214 lines
26 KiB
XML
214 lines
26 KiB
XML
<document id="E75E572372DB031C2B7B8A2AD7C6230B" ID-DOI="10.5281/zenodo.5356887" ID-ISSN="2345-7600" ID-Zenodo-Dep="5356887" ID-ZooBank="9EB66A01-DC75-4502-9DD0-56A7CFA4B7BD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630218432546" checkinUser="valdenar" docAuthor="Aspe, Nonillon M. & James, Samuel W." docDate="2017" docId="5443705CFFF9793ED591FB5CFBB2F895" docLanguage="en" docName="65rbz357-372.pdf" docOrigin="Raffles Bulletin of Zoology 65" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Amynthas apongensis Aspe & James 2017, new species" docType="treatment" docUuid="83663A13-1176-4F98-A9EC-B76F2238A5EE" docUuidSource="ZooBank" docVersion="5" lastPageNumber="358" masterDocId="A87A0824FFF8793FD57CFFE1FF88FFA0" masterDocTitle="Pheretimoid earthworms (Clitellata: Megascolecidae) from Mt. Apo, Mindanao Island, Philippines with description of eight new species" masterLastPageNumber="372" masterPageNumber="357" pageNumber="358" updateTime="1699190590497" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="47C710FC2D462D2E3B710F63FCBDEA8C">Pheretimoid earthworms (Clitellata: Megascolecidae) from Mt. Apo, Mindanao Island, Philippines with description of eight new species</mods:title>
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<mods:namePart id="926106CA9316E3D97E08FC1865C305A2">Aspe, Nonillon M.</mods:namePart>
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<mods:namePart id="9C2C950ABE3456C003A661D12CFB5812">James, Samuel W.</mods:namePart>
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<mods:typeOfResource id="2290453729CA7C31C94024C4791A61D8">text</mods:typeOfResource>
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<mods:title id="64C1CA7D3B78E9F3ABDEA23EFF0947FE">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="7D82788F833046B42218F451BECB287A">
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<mods:date id="B338ED6B8665FF02884A9C252FC00BEE">2017</mods:date>
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<mods:number id="24FDB84F54DCA08BAD1B37D212C47748">2017-08-07</mods:number>
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<mods:detail id="E027DCB7B136C412DED025CDA039E94A" type="volume">
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<mods:number id="36F0210F2C584C4771840D227572E61E">65</mods:number>
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<mods:start id="34C05E4A8EC4F12EDBF9F315F1211E2D">357</mods:start>
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<mods:classification id="71A558CE425DD1DBE04CAEA9AEB37B53">journal article</mods:classification>
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<mods:identifier id="74C9AF499F1CA5BA7ED34CF656259E13" type="DOI">10.5281/zenodo.5356887</mods:identifier>
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<mods:identifier id="55A298039248038FA7A248F47A1E5EE9" type="ISSN">2345-7600</mods:identifier>
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<treatment id="5443705CFFF9793ED591FB5CFBB2F895" ID-DOI="http://doi.org/10.5281/zenodo.5453261" ID-Zenodo-Dep="5453261" LSID="urn:lsid:zoobank.org:act:83663A13-1176-4F98-A9EC-B76F2238A5EE" httpUri="http://treatment.plazi.org/id/5443705CFFF9793ED591FB5CFBB2F895" lastPageNumber="358" pageId="1" pageNumber="358">
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<subSubSection id="94F092C1FFF9793ED591FB5CFDFCFB75" box="[237,628,1213,1237]" pageId="1" pageNumber="358" type="nomenclature">
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<paragraph id="DC55C14AFFF9793ED591FB5CFDFCFB75" blockId="1.[237,628,1213,1269]" box="[237,628,1213,1237]" pageId="1" pageNumber="358">
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<heading id="871D7626FFF9793ED591FB5CFDFCFB75" bold="true" box="[237,628,1213,1237]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="358" reason="2">
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<emphasis id="EE9E1D58FFF9793ED591FB5CFDFCFB75" bold="true" box="[237,628,1213,1237]" pageId="1" pageNumber="358">
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<taxonomicName id="1BEABAC9FFF9793ED591FB5CFE68FB75" authority="Aspe & James, 2017" authorityName="Aspe & James" authorityYear="2017" box="[237,480,1213,1237]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="358" phylum="Annelida" rank="species" species="apongensis" status="sp. nov.">
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<emphasis id="EE9E1D58FFF9793ED591FB5CFE68FB75" bold="true" box="[237,480,1213,1237]" italics="true" pageId="1" pageNumber="358">Amynthas apongensis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="F5ADA023FFF9793ED493FB5CFDFCFB75" box="[495,628,1213,1237]" pageId="1" pageNumber="358" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="94F092C1FFF9793ED411FB3CFE7BFB55" box="[365,499,1245,1269]" pageId="1" pageNumber="358" type="description">
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<paragraph id="DC55C14AFFF9793ED411FB3CFE7BFB55" blockId="1.[237,628,1213,1269]" box="[365,499,1245,1269]" pageId="1" pageNumber="358">
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(
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<figureCitation id="44D1DDCFFFF9793ED409FB3CFE63FB55" box="[373,491,1245,1269]" captionStart="Fig" captionStartId="2.[135,170,1943,1965]" captionTargetBox="[130,1474,254,1929]" captionTargetId="figure-89@2.[121,1487,245,1939]" captionTargetPageId="2" captionText="Fig. 1. A, Diagrammatic ventral view of Amynthas apongensis, new species showing the spermathecal pores (sp), clitellum (cl), male pores (m), and the pattern of the genital markings (gm). Schematic drawings of the internal morphology in dorsal view: B, A. apongensis, new species; C, Pheretima floresi, new species; D, P. baracatanensis, new species; E, P. solisi, new species. Abbreviations: s, spermatheca; h, heart; p, prostate gland; cb, copulatory bursa; c, caecum. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5356889" httpUri="https://zenodo.org/record/5356889/files/figure.png" pageId="1" pageNumber="358">Fig. 1A, B</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="94F092C1FFF9793ED50DFAFCFD98FA35" pageId="1" pageNumber="358" type="materials_examined">
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<paragraph id="DC55C14AFFF9793ED50DFAFCFD98FA35" blockId="1.[113,751,1309,1430]" pageId="1" pageNumber="358">
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<emphasis id="EE9E1D58FFF9793ED50DFAFCFED0FA95" bold="true" box="[113,344,1309,1333]" pageId="1" pageNumber="358">Material examined.</emphasis>
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<materialsCitation id="6C82CB17FFF9793ED41EFAFCFD98FA35" collectingDate="2003-12-11" collectingDateMax="2003-12-14" collectingDateMin="2003-12-11" collectionCode="NMP" collectorName="N. Aspe" country="Philippines" county="Mindanao Island" elevation="1524" latitude="7.001111" location="Brgy Baracatan" longLatPrecision="21" longitude="125.36527" municipality="Mt. Apo National Park" pageId="1" pageNumber="358" specimenCode="NMP 4628" specimenCount="1" stateProvince="Davao" typeStatus="holotype">
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<typeStatus id="03517FE8FFF9793ED41EFAFCFE44FA95" box="[354,460,1309,1333]" pageId="1" pageNumber="358" type="holotype">Holotype</typeStatus>
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: adult (
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<specimenCode id="8C4C6931FFF9793ED75EFAFCFD2AFA95" box="[546,674,1309,1333]" collectionCode="NMP" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="1" pageNumber="358">NMP 4628</specimenCode>
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),
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<location id="D9359791FFF9793ED7C5FAFFFF56FAF5" LSID="urn:lsid:plazi:treatment:5443705CFFF9793ED591FB5CFBB2F895:D9359791FFF9793ED7C5FAFFFF56FAF5" country="Philippines" county="Mindanao Island" latitude="7.001111" longLatPrecision="21" longitude="125.36527" municipality="Mt. Apo National Park" name="Brgy Baracatan" pageId="1" pageNumber="358" stateProvince="Davao">Brgy Baracatan</location>
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,
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<collectingRegion id="1E2E0FA8FFF9793ED596FADFFEB8FAF5" box="[234,304,1342,1365]" country="Philippines" name="Davao" pageId="1" pageNumber="358">Davao</collectingRegion>
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<location id="D9359791FFF9793ED44AFADCFEEFFAF5" LSID="urn:lsid:plazi:treatment:5443705CFFF9793ED591FB5CFBB2F895:D9359791FFF9793ED44AFADCFEEFFAF5" box="[310,359,1341,1365]" country="Philippines" county="Mindanao Island" latitude="7.001111" longLatPrecision="21" longitude="125.36527" municipality="Mt. Apo National Park" name="City" pageId="1" pageNumber="358" stateProvince="Davao">City</location>
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,
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<collectingMunicipality id="3C315B30FFF9793ED40EFADFFDEFFAF5" box="[370,615,1341,1366]" pageId="1" pageNumber="358">Mt. Apo National Park</collectingMunicipality>
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(
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<geoCoordinate id="B9DEA78DFFF9793ED70AFADDFD63FAF5" box="[630,747,1340,1365]" degrees="7" direction="north" minutes="00" orientation="latitude" pageId="1" pageNumber="358" precision="15" seconds="04" value="7.001111">7°00′04″N</geoCoordinate>
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,
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<geoCoordinate id="B9DEA78DFFF9793ED50DFABDFF76FAD5" box="[113,254,1372,1397]" degrees="125" direction="east" minutes="21" orientation="longitude" pageId="1" pageNumber="358" precision="15" seconds="55" value="125.36527">125°21′55″E</geoCoordinate>
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),
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<elevation id="57C72679FFF9793ED469FABCFE12FAD5" box="[277,410,1373,1397]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="1" pageNumber="358" unit="m" value="1524.0">
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<quantity id="1B126CAFFFF9793ED469FABCFEF8FAD5" box="[277,368,1373,1397]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="1" pageNumber="358" unit="m" value="1524.0">1,524 m</quantity>
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asl
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</elevation>
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,
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<collectingCounty id="3534B9C6FFF9793ED4D4FABCFDEAFAD5" box="[424,610,1373,1397]" pageId="1" pageNumber="358">Mindanao Island</collectingCounty>
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||
,
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<collectingCountry id="A4FD81DAFFF9793ED70CFABCFD63FAD5" box="[624,747,1373,1397]" name="Philippines" pageId="1" pageNumber="358">Philippines</collectingCountry>
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, coll.
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<collectorName id="711FA49CFFF9793ED5D5FA9FFE8CFA36" box="[169,260,1406,1430]" pageId="1" pageNumber="358">N. Aspe</collectorName>
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,
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<date id="A854E78AFFF9793ED46DFA9CFD84FA35" box="[273,524,1405,1429]" pageId="1" pageNumber="358" value="2003-12-11" valueMax="2003-12-14" valueMin="2003-12-11">
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||
<collectingDate id="B8101E62FFF9793ED46DFA9CFD84FA35" box="[273,524,1405,1429]" pageId="1" pageNumber="358" value="2003-12-11" valueMax="2003-12-14" valueMin="2003-12-11">11–14 December 2003</collectingDate>
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</date>
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||
.
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||
</materialsCitation>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="94F092C1FFF9793ED50DFA5CFD67FA55" pageId="1" pageNumber="358" type="etymology">
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<paragraph id="DC55C14AFFF9793ED50DFA5CFD67FA55" blockId="1.[112,752,1469,1526]" pageId="1" pageNumber="358">
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<emphasis id="EE9E1D58FFF9793ED50DFA5CFF79FA75" bold="true" box="[113,241,1469,1493]" pageId="1" pageNumber="358">Etymology.</emphasis>
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The species is named after Apong, a nobleman of whom the name of Mt. Apo was said to have derived from.
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</paragraph>
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</subSubSection>
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<subSubSection id="94F092C1FFF9793ED50DF9FCFD63F8B5" pageId="1" pageNumber="358" type="diagnosis">
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<paragraph id="DC55C14AFFF9793ED50DF9FCFD63F8B5" blockId="1.[112,752,1565,1813]" pageId="1" pageNumber="358">
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<emphasis id="EE9E1D58FFF9793ED50DF9FCFF6EF995" bold="true" box="[113,230,1565,1589]" pageId="1" pageNumber="358">Diagnosis.</emphasis>
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Reddish brown worm with body dimension
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<quantity id="1B126CAFFFF9793ED7A9F9FCFF13F9F5" metricMagnitude="-2" metricUnit="m" metricValue="6.1" pageId="1" pageNumber="358" unit="mm" value="61.0">61 mm</quantity>
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×
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<quantity id="1B126CAFFFF9793ED5C3F9DCFEBBF9F5" box="[191,307,1597,1621]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.7" metricValueMin="4.0" pageId="1" pageNumber="358" unit="mm" value="4.35" valueMax="4.7" valueMin="4.0">4–4.7 mm</quantity>
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; one pair of spermathecal pores at 7/8; pairs of genital markings in viii and in ix, those in viii closely positioned to the spermathecal pores; pairs of genital markings in xviii and xix, those in xviii presetal, medial to male pores, those in xix mid-ventral on setal line; male sexual system proandric; spermatheca with a small and round ampulla; single, dense, racemose prostates in xvii to xviii.
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</paragraph>
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</subSubSection>
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<subSubSection id="94F092C1FFF9793ED50DF8DCFB18FB55" pageId="1" pageNumber="358" type="description">
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<paragraph id="DC55C14AFFF9793ED50DF8DCFC30FE15" blockId="1.[112,753,1853,2101]" lastBlockId="1.[812,1454,189,437]" pageId="1" pageNumber="358">
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<emphasis id="EE9E1D58FFF9793ED50DF8DCFE8DF8F5" bold="true" box="[113,261,1853,1877]" pageId="1" pageNumber="358">Description.</emphasis>
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Reddish brown dorsum, lighter ventrum, equators pigmented. Length
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<quantity id="1B126CAFFFF9793ED4D1F8BCFE74F8D5" box="[429,508,1885,1909]" metricMagnitude="-2" metricUnit="m" metricValue="6.1" pageId="1" pageNumber="358" unit="mm" value="61.0">61 mm</quantity>
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(n=
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<specimenCount id="CAEC0AC3FFF9793ED753F8BCFDF4F8D5" box="[559,636,1885,1909]" pageId="1" pageNumber="358" type="adult">1 adult</specimenCount>
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); diameter
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<quantity id="1B126CAFFFF9793ED50DF89CFF27F835" box="[113,175,1917,1941]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="358" unit="mm" value="4.0">4 mm</quantity>
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at x,
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<quantity id="1B126CAFFFF9793ED591F89CFEB7F835" box="[237,319,1917,1941]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="1" pageNumber="358" unit="mm" value="4.7">4.7 mm</quantity>
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at xx; body cylindrical in cross-section, tail tapering; 77 segments. First dorsal pore 12/13. One pair of spermathecal pores at 7/8, distance between spermathecal pores
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<quantity id="1B126CAFFFF9793ED5CEF83CFE8CF855" box="[178,260,2013,2037]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="358" unit="mm" value="1.7">1.7 mm</quantity>
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(0.13 circumference ventrally apart). Female pore single in xiv, male pore openings paired in xviii, distance between pores
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<quantity id="1B126CAFFFF9793ED473F7FCFEE8F795" box="[271,352,2077,2101]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="358" unit="mm" value="1.8">1.8 mm</quantity>
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(0.12 circumference apart ventrally), seven setae between male pores. Clitellum annular, from xiv to xvi. Setae unevenly distributed around equators in some segments; 44 setae on vii, 52 setae on xx, dorsal and ventral setal gaps present. Pairs of genital markings in viii and in ix, those in viii closely positioned to the spermathecal pores; pairs of genital markings in xviii and xix, those in xviii presetal, medial to male pores, those in xix mid-ventral on setal line.
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||
</paragraph>
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<paragraph id="DC55C14AFFF9793ED651FE3CFA24FD75" blockId="1.[812,1453,477,725]" pageId="1" pageNumber="358">Septa 4/5–8/9, 10/11–13/14 thin, 9/10 lacking. Dense tufts of nephridia on anterior faces of 5/6 and 6/7; nephridia of intestinal segments located mainly on body near septum/ body wall junction. Large gizzard in viii–x, esophagus with low vertical lamellae x–xiii, intestinal origin in xv, caeca simple originating in xxvii, extending forward to xxv. Typhlosole originating in xxvii, simple fold, 1/3 lumen diameter. Intestinal wall with 32 longitudinal blood vessels.</paragraph>
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<paragraph id="DC55C14AFFF9793ED651FD1FFB18FB55" blockId="1.[812,1454,765,1269]" pageId="1" pageNumber="358">Hearts in x to xiii, esophageal; commissural vessels in vi, vii, and ix lateral; those in viii extending to gizzard. Ovaries and funnels free in xiii. Spermathecae paired, postseptal in viii, nephridia on ducts lacking; each spermatheca with a small, round ampulla and a long, slender duct, diverticulum attached ectally to the left face of the right spermathecal duct, and to the right face of the left spermathecal duct, terminating in an elongated receptacle, stalk longer than ampulla. Male sexual system proandric, testes and funnels enclosed in paired sacs in x; seminal vesicles in xi, each with a digitate dorsal lobe, enclosed in testis sacs; vasa deferentia slender, free from body wall en route to ental end of prostatic ducts; prostates in xvii to xviii; each prostate a single, dense, racemose mass; stout muscular duct widened towards body wall, then slightly narrowed just before body wall. Copulatory bursae lacking.</paragraph>
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</subSubSection>
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<subSubSection id="94F092C1FFF9793ED651FAFCFBB2F895" pageId="1" pageNumber="358" type="discussion">
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<paragraph id="DC55C14AFFF9793ED651FAFCFBB2F895" blockId="1.[812,1452,1309,1845]" pageId="1" pageNumber="358">
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||
<emphasis id="EE9E1D58FFF9793ED651FAFCFC11FA95" bold="true" box="[813,921,1309,1333]" pageId="1" pageNumber="358">Remarks.</emphasis>
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Currently, there is a total of 17 species of
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<taxonomicName id="1BEABAC9FFF9793ED03FFAFCFA22FA95" baseAuthorityName="Kindberg" baseAuthorityYear="1867" box="[1347,1450,1309,1333]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="358" phylum="Annelida" rank="genus">
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<emphasis id="EE9E1D58FFF9793ED03FFAFCFA22FA95" box="[1347,1450,1309,1333]" italics="true" pageId="1" pageNumber="358">Amynthas</emphasis>
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||
</taxonomicName>
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in the
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<collectingCountry id="A4FD81DAFFF9793ED605FADCFC70FAF5" box="[889,1016,1341,1365]" name="Philippines" pageId="1" pageNumber="358">Philippines</collectingCountry>
|
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, including
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<taxonomicName id="1BEABAC9FFF9793ED106FADCFAE4FAF5" authority="Aspe & James, 2017" authorityName="Aspe & James" authorityYear="2017" box="[1146,1388,1341,1365]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="358" phylum="Annelida" rank="species" species="apongensis" status="sp. nov.">
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||
<emphasis id="EE9E1D58FFF9793ED106FADCFAE4FAF5" box="[1146,1388,1341,1365]" italics="true" pageId="1" pageNumber="358">Amynthas apongensis</emphasis>
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</taxonomicName>
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,
|
||
<taxonomicNameLabel id="F5ADA023FFF9793ED002FADFFCF5FAD5" pageId="1" pageNumber="358" rank="species">new species</taxonomicNameLabel>
|
||
. While
|
||
<taxonomicName id="1BEABAC9FFF9793ED6B0FABCFB01FAD5" box="[972,1161,1373,1397]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="358" phylum="Annelida" rank="species" species="undetermined">
|
||
<emphasis id="EE9E1D58FFF9793ED6B0FABCFBBDFAD5" box="[972,1077,1373,1397]" italics="true" pageId="1" pageNumber="358">Amynthas</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
vary in the number of pairs of spermathecae (one to five pairs except
|
||
<taxonomicName id="1BEABAC9FFF9793ED07FFA9FFA24FA35" baseAuthorityName="Aspe & James" baseAuthorityYear="2016" box="[1283,1452,1405,1429]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="358" phylum="Annelida" rank="species" species="dinagatensis">
|
||
<emphasis id="EE9E1D58FFF9793ED07FFA9FFA24FA35" box="[1283,1452,1405,1429]" italics="true" pageId="1" pageNumber="358">A. dinagatensis</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<bibRefCitation id="B87BBCBBFFF9793ED64FFA7FFB91FA15" author="Aspe NM & James SW" box="[819,1049,1437,1462]" pageId="1" pageNumber="358" pagination="8" refId="ref14648" refString="Aspe NM & James SW (2016) New species of Pheretima, Amynthas, Polypheretima, and Pithemera (Clitellata: Megascolecidae) from Mindanao and associated islands, Philippines. Zoological Studies, 55: 8. doi: 10.6620 / ZS. 2016.55 - 08." type="journal article" year="2016">Aspe & James, 2016</bibRefCitation>
|
||
], which is athecate),
|
||
<taxonomicName id="1BEABAC9FFF9793ED181FA7FFA1BFA15" authorityName="Aspe & James" authorityYear="2017" box="[1277,1427,1438,1461]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="358" phylum="Annelida" rank="species" species="apongensis">
|
||
<emphasis id="EE9E1D58FFF9793ED181FA7FFA1BFA15" box="[1277,1427,1438,1461]" italics="true" pageId="1" pageNumber="358">A. apongensis</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="1BEABAC9FFF9793ED6EEFA5FFABCFA75" authority="(Hong & James, 2004)" baseAuthorityName="Hong & James" baseAuthorityYear="2004" box="[914,1332,1469,1493]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="358" phylum="Annelida" rank="species" species="philippinensis">
|
||
<emphasis id="EE9E1D58FFF9793ED6EEFA5FFBC9FA75" box="[914,1089,1469,1493]" italics="true" pageId="1" pageNumber="358">A. philippinensis</emphasis>
|
||
(
|
||
<bibRefCitation id="B87BBCBBFFF9793ED133FA5CFAA5FA75" author="James SW & Hong Y & Kim TH" box="[1103,1325,1469,1493]" pageId="1" pageNumber="358" pagination="3 - 10" refId="ref15441" refString="James SW, Hong Y & Kim TH (2004) New earthworms of Pheretima and Pithemera (Oligochaeta: Megascolecidae) from Mt. Arayat, Luzon Island, Philippines. Revue Suisse de Zoologie, 111: 3 - 10." type="journal article" year="2004">Hong & James, 2004</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="1BEABAC9FFF9793ED040FA5FFC4AFA55" authority="(James, 2004)" baseAuthorityName="James" baseAuthorityYear="2004" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="358" phylum="Annelida" rank="species" species="heaneyi">
|
||
<emphasis id="EE9E1D58FFF9793ED040FA5FFA22FA75" box="[1340,1450,1469,1493]" italics="true" pageId="1" pageNumber="358">A. heaneyi</emphasis>
|
||
(James, 2004)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="1BEABAC9FFF9793ED688FA3FFA23FA55" authority="(Aspe & James, 2016)" baseAuthorityName="Aspe & James" baseAuthorityYear="2016" box="[1012,1451,1501,1525]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="358" phylum="Annelida" rank="species" species="cagdianaoensis">
|
||
<emphasis id="EE9E1D58FFF9793ED688FA3FFB30FA55" box="[1012,1208,1501,1525]" italics="true" pageId="1" pageNumber="358">A. cagdianaoensis</emphasis>
|
||
(
|
||
<bibRefCitation id="B87BBCBBFFF9793ED1B9FA3CFA2CFA55" author="Aspe NM & James SW" box="[1221,1444,1501,1525]" pageId="1" pageNumber="358" pagination="8" refId="ref14648" refString="Aspe NM & James SW (2016) New species of Pheretima, Amynthas, Polypheretima, and Pithemera (Clitellata: Megascolecidae) from Mindanao and associated islands, Philippines. Zoological Studies, 55: 8. doi: 10.6620 / ZS. 2016.55 - 08." type="journal article" year="2016">Aspe & James, 2016</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
in having only one pair. However,
|
||
<taxonomicName id="1BEABAC9FFF9793ED1D7FA1FFAE8F9B5" baseAuthorityName="Hong & James" baseAuthorityYear="2004" box="[1195,1376,1533,1557]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="358" phylum="Annelida" rank="species" species="philippinensis">
|
||
<emphasis id="EE9E1D58FFF9793ED1D7FA1FFAE8F9B5" box="[1195,1376,1533,1557]" italics="true" pageId="1" pageNumber="358">A. philippinensis</emphasis>
|
||
</taxonomicName>
|
||
has its pair of spermathecal pores at 4/5 while
|
||
<taxonomicName id="1BEABAC9FFF9793ED199F9FFFA23F995" baseAuthorityName="Aspe & James" baseAuthorityYear="2016" box="[1253,1451,1565,1589]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="358" phylum="Annelida" rank="species" species="cagdianaoensis">
|
||
<emphasis id="EE9E1D58FFF9793ED199F9FFFA23F995" box="[1253,1451,1565,1589]" italics="true" pageId="1" pageNumber="358">A. cagdianaoensis</emphasis>
|
||
</taxonomicName>
|
||
has its pair at 8/9.
|
||
<taxonomicName id="1BEABAC9FFF9793ED174F9DCFB71F9F5" authorityName="Aspe & James" authorityYear="2017" box="[1032,1273,1597,1621]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="358" phylum="Annelida" rank="species" species="apongensis">
|
||
<emphasis id="EE9E1D58FFF9793ED174F9DCFB71F9F5" box="[1032,1273,1597,1621]" italics="true" pageId="1" pageNumber="358">Amynthas apongensis</emphasis>
|
||
</taxonomicName>
|
||
is most similar to
|
||
<taxonomicName id="1BEABAC9FFF9793ED637F9BFFC36F9D5" baseAuthorityName="James" baseAuthorityYear="2004" box="[843,958,1629,1653]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="358" phylum="Annelida" rank="species" species="heaneyi">
|
||
<emphasis id="EE9E1D58FFF9793ED637F9BFFC36F9D5" box="[843,958,1629,1653]" italics="true" pageId="1" pageNumber="358">A. heaneyi</emphasis>
|
||
</taxonomicName>
|
||
in the location of the spermathecal pores, in the shape of the spermathecae and in having a proandric male sexual system. However,
|
||
<taxonomicName id="1BEABAC9FFF9793ED100F97FFB65F915" baseAuthorityName="James" baseAuthorityYear="2004" box="[1148,1261,1693,1717]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="358" phylum="Annelida" rank="species" species="heaneyi">
|
||
<emphasis id="EE9E1D58FFF9793ED100F97FFB65F915" box="[1148,1261,1693,1717]" italics="true" pageId="1" pageNumber="358">A. heaneyi</emphasis>
|
||
</taxonomicName>
|
||
is larger (
|
||
<quantity id="1B126CAFFFF9793ED026F97CFCDEF975" metricMagnitude="-2" metricUnit="m" metricValue="9.75" metricValueMax="11.1" metricValueMin="8.4" pageId="1" pageNumber="358" unit="mm" value="97.5" valueMax="111.0" valueMin="84.0">84–111 mm</quantity>
|
||
×
|
||
<quantity id="1B126CAFFFF9793ED606F95CFB89F975" box="[890,1025,1725,1749]" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="5.7" metricValueMin="4.6" pageId="1" pageNumber="358" unit="mm" value="5.15" valueMax="5.7" valueMin="4.6">4.6–5.7 mm</quantity>
|
||
), has dark gray to black pigmentation, has wider distance between the male pores (0.14–0.16 circumference apart), and has a different pattern of genital markings (James, 2004).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |