Regev 
<emphasis id="EE8CF413FF82FFC709D5FA3E0E01E9E1" box="[973,1026,1439,1465]" italics="true" pageId="18" pageNumber="167">et al</emphasis>
. 2022
</bibRefCitation>
). Although 
<taxonomicName id="1BF85382FF82FFC70EC2FA010B1DE985" authority="Lehtinen, 1967" authorityName="Lehtinen" authorityYear="1967" class="Arachnida" family="Filistatidae" genus="Pritha" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="167" phylum="Arthropoda" rank="genus">
<emphasis id="EE8CF413FF82FFC70EC2FA010F26E9E1" box="[1242,1317,1439,1465]" italics="true" pageId="18" pageNumber="167">Pritha</emphasis>
<bibRefCitation id="B86955F0FF82FFC70F35FA010B1DE985" author="Lehtinen P. T." pageId="18" pageNumber="167" pagination="199 - 468" refId="ref14139" refString="Lehtinen P. T. 1967. Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici 4: 199 - 468." type="journal article" year="1967">Lehtinen, 1967</bibRefCitation>
</taxonomicName>
has not been thoroughly revised, valuable taxonomic information for several species is available in scattered works (
<bibRefCitation id="B86955F0FF82FFC70BB2FA780857EA59" author="Brignoli P. M." box="[426,596,1510,1537]" pageId="18" pageNumber="167" pagination="65 - 75" refId="ref13326" refString="Brignoli P. M. 1982. Contribution a la connaissance des Filistatidae palearctiques (Araneae). Revue arachnologique 4 (4): 65 - 75." type="journal article" year="1982">Brignoli 1982</bibRefCitation>
; 
<bibRefCitation id="B86955F0FF82FFC70879FA780994EA59" author="Marusik Y. M. &amp; Zamani A." box="[609,919,1510,1537]" pageId="18" pageNumber="167" pagination="429 - 435" refId="ref14637" refString="Marusik Y. M. &amp; Zamani A. 2015 b. Additional new species of Filistatidae (Aranei) from Iran. Arthropoda Selecta 24 (4): 429 - 435. https: // doi. org / 10.15298 / arthsel. 24.4.05" type="journal article" year="2015">Marusik &amp; Zamani 2015b</bibRefCitation>
; 
<bibRefCitation id="B86955F0FF82FFC709BCFA780EA2EA59" author="Legittimo C. M. &amp; Simeon E. &amp; Pompeo P. D. I. &amp; Kulczycki A." box="[932,1185,1510,1537]" pageId="18" pageNumber="167" pagination="201 - 248" refId="ref14073" refString="Legittimo C. M., Simeon E., Pompeo P. D. I. &amp; Kulczycki A. 2017. The Italian species of Pritha (Araneae, Filistatidae): a critical revision and description of two new species. Zootaxa 4243 (2): 201 - 248. https: // doi. org / 10.11646 / zootaxa. 4243.2.1" type="journal article" year="2017">
Legittimo 
<emphasis id="EE8CF413FF82FFC70E07FA790E50EA58" box="[1055,1107,1510,1536]" italics="true" pageId="18" pageNumber="167">et al</emphasis>
. 2017
</bibRefCitation>
; 
<bibRefCitation id="B86955F0FF82FFC70EB5FA780B20EA7C" author="Zonstein S. &amp; Marusik Y. M." pageId="18" pageNumber="167" pagination="53 - 93" refId="ref15464" refString="Zonstein S. &amp; Marusik Y. M. 2019. A revision of the spider genus Filistata (Araneae: Filistatidae). Arachnology 18 (2): 53 - 93. https: // doi. org / 10.13156 / arac. 2018.18.2.53" type="journal article" year="2019">Zonstein &amp; Marusik 2019</bibRefCitation>
).
</paragraph>
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<keyLead id="67098B34FF82FFC70AFBF9D20F9EEB42" pageId="18" pageNumber="167">
1. Tarsi without macrosetae; abdomen dorsally with white setae (in males, in one or two large patches) (
<figureCitation id="44C33484FF82FFC70B01F9EE0BA1EAD3" box="[281,418,1648,1675]" captionStart="Fig" captionStartId="19.[227,270,1684,1710]" captionTargetBox="[291,1376,303,1648]" captionTargetId="figure-0@19.[285,1377,302,1651]" captionTargetPageId="19" captionText="Fig. 10. Filistatid morphology. A. Filistata insidiatrix (Forsskål, 1775), ♀, leg tarsus, arrow points to macrosetae. B. Pritha nana (Simon, 1968), arrow points to right calamistrum. C. Sahastata aravaensis Ganem et al., 2022, left calamistrum. D. Zaitunia schmitzi Kulczyński, 1911, left calamistrum, arrow points to gap between setae. E. Filistata insidiatrix, left calamistrum. F. Filistata lubinae Zonstein &amp; Marusik, 2019, ♂, tarsus I. G. Sahastata aravaensis, ♀. H. Pritha IFMsp185, ♀. I. Pritha sp., male abdomen, dorsal view. J. Zaitunia schmitzi, ♂. K. Zaitunia schmitzi, spermathecae ventral view. L. Pritha tenuispina (Strand, 1914), ♂, left pedipalp, prolateral view, arrow points to cymbium. M. Filistata lubinae, left copulatory bulb, prolateral view. N. Zaitunia schmitzi, ♂, left pedipalp, prolateral view, arrow points to cymbium. O. Filistata albens Zonstein &amp; Marusik, 2019, left copulatory bulb, prolateral view. P. Zaitunia schmitzi, male clypeus, lateral view, arrow points to stout setae. Q. Filistata insidiatrix, female clypeus, lateral view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6916970" httpUri="https://zenodo.org/record/6916970/files/figure.png" pageId="18" pageNumber="167">Fig. 10H, I</figureCitation>
); female with calamistrum with three rows of inconspicuous setae (
<figureCitation id="44C33484FF82FFC70ECAF9EE0F43EAD2" box="[1234,1344,1648,1674]" captionStart="Fig" captionStartId="19.[227,270,1684,1710]" captionTargetBox="[291,1376,303,1648]" captionTargetId="figure-0@19.[285,1377,302,1651]" captionTargetPageId="19" captionText="Fig. 10. Filistatid morphology. A. Filistata insidiatrix (Forsskål, 1775), ♀, leg tarsus, arrow points to macrosetae. B. Pritha nana (Simon, 1968), arrow points to right calamistrum. C. Sahastata aravaensis Ganem et al., 2022, left calamistrum. D. Zaitunia schmitzi Kulczyński, 1911, left calamistrum, arrow points to gap between setae. E. Filistata insidiatrix, left calamistrum. F. Filistata lubinae Zonstein &amp; Marusik, 2019, ♂, tarsus I. G. Sahastata aravaensis, ♀. H. Pritha IFMsp185, ♀. I. Pritha sp., male abdomen, dorsal view. J. Zaitunia schmitzi, ♂. K. Zaitunia schmitzi, spermathecae ventral view. L. Pritha tenuispina (Strand, 1914), ♂, left pedipalp, prolateral view, arrow points to cymbium. M. Filistata lubinae, left copulatory bulb, prolateral view. N. Zaitunia schmitzi, ♂, left pedipalp, prolateral view, arrow points to cymbium. O. Filistata albens Zonstein &amp; Marusik, 2019, left copulatory bulb, prolateral view. P. Zaitunia schmitzi, male clypeus, lateral view, arrow points to stout setae. Q. Filistata insidiatrix, female clypeus, lateral view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6916970" httpUri="https://zenodo.org/record/6916970/files/figure.png" pageId="18" pageNumber="167">Fig. 10B</figureCitation>
); males with rigid leg tarsi, not flexible; male cymbium shorter than wide (
<figureCitation id="44C33484FF82FFC70E07F90A0E8AEAF6" box="[1055,1161,1684,1710]" captionStart="Fig" captionStartId="19.[227,270,1684,1710]" captionTargetBox="[291,1376,303,1648]" captionTargetId="figure-0@19.[285,1377,302,1651]" captionTargetPageId="19" captionText="Fig. 10. Filistatid morphology. A. Filistata insidiatrix (Forsskål, 1775), ♀, leg tarsus, arrow points to macrosetae. B. Pritha nana (Simon, 1968), arrow points to right calamistrum. C. Sahastata aravaensis Ganem et al., 2022, left calamistrum. D. Zaitunia schmitzi Kulczyński, 1911, left calamistrum, arrow points to gap between setae. E. Filistata insidiatrix, left calamistrum. F. Filistata lubinae Zonstein &amp; Marusik, 2019, ♂, tarsus I. G. Sahastata aravaensis, ♀. H. Pritha IFMsp185, ♀. I. Pritha sp., male abdomen, dorsal view. J. Zaitunia schmitzi, ♂. K. Zaitunia schmitzi, spermathecae ventral view. L. Pritha tenuispina (Strand, 1914), ♂, left pedipalp, prolateral view, arrow points to cymbium. M. Filistata lubinae, left copulatory bulb, prolateral view. N. Zaitunia schmitzi, ♂, left pedipalp, prolateral view, arrow points to cymbium. O. Filistata albens Zonstein &amp; Marusik, 2019, left copulatory bulb, prolateral view. P. Zaitunia schmitzi, male clypeus, lateral view, arrow points to stout setae. Q. Filistata insidiatrix, female clypeus, lateral view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6916970" httpUri="https://zenodo.org/record/6916970/files/figure.png" pageId="18" pageNumber="167">Fig. 10L</figureCitation>
); male sperm duct with a single coil (
<figureCitation id="44C33484FF82FFC70BB6F9260815EA8A" box="[430,534,1720,1746]" captionStart="Fig" captionStartId="19.[227,270,1684,1710]" captionTargetBox="[291,1376,303,1648]" captionTargetId="figure-0@19.[285,1377,302,1651]" captionTargetPageId="19" captionText="Fig. 10. Filistatid morphology. A. Filistata insidiatrix (Forsskål, 1775), ♀, leg tarsus, arrow points to macrosetae. B. Pritha nana (Simon, 1968), arrow points to right calamistrum. C. Sahastata aravaensis Ganem et al., 2022, left calamistrum. D. Zaitunia schmitzi Kulczyński, 1911, left calamistrum, arrow points to gap between setae. E. Filistata insidiatrix, left calamistrum. F. Filistata lubinae Zonstein &amp; Marusik, 2019, ♂, tarsus I. G. Sahastata aravaensis, ♀. H. Pritha IFMsp185, ♀. I. Pritha sp., male abdomen, dorsal view. J. Zaitunia schmitzi, ♂. K. Zaitunia schmitzi, spermathecae ventral view. L. Pritha tenuispina (Strand, 1914), ♂, left pedipalp, prolateral view, arrow points to cymbium. M. Filistata lubinae, left copulatory bulb, prolateral view. N. Zaitunia schmitzi, ♂, left pedipalp, prolateral view, arrow points to cymbium. O. Filistata albens Zonstein &amp; Marusik, 2019, left copulatory bulb, prolateral view. P. Zaitunia schmitzi, male clypeus, lateral view, arrow points to stout setae. Q. Filistata insidiatrix, female clypeus, lateral view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6916970" httpUri="https://zenodo.org/record/6916970/files/figure.png" pageId="18" pageNumber="167">Fig. 10L</figureCitation>
)............. 
<emphasis id="EE8CF413FF82FFC7089AF9290902EA89" bold="true" box="[642,769,1719,1745]" pageId="18" pageNumber="167">Prithinae:</emphasis>
<taxonomicName id="1BF85382FF82FFC70910F9290E0DEA8A" ID-CoL="6VYD" authority="Lehtinen, 1967" authorityName="Lehtinen" authorityYear="1967" box="[776,1038,1719,1746]" class="Arachnida" family="Filistatidae" genus="Pritha" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="167" phylum="Arthropoda" rank="genus">
<emphasis id="EE8CF413FF82FFC70910F9290956EA8A" bold="true" box="[776,853,1719,1746]" italics="true" pageId="18" pageNumber="167">Pritha</emphasis>
<bibRefCitation id="B86955F0FF82FFC70944F9260E0DEA8A" author="Lehtinen P. T." box="[860,1038,1720,1746]" pageId="18" pageNumber="167" pagination="199 - 468" refId="ref14139" refString="Lehtinen P. T. 1967. Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici 4: 199 - 468." type="journal article" year="1967">Lehtinen, 1967</bibRefCitation>
</taxonomicName>
[three described species in 
<collectingCountry id="A4EF6891FF82FFC70F4BF9260F9AEA8A" box="[1363,1433,1720,1746]" name="Israel" pageId="18" pageNumber="167">Israel</collectingCountry>
: 
<taxonomicName id="1BF85382FF82FFC70B08F9450974EAAE" ID-CoL="4MKRM" authority="(O. Pickard-Cambridge, 1872)" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[272,887,1755,1782]" class="Arachnida" family="Filistatidae" genus="Pritha" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="167" phylum="Arthropoda" rank="species" species="albimaculata">
<emphasis id="EE8CF413FF82FFC70B08F9450BFCEAAD" box="[272,511,1755,1781]" italics="true" pageId="18" pageNumber="167">Pritha albimaculata</emphasis>
(O. Pickard-Cambridge, 1872)
</taxonomicName>
, 
<taxonomicName id="1BF85382FF82FFC7099AF9450F9FEAAE" ID-CoL="99BRT" authority="(O. Pickard-Cambridge, 1872)" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[898,1436,1755,1782]" class="Arachnida" family="Filistatidae" genus="Pritha" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="167" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="EE8CF413FF82FFC7099AF9450E2AEAAD" box="[898,1065,1755,1781]" italics="true" pageId="18" pageNumber="167">Pritha hirsuta</emphasis>
(O. Pickard-Cambridge, 1872)
</taxonomicName>
and 
<taxonomicName id="1BF85382FF82FFC70B59F96108C3EB41" ID-CoL="4MKSD" authority="(Strand, 1814)" baseAuthorityName="Strand" baseAuthorityYear="1814" box="[321,704,1791,1817]" class="Arachnida" family="Filistatidae" genus="Pritha" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="167" phylum="Arthropoda" rank="species" species="tenuispina">
<emphasis id="EE8CF413FF82FFC70B59F961080EEB41" box="[321,525,1791,1817]" italics="true" pageId="18" pageNumber="167">Pritha tenuispina</emphasis>
(Strand, 1814)
</taxonomicName>
, in addition to undescribed species; genus in need of revision].
</keyLead>
</paragraph>
<paragraph id="DC472801FF82FFC70AFFF8BA0F9FEBF1" blockId="18.[227,1437,1612,2063]" pageId="18" pageNumber="167">
<keyLead id="67098B34FF82FFC70AFFF8BA0F9FEBF1" pageId="18" pageNumber="167">
– Tarsi with macroseta (
<figureCitation id="44C33484FF82FFC70815F8BD087BEB65" box="[525,632,1827,1853]" captionStart="Fig" captionStartId="19.[227,270,1684,1710]" captionTargetBox="[291,1376,303,1648]" captionTargetId="figure-0@19.[285,1377,302,1651]" captionTargetPageId="19" captionText="Fig. 10. Filistatid morphology. A. Filistata insidiatrix (Forsskål, 1775), ♀, leg tarsus, arrow points to macrosetae. B. Pritha nana (Simon, 1968), arrow points to right calamistrum. C. Sahastata aravaensis Ganem et al., 2022, left calamistrum. D. Zaitunia schmitzi Kulczyński, 1911, left calamistrum, arrow points to gap between setae. E. Filistata insidiatrix, left calamistrum. F. Filistata lubinae Zonstein &amp; Marusik, 2019, ♂, tarsus I. G. Sahastata aravaensis, ♀. H. Pritha IFMsp185, ♀. I. Pritha sp., male abdomen, dorsal view. J. Zaitunia schmitzi, ♂. K. Zaitunia schmitzi, spermathecae ventral view. L. Pritha tenuispina (Strand, 1914), ♂, left pedipalp, prolateral view, arrow points to cymbium. M. Filistata lubinae, left copulatory bulb, prolateral view. N. Zaitunia schmitzi, ♂, left pedipalp, prolateral view, arrow points to cymbium. O. Filistata albens Zonstein &amp; Marusik, 2019, left copulatory bulb, prolateral view. P. Zaitunia schmitzi, male clypeus, lateral view, arrow points to stout setae. Q. Filistata insidiatrix, female clypeus, lateral view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6916970" httpUri="https://zenodo.org/record/6916970/files/figure.png" pageId="18" pageNumber="167">Fig. 10A</figureCitation>
; difficult to see in 
<taxonomicName id="1BF85382FF82FFC7095DF8BA09ABEB65" ID-CoL="63X5V" authorityName="Lehtinen" authorityYear="1967" box="[837,936,1828,1853]" class="Arachnida" family="Filistatidae" genus="Zaitunia" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="167" phylum="Arthropoda" rank="genus">
<emphasis id="EE8CF413FF82FFC7095DF8BA09ABEB65" box="[837,936,1828,1853]" italics="true" pageId="18" pageNumber="167">Zaitunia</emphasis>
</taxonomicName>
); white setae absent from abdomen; female calamistrum with conspicuous setae (
<figureCitation id="44C33484FF82FFC708CBF8D90967EB39" box="[723,868,1863,1889]" captionStart="Fig" captionStartId="19.[227,270,1684,1710]" captionTargetBox="[291,1376,303,1648]" captionTargetId="figure-0@19.[285,1377,302,1651]" captionTargetPageId="19" captionText="Fig. 10. Filistatid morphology. A. Filistata insidiatrix (Forsskål, 1775), ♀, leg tarsus, arrow points to macrosetae. B. Pritha nana (Simon, 1968), arrow points to right calamistrum. C. Sahastata aravaensis Ganem et al., 2022, left calamistrum. D. Zaitunia schmitzi Kulczyński, 1911, left calamistrum, arrow points to gap between setae. E. Filistata insidiatrix, left calamistrum. F. Filistata lubinae Zonstein &amp; Marusik, 2019, ♂, tarsus I. G. Sahastata aravaensis, ♀. H. Pritha IFMsp185, ♀. I. Pritha sp., male abdomen, dorsal view. J. Zaitunia schmitzi, ♂. K. Zaitunia schmitzi, spermathecae ventral view. L. Pritha tenuispina (Strand, 1914), ♂, left pedipalp, prolateral view, arrow points to cymbium. M. Filistata lubinae, left copulatory bulb, prolateral view. N. Zaitunia schmitzi, ♂, left pedipalp, prolateral view, arrow points to cymbium. O. Filistata albens Zonstein &amp; Marusik, 2019, left copulatory bulb, prolateral view. P. Zaitunia schmitzi, male clypeus, lateral view, arrow points to stout setae. Q. Filistata insidiatrix, female clypeus, lateral view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6916970" httpUri="https://zenodo.org/record/6916970/files/figure.png" pageId="18" pageNumber="167">Fig. 10C–E</figureCitation>
); males with flexible leg tarsi (
<figureCitation id="44C33484FF82FFC70EC7F8D90F48EB39" box="[1247,1355,1863,1889]" captionStart="Fig" captionStartId="19.[227,270,1684,1710]" captionTargetBox="[291,1376,303,1648]" captionTargetId="figure-0@19.[285,1377,302,1651]" captionTargetPageId="19" captionText="Fig. 10. Filistatid morphology. A. Filistata insidiatrix (Forsskål, 1775), ♀, leg tarsus, arrow points to macrosetae. B. Pritha nana (Simon, 1968), arrow points to right calamistrum. C. Sahastata aravaensis Ganem et al., 2022, left calamistrum. D. Zaitunia schmitzi Kulczyński, 1911, left calamistrum, arrow points to gap between setae. E. Filistata insidiatrix, left calamistrum. F. Filistata lubinae Zonstein &amp; Marusik, 2019, ♂, tarsus I. G. Sahastata aravaensis, ♀. H. Pritha IFMsp185, ♀. I. Pritha sp., male abdomen, dorsal view. J. Zaitunia schmitzi, ♂. K. Zaitunia schmitzi, spermathecae ventral view. L. Pritha tenuispina (Strand, 1914), ♂, left pedipalp, prolateral view, arrow points to cymbium. M. Filistata lubinae, left copulatory bulb, prolateral view. N. Zaitunia schmitzi, ♂, left pedipalp, prolateral view, arrow points to cymbium. O. Filistata albens Zonstein &amp; Marusik, 2019, left copulatory bulb, prolateral view. P. Zaitunia schmitzi, male clypeus, lateral view, arrow points to stout setae. Q. Filistata insidiatrix, female clypeus, lateral view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6916970" httpUri="https://zenodo.org/record/6916970/files/figure.png" pageId="18" pageNumber="167">Fig. 10F</figureCitation>
); male cymbium longer than wide (
<figureCitation id="44C33484FF82FFC70842F8F408F7EBDD" box="[602,756,1898,1925]" captionStart="Fig" captionStartId="19.[227,270,1684,1710]" captionTargetBox="[291,1376,303,1648]" captionTargetId="figure-0@19.[285,1377,302,1651]" captionTargetPageId="19" captionText="Fig. 10. Filistatid morphology. A. Filistata insidiatrix (Forsskål, 1775), ♀, leg tarsus, arrow points to macrosetae. B. Pritha nana (Simon, 1968), arrow points to right calamistrum. C. Sahastata aravaensis Ganem et al., 2022, left calamistrum. D. Zaitunia schmitzi Kulczyński, 1911, left calamistrum, arrow points to gap between setae. E. Filistata insidiatrix, left calamistrum. F. Filistata lubinae Zonstein &amp; Marusik, 2019, ♂, tarsus I. G. Sahastata aravaensis, ♀. H. Pritha IFMsp185, ♀. I. Pritha sp., male abdomen, dorsal view. J. Zaitunia schmitzi, ♂. K. Zaitunia schmitzi, spermathecae ventral view. L. Pritha tenuispina (Strand, 1914), ♂, left pedipalp, prolateral view, arrow points to cymbium. M. Filistata lubinae, left copulatory bulb, prolateral view. N. Zaitunia schmitzi, ♂, left pedipalp, prolateral view, arrow points to cymbium. O. Filistata albens Zonstein &amp; Marusik, 2019, left copulatory bulb, prolateral view. P. Zaitunia schmitzi, male clypeus, lateral view, arrow points to stout setae. Q. Filistata insidiatrix, female clypeus, lateral view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6916970" httpUri="https://zenodo.org/record/6916970/files/figure.png" pageId="18" pageNumber="167">Figs 10M, N</figureCitation>
, 
<figureCitation id="44C33484FF82FFC708E7F8F5090CEBDD" box="[767,783,1899,1925]" captionStart="Fig" captionStartId="16.[227,270,1901,1927]" captionTargetBox="[233,1434,365,1831]" captionTargetId="figure-70@16.[227,1437,352,1846]" captionTargetPageId="16" captionText="Fig. 8. A–G. Filistata betarif sp. nov., holotype, ♂ (HUJ -INVAr 21058), left pedipalp. A–B, E. Prolateral view. Arrow points to prolateral strong setae in cymbium. C, F–G. Retrolateral view. Arrow points to embolic keel. D. Dorsal view. — H. Filistata insidiatrix (Forsskål, 1775) (MACN-Ar 39461), bulb, retrolateral view. Arrow points to embolic keel. Scale bars: A–F = 0.2 mm; G–H = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6916964" httpUri="https://zenodo.org/record/6916964/files/figure.png" pageId="18" pageNumber="167">8</figureCitation>
); male sperm duct with at least two coils (
<figureCitation id="44C33484FF82FFC70EE6F8F40F94EBDD" box="[1278,1431,1898,1925]" captionStart="Fig" captionStartId="19.[227,270,1684,1710]" captionTargetBox="[291,1376,303,1648]" captionTargetId="figure-0@19.[285,1377,302,1651]" captionTargetPageId="19" captionText="Fig. 10. Filistatid morphology. A. Filistata insidiatrix (Forsskål, 1775), ♀, leg tarsus, arrow points to macrosetae. B. Pritha nana (Simon, 1968), arrow points to right calamistrum. C. Sahastata aravaensis Ganem et al., 2022, left calamistrum. D. Zaitunia schmitzi Kulczyński, 1911, left calamistrum, arrow points to gap between setae. E. Filistata insidiatrix, left calamistrum. F. Filistata lubinae Zonstein &amp; Marusik, 2019, ♂, tarsus I. G. Sahastata aravaensis, ♀. H. Pritha IFMsp185, ♀. I. Pritha sp., male abdomen, dorsal view. J. Zaitunia schmitzi, ♂. K. Zaitunia schmitzi, spermathecae ventral view. L. Pritha tenuispina (Strand, 1914), ♂, left pedipalp, prolateral view, arrow points to cymbium. M. Filistata lubinae, left copulatory bulb, prolateral view. N. Zaitunia schmitzi, ♂, left pedipalp, prolateral view, arrow points to cymbium. O. Filistata albens Zonstein &amp; Marusik, 2019, left copulatory bulb, prolateral view. P. Zaitunia schmitzi, male clypeus, lateral view, arrow points to stout setae. Q. Filistata insidiatrix, female clypeus, lateral view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6916970" httpUri="https://zenodo.org/record/6916970/files/figure.png" pageId="18" pageNumber="167">Figs 10M–N</figureCitation>
, 
<figureCitation id="44C33484FF82FFC70B08F8100B1CEBF0" box="[272,287,1934,1960]" captionStart="Fig" captionStartId="16.[227,270,1901,1927]" captionTargetBox="[233,1434,365,1831]" captionTargetId="figure-70@16.[227,1437,352,1846]" captionTargetPageId="16" captionText="Fig. 8. A–G. Filistata betarif sp. nov., holotype, ♂ (HUJ -INVAr 21058), left pedipalp. A–B, E. Prolateral view. Arrow points to prolateral strong setae in cymbium. C, F–G. Retrolateral view. Arrow points to embolic keel. D. Dorsal view. — H. Filistata insidiatrix (Forsskål, 1775) (MACN-Ar 39461), bulb, retrolateral view. Arrow points to embolic keel. Scale bars: A–F = 0.2 mm; G–H = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6916964" httpUri="https://zenodo.org/record/6916964/files/figure.png" pageId="18" pageNumber="167">8</figureCitation>
)............................................................................................................ 2 (
<taxonomicName id="1BF85382FF82FFC70E71F8100F97EBF1" authority="Simon, 1864" authorityName="Simon" authorityYear="1864" box="[1129,1428,1934,1961]" class="Arachnida" family="Filistatidae" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="155" phylum="Arthropoda" rank="subFamily" subFamily="Filistatinae">
<emphasis id="EE8CF413FF82FFC70E71F8100EF6EBF0" bold="true" box="[1129,1269,1934,1960]" pageId="18" pageNumber="167">Filistatinae</emphasis>
<bibRefCitation id="B86955F0FF82FFC70EE5F8100F97EBF1" author="Simon E." box="[1277,1428,1934,1961]" pageId="18" pageNumber="167" refId="ref14918" refString="Simon E. 1864. Histoire naturelle des araignees (araneides). Librairie encyclopedique de Roret, Paris. https: // doi. org / 10.5962 / bhl. title. 47654" type="book" year="1864">Simon, 1864</bibRefCitation>
</taxonomicName>
)
</keyLead>
</paragraph>
</keyStep>
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<paragraph id="DC472801FF82FFC10AFBF84E0F9EEDEB" blockId="18.[227,1437,1612,2063]" lastBlockId="20.[227,1437,303,1402]" lastPageId="20" lastPageNumber="169" pageId="18" pageNumber="167">
<keyLead id="67098B34FF82FFC10AFBF84E0F9EEDEB" lastPageId="20" lastPageNumber="169" pageId="18" pageNumber="167">
2. Females with continuous calamistrum, with three rows with many setae (15 or more) (
<figureCitation id="44C33484FF82FFC70F38F84E0F8DEBB2" box="[1312,1422,2000,2026]" captionStart="Fig" captionStartId="19.[227,270,1684,1710]" captionTargetBox="[291,1376,303,1648]" captionTargetId="figure-0@19.[285,1377,302,1651]" captionTargetPageId="19" captionText="Fig. 10. Filistatid morphology. A. Filistata insidiatrix (Forsskål, 1775), ♀, leg tarsus, arrow points to macrosetae. B. Pritha nana (Simon, 1968), arrow points to right calamistrum. C. Sahastata aravaensis Ganem et al., 2022, left calamistrum. D. Zaitunia schmitzi Kulczyński, 1911, left calamistrum, arrow points to gap between setae. E. Filistata insidiatrix, left calamistrum. F. Filistata lubinae Zonstein &amp; Marusik, 2019, ♂, tarsus I. G. Sahastata aravaensis, ♀. H. Pritha IFMsp185, ♀. I. Pritha sp., male abdomen, dorsal view. J. Zaitunia schmitzi, ♂. K. Zaitunia schmitzi, spermathecae ventral view. L. Pritha tenuispina (Strand, 1914), ♂, left pedipalp, prolateral view, arrow points to cymbium. M. Filistata lubinae, left copulatory bulb, prolateral view. N. Zaitunia schmitzi, ♂, left pedipalp, prolateral view, arrow points to cymbium. O. Filistata albens Zonstein &amp; Marusik, 2019, left copulatory bulb, prolateral view. P. Zaitunia schmitzi, male clypeus, lateral view, arrow points to stout setae. Q. Filistata insidiatrix, female clypeus, lateral view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6916970" httpUri="https://zenodo.org/record/6916970/files/figure.png" pageId="18" pageNumber="167">Fig. 10C</figureCitation>
); large spiders (females with body length around 
<quantity id="1B0085E4FF82FFC70956F86A09ABE457" box="[846,936,2036,2063]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="167" unit="mm" value="10.0">10 mm</quantity>
); females dark brown to black (
<figureCitation id="44C33484FF82FFC70F3FF86A0F97E456" box="[1319,1428,2036,2062]" captionStart="Fig" captionStartId="19.[227,270,1684,1710]" captionTargetBox="[291,1376,303,1648]" captionTargetId="figure-0@19.[285,1377,302,1651]" captionTargetPageId="19" captionText="Fig. 10. Filistatid morphology. A. Filistata insidiatrix (Forsskål, 1775), ♀, leg tarsus, arrow points to macrosetae. B. Pritha nana (Simon, 1968), arrow points to right calamistrum. C. Sahastata aravaensis Ganem et al., 2022, left calamistrum. D. Zaitunia schmitzi Kulczyński, 1911, left calamistrum, arrow points to gap between setae. E. Filistata insidiatrix, left calamistrum. F. Filistata lubinae Zonstein &amp; Marusik, 2019, ♂, tarsus I. G. Sahastata aravaensis, ♀. H. Pritha IFMsp185, ♀. I. Pritha sp., male abdomen, dorsal view. J. Zaitunia schmitzi, ♂. K. Zaitunia schmitzi, spermathecae ventral view. L. Pritha tenuispina (Strand, 1914), ♂, left pedipalp, prolateral view, arrow points to cymbium. M. Filistata lubinae, left copulatory bulb, prolateral view. N. Zaitunia schmitzi, ♂, left pedipalp, prolateral view, arrow points to cymbium. O. Filistata albens Zonstein &amp; Marusik, 2019, left copulatory bulb, prolateral view. P. Zaitunia schmitzi, male clypeus, lateral view, arrow points to stout setae. Q. Filistata insidiatrix, female clypeus, lateral view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6916970" httpUri="https://zenodo.org/record/6916970/files/figure.png" pageId="18" pageNumber="167">Fig. 10G</figureCitation>
) and males light cream; males of Israeli species unknown, but based on congeners, should have pedipalpal femur with dorsal and ventral macrosetae ......................................................................... ...................................... 
<taxonomicName id="1BF85382FF84FFC10829FEEB0944EDD7" ID-CoL="7BPH" authority="Benoit, 1968" authorityName="Benoit" authorityYear="1968" box="[561,839,373,400]" class="Arachnida" family="Filistatidae" genus="Sahastata" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="EE8CF413FF84FFC10829FEEB08ABEDC8" bold="true" box="[561,680,373,400]" italics="true" pageId="20" pageNumber="169">Sahastata</emphasis>
<bibRefCitation id="B86955F0FF84FFC108A8FEEB0944EDD7" author="Benoit P. L. G." box="[688,839,373,399]" pageId="20" pageNumber="169" pagination="92 - 102" refId="ref13241" refString="Benoit P. L. G. 1968. Synopsis des Filistatidae africains. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot; 77: 92 - 102." type="journal article" year="1968">Benoit, 1968</bibRefCitation>
</taxonomicName>
[a single known species in 
<collectingCountry id="A4EF6891FF84FFC10E93FEEB0ECCEDD7" box="[1163,1231,373,399]" name="Israel" pageId="20" pageNumber="169">Israel</collectingCountry>
, known currently only from females, 
<taxonomicName id="1BF85382FF84FFC1081AFE060F92EDEB" ID-CoL="99BJ5" authority="Ganem, Magalhaes, Zonstein &amp; Gavish-Regev, 2022" authorityName="Ganem, Magalhaes, Zonstein &amp; Gavish-Regev" authorityYear="2022" box="[514,1425,408,435]" class="Arachnida" family="Filistatidae" genus="Sahastata" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="169" phylum="Arthropoda" rank="species" species="aravaensis">
<emphasis id="EE8CF413FF84FFC1081AFE060903EDEA" box="[514,768,408,434]" italics="true" pageId="20" pageNumber="169">Sahastata aravaensis</emphasis>
Ganem, Magalhaes, Zonstein &amp; 
<bibRefCitation id="B86955F0FF84FFC10E80FE060F92EDEB" author="Gavish-Regev E. &amp; Steinpress I. A. &amp; Salman I. N. A. &amp; Segev N. &amp; Uzan A. &amp; Byun Y. &amp; Levy T. &amp; Aharon S. &amp; Zvik Y. &amp; Shtuhin R. &amp; Shapira Y. &amp; Majer M. &amp; Ganem Z. &amp; Zonstein S. &amp; Magalhaes I. L. F. &amp; Lubin Y." box="[1176,1425,408,435]" pageId="20" pageNumber="169" pagination="101" refId="ref13752" refString="Gavish-Regev E., Steinpress I. A., Salman I. N. A., Segev N., Uzan A., Byun Y., Levy T., Aharon S., Zvik Y., Shtuhin R., Shapira Y., Majer M., Ganem Z., Zonstein S., Magalhaes I. L. F. &amp; Lubin Y. 2022. Five-year monitoring of a desert burrow-dwelling spider following an environmental disaster indicates long-term impacts. Insects 13: 101. https: // doi. org / 10.3390 / insects 13010101" type="journal article" year="2022">Gavish-Regev, 2022</bibRefCitation>
</taxonomicName>
].
</keyLead>
</paragraph>
<caption id="88877889FF83FFC60AFBF90A0932E457" ID-DOI="http://doi.org/10.5281/zenodo.6916970" ID-Zenodo-Dep="6916970" httpUri="https://zenodo.org/record/6916970/files/figure.png" pageId="19" pageNumber="168" startId="19.[227,270,1684,1710]" targetBox="[291,1376,303,1648]" targetPageId="19">
<paragraph id="DC472801FF83FFC60AFBF90A0932E457" blockId="19.[227,1436,1684,2063]" pageId="19" pageNumber="168">
<emphasis id="EE8CF413FF83FFC60AFBF90A0B42EAF6" bold="true" box="[227,321,1684,1710]" pageId="19" pageNumber="168">Fig. 10.</emphasis>
Filistatid morphology. 
<emphasis id="EE8CF413FF83FFC60840F90A086EEAF6" bold="true" box="[600,621,1684,1710]" pageId="19" pageNumber="168">A</emphasis>
. 
<taxonomicName id="1BF85382FF83FFC60865F90A0E30EAF7" ID-CoL="6J2PS" authority="(Forsskal, 1775)" baseAuthorityName="Forsskal" baseAuthorityYear="1775" box="[637,1075,1684,1711]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="168" phylum="Arthropoda" rank="species" species="insidiatrix">
<emphasis id="EE8CF413FF83FFC60865F90A0960EAF6" box="[637,867,1684,1710]" italics="true" pageId="19" pageNumber="168">Filistata insidiatrix</emphasis>
(
<bibRefCitation id="B86955F0FF83FFC6096EF90A0E2AEAF7" author="Forsskal P." box="[886,1065,1684,1711]" pageId="19" pageNumber="168" refId="ref13586" refString="Forsskal P. 1775. Descriptiones animalium avium, amphibiorum, piscium, insectorum, vermium; quae in itinere orientali observavit Petrus Forskal [sic]. Moller, Havniae [= Copenhagen]. https: // doi. org / 10.5962 / bhl. title. 2154" type="book" year="1775">Forsskål, 1775</bibRefCitation>
)
</taxonomicName>
, ♀, leg tarsus, arrow points to macrosetae. 
<emphasis id="EE8CF413FF83FFC60B6EF9260B89EA8A" bold="true" box="[374,394,1720,1746]" pageId="19" pageNumber="168">B</emphasis>
. 
<taxonomicName id="1BF85382FF83FFC60B80F92908DAEA8A" ID-CoL="4MKS3" authority="(Simon, 1968)" baseAuthorityName="Simon" baseAuthorityYear="1968" box="[408,729,1719,1746]" class="Arachnida" family="Filistatidae" genus="Pritha" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="168" phylum="Arthropoda" rank="species" species="nana">
<emphasis id="EE8CF413FF83FFC60B80F9290825EA89" box="[408,550,1719,1745]" italics="true" pageId="19" pageNumber="168">Pritha nana</emphasis>
(Simon, 1968)
</taxonomicName>
, arrow points to right calamistrum. 
<emphasis id="EE8CF413FF83FFC60E66F9290E90EA89" bold="true" box="[1150,1171,1719,1745]" pageId="19" pageNumber="168">C</emphasis>
. 
<taxonomicName id="1BF85382FF83FFC60EBAF9290BCFEAAD" ID-CoL="99BJ5" authority="Ganem et al., 2022" authorityName="Ganem" authorityYear="2022" class="Arachnida" family="Filistatidae" genus="Sahastata" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="168" phylum="Arthropoda" rank="species" species="aravaensis">
<emphasis id="EE8CF413FF83FFC60EBAF9290F9FEA89" box="[1186,1436,1719,1745]" italics="true" pageId="19" pageNumber="168">Sahastata aravaensis</emphasis>
Ganem 
<emphasis id="EE8CF413FF83FFC60B59F9450B7EEAAD" box="[321,381,1755,1781]" italics="true" pageId="19" pageNumber="168">et al.</emphasis>
, 2022
</taxonomicName>
, left calamistrum. 
<emphasis id="EE8CF413FF83FFC608B2F94508BCEAAD" bold="true" box="[682,703,1755,1781]" pageId="19" pageNumber="168">D</emphasis>
. 
<taxonomicName id="1BF85382FF83FFC608C8F9450E7BEAAD" ID-CoL="7GBLC" authority="Kulczynski, 1911" authorityName="Kulczynski" authorityYear="1911" box="[720,1144,1755,1781]" class="Arachnida" family="Filistatidae" genus="Zaitunia" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="168" phylum="Arthropoda" rank="species" species="schmitzi">
<emphasis id="EE8CF413FF83FFC608C8F945099EEAAD" box="[720,925,1755,1781]" italics="true" pageId="19" pageNumber="168">Zaitunia schmitzi</emphasis>
Kulczyński, 1911
</taxonomicName>
, left calamistrum, arrow points to gap between setae. 
<emphasis id="EE8CF413FF83FFC60824F9600853EB40" bold="true" box="[572,592,1790,1816]" pageId="19" pageNumber="168">E</emphasis>
. 
<taxonomicName id="1BF85382FF83FFC60878F9600946EB40" ID-CoL="6J2PS" baseAuthorityName="Forsskal" baseAuthorityYear="1775" box="[608,837,1790,1816]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="168" phylum="Arthropoda" rank="species" species="insidiatrix">
<emphasis id="EE8CF413FF83FFC60878F9600946EB40" box="[608,837,1790,1816]" italics="true" pageId="19" pageNumber="168">Filistata insidiatrix</emphasis>
</taxonomicName>
, left calamistrum. 
<emphasis id="EE8CF413FF83FFC60E30F9600E3BEB40" bold="true" box="[1064,1080,1790,1816]" pageId="19" pageNumber="168">F</emphasis>
. 
<taxonomicName id="1BF85382FF83FFC60E51F9600B96EB63" ID-CoL="99CSP" authority="Zonstein &amp; Marusik, 2019" authorityName="Zonstein &amp; Marusik" authorityYear="2019" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="168" phylum="Arthropoda" rank="species" species="lubinae">
<emphasis id="EE8CF413FF83FFC60E51F9600F0EEB40" box="[1097,1293,1790,1816]" italics="true" pageId="19" pageNumber="168">Filistata lubinae</emphasis>
<bibRefCitation id="B86955F0FF83FFC60F0EF9600B96EB63" author="Zonstein S. &amp; Marusik Y. M." pageId="19" pageNumber="168" pagination="53 - 93" refId="ref15464" refString="Zonstein S. &amp; Marusik Y. M. 2019. A revision of the spider genus Filistata (Araneae: Filistatidae). Arachnology 18 (2): 53 - 93. https: // doi. org / 10.13156 / arac. 2018.18.2.53" type="journal article" year="2019">Zonstein &amp; Marusik, 2019</bibRefCitation>
</taxonomicName>
, ♂, tarsus I. 
<emphasis id="EE8CF413FF83FFC6082AF8BF084AEB63" bold="true" box="[562,585,1825,1851]" pageId="19" pageNumber="168">G</emphasis>
. 
<taxonomicName id="1BF85382FF83FFC60842F8BF0954EB63" ID-CoL="99BJ5" box="[602,855,1825,1851]" class="Arachnida" family="Filistatidae" genus="Sahastata" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="168" phylum="Arthropoda" rank="species" species="aravaensis">
<emphasis id="EE8CF413FF83FFC60842F8BF0954EB63" box="[602,855,1825,1851]" italics="true" pageId="19" pageNumber="168">Sahastata aravaensis</emphasis>
</taxonomicName>
, ♀. 
<emphasis id="EE8CF413FF83FFC60997F8BF09A5EB63" bold="true" box="[911,934,1825,1851]" pageId="19" pageNumber="168">H</emphasis>
. 
<taxonomicName id="1BF85382FF83FFC609AFF8BF0E01EB63" ID-CoL="6VYD" authorityName="Lehtinen" authorityYear="1967" box="[951,1026,1825,1851]" class="Arachnida" family="Filistatidae" genus="Pritha" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="168" phylum="Arthropoda" rank="genus">
<emphasis id="EE8CF413FF83FFC609AFF8BF0E01EB63" box="[951,1026,1825,1851]" italics="true" pageId="19" pageNumber="168">Pritha</emphasis>
</taxonomicName>
IFMsp185, ♀. 
<emphasis id="EE8CF413FF83FFC60EA7F8BF0EC9EB63" bold="true" box="[1215,1226,1825,1851]" pageId="19" pageNumber="168">I</emphasis>
. 
<taxonomicName id="1BF85382FF83FFC60EC4F8BF0F57EB64" box="[1244,1364,1825,1852]" class="Arachnida" family="Filistatidae" genus="Pritha" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="168" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="EE8CF413FF83FFC60EC4F8BF0F24EB63" box="[1244,1319,1825,1851]" italics="true" pageId="19" pageNumber="168">Pritha</emphasis>
sp.
</taxonomicName>
, male abdomen, dorsal view. 
<emphasis id="EE8CF413FF83FFC60BF2F8DA0BFAEB06" bold="true" box="[490,505,1860,1886]" pageId="19" pageNumber="168">J</emphasis>
. 
<taxonomicName id="1BF85382FF83FFC6081DF8DB08CEEB06" ID-CoL="7GBLC" authorityName="Kulczynski" authorityYear="1911" box="[517,717,1860,1886]" class="Arachnida" family="Filistatidae" genus="Zaitunia" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="168" phylum="Arthropoda" rank="species" species="schmitzi">
<emphasis id="EE8CF413FF83FFC6081DF8DB08CEEB06" box="[517,717,1860,1886]" italics="true" pageId="19" pageNumber="168">Zaitunia schmitzi</emphasis>
</taxonomicName>
, ♂. 
<emphasis id="EE8CF413FF83FFC608E4F8DA0910EB06" bold="true" box="[764,787,1860,1886]" pageId="19" pageNumber="168">K</emphasis>
. 
<taxonomicName id="1BF85382FF83FFC60907F8DB09E4EB06" ID-CoL="7GBLC" authorityName="Kulczynski" authorityYear="1911" box="[799,999,1860,1886]" class="Arachnida" family="Filistatidae" genus="Zaitunia" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="168" phylum="Arthropoda" rank="species" species="schmitzi">
<emphasis id="EE8CF413FF83FFC60907F8DB09E4EB06" box="[799,999,1860,1886]" italics="true" pageId="19" pageNumber="168">Zaitunia schmitzi</emphasis>
</taxonomicName>
, spermathecae ventral view. 
<emphasis id="EE8CF413FF83FFC60F2AF8DA0F45EB06" bold="true" box="[1330,1350,1860,1886]" pageId="19" pageNumber="168">L</emphasis>
. 
<taxonomicName id="1BF85382FF83FFC60F49F8DA081AEBDA" ID-CoL="4MKSD" authority="(Strand, 1914)" baseAuthorityName="Strand" baseAuthorityYear="1914" class="Arachnida" family="Filistatidae" genus="Pritha" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="168" phylum="Arthropoda" rank="species" species="tenuispina">
<emphasis id="EE8CF413FF83FFC60F49F8DA0B5EEBD9" italics="true" pageId="19" pageNumber="168">Pritha tenuispina</emphasis>
(Strand, 1914)
</taxonomicName>
, ♂, left pedipalp, prolateral view, arrow points to cymbium. 
<emphasis id="EE8CF413FF83FFC60F12F8F60F25EBDA" bold="true" box="[1290,1318,1896,1922]" pageId="19" pageNumber="168">M</emphasis>
. 
<taxonomicName id="1BF85382FF83FFC60F21F8F90B38EBFD" ID-CoL="99CSP" authorityName="Zonstein &amp; Marusik" authorityYear="2019" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="168" phylum="Arthropoda" rank="species" species="lubinae">
<emphasis id="EE8CF413FF83FFC60F21F8F90B38EBFD" italics="true" pageId="19" pageNumber="168">Filistata lubinae</emphasis>
</taxonomicName>
, left copulatory bulb, prolateral view. 
<emphasis id="EE8CF413FF83FFC60916F8150920EBFD" bold="true" box="[782,803,1931,1957]" pageId="19" pageNumber="168">N</emphasis>
. 
<taxonomicName id="1BF85382FF83FFC6092DF8150E01EBFD" ID-CoL="7GBLC" authorityName="Kulczynski" authorityYear="1911" box="[821,1026,1931,1957]" class="Arachnida" family="Filistatidae" genus="Zaitunia" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="168" phylum="Arthropoda" rank="species" species="schmitzi">
<emphasis id="EE8CF413FF83FFC6092DF8150E01EBFD" box="[821,1026,1931,1957]" italics="true" pageId="19" pageNumber="168">Zaitunia schmitzi</emphasis>
</taxonomicName>
, ♂, left pedipalp, prolateral view, arrow points to cymbium. 
<emphasis id="EE8CF413FF83FFC6080DF833082FEB9F" bold="true" box="[533,556,1965,1991]" pageId="19" pageNumber="168">O</emphasis>
. 
<taxonomicName id="1BF85382FF83FFC60822F8300E2DEB90" ID-CoL="99BS5" authority="Zonstein &amp; Marusik, 2019" authorityName="Zonstein &amp; Marusik" authorityYear="2019" box="[570,1070,1966,1992]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="168" phylum="Arthropoda" rank="species" species="albens">
<emphasis id="EE8CF413FF83FFC60822F83008F2EB90" box="[570,753,1966,1992]" italics="true" pageId="19" pageNumber="168">Filistata albens</emphasis>
<bibRefCitation id="B86955F0FF83FFC608EFF8300E2DEB90" author="Zonstein S. &amp; Marusik Y. M." box="[759,1070,1966,1992]" pageId="19" pageNumber="168" pagination="53 - 93" refId="ref15464" refString="Zonstein S. &amp; Marusik Y. M. 2019. A revision of the spider genus Filistata (Araneae: Filistatidae). Arachnology 18 (2): 53 - 93. https: // doi. org / 10.13156 / arac. 2018.18.2.53" type="journal article" year="2019">Zonstein &amp; Marusik, 2019</bibRefCitation>
</taxonomicName>
, left copulatory bulb, prolateral view. 
<emphasis id="EE8CF413FF83FFC60B3FF84F0B3AEBB3" bold="true" box="[295,313,2001,2027]" pageId="19" pageNumber="168">P</emphasis>
. 
<taxonomicName id="1BF85382FF83FFC60B5EF84C080CEBB3" ID-CoL="7GBLC" authorityName="Kulczynski" authorityYear="1911" box="[326,527,2001,2027]" class="Arachnida" family="Filistatidae" genus="Zaitunia" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="168" phylum="Arthropoda" rank="species" species="schmitzi">
<emphasis id="EE8CF413FF83FFC60B5EF84C080CEBB3" box="[326,527,2001,2027]" italics="true" pageId="19" pageNumber="168">Zaitunia schmitzi</emphasis>
</taxonomicName>
, male clypeus, lateral view, arrow points to stout setae. 
<emphasis id="EE8CF413FF83FFC60E97F84F0EA5EBB3" bold="true" box="[1167,1190,2001,2027]" pageId="19" pageNumber="168">Q</emphasis>
. 
<taxonomicName id="1BF85382FF83FFC60EABF84F0F96EBB3" ID-CoL="6J2PS" baseAuthorityName="Forsskal" baseAuthorityYear="1775" box="[1203,1429,2001,2027]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="168" phylum="Arthropoda" rank="species" species="insidiatrix">
<emphasis id="EE8CF413FF83FFC60EABF84F0F96EBB3" box="[1203,1429,2001,2027]" italics="true" pageId="19" pageNumber="168">Filistata insidiatrix</emphasis>
</taxonomicName>
, female clypeus, lateral view. Scale bars = 0.2 mm.
</paragraph>
</caption>
<paragraph id="DC472801FF84FFC10AFFFE220F9EEE18" blockId="20.[227,1437,303,1402]" pageId="20" pageNumber="169">
<keyLead id="67098B34FF84FFC10AFFFE220F9EEE18" pageId="20" pageNumber="169">
– Females with discontinuous calamistrum interrupted by a medial gap, with staggered rows with few setae (10 or less) (
<figureCitation id="44C33484FF84FFC10BFDFE410877EDA1" box="[485,628,479,505]" captionStart="Fig" captionStartId="19.[227,270,1684,1710]" captionTargetBox="[291,1376,303,1648]" captionTargetId="figure-0@19.[285,1377,302,1651]" captionTargetPageId="19" captionText="Fig. 10. Filistatid morphology. A. Filistata insidiatrix (Forsskål, 1775), ♀, leg tarsus, arrow points to macrosetae. B. Pritha nana (Simon, 1968), arrow points to right calamistrum. C. Sahastata aravaensis Ganem et al., 2022, left calamistrum. D. Zaitunia schmitzi Kulczyński, 1911, left calamistrum, arrow points to gap between setae. E. Filistata insidiatrix, left calamistrum. F. Filistata lubinae Zonstein &amp; Marusik, 2019, ♂, tarsus I. G. Sahastata aravaensis, ♀. H. Pritha IFMsp185, ♀. I. Pritha sp., male abdomen, dorsal view. J. Zaitunia schmitzi, ♂. K. Zaitunia schmitzi, spermathecae ventral view. L. Pritha tenuispina (Strand, 1914), ♂, left pedipalp, prolateral view, arrow points to cymbium. M. Filistata lubinae, left copulatory bulb, prolateral view. N. Zaitunia schmitzi, ♂, left pedipalp, prolateral view, arrow points to cymbium. O. Filistata albens Zonstein &amp; Marusik, 2019, left copulatory bulb, prolateral view. P. Zaitunia schmitzi, male clypeus, lateral view, arrow points to stout setae. Q. Filistata insidiatrix, female clypeus, lateral view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6916970" httpUri="https://zenodo.org/record/6916970/files/figure.png" pageId="20" pageNumber="169">Fig. 10D–E</figureCitation>
); small to medium spiders (adults with body length around 
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); females brown or light cream (
<figureCitation id="44C33484FF84FFC10860FD9C08C6EE45" box="[632,709,514,541]" captionStart="Fig" captionStartId="15.[227,270,1901,1927]" captionTargetBox="[237,1430,362,1838]" captionTargetId="figure-59@15.[227,1437,352,1846]" captionTargetPageId="15" captionText="Fig. 7. Filistata betarif sp. nov., habitus. A–C. Holotype, ♂, from Tinshemet cave (HUJ -INVAr 21058). A. Dorsal view. B. Lateral view. C. Ventral view. — D–F. Paratype, ♀, from the same locality (MACN- Ar 41222). D. Lateral view. E. Dorsal view. F. Ventral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6916960" httpUri="https://zenodo.org/record/6916960/files/figure.png" pageId="20" pageNumber="169">Figs 7</figureCitation>
, 
<figureCitation id="44C33484FF84FFC108C9FD9C08FEEE44" box="[721,765,514,540]" captionStart="Fig" captionStartId="19.[227,270,1684,1710]" captionTargetBox="[291,1376,303,1648]" captionTargetId="figure-0@19.[285,1377,302,1651]" captionTargetPageId="19" captionText="Fig. 10. Filistatid morphology. A. Filistata insidiatrix (Forsskål, 1775), ♀, leg tarsus, arrow points to macrosetae. B. Pritha nana (Simon, 1968), arrow points to right calamistrum. C. Sahastata aravaensis Ganem et al., 2022, left calamistrum. D. Zaitunia schmitzi Kulczyński, 1911, left calamistrum, arrow points to gap between setae. E. Filistata insidiatrix, left calamistrum. F. Filistata lubinae Zonstein &amp; Marusik, 2019, ♂, tarsus I. G. Sahastata aravaensis, ♀. H. Pritha IFMsp185, ♀. I. Pritha sp., male abdomen, dorsal view. J. Zaitunia schmitzi, ♂. K. Zaitunia schmitzi, spermathecae ventral view. L. Pritha tenuispina (Strand, 1914), ♂, left pedipalp, prolateral view, arrow points to cymbium. M. Filistata lubinae, left copulatory bulb, prolateral view. N. Zaitunia schmitzi, ♂, left pedipalp, prolateral view, arrow points to cymbium. O. Filistata albens Zonstein &amp; Marusik, 2019, left copulatory bulb, prolateral view. P. Zaitunia schmitzi, male clypeus, lateral view, arrow points to stout setae. Q. Filistata insidiatrix, female clypeus, lateral view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6916970" httpUri="https://zenodo.org/record/6916970/files/figure.png" pageId="20" pageNumber="169">10J</figureCitation>
); males with pedipalpal femur without dorsal macrosetae (ventral macrosetae present in 
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<emphasis id="EE8CF413FF84FFC10860FDBB08D8EE67" box="[632,731,549,575]" italics="true" pageId="20" pageNumber="169">Filistata</emphasis>
</taxonomicName>
, difficult to see in 
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<emphasis id="EE8CF413FF84FFC109AFFDB80E19EE67" box="[951,1050,550,575]" italics="true" pageId="20" pageNumber="169">Zaitunia</emphasis>
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)................................................ 3
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</paragraph>
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<keyStep id="670C30A4FF84FFC10AFBFDF90F9EEFBA" pageId="20" pageNumber="169">
<paragraph id="DC472801FF84FFC10AFBFDF90F9FEEB3" blockId="20.[227,1437,303,1402]" pageId="20" pageNumber="169">
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3. Brown spiders, with subrounded carapace (
<figureCitation id="44C33484FF84FFC108E7FDF90963EED9" box="[767,864,615,641]" captionStart="Fig" captionStartId="19.[227,270,1684,1710]" captionTargetBox="[291,1376,303,1648]" captionTargetId="figure-0@19.[285,1377,302,1651]" captionTargetPageId="19" captionText="Fig. 10. Filistatid morphology. A. Filistata insidiatrix (Forsskål, 1775), ♀, leg tarsus, arrow points to macrosetae. B. Pritha nana (Simon, 1968), arrow points to right calamistrum. C. Sahastata aravaensis Ganem et al., 2022, left calamistrum. D. Zaitunia schmitzi Kulczyński, 1911, left calamistrum, arrow points to gap between setae. E. Filistata insidiatrix, left calamistrum. F. Filistata lubinae Zonstein &amp; Marusik, 2019, ♂, tarsus I. G. Sahastata aravaensis, ♀. H. Pritha IFMsp185, ♀. I. Pritha sp., male abdomen, dorsal view. J. Zaitunia schmitzi, ♂. K. Zaitunia schmitzi, spermathecae ventral view. L. Pritha tenuispina (Strand, 1914), ♂, left pedipalp, prolateral view, arrow points to cymbium. M. Filistata lubinae, left copulatory bulb, prolateral view. N. Zaitunia schmitzi, ♂, left pedipalp, prolateral view, arrow points to cymbium. O. Filistata albens Zonstein &amp; Marusik, 2019, left copulatory bulb, prolateral view. P. Zaitunia schmitzi, male clypeus, lateral view, arrow points to stout setae. Q. Filistata insidiatrix, female clypeus, lateral view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6916970" httpUri="https://zenodo.org/record/6916970/files/figure.png" pageId="20" pageNumber="169">Fig. 10J</figureCitation>
); steep clypeus with strong, erect setae (
<figureCitation id="44C33484FF84FFC10F30FDF90F8DEED9" box="[1320,1422,615,641]" captionStart="Fig" captionStartId="19.[227,270,1684,1710]" captionTargetBox="[291,1376,303,1648]" captionTargetId="figure-0@19.[285,1377,302,1651]" captionTargetPageId="19" captionText="Fig. 10. Filistatid morphology. A. Filistata insidiatrix (Forsskål, 1775), ♀, leg tarsus, arrow points to macrosetae. B. Pritha nana (Simon, 1968), arrow points to right calamistrum. C. Sahastata aravaensis Ganem et al., 2022, left calamistrum. D. Zaitunia schmitzi Kulczyński, 1911, left calamistrum, arrow points to gap between setae. E. Filistata insidiatrix, left calamistrum. F. Filistata lubinae Zonstein &amp; Marusik, 2019, ♂, tarsus I. G. Sahastata aravaensis, ♀. H. Pritha IFMsp185, ♀. I. Pritha sp., male abdomen, dorsal view. J. Zaitunia schmitzi, ♂. K. Zaitunia schmitzi, spermathecae ventral view. L. Pritha tenuispina (Strand, 1914), ♂, left pedipalp, prolateral view, arrow points to cymbium. M. Filistata lubinae, left copulatory bulb, prolateral view. N. Zaitunia schmitzi, ♂, left pedipalp, prolateral view, arrow points to cymbium. O. Filistata albens Zonstein &amp; Marusik, 2019, left copulatory bulb, prolateral view. P. Zaitunia schmitzi, male clypeus, lateral view, arrow points to stout setae. Q. Filistata insidiatrix, female clypeus, lateral view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6916970" httpUri="https://zenodo.org/record/6916970/files/figure.png" pageId="20" pageNumber="169">Fig. 10P</figureCitation>
); females with two separate pairs of spermathecae (
<figureCitation id="44C33484FF84FFC1094EFD1409C0EEFC" box="[854,963,650,676]" captionStart="Fig" captionStartId="19.[227,270,1684,1710]" captionTargetBox="[291,1376,303,1648]" captionTargetId="figure-0@19.[285,1377,302,1651]" captionTargetPageId="19" captionText="Fig. 10. Filistatid morphology. A. Filistata insidiatrix (Forsskål, 1775), ♀, leg tarsus, arrow points to macrosetae. B. Pritha nana (Simon, 1968), arrow points to right calamistrum. C. Sahastata aravaensis Ganem et al., 2022, left calamistrum. D. Zaitunia schmitzi Kulczyński, 1911, left calamistrum, arrow points to gap between setae. E. Filistata insidiatrix, left calamistrum. F. Filistata lubinae Zonstein &amp; Marusik, 2019, ♂, tarsus I. G. Sahastata aravaensis, ♀. H. Pritha IFMsp185, ♀. I. Pritha sp., male abdomen, dorsal view. J. Zaitunia schmitzi, ♂. K. Zaitunia schmitzi, spermathecae ventral view. L. Pritha tenuispina (Strand, 1914), ♂, left pedipalp, prolateral view, arrow points to cymbium. M. Filistata lubinae, left copulatory bulb, prolateral view. N. Zaitunia schmitzi, ♂, left pedipalp, prolateral view, arrow points to cymbium. O. Filistata albens Zonstein &amp; Marusik, 2019, left copulatory bulb, prolateral view. P. Zaitunia schmitzi, male clypeus, lateral view, arrow points to stout setae. Q. Filistata insidiatrix, female clypeus, lateral view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6916970" httpUri="https://zenodo.org/record/6916970/files/figure.png" pageId="20" pageNumber="169">Fig. 10K</figureCitation>
); male cymbium without two strong and elongated prolateral setae; male pedipalpal tibia short, at most 2 times as long as wide (
<figureCitation id="44C33484FF84FFC10F08FD330F78EE90" box="[1296,1403,685,712]" captionStart="Fig" captionStartId="19.[227,270,1684,1710]" captionTargetBox="[291,1376,303,1648]" captionTargetId="figure-0@19.[285,1377,302,1651]" captionTargetPageId="19" captionText="Fig. 10. Filistatid morphology. A. Filistata insidiatrix (Forsskål, 1775), ♀, leg tarsus, arrow points to macrosetae. B. Pritha nana (Simon, 1968), arrow points to right calamistrum. C. Sahastata aravaensis Ganem et al., 2022, left calamistrum. D. Zaitunia schmitzi Kulczyński, 1911, left calamistrum, arrow points to gap between setae. E. Filistata insidiatrix, left calamistrum. F. Filistata lubinae Zonstein &amp; Marusik, 2019, ♂, tarsus I. G. Sahastata aravaensis, ♀. H. Pritha IFMsp185, ♀. I. Pritha sp., male abdomen, dorsal view. J. Zaitunia schmitzi, ♂. K. Zaitunia schmitzi, spermathecae ventral view. L. Pritha tenuispina (Strand, 1914), ♂, left pedipalp, prolateral view, arrow points to cymbium. M. Filistata lubinae, left copulatory bulb, prolateral view. N. Zaitunia schmitzi, ♂, left pedipalp, prolateral view, arrow points to cymbium. O. Filistata albens Zonstein &amp; Marusik, 2019, left copulatory bulb, prolateral view. P. Zaitunia schmitzi, male clypeus, lateral view, arrow points to stout setae. Q. Filistata insidiatrix, female clypeus, lateral view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6916970" httpUri="https://zenodo.org/record/6916970/files/figure.png" pageId="20" pageNumber="169">Fig. 10N</figureCitation>
) ... 
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<emphasis id="EE8CF413FF84FFC10B3AFD4E0B89EEB2" bold="true" box="[290,394,720,746]" italics="true" pageId="20" pageNumber="169">Zaitunia</emphasis>
<bibRefCitation id="B86955F0FF84FFC10B8AFD4F0847EEB2" author="Lehtinen P. T." box="[402,580,720,747]" pageId="20" pageNumber="169" pagination="199 - 468" refId="ref14139" refString="Lehtinen P. T. 1967. Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici 4: 199 - 468." type="journal article" year="1967">Lehtinen, 1967</bibRefCitation>
</taxonomicName>
[a single known species in 
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, 
<taxonomicName id="1BF85382FF84FFC109CFFD4F0F92EEB3" ID-CoL="7GBLC" authority="(Kulczynski, 1911)" baseAuthorityName="Kulczynski" baseAuthorityYear="1911" box="[983,1425,720,747]" class="Arachnida" family="Filistatidae" genus="Zaitunia" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="169" phylum="Arthropoda" rank="species" species="schmitzi">
<emphasis id="EE8CF413FF84FFC109CFFD4F0EA1EEB3" box="[983,1186,721,747]" italics="true" pageId="20" pageNumber="169">Zaitunia schmitzi</emphasis>
(Kulczyński, 1911)
</taxonomicName>
].
</keyLead>
</paragraph>
<paragraph id="DC472801FF84FFC10AFFFD6A0F9EEFBA" blockId="20.[227,1437,303,1402]" pageId="20" pageNumber="169">
<keyLead id="67098B34FF84FFC10AFFFD6A0F9EEFBA" pageId="20" pageNumber="169">
– Light cream spiders, with carapace longer than wide (
<figureCitation id="44C33484FF84FFC10961FD6A09C3EF56" box="[889,960,756,782]" captionStart="Fig" captionStartId="15.[227,270,1901,1927]" captionTargetBox="[237,1430,362,1838]" captionTargetId="figure-59@15.[227,1437,352,1846]" captionTargetPageId="15" captionText="Fig. 7. Filistata betarif sp. nov., habitus. A–C. Holotype, ♂, from Tinshemet cave (HUJ -INVAr 21058). A. Dorsal view. B. Lateral view. C. Ventral view. — D–F. Paratype, ♀, from the same locality (MACN- Ar 41222). D. Lateral view. E. Dorsal view. F. Ventral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6916960" httpUri="https://zenodo.org/record/6916960/files/figure.png" pageId="20" pageNumber="169">Fig. 7</figureCitation>
); flat clypeus, without particularly strong setae (
<figureCitation id="44C33484FF84FFC10B43FC890BCAEF69" box="[347,457,791,817]" captionStart="Fig" captionStartId="19.[227,270,1684,1710]" captionTargetBox="[291,1376,303,1648]" captionTargetId="figure-0@19.[285,1377,302,1651]" captionTargetPageId="19" captionText="Fig. 10. Filistatid morphology. A. Filistata insidiatrix (Forsskål, 1775), ♀, leg tarsus, arrow points to macrosetae. B. Pritha nana (Simon, 1968), arrow points to right calamistrum. C. Sahastata aravaensis Ganem et al., 2022, left calamistrum. D. Zaitunia schmitzi Kulczyński, 1911, left calamistrum, arrow points to gap between setae. E. Filistata insidiatrix, left calamistrum. F. Filistata lubinae Zonstein &amp; Marusik, 2019, ♂, tarsus I. G. Sahastata aravaensis, ♀. H. Pritha IFMsp185, ♀. I. Pritha sp., male abdomen, dorsal view. J. Zaitunia schmitzi, ♂. K. Zaitunia schmitzi, spermathecae ventral view. L. Pritha tenuispina (Strand, 1914), ♂, left pedipalp, prolateral view, arrow points to cymbium. M. Filistata lubinae, left copulatory bulb, prolateral view. N. Zaitunia schmitzi, ♂, left pedipalp, prolateral view, arrow points to cymbium. O. Filistata albens Zonstein &amp; Marusik, 2019, left copulatory bulb, prolateral view. P. Zaitunia schmitzi, male clypeus, lateral view, arrow points to stout setae. Q. Filistata insidiatrix, female clypeus, lateral view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6916970" httpUri="https://zenodo.org/record/6916970/files/figure.png" pageId="20" pageNumber="169">Fig. 10Q</figureCitation>
); females with a single pair of spermathecae (
<figureCitation id="44C33484FF84FFC109FDFC890E2EEF69" box="[997,1069,791,817]" captionStart="Fig" captionStartId="17.[227,270,1965,1991]" captionTargetBox="[227,1437,1209,1925]" captionTargetId="figure-320@17.[227,1437,1209,1925]" captionTargetPageId="17" captionText="Fig. 9. Filistata betarif sp. nov., female spermathecae, ventral. A. Oah cave (HUJ-INVAr 21055). B. Tinshemet cave (MACN-Ar 41222). C–D. Tinshemet cave (MACN-Ar 41791). Scale bars: A–C = 0.2 mm; D = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6916968" httpUri="https://zenodo.org/record/6916968/files/figure.png" pageId="20" pageNumber="169">Fig. 9</figureCitation>
), or if with two pairs, these are partly fused (
<figureCitation id="44C33484FF84FFC10BB2FCA4082AEF0C" box="[426,553,826,852]" captionStart="Fig" captionStartId="8.[227,270,1754,1780]" captionTargetBox="[273,1385,302,1702]" captionTargetId="figure-137@8.[264,1399,302,1703]" captionTargetPageId="8" captionText="Fig. 3. A–G. Endogynes Filistata Latreille, 1810, ventral view. A. Filistata wunderlichi Zonstein &amp; Marusik, 2019 from Spain, Boca de los Frailes (CRBA IFM-2197). B. Filistata IFMsp191 from Israel, Odem (MACN-Ar 41262). C. Filistata insidiatrix (Forsskål, 1775) from France, Gonfaron (MACN-Ar 39461). D. Filistata insidiatrix from Israel, Beriniki (HUJ-INVAr 21052). E. Filistata betarif sp. nov. from Israel, Oah cave (HUJ-INVAr 21055). F. Filistata betarif sp. nov. from Israel, Tinshemet cave (MACN-Ar 41222). G. Filistata IFMsp99 (subadult ♀?) from Palestine, En Fashkha (HUJ-INVAr 20277). — H–I. ♂♂, left pedipalp, prolateral. H. Filistata betarif sp. nov., holotype from Israel, Tinshemet cave (HUJ -INVAr 21058). I. Filistata insidiatrix from Spain, Mallorca (MACN-Ar 42346). Scale bar = 0.1 mm, all figures to scale. Colours correspond to clades in Fig. 2." figureDoi="http://doi.org/10.5281/zenodo.6916944" httpUri="https://zenodo.org/record/6916944/files/figure.png" pageId="20" pageNumber="169">Fig. 3B–C</figureCitation>
); male cymbium with two strong prolateral setae (
<figureCitation id="44C33484FF84FFC10E69FCA40ECFEF0C" box="[1137,1228,826,852]" captionStart="Fig" captionStartId="16.[227,270,1901,1927]" captionTargetBox="[233,1434,365,1831]" captionTargetId="figure-70@16.[227,1437,352,1846]" captionTargetPageId="16" captionText="Fig. 8. A–G. Filistata betarif sp. nov., holotype, ♂ (HUJ -INVAr 21058), left pedipalp. A–B, E. Prolateral view. Arrow points to prolateral strong setae in cymbium. C, F–G. Retrolateral view. Arrow points to embolic keel. D. Dorsal view. — H. Filistata insidiatrix (Forsskål, 1775) (MACN-Ar 39461), bulb, retrolateral view. Arrow points to embolic keel. Scale bars: A–F = 0.2 mm; G–H = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6916964" httpUri="https://zenodo.org/record/6916964/files/figure.png" pageId="20" pageNumber="169">Fig. 8B</figureCitation>
); male pedipalpal tibia long, at least three times as long as wide (
<figureCitation id="44C33484FF84FFC1092DFCC30992EF20" box="[821,913,861,888]" captionStart="Fig" captionStartId="16.[227,270,1901,1927]" captionTargetBox="[233,1434,365,1831]" captionTargetId="figure-70@16.[227,1437,352,1846]" captionTargetPageId="16" captionText="Fig. 8. A–G. Filistata betarif sp. nov., holotype, ♂ (HUJ -INVAr 21058), left pedipalp. A–B, E. Prolateral view. Arrow points to prolateral strong setae in cymbium. C, F–G. Retrolateral view. Arrow points to embolic keel. D. Dorsal view. — H. Filistata insidiatrix (Forsskål, 1775) (MACN-Ar 39461), bulb, retrolateral view. Arrow points to embolic keel. Scale bars: A–F = 0.2 mm; G–H = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6916964" httpUri="https://zenodo.org/record/6916964/files/figure.png" pageId="20" pageNumber="169">Fig. 8A</figureCitation>
)..................................................................... ........................................................................................ 4 (
<taxonomicName id="1BF85382FF84FFC109D8FC1E0ED9EFC3" ID-CoL="4K4M" authority="Latreille, 1810" authorityName="Latreille" authorityYear="1810" box="[960,1242,896,923]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="EE8CF413FF84FFC109D8FC1E0E26EFC3" bold="true" box="[960,1061,896,923]" italics="true" pageId="20" pageNumber="169">Filistata</emphasis>
<bibRefCitation id="B86955F0FF84FFC10E34FC1F0ED9EFC3" author="Latreille P. A." box="[1068,1242,897,923]" pageId="20" pageNumber="169" refId="ref13962" refString="Latreille P. A. 1810. Considerations generales sur l'ordre naturel des crustaces, des arachnides et des insectes. F. Schoell, Paris. https: // doi. org / 10.5962 / bhl. title. 39620" type="book" year="1810">Latreille, 1810</bibRefCitation>
</taxonomicName>
) [four described species in 
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, of those four species, only the females of 
<taxonomicName id="1BF85382FF84FFC109CAFC3B0E6AEFE6" ID-CoL="6J2PS" baseAuthorityName="Forsskal" baseAuthorityYear="1775" box="[978,1129,932,958]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="169" phylum="Arthropoda" rank="species" species="insidiatrix">
<emphasis id="EE8CF413FF84FFC109CAFC3B0E6AEFE6" box="[978,1129,932,958]" italics="true" pageId="20" pageNumber="169">F. insidiatrix</emphasis>
</taxonomicName>
and 
<taxonomicName id="1BF85382FF84FFC10EBCFC3B0F13EFE6" authorityName="Magalhaes &amp; Aharon &amp; Ganem &amp; Gavish-Regev" authorityYear="2022" box="[1188,1296,932,958]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="169" phylum="Arthropoda" rank="species" species="betarif">
<emphasis id="EE8CF413FF84FFC10EBCFC3B0F13EFE6" box="[1188,1296,932,958]" italics="true" pageId="20" pageNumber="169">F. betarif</emphasis>
</taxonomicName>
are known, and are not morphologically diagnosable; thus, the remainder of the key refers only to males].
</keyLead>
</paragraph>
</keyStep>
<keyStep id="670C30A4FF84FFC10AFBFB970F9EE832" pageId="20" pageNumber="169">
<paragraph id="DC472801FF84FFC10AFBFB970F9EE81E" blockId="20.[227,1437,303,1402]" pageId="20" pageNumber="169">
<keyLead id="67098B34FF84FFC10AFBFB970F9EE81E" pageId="20" pageNumber="169">
4. Male copulatory bulb short and stout, with long and coiled embolus (
<figureCitation id="44C33484FF84FFC10E84FB970F1BE87B" box="[1180,1304,1033,1059]" captionStart="Fig" captionStartId="19.[227,270,1684,1710]" captionTargetBox="[291,1376,303,1648]" captionTargetId="figure-0@19.[285,1377,302,1651]" captionTargetPageId="19" captionText="Fig. 10. Filistatid morphology. A. Filistata insidiatrix (Forsskål, 1775), ♀, leg tarsus, arrow points to macrosetae. B. Pritha nana (Simon, 1968), arrow points to right calamistrum. C. Sahastata aravaensis Ganem et al., 2022, left calamistrum. D. Zaitunia schmitzi Kulczyński, 1911, left calamistrum, arrow points to gap between setae. E. Filistata insidiatrix, left calamistrum. F. Filistata lubinae Zonstein &amp; Marusik, 2019, ♂, tarsus I. G. Sahastata aravaensis, ♀. H. Pritha IFMsp185, ♀. I. Pritha sp., male abdomen, dorsal view. J. Zaitunia schmitzi, ♂. K. Zaitunia schmitzi, spermathecae ventral view. L. Pritha tenuispina (Strand, 1914), ♂, left pedipalp, prolateral view, arrow points to cymbium. M. Filistata lubinae, left copulatory bulb, prolateral view. N. Zaitunia schmitzi, ♂, left pedipalp, prolateral view, arrow points to cymbium. O. Filistata albens Zonstein &amp; Marusik, 2019, left copulatory bulb, prolateral view. P. Zaitunia schmitzi, male clypeus, lateral view, arrow points to stout setae. Q. Filistata insidiatrix, female clypeus, lateral view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6916970" httpUri="https://zenodo.org/record/6916970/files/figure.png" pageId="20" pageNumber="169">Fig. 10M</figureCitation>
); carapace subrounded..................................................................... 
<taxonomicName id="1BF85382FF84FFC1098EFBB50F9EE81E" ID-CoL="99CSP" authority="Zonstein &amp; Marusik, 2019" authorityName="Zonstein &amp; Marusik" authorityYear="2019" box="[918,1437,1067,1094]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="169" phylum="Arthropoda" rank="species" species="lubinae">
<emphasis id="EE8CF413FF84FFC1098EFBB50E5EE81E" bold="true" box="[918,1117,1067,1094]" italics="true" pageId="20" pageNumber="169">Filistata lubinae</emphasis>
<bibRefCitation id="B86955F0FF84FFC10E7DFBB20F9EE81E" author="Zonstein S. &amp; Marusik Y. M." box="[1125,1437,1068,1094]" pageId="20" pageNumber="169" pagination="53 - 93" refId="ref15464" refString="Zonstein S. &amp; Marusik Y. M. 2019. A revision of the spider genus Filistata (Araneae: Filistatidae). Arachnology 18 (2): 53 - 93. https: // doi. org / 10.13156 / arac. 2018.18.2.53" type="journal article" year="2019">Zonstein &amp; Marusik, 2019</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="DC472801FF84FFC10AFFFBCE0F9EE832" blockId="20.[227,1437,303,1402]" box="[231,1437,1103,1130]" pageId="20" pageNumber="169">
<keyLead id="67098B34FF84FFC10AFFFBCE0F9EE832" box="[231,1437,1103,1130]" pageId="20" pageNumber="169">
– Male copulatory bulb subconical (
<figureCitation id="44C33484FF84FFC108B9FBD108EFE831" box="[673,748,1103,1129]" captionStart="Fig" captionStartId="16.[227,270,1901,1927]" captionTargetBox="[233,1434,365,1831]" captionTargetId="figure-70@16.[227,1437,352,1846]" captionTargetPageId="16" captionText="Fig. 8. A–G. Filistata betarif sp. nov., holotype, ♂ (HUJ -INVAr 21058), left pedipalp. A–B, E. Prolateral view. Arrow points to prolateral strong setae in cymbium. C, F–G. Retrolateral view. Arrow points to embolic keel. D. Dorsal view. — H. Filistata insidiatrix (Forsskål, 1775) (MACN-Ar 39461), bulb, retrolateral view. Arrow points to embolic keel. Scale bars: A–F = 0.2 mm; G–H = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6916964" httpUri="https://zenodo.org/record/6916964/files/figure.png" pageId="20" pageNumber="169">Figs 8</figureCitation>
, 
<figureCitation id="44C33484FF84FFC108E2FBD1092CE831" box="[762,815,1103,1129]" captionStart="Fig" captionStartId="19.[227,270,1684,1710]" captionTargetBox="[291,1376,303,1648]" captionTargetId="figure-0@19.[285,1377,302,1651]" captionTargetPageId="19" captionText="Fig. 10. Filistatid morphology. A. Filistata insidiatrix (Forsskål, 1775), ♀, leg tarsus, arrow points to macrosetae. B. Pritha nana (Simon, 1968), arrow points to right calamistrum. C. Sahastata aravaensis Ganem et al., 2022, left calamistrum. D. Zaitunia schmitzi Kulczyński, 1911, left calamistrum, arrow points to gap between setae. E. Filistata insidiatrix, left calamistrum. F. Filistata lubinae Zonstein &amp; Marusik, 2019, ♂, tarsus I. G. Sahastata aravaensis, ♀. H. Pritha IFMsp185, ♀. I. Pritha sp., male abdomen, dorsal view. J. Zaitunia schmitzi, ♂. K. Zaitunia schmitzi, spermathecae ventral view. L. Pritha tenuispina (Strand, 1914), ♂, left pedipalp, prolateral view, arrow points to cymbium. M. Filistata lubinae, left copulatory bulb, prolateral view. N. Zaitunia schmitzi, ♂, left pedipalp, prolateral view, arrow points to cymbium. O. Filistata albens Zonstein &amp; Marusik, 2019, left copulatory bulb, prolateral view. P. Zaitunia schmitzi, male clypeus, lateral view, arrow points to stout setae. Q. Filistata insidiatrix, female clypeus, lateral view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6916970" httpUri="https://zenodo.org/record/6916970/files/figure.png" pageId="20" pageNumber="169">10O</figureCitation>
), embolus not coiled; carapace longer than wide.... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="670C30A4FF84FFC10AFBFB0F0F9EE94C" pageId="20" pageNumber="169">
<paragraph id="DC472801FF84FFC10AFBFB0F0F9EE896" blockId="20.[227,1437,303,1402]" pageId="20" pageNumber="169">
<keyLead id="67098B34FF84FFC10AFBFB0F0F9EE896" pageId="20" pageNumber="169">
5. Sperm duct loosely packed, with space between coils; copulatory bulb shorter (ca 80% of cymbium length), with ventral side nearly straight (
<figureCitation id="44C33484FF84FFC108EDFB2A0963E896" box="[757,864,1204,1230]" captionStart="Fig" captionStartId="19.[227,270,1684,1710]" captionTargetBox="[291,1376,303,1648]" captionTargetId="figure-0@19.[285,1377,302,1651]" captionTargetPageId="19" captionText="Fig. 10. Filistatid morphology. A. Filistata insidiatrix (Forsskål, 1775), ♀, leg tarsus, arrow points to macrosetae. B. Pritha nana (Simon, 1968), arrow points to right calamistrum. C. Sahastata aravaensis Ganem et al., 2022, left calamistrum. D. Zaitunia schmitzi Kulczyński, 1911, left calamistrum, arrow points to gap between setae. E. Filistata insidiatrix, left calamistrum. F. Filistata lubinae Zonstein &amp; Marusik, 2019, ♂, tarsus I. G. Sahastata aravaensis, ♀. H. Pritha IFMsp185, ♀. I. Pritha sp., male abdomen, dorsal view. J. Zaitunia schmitzi, ♂. K. Zaitunia schmitzi, spermathecae ventral view. L. Pritha tenuispina (Strand, 1914), ♂, left pedipalp, prolateral view, arrow points to cymbium. M. Filistata lubinae, left copulatory bulb, prolateral view. N. Zaitunia schmitzi, ♂, left pedipalp, prolateral view, arrow points to cymbium. O. Filistata albens Zonstein &amp; Marusik, 2019, left copulatory bulb, prolateral view. P. Zaitunia schmitzi, male clypeus, lateral view, arrow points to stout setae. Q. Filistata insidiatrix, female clypeus, lateral view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6916970" httpUri="https://zenodo.org/record/6916970/files/figure.png" pageId="20" pageNumber="169">Fig. 10O</figureCitation>
) ....... 
<taxonomicName id="1BF85382FF84FFC109BBFB2D0F9EE896" ID-CoL="99BS5" authority="Zonstein &amp; Marusik, 2019" authorityName="Zonstein &amp; Marusik" authorityYear="2019" box="[931,1437,1203,1230]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="169" phylum="Arthropoda" rank="species" species="albens">
<emphasis id="EE8CF413FF84FFC109BBFB2D0E5EE896" bold="true" box="[931,1117,1203,1230]" italics="true" pageId="20" pageNumber="169">Filistata albens</emphasis>
<bibRefCitation id="B86955F0FF84FFC10E7DFB2A0F9EE896" author="Zonstein S. &amp; Marusik Y. M." box="[1125,1437,1204,1230]" pageId="20" pageNumber="169" pagination="53 - 93" refId="ref15464" refString="Zonstein S. &amp; Marusik Y. M. 2019. A revision of the spider genus Filistata (Araneae: Filistatidae). Arachnology 18 (2): 53 - 93. https: // doi. org / 10.13156 / arac. 2018.18.2.53" type="journal article" year="2019">Zonstein &amp; Marusik, 2019</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="DC472801FF84FFC10AFFFB460F9EE94C" blockId="20.[227,1437,303,1402]" pageId="20" pageNumber="169">
<keyLead id="67098B34FF84FFC10AFFFB460F9EE94C" pageId="20" pageNumber="169">
– Sperm duct tightly packed (
<figureCitation id="44C33484FF84FFC1084FFB49089DE8A9" box="[599,670,1239,1265]" captionStart="Fig" captionStartId="16.[227,270,1901,1927]" captionTargetBox="[233,1434,365,1831]" captionTargetId="figure-70@16.[227,1437,352,1846]" captionTargetPageId="16" captionText="Fig. 8. A–G. Filistata betarif sp. nov., holotype, ♂ (HUJ -INVAr 21058), left pedipalp. A–B, E. Prolateral view. Arrow points to prolateral strong setae in cymbium. C, F–G. Retrolateral view. Arrow points to embolic keel. D. Dorsal view. — H. Filistata insidiatrix (Forsskål, 1775) (MACN-Ar 39461), bulb, retrolateral view. Arrow points to embolic keel. Scale bars: A–F = 0.2 mm; G–H = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6916964" httpUri="https://zenodo.org/record/6916964/files/figure.png" pageId="20" pageNumber="169">Fig. 8</figureCitation>
), without space between coils; copulatory bulb longer (almost as long as cymbium), with ventral side curved inwards ........................................................................ 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="670C30A4FF84FFC10AFBFAA20F9FE922" pageId="20" pageNumber="169">
<paragraph id="DC472801FF84FFC10AFBFAA20F9EE90E" blockId="20.[227,1437,303,1402]" box="[227,1437,1339,1367]" pageId="20" pageNumber="169">
<keyLead id="67098B34FF84FFC10AFBFAA20F9EE90E" box="[227,1437,1339,1367]" pageId="20" pageNumber="169">
6. Embolic keel conspicuous (
<figureCitation id="44C33484FF84FFC1084CFAA208B6E90E" box="[596,693,1340,1366]" captionStart="Fig" captionStartId="8.[227,270,1754,1780]" captionTargetBox="[273,1385,302,1702]" captionTargetId="figure-137@8.[264,1399,302,1703]" captionTargetPageId="8" captionText="Fig. 3. A–G. Endogynes Filistata Latreille, 1810, ventral view. A. Filistata wunderlichi Zonstein &amp; Marusik, 2019 from Spain, Boca de los Frailes (CRBA IFM-2197). B. Filistata IFMsp191 from Israel, Odem (MACN-Ar 41262). C. Filistata insidiatrix (Forsskål, 1775) from France, Gonfaron (MACN-Ar 39461). D. Filistata insidiatrix from Israel, Beriniki (HUJ-INVAr 21052). E. Filistata betarif sp. nov. from Israel, Oah cave (HUJ-INVAr 21055). F. Filistata betarif sp. nov. from Israel, Tinshemet cave (MACN-Ar 41222). G. Filistata IFMsp99 (subadult ♀?) from Palestine, En Fashkha (HUJ-INVAr 20277). — H–I. ♂♂, left pedipalp, prolateral. H. Filistata betarif sp. nov., holotype from Israel, Tinshemet cave (HUJ -INVAr 21058). I. Filistata insidiatrix from Spain, Mallorca (MACN-Ar 42346). Scale bar = 0.1 mm, all figures to scale. Colours correspond to clades in Fig. 2." figureDoi="http://doi.org/10.5281/zenodo.6916944" httpUri="https://zenodo.org/record/6916944/files/figure.png" pageId="20" pageNumber="169">Figs 3H</figureCitation>
, 
<figureCitation id="44C33484FF84FFC108DAFAA208EBE90E" box="[706,744,1340,1366]" captionStart="Fig" captionStartId="16.[227,270,1901,1927]" captionTargetBox="[233,1434,365,1831]" captionTargetId="figure-70@16.[227,1437,352,1846]" captionTargetPageId="16" captionText="Fig. 8. A–G. Filistata betarif sp. nov., holotype, ♂ (HUJ -INVAr 21058), left pedipalp. A–B, E. Prolateral view. Arrow points to prolateral strong setae in cymbium. C, F–G. Retrolateral view. Arrow points to embolic keel. D. Dorsal view. — H. Filistata insidiatrix (Forsskål, 1775) (MACN-Ar 39461), bulb, retrolateral view. Arrow points to embolic keel. Scale bars: A–F = 0.2 mm; G–H = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6916964" httpUri="https://zenodo.org/record/6916964/files/figure.png" pageId="20" pageNumber="169">8G</figureCitation>
, 
<figureCitation id="44C33484FF84FFC108ECFAA2091AE90F" box="[756,793,1340,1367]" captionStart="Fig" captionStartId="13.[227,270,1928,1954]" captionTargetBox="[227,1437,808,1883]" captionTargetId="figure-274@13.[227,1437,808,1883]" captionTargetPageId="13" captionText="Fig. 5. A, C, E. Filistata betarif sp. nov., paratype, ♂, from Israel, Tinshemet cave, (MACN-Ar 41793), left pedipalp. B, D, F. Filistata insidiatrix (Forsskål, 1775), ♂, from Spain, Mallorca (MACN-Ar 42347), right pedipalp, mirrored. A–B. Prolateral view. C–D. Dorsal view. E–F. Retrolateral view. Abbreviations: k = embolic keel; ps = prolateral setae. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6916952" httpUri="https://zenodo.org/record/6916952/files/figure.png" pageId="20" pageNumber="169">5A</figureCitation>
) .............................................. 
<taxonomicName id="1BF85382FF84FFC10E66FAA50F38E90E" authority="Magalhaes &amp; Aharon &amp; Ganem &amp; Gavish-Regev, 2022" authorityName="Magalhaes &amp; Aharon &amp; Ganem &amp; Gavish-Regev" authorityYear="2022" box="[1150,1339,1339,1366]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="169" phylum="Arthropoda" rank="species" species="betarif" status="sp. nov.">
<emphasis id="EE8CF413FF84FFC10E66FAA50F38E90E" bold="true" box="[1150,1339,1339,1366]" italics="true" pageId="20" pageNumber="169">Filistata betarif</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F5BF4968FF84FFC10F5AFAA20F9EE90E" box="[1346,1437,1340,1366]" pageId="20" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="DC472801FF84FFC10AFFFAFE0F9FE922" blockId="20.[227,1437,303,1402]" box="[231,1436,1375,1402]" pageId="20" pageNumber="169">
<keyLead id="67098B34FF84FFC10AFFFAFE0F9FE922" box="[231,1436,1375,1402]" pageId="20" pageNumber="169">
– Embolic keel inconspicuous (
<figureCitation id="44C33484FF84FFC10873FAC108BFE921" box="[619,700,1375,1401]" captionStart="Fig" captionStartId="8.[227,270,1754,1780]" captionTargetBox="[273,1385,302,1702]" captionTargetId="figure-137@8.[264,1399,302,1703]" captionTargetPageId="8" captionText="Fig. 3. A–G. Endogynes Filistata Latreille, 1810, ventral view. A. Filistata wunderlichi Zonstein &amp; Marusik, 2019 from Spain, Boca de los Frailes (CRBA IFM-2197). B. Filistata IFMsp191 from Israel, Odem (MACN-Ar 41262). C. Filistata insidiatrix (Forsskål, 1775) from France, Gonfaron (MACN-Ar 39461). D. Filistata insidiatrix from Israel, Beriniki (HUJ-INVAr 21052). E. Filistata betarif sp. nov. from Israel, Oah cave (HUJ-INVAr 21055). F. Filistata betarif sp. nov. from Israel, Tinshemet cave (MACN-Ar 41222). G. Filistata IFMsp99 (subadult ♀?) from Palestine, En Fashkha (HUJ-INVAr 20277). — H–I. ♂♂, left pedipalp, prolateral. H. Filistata betarif sp. nov., holotype from Israel, Tinshemet cave (HUJ -INVAr 21058). I. Filistata insidiatrix from Spain, Mallorca (MACN-Ar 42346). Scale bar = 0.1 mm, all figures to scale. Colours correspond to clades in Fig. 2." figureDoi="http://doi.org/10.5281/zenodo.6916944" httpUri="https://zenodo.org/record/6916944/files/figure.png" pageId="20" pageNumber="169">Fig. 3I</figureCitation>
, 
<figureCitation id="44C33484FF84FFC108D1FAC108ECE921" box="[713,751,1375,1401]" captionStart="Fig" captionStartId="16.[227,270,1901,1927]" captionTargetBox="[233,1434,365,1831]" captionTargetId="figure-70@16.[227,1437,352,1846]" captionTargetPageId="16" captionText="Fig. 8. A–G. Filistata betarif sp. nov., holotype, ♂ (HUJ -INVAr 21058), left pedipalp. A–B, E. Prolateral view. Arrow points to prolateral strong setae in cymbium. C, F–G. Retrolateral view. Arrow points to embolic keel. D. Dorsal view. — H. Filistata insidiatrix (Forsskål, 1775) (MACN-Ar 39461), bulb, retrolateral view. Arrow points to embolic keel. Scale bars: A–F = 0.2 mm; G–H = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6916964" httpUri="https://zenodo.org/record/6916964/files/figure.png" pageId="20" pageNumber="169">8H</figureCitation>
, 
<figureCitation id="44C33484FF84FFC108E3FAC1091DE922" box="[763,798,1375,1402]" captionStart="Fig" captionStartId="13.[227,270,1928,1954]" captionTargetBox="[227,1437,808,1883]" captionTargetId="figure-274@13.[227,1437,808,1883]" captionTargetPageId="13" captionText="Fig. 5. A, C, E. Filistata betarif sp. nov., paratype, ♂, from Israel, Tinshemet cave, (MACN-Ar 41793), left pedipalp. B, D, F. Filistata insidiatrix (Forsskål, 1775), ♂, from Spain, Mallorca (MACN-Ar 42347), right pedipalp, mirrored. A–B. Prolateral view. C–D. Dorsal view. E–F. Retrolateral view. Abbreviations: k = embolic keel; ps = prolateral setae. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6916952" httpUri="https://zenodo.org/record/6916952/files/figure.png" pageId="20" pageNumber="169">5B</figureCitation>
) .......................... 
<taxonomicName id="1BF85382FF84FFC109F4FAC10F9FE922" ID-CoL="6J2PS" authority="(Forsskal, 1775)" baseAuthorityName="Forsskal" baseAuthorityYear="1775" box="[1004,1436,1375,1402]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="169" phylum="Arthropoda" rank="species" species="insidiatrix">
<emphasis id="EE8CF413FF84FFC109F4FAC10ED6E922" bold="true" box="[1004,1237,1375,1402]" italics="true" pageId="20" pageNumber="169">Filistata insidiatrix</emphasis>
(
<bibRefCitation id="B86955F0FF84FFC10EFCFAC10F97E922" author="Forsskal P." box="[1252,1428,1375,1402]" pageId="20" pageNumber="169" refId="ref13586" refString="Forsskal P. 1775. Descriptiones animalium avium, amphibiorum, piscium, insectorum, vermium; quae in itinere orientali observavit Petrus Forskal [sic]. Moller, Havniae [= Copenhagen]. https: // doi. org / 10.5962 / bhl. title. 2154" type="book" year="1775">Forsskål, 1775</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>