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<document id="1D318F39F5DE4CF9C3828D02B112418C" ID-CLB-Dataset="133200" ID-DOI="10.1649/0010-065X-73.3.629" ID-GBIF-Dataset="99bf6930-d483-4789-886e-2aaf5b44d9b1" ID-ISSN="1938-4394" ID-Zenodo-Dep="5402311" ID-ZooBank="urn:lsid:zoobank.org:pub:48CFB8E9-AA28-413D-98CB-1841B65ED236" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630336494997" checkinUser="valdenar" docAuthor="Will, Kipling" docDate="2019" docId="543F8F0FFFEDF8561A9D1941FB4AF5C5" docLanguage="en" docName="TheColeopBull.73.3.629-645.pdf" docOrigin="The Coleopterists Bulletin 73 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-73.3.629" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Pediomorphus CHAUDOIR 1878" docType="treatment" docVersion="7" lastPageNumber="644" masterDocId="A806F777FFEFF84618611B1BFFD9F64C" masterDocTitle="Taxonomic Review of the Australian Genus Pediomorphus Chaudoir, 1878 (Coleoptera: Carabidae: Abacetini)" masterLastPageNumber="645" masterPageNumber="629" pageNumber="630" updateTime="1699242009079" updateUser="ExternalLinkService">
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<mods:title id="7B3B16B8B0E2134A11ECB33AAF590963">Taxonomic Review of the Australian Genus Pediomorphus Chaudoir, 1878 (Coleoptera: Carabidae: Abacetini)</mods:title>
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<mods:namePart id="5FAA782EA32F5AC32DC0DAB79F5FE99A">Will, Kipling</mods:namePart>
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<subSubSection id="948C6D92FFEDF8441A9D1941FB9CF43C" box="[764,1093,602,624]" pageId="2" pageNumber="630" type="nomenclature">
<paragraph id="DC293E19FFEDF8441A9D1941FB9CF43C" blockId="2.[762,1095,550,624]" box="[764,1093,602,624]" pageId="2" pageNumber="630">
<taxonomicName id="1B96459AFFEDF8441A9D1941FB9CF43C" ID-CoL="8KYBR" authority="Chaudoir 1878: 28" authorityName="CHAUDOIR" authorityPageNumber="28" authorityYear="1878" box="[764,1093,602,624]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="630" phylum="Arthropoda" rank="genus">
<emphasis id="EEE2E20BFFEDF8441A9D1941FC5EF43C" box="[764,903,602,624]" italics="true" pageId="2" pageNumber="630">Pediomorphus</emphasis>
<bibRefCitation id="B80743E8FFEDF8441BEE1940FB9CF43C" author="Chaudoir, M. d." box="[911,1093,603,624]" firstAuthor="Chaudoir" journalOrPublisher="Bulletin de la Societe imperiale des naturalistes de Moscou" page="28" pageId="2" pageNumber="630" pagination="1 - 80" part="53" refId="ref10205" refString="Chaudoir, M. d. 1878. Descriptions de genres nouveaux et d' especes inedites de la famille des carabiques. Bulletin de la Societe imperiale des naturalistes de Moscou 53: 1 - 80." title="Descriptions de genres nouveaux et d' especes inedites de la famille des carabiques" type="journal article" year="1878">Chaudoir 1878: 28</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="948C6D92FFEDF8441AA4199EFC40F4F9" pageId="2" pageNumber="630" type="type_taxon">
<paragraph id="DC293E19FFEDF8441AA4199EFC40F4F9" blockId="2.[685,1172,645,879]" pageId="2" pageNumber="630">
<emphasis id="EEE2E20BFFEDF8441AA4199EFC93F4D6" bold="true" box="[709,842,645,666]" pageId="2" pageNumber="630">
<typeStatus id="032D80BBFFEDF8441AA4199EFD2CF4D6" box="[709,757,645,666]" pageId="2" pageNumber="630">Type</typeStatus>
Species.
</emphasis>
<taxonomicName id="1B96459AFFEDF8441B32199EFCD7F4F9" ID-CoL="75Y5Q" authority="Chaudoir, 1878" authorityName="Chaudoir" authorityYear="1878" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="630" phylum="Arthropoda" rank="species" species="planiusculus">
<emphasis id="EEE2E20BFFEDF8441B32199EFB8AF4D7" box="[851,1107,645,667]" italics="true" pageId="2" pageNumber="630">Pediomorphus planiusculus</emphasis>
<bibRefCitation id="B80743E8FFEDF8441C3D199EFCD7F4F9" author="Chaudoir, M. d." firstAuthor="Chaudoir" journalOrPublisher="Bulletin de la Societe imperiale des naturalistes de Moscou" pageId="2" pageNumber="630" pagination="1 - 80" part="53" refId="ref10205" refString="Chaudoir, M. d. 1878. Descriptions de genres nouveaux et d' especes inedites de la famille des carabiques. Bulletin de la Societe imperiale des naturalistes de Moscou 53: 1 - 80." title="Descriptions de genres nouveaux et d' especes inedites de la famille des carabiques" type="journal article" year="1878">Chaudoir, 1878</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="948C6D92FFEDF8441AA419A1FC67F576" pageId="2" pageNumber="630" type="reference_group">
<paragraph id="DC293E19FFEDF8441AA419A1FC67F576" blockId="2.[685,1172,645,879]" pageId="2" pageNumber="630">
<emphasis id="EEE2E20BFFEDF8441AA419A1FC70F483" bold="true" box="[709,937,698,719]" pageId="2" pageNumber="630">References to Genus.</emphasis>
<bibRefCitation id="B80743E8FFEDF8441BD619A1FB98F483" author="Masters, G." box="[951,1089,698,719]" firstAuthor="Masters" journalOrPublisher="Proceedings of the Linnean Society of New South Wales" pageId="2" pageNumber="630" pagination="359 - 444" part="10" refId="ref10424" refString="Masters, G. 1885. Catalogue of the described Coleoptera of Australia. Part I. Proceedings of the Linnean Society of New South Wales 10: 359 - 444." title="Catalogue of the described Coleoptera of Australia. Part I" type="journal article" year="1885">Masters 1885</bibRefCitation>
; 
<bibRefCitation id="B80743E8FFEDF8441C3219A1FD07F4A6" author="Sloane, T. G." firstAuthor="Sloane" journalOrPublisher="Proceedings of the Linnean Society of New South Wales" pageId="2" pageNumber="630" pagination="393 - 455" part="9" refId="ref10695" refString="Sloane, T. G. 1895. Studies in Australian entomology. No. VII. New genera and species of Carabidae (including some notes on previously described species, and synoptic lists of genera and species). Proceedings of the Linnean Society of New South Wales 9: 393 - 455." title="Studies in Australian entomology. No. VII. New genera and species of Carabidae (including some notes on previously described species, and synoptic lists of genera and species)" type="journal article" year="1895">Sloane 1895</bibRefCitation>
, 
<bibRefCitation id="B80743E8FFEDF8441A8819CEFCC3F4A6" author="Sloane, T. G." box="[745,794,725,746]" firstAuthor="Sloane" journalOrPublisher="Proceedings of the Linnean Society of New South Wales" pageId="2" pageNumber="630" pagination="444 - 520" part="23" refId="ref10751" refString="Sloane, T. G. 1898. On Carabidae from west Australia, sent by Mr. A. M. Lea (with descriptions of new genera and species, synoptic tables, &amp; c.). Proceedings of the Linnean Society of New South Wales 23: 444 - 520." title="On Carabidae from west Australia, sent by Mr. A. M. Lea (with descriptions of new genera and species, synoptic tables, &amp; c.)" type="journal article" year="1898">1898</bibRefCitation>
, 
<bibRefCitation id="B80743E8FFEDF8441B4719CEFC8EF4A6" author="Sloane, T. G." box="[806,855,725,746]" firstAuthor="Sloane" journalOrPublisher="Proceedings of the Linnean Society of New South Wales" pageId="2" pageNumber="630" pagination="553 - 584" part="24" refId="ref10806" refString="Sloane, T. G. 1900. Studies in Australian Entomology. No. IX. New Speceis of Carabidae (with notes on some previously described species, and synoptic lists of species). Proceedings of the Linnean Society of New South Wales 24: 553 - 584." title="Studies in Australian Entomology. No. IX. New Speceis of Carabidae (with notes on some previously described species, and synoptic lists of species)" type="journal article" year="1900">1900</bibRefCitation>
, 
<bibRefCitation id="B80743E8FFEDF8441B0319CEFC4AF4A6" author="Sloane, T. G." box="[866,915,725,746]" firstAuthor="Sloane" journalOrPublisher="Proceedings of the Linnean Society of New South Wales" pageId="2" pageNumber="630" pagination="568 - 614" part="39" refId="ref10858" refString="Sloane, T. G. 1915. Revisional notes on Australian Carabidae. Part V. Proceedings of the Linnean Society of New South Wales 39: 568 - 614." title="Revisional notes on Australian Carabidae. Part V" type="journal article" year="1915">1915</bibRefCitation>
; 
<bibRefCitation id="B80743E8FFEDF8441BFF19CEFB85F4A6" author="Tschitscherine, T." box="[926,1116,725,746]" firstAuthor="Tschitscherine" journalOrPublisher="Horae Societatis Entomologicae Rossicae" pageId="2" pageNumber="630" pagination="502 - 534" part="35" refId="ref10915" refString="Tschitscherine, T. 1902. Notes sur les Platysmatini de l' Australie. Horae Societatis Entomologicae Rossicae 35: 502 - 534." title="Notes sur les Platysmatini de l' Australie" type="journal article" year="1902">Tschitscherine 1902</bibRefCitation>
; 
<bibRefCitation id="B80743E8FFEDF8441C0619CEFCD3F549" author="Straneo, S. L." firstAuthor="Straneo" pageId="2" pageNumber="630" pagination="416 - 428" part="11" refId="ref10890" refString="Straneo, S. L. 1960. Elenco di Carabidi entrati recentemente nelle Collezioni del Museo Frey. 11: 416 - 428." title="Elenco di Carabidi entrati recentemente nelle Collezioni del Museo Frey" type="journal article" year="1960">Straneo 1960</bibRefCitation>
; 
<bibRefCitation id="B80743E8FFEDF8441B7619EBFC49F549" author="Moore, B. P." box="[791,912,752,773]" firstAuthor="Moore" journalOrPublisher="Transactions of the Royal Entomological Society of London" pageId="2" pageNumber="630" pagination="1 - 32" part="117" refId="ref10510" refString="Moore, B. P. 1965. Studies on Australian Carabidae (Coleoptera) 4. - The Pterostichinae. Transactions of the Royal Entomological Society of London 117: 1 - 32." title="Studies on Australian Carabidae (Coleoptera) 4. - The Pterostichinae" type="journal article" year="1965">Moore 1965</bibRefCitation>
, 
<bibRefCitation id="B80743E8FFEDF8441BFC19EBFC17F549" author="Moore, B. P." box="[925,974,752,773]" firstAuthor="Moore" journalOrPublisher="Entomologische Arbeiten aus dem Museum G. Frey, Tutzing bei Munchen" pageId="2" pageNumber="630" pagination="308 - 322" part="18" refId="ref10545" refString="Moore, B. P. 1967. New or little known Australian Carabidae in the Frey Museum (Col.). Entomologische Arbeiten aus dem Museum G. Frey, Tutzing bei Munchen 18: 308 - 322." title="New or little known Australian Carabidae in the Frey Museum (Col.)" type="journal article" year="1967">1967</bibRefCitation>
; 
<bibRefCitation id="B80743E8FFEDF8441BBD19EBFB56F549" author="Moore, B. P. &amp; T. A. Weir &amp; J. E. Pyke" box="[988,1167,751,773]" editor="D. W. Walton" etAl="et al." firstAuthor="Moore" journalOrPublisher="Australian Government Publishing Service, Canberra, Australia" pageId="2" pageNumber="630" pagination="20 - 230" refId="ref10586" refString="Moore, B. P., T. A. Weir, and J. E. Pyke. 1987. Rhysodidae and Carabidae [pp. 20 - 230]. In: Zoological Catalogue of Australia, vol. 4 (Coleoptera: Archostemata, Myxophaga and Adephaga) (D. W. Walton, editor). Australian Government Publishing Service, Canberra, Australia." title="Rhysodidae and Carabidae" type="book chapter" volumeTitle="Zoological Catalogue of Australia, vol. 4 (Coleoptera: Archostemata, Myxophaga and Adephaga)" year="1987">
Moore 
<emphasis id="EEE2E20BFFEDF8441C4219EBFB8FF549" box="[1059,1110,751,773]" italics="true" pageId="2" pageNumber="630">et al.</emphasis>
1987
</bibRefCitation>
; 
<bibRefCitation id="B80743E8FFEDF8441ACC1811FC56F553" author="Liebherr, J. K. &amp; K. W. Will" box="[685,911,778,799]" editor="G. E. Ball &amp; A. Casale &amp; A. V. Taglianti" firstAuthor="Liebherr" journalOrPublisher="XX International Congress of Entomology. Atti Museo Regionale di Scienze Naturali di Torino, Italy" pageId="2" pageNumber="630" pagination="107 - 170" refId="ref10242" refString="Liebherr, J. K., and K. W. Will. 1998. Inferring phylogenetic relationships within Carabidae (Insecta, Coleoptera) from characters of the female reproductive tract [pp. 107 - 170]. In: Proceedings of a Symposium (28 August 1996, Florence, Italy) (G. E. Ball, A. Casale, and A. V. Taglianti, editors), XX International Congress of Entomology. Atti Museo Regionale di Scienze Naturali di Torino, Italy." title="Inferring phylogenetic relationships within Carabidae (Insecta, Coleoptera) from characters of the female reproductive tract" type="proceedings paper" volumeTitle="Proceedings of a Symposium" year="1998">Liebherr and Will 1998</bibRefCitation>
; 
<bibRefCitation id="B80743E8FFEDF8441BF81811FBC4F553" author="Lorenz, W." box="[921,1053,778,799]" firstAuthor="Lorenz" journalOrPublisher="Privately published, Tutzing, Germany" pageId="2" pageNumber="630" refId="ref10336" refString="Lorenz, W. 2005 a. A Systematic List of Extant Ground Beetles of the World (Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae, incl. Paussinae, Cicindelinae, Rhysodinae). Privately published, Tutzing, Germany." title="A Systematic List of Extant Ground Beetles of the World (Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae, incl. Paussinae, Cicindelinae, Rhysodinae)" type="book" year="2005" yearSuffix="a">Lorenz 2005a</bibRefCitation>
, b; 
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; 
<bibRefCitation id="B80743E8FFEDF8441B7D183EFCA4F576" author="Will, K. W." box="[796,893,805,826]" firstAuthor="Will" journalOrPublisher="The Coleopterists Bulletin" pageId="2" pageNumber="630" pagination="91" part="64" refId="ref10939" refString="Will, K. W. 2010. Synonymy in Sugimotoa Habu (Coleoptera: Carabidae: Lebiini). The Coleopterists Bulletin 64: 91." title="Synonymy in Sugimotoa Habu (Coleoptera: Carabidae: Lebiini)" type="journal article" year="2010">Will 2010</bibRefCitation>
, 
<bibRefCitation id="B80743E8FFEDF8441BE9183EFC63F576" author="Will, K. W." box="[904,954,805,826]" firstAuthor="Will" journalOrPublisher="ZooKeys" pageId="2" pageNumber="630" pagination="131 - 137" part="545" refId="ref10966" refString="Will, K. W. 2015. Resolution of taxonomic problems in Australian Harpalini, Abacetini, Pterostichini, and Oodini (Coleoptera, Carabidae). ZooKeys 545: 131 - 137." title="Resolution of taxonomic problems in Australian Harpalini, Abacetini, Pterostichini, and Oodini (Coleoptera, Carabidae)" type="journal article" year="2015">2015</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="948C6D92FFEDF8451AA41824FE91F70D" lastPageId="3" lastPageNumber="631" pageId="2" pageNumber="630" type="diagnosis">
<paragraph id="DC293E19FFEDF8451AA41824FE91F70D" blockId="2.[685,1172,645,879]" lastBlockId="3.[173,659,194,932]" lastPageId="3" lastPageNumber="631" pageId="2" pageNumber="630">
<emphasis id="EEE2E20BFFEDF8441AA41824FC5FF518" bold="true" box="[709,902,831,852]" pageId="2" pageNumber="630">Generic Diagnosis.</emphasis>
Typical Loxandrina recognizable by the combination of the enlarged apical labial palpomeres with a setose, ventral sensorium (
<figureCitation id="44AD229CFFECF84518D51BC7FF23F6BD" box="[180,250,220,241]" captionStart="Figs" captionStartId="12.[175,214,1228,1247]" captionTargetBox="[156,1167,196,1210]" captionTargetId="figure-183@12.[154,1169,194,1212]" captionTargetPageId="12" captionText="Figs. 14–18. Pediomorphus species. P. planiusculus: 14) Labial palp; 15) Base of right elytron. P. crenulatus: 16) Base of right elytron. P. macleayi: 17) Head, showing large antennomere 1. P. robustus: 18) Ventral view of head, arrow indicating subocular carina." figureDoi="http://doi.org/10.5281/zenodo.5402321" httpUri="https://zenodo.org/record/5402321/files/figure.png" pageId="3" pageNumber="631">Fig. 14</figureCitation>
), subocular carina (
<figureCitation id="44AD229CFFECF84519D21BC7FE20F6BD" box="[435,505,220,241]" captionStart="Figs" captionStartId="12.[175,214,1228,1247]" captionTargetBox="[156,1167,196,1210]" captionTargetId="figure-183@12.[154,1169,194,1212]" captionTargetPageId="12" captionText="Figs. 14–18. Pediomorphus species. P. planiusculus: 14) Labial palp; 15) Base of right elytron. P. crenulatus: 16) Base of right elytron. P. macleayi: 17) Head, showing large antennomere 1. P. robustus: 18) Ventral view of head, arrow indicating subocular carina." figureDoi="http://doi.org/10.5281/zenodo.5402321" httpUri="https://zenodo.org/record/5402321/files/figure.png" pageId="3" pageNumber="631">Fig. 18</figureCitation>
), presence of an elytral plica, and lack of elytral discal setae. The characteristic labial palps are not known from any other Abacetini.
</paragraph>
</subSubSection>
<caption id="88E96E91FFEDF84418CE1DB4FB9AF08E" ID-DOI="http://doi.org/10.5281/zenodo.5402313" ID-Zenodo-Dep="5402313" box="[175,1091,1710,1730]" httpUri="https://zenodo.org/record/5402313/files/figure.png" pageId="2" pageNumber="630" startId="2.[175,214,1711,1730]" targetBox="[156,1167,936,1693]" targetPageId="2">
<paragraph id="DC293E19FFEDF84418CE1DB4FB9AF08E" blockId="2.[175,1091,1710,1730]" box="[175,1091,1710,1730]" pageId="2" pageNumber="630">
<emphasis id="EEE2E20BFFEDF84418CE1DB4FEDFF08E" bold="true" box="[175,262,1710,1730]" pageId="2" pageNumber="630">Figs. 1–2.</emphasis>
<taxonomicName id="1B96459AFFEDF844197A1DB5FE05F08E" box="[283,476,1710,1730]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="630" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="EEE2E20BFFEDF844197A1DB5FE4FF08D" box="[283,406,1710,1729]" italics="true" pageId="2" pageNumber="630">Pediomorphus</emphasis>
species
</taxonomicName>
, dorsal habitus. 
<emphasis id="EEE2E20BFFEDF8441A091DB4FDA3F08E" bold="true" box="[616,634,1711,1730]" pageId="2" pageNumber="630">1)</emphasis>
<taxonomicName id="1B96459AFFEDF8441AE11DB4FCD8F08D" authorityName="Chaudoir" authorityYear="1878" box="[640,769,1710,1729]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="630" phylum="Arthropoda" rank="species" species="planiusculus">
<emphasis id="EEE2E20BFFEDF8441AE11DB4FCD8F08D" box="[640,769,1710,1729]" italics="true" pageId="2" pageNumber="630">P. planiusculus</emphasis>
</taxonomicName>
; 
<emphasis id="EEE2E20BFFEDF8441B6F1DB4FCC6F08E" bold="true" box="[782,799,1711,1730]" pageId="2" pageNumber="630">2)</emphasis>
<taxonomicName id="1B96459AFFEDF8441B471DB4FC57F08D" authorityName="Sloane" authorityYear="1898" box="[806,910,1710,1729]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="630" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="EEE2E20BFFEDF8441B471DB4FC57F08D" box="[806,910,1710,1729]" italics="true" pageId="2" pageNumber="630">P. elongatus</emphasis>
</taxonomicName>
. Scale bars = 1 mm.
</paragraph>
</caption>
<subSubSection id="948C6D92FFECF84518A41A5DFDEBF5E8" pageId="3" pageNumber="631" type="description">
<paragraph id="DC293E19FFECF84518A41A5DFDEBF5E8" blockId="3.[173,659,194,932]" pageId="3" pageNumber="631">
<emphasis id="EEE2E20BFFECF84518A41A5DFE76F717" bold="true" box="[197,431,326,347]" pageId="3" pageNumber="631">Genus Characteristics.</emphasis>
Modified from 
<bibRefCitation id="B80743E8FFECF8451A351A5DFF37F73A" author="Moore, B. P." firstAuthor="Moore" journalOrPublisher="Transactions of the Royal Entomological Society of London" pageId="3" pageNumber="631" pagination="1 - 32" part="117" refId="ref10510" refString="Moore, B. P. 1965. Studies on Australian Carabidae (Coleoptera) 4. - The Pterostichinae. Transactions of the Royal Entomological Society of London 117: 1 - 32." title="Studies on Australian Carabidae (Coleoptera) 4. - The Pterostichinae" type="journal article" year="1965">Moore (1965)</bibRefCitation>
. Small to medium size beetles (
<quantity id="1B6E93FCFFECF8451A721A7AFD50F73A" box="[531,649,353,375]" metricMagnitude="-3" metricUnit="m" metricValue="5.85" metricValueMax="8.2" metricValueMin="3.5" pageId="3" pageNumber="631" unit="mm" value="5.85" valueMax="8.2" valueMin="3.5">3.5–8.2 mm</quantity>
); pale brown, castaneous, or rarely darker, nearly piceous; parallel-sided, somewhat or extremely dorso-ventrally depressed body form; relatively large pronotum (
<figureCitation id="44AD229CFFECF84519391AD0FE1FF7AC" box="[344,454,459,481]" captionStart-0="Figs" captionStart-1="Figs" captionStart-2="Figs" captionStart-3="Figs" captionStartId-0="2.[175,214,1711,1730]" captionStartId-1="4.[175,214,1681,1700]" captionStartId-2="7.[197,236,1640,1659]" captionStartId-3="11.[197,236,1681,1700]" captionTargetBox-0="[156,1167,936,1693]" captionTargetBox-1="[235,1088,202,1660]" captionTargetBox-2="[177,1183,207,1622]" captionTargetBox-3="[240,1122,207,1664]" captionTargetId-0="figure-367@2.[154,1169,934,1695]" captionTargetId-1="figure-0@4.[233,1090,194,1665]" captionTargetId-2="figure-0@7.[175,1190,194,1624]" captionTargetId-3="figure-0@11.[238,1129,194,1665]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="7" captionTargetPageId-3="11" captionText-0="Figs. 1–2. Pediomorphus species, dorsal habitus. 1) P. planiusculus; 2) P. elongatus. Scale bars = 1 mm." captionText-1="Figs. 3–6. Pediomorphus species, dorsal habitus. 3) P. macleayi; 4) P. ruficollis; 5) P. variabilis; 6) P. maximus, holotype. Scale bars = 1 mm." captionText-2="Figs. 7–10. Pediomorphus species, dorsal habitus. 7) P. crenulatus, holotype; 8) P. minor, holotype; 9) P. semilaevis, paratype; 10) P. punctatus, paratype. Scale bars = 1 mm." captionText-3="Figs. 11–13. Pediomorphus species, dorsal habitus. 11) P. robustus, paratype; 12) P. storeyi, holotype; 13) P. obtusus, paratype. Scale bars = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5402313" figureDoi-1="http://doi.org/10.5281/zenodo.5402315" figureDoi-2="http://doi.org/10.5281/zenodo.5402317" figureDoi-3="http://doi.org/10.5281/zenodo.5402319" httpUri-0="https://zenodo.org/record/5402313/files/figure.png" httpUri-1="https://zenodo.org/record/5402315/files/figure.png" httpUri-2="https://zenodo.org/record/5402317/files/figure.png" httpUri-3="https://zenodo.org/record/5402319/files/figure.png" pageId="3" pageNumber="631">Figs. 1–13</figureCitation>
); apical segment of labial palpi enlarged, subglobose, plurisetose, with a setose, ventral sensorium (
<figureCitation id="44AD229CFFECF84519D8191BFDDCF459" box="[441,517,512,533]" captionStart="Figs" captionStartId="12.[175,214,1228,1247]" captionTargetBox="[156,1167,196,1210]" captionTargetId="figure-183@12.[154,1169,194,1212]" captionTargetPageId="12" captionText="Figs. 14–18. Pediomorphus species. P. planiusculus: 14) Labial palp; 15) Base of right elytron. P. crenulatus: 16) Base of right elytron. P. macleayi: 17) Head, showing large antennomere 1. P. robustus: 18) Ventral view of head, arrow indicating subocular carina." figureDoi="http://doi.org/10.5281/zenodo.5402321" httpUri="https://zenodo.org/record/5402321/files/figure.png" pageId="3" pageNumber="631">Fig. 14</figureCitation>
); mentum triangular, shallowly emarginate; epilobes long and narrow, not prominent; median tooth prominent and entire or slightly emarginate, not reaching tips of lobes, paramedial pits absent; paraglossae short, glabrous; antennae long, filiform, with 3 basal segments glabrous; postocular orbits not pronounced; elytra free, fully striate, border entire across base, epipleural plica present, but often very small, parascutellar stria present, joined to stria 1, angular base of stria 1 absent, no discal punctures, 13–15 umbilicate setae in stria 8, humeri rounded; hind wings full size; male anterior tarsi with 3 basal segments dilated and squamose beneath; aedeagus (
<figureCitation id="44AD229CFFECF84518D5186FFEF3F5C5" box="[180,298,884,906]" captionStart="Figs" captionStartId="15.[197,236,1579,1598]" captionTargetBox="[185,645,195,1561]" captionTargetId="figure-359@15.[184,647,194,1563]" captionTargetPageId="15" captionText="Figs. 21–32. Pediomorphus species, aedeagi, left lateral view (A) and ventral view of tip (B). 21) P. planiusculus; 22) P. elongatus; 23) P. macleayi; 24) P. ruficollis; 25) P. variabilis; 26) P. crenulatus; 27) P. minor; 28) P. semilaevis; 29) P. punctatus; 30) P. robustus; 31) P. storeyi; 32) P. obtusus." figureDoi="http://doi.org/10.5281/zenodo.5402327" httpUri="https://zenodo.org/record/5402327/files/figure.png" pageId="3" pageNumber="631">Figs. 21–32</figureCitation>
) with orifice on dorsum; parameres conchoid, the right smaller than the left.
</paragraph>
</subSubSection>
<caption id="88E96E91FFEBF84218CE1D8AFE4EF0F0" ID-DOI="http://doi.org/10.5281/zenodo.5402315" ID-Zenodo-Dep="5402315" httpUri="https://zenodo.org/record/5402315/files/figure.png" pageId="4" pageNumber="632" startId="4.[175,214,1681,1700]" targetBox="[235,1088,202,1660]" targetPageId="4">
<paragraph id="DC293E19FFEBF84218CE1D8AFE4EF0F0" blockId="4.[151,1171,1680,1724]" pageId="4" pageNumber="632">
<emphasis id="EEE2E20BFFEBF84218CE1D8AFEDEF0E8" bold="true" box="[175,263,1681,1700]" pageId="4" pageNumber="632">Figs. 3–6.</emphasis>
<taxonomicName id="1B96459AFFEBF842197D1D8BFE07F0E8" box="[284,478,1680,1700]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="632" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="EEE2E20BFFEBF842197D1D8BFE4EF0EF" box="[284,407,1680,1699]" italics="true" pageId="4" pageNumber="632">Pediomorphus</emphasis>
species
</taxonomicName>
, dorsal habitus. 
<emphasis id="EEE2E20BFFEBF8421A0F1D8AFD59F0E8" bold="true" box="[622,640,1681,1700]" pageId="4" pageNumber="632">3)</emphasis>
<taxonomicName id="1B96459AFFEBF8421AE91D8AFD35F0EF" authorityName="Sloane" authorityYear="1900" box="[648,748,1680,1699]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="632" phylum="Arthropoda" rank="species" species="macleayi">
<emphasis id="EEE2E20BFFEBF8421AE91D8AFD35F0EF" box="[648,748,1680,1699]" italics="true" pageId="4" pageNumber="632">P. macleayi</emphasis>
</taxonomicName>
; 
<emphasis id="EEE2E20BFFEBF8421A981D8AFCD3F0E8" bold="true" box="[761,778,1681,1700]" pageId="4" pageNumber="632">4)</emphasis>
<taxonomicName id="1B96459AFFEBF8421B731D8AFCACF0EF" authorityName="Sloane" authorityYear="1900" box="[786,885,1680,1699]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="632" phylum="Arthropoda" rank="species" species="ruficollis">
<emphasis id="EEE2E20BFFEBF8421B731D8AFCACF0EF" box="[786,885,1680,1699]" italics="true" pageId="4" pageNumber="632">P. ruficollis</emphasis>
</taxonomicName>
; 
<emphasis id="EEE2E20BFFEBF8421BE21D8AFC4DF0E8" bold="true" box="[899,916,1681,1700]" pageId="4" pageNumber="632">5)</emphasis>
<taxonomicName id="1B96459AFFEBF8421BFC1D8AFBDDF0EF" baseAuthorityName="Straneo" baseAuthorityYear="1960" box="[925,1028,1680,1699]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="632" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="EEE2E20BFFEBF8421BFC1D8AFBDDF0EF" box="[925,1028,1680,1699]" italics="true" pageId="4" pageNumber="632">P. variabilis</emphasis>
</taxonomicName>
; 
<emphasis id="EEE2E20BFFEBF8421C731D8AFBFAF0E8" bold="true" box="[1042,1059,1681,1700]" pageId="4" pageNumber="632">6)</emphasis>
<taxonomicName id="1B96459AFFEBF8421C4D1D8AFB57F0EF" authorityName="Will" authorityYear="2019" box="[1068,1166,1681,1699]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="632" phylum="Arthropoda" rank="species" species="maximus">
<emphasis id="EEE2E20BFFEBF8421C4D1D8AFB57F0EF" box="[1068,1166,1681,1699]" italics="true" pageId="4" pageNumber="632">P. maximus</emphasis>
</taxonomicName>
, holotype. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="88E96E91FFE8F84118A41D73FD20F0DE" ID-DOI="http://doi.org/10.5281/zenodo.5402317" ID-Zenodo-Dep="5402317" httpUri="https://zenodo.org/record/5402317/files/figure.png" pageId="7" pageNumber="635" startId="7.[197,236,1640,1659]" targetBox="[177,1183,207,1622]" targetPageId="7">
<paragraph id="DC293E19FFE8F84118A41D73FD20F0DE" blockId="7.[173,1194,1639,1682]" pageId="7" pageNumber="635">
<emphasis id="EEE2E20BFFE8F84118A41D73FEF2F037" bold="true" box="[197,299,1639,1659]" pageId="7" pageNumber="635">Figs. 7–10.</emphasis>
<taxonomicName id="1B96459AFFE8F84119211D7CFDDEF037" box="[320,519,1639,1659]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="635" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="EEE2E20BFFE8F84119211D7CFE62F036" box="[320,443,1639,1658]" italics="true" pageId="7" pageNumber="635">Pediomorphus</emphasis>
species
</taxonomicName>
, dorsal habitus. 
<emphasis id="EEE2E20BFFE8F8411AC51D73FD6CF037" bold="true" box="[676,693,1640,1659]" pageId="7" pageNumber="635">7)</emphasis>
<taxonomicName id="1B96459AFFE8F8411AA31D73FCECF036" authorityName="Will" authorityYear="2019" box="[706,821,1639,1658]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="635" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="EEE2E20BFFE8F8411AA31D73FCECF036" box="[706,821,1639,1658]" italics="true" pageId="7" pageNumber="635">P. crenulatus</emphasis>
</taxonomicName>
, holotype; 
<emphasis id="EEE2E20BFFE8F8411BC21D73FC6DF037" bold="true" box="[931,948,1640,1659]" pageId="7" pageNumber="635">8)</emphasis>
<taxonomicName id="1B96459AFFE8F8411BA01D73FBD7F036" authorityName="Will" authorityYear="2019" box="[961,1038,1640,1658]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="635" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="EEE2E20BFFE8F8411BA01D73FBD7F036" box="[961,1038,1640,1658]" italics="true" pageId="7" pageNumber="635">P. minor</emphasis>
</taxonomicName>
, holotype; 
<emphasis id="EEE2E20BFFE8F8411C1D1D73FB54F037" bold="true" box="[1148,1165,1640,1659]" pageId="7" pageNumber="635">9)</emphasis>
<taxonomicName id="1B96459AFFE8F8411CF81D73FEDDF0DE" authorityName="Will" authorityYear="2019" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="635" phylum="Arthropoda" rank="species" species="semilaevis">
<emphasis id="EEE2E20BFFE8F8411CF81D73FEDDF0DE" italics="true" pageId="7" pageNumber="635">P. semilaevis</emphasis>
</taxonomicName>
, paratype; 
<emphasis id="EEE2E20BFFE8F84119041D64FE58F0DE" bold="true" box="[357,385,1663,1682]" pageId="7" pageNumber="635">10)</emphasis>
<taxonomicName id="1B96459AFFE8F84119E91D9BFE29F0DE" authorityName="Will" authorityYear="2019" box="[392,496,1664,1682]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="635" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="EEE2E20BFFE8F84119E91D9BFE29F0DE" box="[392,496,1664,1682]" italics="true" pageId="7" pageNumber="635">P. punctatus</emphasis>
</taxonomicName>
, paratype. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="88E96E91FFE4F84D18A41D8AFDD4F0F0" ID-DOI="http://doi.org/10.5281/zenodo.5402319" ID-Zenodo-Dep="5402319" httpUri="https://zenodo.org/record/5402319/files/figure.png" pageId="11" pageNumber="639" startId="11.[197,236,1681,1700]" targetBox="[240,1122,207,1664]" targetPageId="11">
<paragraph id="DC293E19FFE4F84D18A41D8AFDD4F0F0" blockId="11.[173,1193,1680,1724]" pageId="11" pageNumber="639">
<emphasis id="EEE2E20BFFE4F84D18A41D8AFEEAF0E8" bold="true" box="[197,307,1681,1700]" pageId="11" pageNumber="639">Figs. 11–13.</emphasis>
<taxonomicName id="1B96459AFFE4F84D19281D8BFDD6F0E8" box="[329,527,1680,1700]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="639" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="EEE2E20BFFE4F84D19281D8BFE1DF0EF" box="[329,452,1680,1699]" italics="true" pageId="11" pageNumber="639">Pediomorphus</emphasis>
species
</taxonomicName>
, dorsal habitus. 
<emphasis id="EEE2E20BFFE4F84D1ACD1D8AFD1EF0E8" bold="true" box="[684,711,1681,1700]" pageId="11" pageNumber="639">11)</emphasis>
<emphasis id="EEE2E20BFFE4F84D1AB21D8AFCE4F0EF" box="[723,829,1680,1699]" italics="true" pageId="11" pageNumber="639">
<taxonomicName id="1B96459AFFE4F84D1AB21D8AFCE0F0EF" authorityName="Will" authorityYear="2019" box="[723,825,1680,1699]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="639" phylum="Arthropoda" rank="species" species="robustus">P. robustus</taxonomicName>
,
</emphasis>
paratype; 
<emphasis id="EEE2E20BFFE4F84D1BC21D8AFC66F0E8" bold="true" box="[931,959,1681,1700]" pageId="11" pageNumber="639">12)</emphasis>
<taxonomicName id="1B96459AFFE4F84D1BAA1D8AFBC6F0EF" authorityName="Will" authorityYear="2019" box="[971,1055,1681,1699]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="639" phylum="Arthropoda" rank="species" species="storeyi">
<emphasis id="EEE2E20BFFE4F84D1BAA1D8AFBC6F0EF" box="[971,1055,1681,1699]" italics="true" pageId="11" pageNumber="639">P. storeyi</emphasis>
</taxonomicName>
, holotype; 
<emphasis id="EEE2E20BFFE4F84D1CEC1D8AFB70F0E8" bold="true" box="[1165,1193,1681,1700]" pageId="11" pageNumber="639">13)</emphasis>
<taxonomicName id="1B96459AFFE4F84D18CC1DB2FEDDF0F7" authorityName=", Will" authorityYear="2019" box="[173,260,1704,1723]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="639" phylum="Arthropoda" rank="species" species="obtusus">
<emphasis id="EEE2E20BFFE4F84D18CC1DB2FEDDF0F7" box="[173,260,1704,1723]" italics="true" pageId="11" pageNumber="639">P. obtusus</emphasis>
</taxonomicName>
, paratype. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="88E96E91FFE3F84A18CE1FD7FE65F343" ID-DOI="http://doi.org/10.5281/zenodo.5402321" ID-Zenodo-Dep="5402321" httpUri="https://zenodo.org/record/5402321/files/figure.png" pageId="12" pageNumber="640" startId="12.[175,214,1228,1247]" targetBox="[156,1167,196,1210]" targetPageId="12">
<paragraph id="DC293E19FFE3F84A18CE1FD7FE65F343" blockId="12.[151,1173,1227,1295]" pageId="12" pageNumber="640">
<emphasis id="EEE2E20BFFE3F84A18CE1FD7FEC2F293" bold="true" box="[175,283,1228,1247]" pageId="12" pageNumber="640">Figs. 14–18.</emphasis>
<taxonomicName id="1B96459AFFE3F84A19511FD0FE2FF293" box="[304,502,1227,1247]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="640" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="EEE2E20BFFE3F84A19511FD0FE72F292" box="[304,427,1227,1246]" italics="true" pageId="12" pageNumber="640">Pediomorphus</emphasis>
species.
</taxonomicName>
<taxonomicName id="1B96459AFFE3F84A199D1FD7FDA5F292" authorityName="Chaudoir" authorityYear="1878" box="[508,636,1227,1246]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="640" phylum="Arthropoda" rank="species" species="planiusculus">
<emphasis id="EEE2E20BFFE3F84A199D1FD7FDA5F292" box="[508,636,1227,1246]" italics="true" pageId="12" pageNumber="640">P. planiusculus</emphasis>
</taxonomicName>
: 
<emphasis id="EEE2E20BFFE3F84A1AE81FD7FD7CF293" bold="true" box="[649,677,1228,1247]" pageId="12" pageNumber="640">14)</emphasis>
Labial palp; 
<emphasis id="EEE2E20BFFE3F84A1B7B1FD7FCEFF293" bold="true" box="[794,822,1228,1247]" pageId="12" pageNumber="640">15)</emphasis>
Base of right elytron. 
<taxonomicName id="1B96459AFFE3F84A1B9D1FD7FBB2F292" authorityName="Will" authorityYear="2019" box="[1020,1131,1227,1246]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="640" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="EEE2E20BFFE3F84A1B9D1FD7FBB2F292" box="[1020,1131,1227,1246]" italics="true" pageId="12" pageNumber="640">P. crenulatus</emphasis>
</taxonomicName>
: 
<emphasis id="EEE2E20BFFE3F84A1C191FD7FB4AF293" bold="true" box="[1144,1171,1228,1247]" pageId="12" pageNumber="640">16)</emphasis>
Base of right elytron. 
<taxonomicName id="1B96459AFFE3F84A193F1FFFFE1BF2BA" box="[350,450,1251,1270]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="640" phylum="Arthropoda" rank="species" species="macleayi">
<emphasis id="EEE2E20BFFE3F84A193F1FFFFE1BF2BA" box="[350,450,1251,1270]" italics="true" pageId="12" pageNumber="640">P. macleayi</emphasis>
</taxonomicName>
: 
<emphasis id="EEE2E20BFFE3F84A19B11FFFFE35F2BB" bold="true" box="[464,492,1252,1271]" pageId="12" pageNumber="640">17)</emphasis>
Head, showing large antennomere 1. 
<taxonomicName id="1B96459AFFE3F84A1B201FFFFC78F2BA" authorityName="Will" authorityYear="2019" box="[833,929,1251,1270]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="640" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="EEE2E20BFFE3F84A1B201FFFFC78F2BA" box="[833,929,1251,1270]" italics="true" pageId="12" pageNumber="640">P. robustus</emphasis>
</taxonomicName>
: 
<emphasis id="EEE2E20BFFE3F84A1BD11FFFFC15F2BB" bold="true" box="[944,972,1252,1271]" pageId="12" pageNumber="640">18)</emphasis>
Ventral view of head, arrow indicating subocular carina.
</paragraph>
</caption>
<caption id="88E96E91FFE2F84B18A41F92FC34F2D0" ID-DOI="http://doi.org/10.5281/zenodo.5402323" ID-Zenodo-Dep="5402323" box="[197,1005,1161,1180]" httpUri="https://zenodo.org/record/5402323/files/figure.png" pageId="13" pageNumber="641" startId="13.[197,228,1161,1180]" targetBox="[176,1188,195,1144]" targetPageId="13">
<paragraph id="DC293E19FFE2F84B18A41F92FC34F2D0" blockId="13.[197,1005,1161,1180]" box="[197,1005,1161,1180]" pageId="13" pageNumber="641">
<emphasis id="EEE2E20BFFE2F84B18A41F92FED1F2D0" bold="true" box="[197,264,1161,1180]" pageId="13" pageNumber="641">Fig. 19.</emphasis>
Localities of 
<taxonomicName id="1B96459AFFE2F84B19EE1F92FD97F2D0" box="[399,590,1161,1180]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="641" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="EEE2E20BFFE2F84B19EE1F92FDD3F2D0" box="[399,522,1161,1180]" italics="true" pageId="13" pageNumber="641">Pediomorphus</emphasis>
species
</taxonomicName>
based on material examined and type localities.
</paragraph>
</caption>
<caption id="88E96E91FFE1F84818CE1F93FC0EF2D7" ID-DOI="http://doi.org/10.5281/zenodo.5402325" ID-Zenodo-Dep="5402325" box="[175,983,1159,1179]" httpUri="https://zenodo.org/record/5402325/files/figure.png" pageId="14" pageNumber="642" startId="14.[175,206,1160,1179]" targetBox="[156,1167,196,1142]" targetPageId="14">
<paragraph id="DC293E19FFE1F84818CE1F93FC0EF2D7" blockId="14.[175,983,1159,1179]" box="[175,983,1159,1179]" pageId="14" pageNumber="642">
<emphasis id="EEE2E20BFFE1F84818CE1F93FF2AF2D7" bold="true" box="[175,243,1160,1179]" pageId="14" pageNumber="642">Fig. 20.</emphasis>
Localities of 
<taxonomicName id="1B96459AFFE1F848191B1F9CFDE1F2D7" box="[378,568,1159,1179]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="642" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="EEE2E20BFFE1F848191B1F9CFE2CF2D6" box="[378,501,1159,1178]" italics="true" pageId="14" pageNumber="642">Pediomorphus</emphasis>
species
</taxonomicName>
based on material examined and type localities.
</paragraph>
</caption>
<subSubSection id="948C6D92FFE0F8491AA21925FB71F418" box="[707,1192,574,598]" pageId="15" pageNumber="643" type="discussion">
<paragraph id="DC293E19FFE0F8491AA21925FB71F418" blockId="15.[707,1192,574,598]" box="[707,1192,574,598]" pageId="15" pageNumber="643">
<heading id="87618975FFE0F8491AA21925FB71F418" bold="true" box="[707,1192,574,598]" centered="true" fontSize="8" level="1" pageId="15" pageNumber="643" reason="1">
<emphasis id="EEE2E20BFFE0F8491AA21925FBDEF41F" bold="true" box="[707,1031,574,597]" pageId="15" pageNumber="643">KEY TO THE SPECIES OF ADULT</emphasis>
<taxonomicName id="1B96459AFFE0F8491C6F1925FB71F418" authorityName="CHAUDOIR" authorityYear="1878" box="[1038,1192,574,598]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="643" phylum="Arthropoda" rank="genus">
<emphasis id="EEE2E20BFFE0F8491C6F1925FB71F418" bold="true" box="[1038,1192,574,598]" italics="true" pageId="15" pageNumber="643">PEDIOMORPHUS</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="948C6D92FFE0F8561AA21977FB4AF5C5" lastPageId="16" lastPageNumber="644" pageId="15" pageNumber="643" type="key">
<key id="A8076928FFE0F8561AA21977FB4AF5C5" lastPageId="16" lastPageNumber="644" pageId="15" pageNumber="643">
<keyStep id="676226BCFFE0F8491AA21977FB70F5E9" pageId="15" pageNumber="643">
<paragraph id="DC293E19FFE0F8491AA21977FB70F56C" blockId="15.[707,1194,620,1730]" pageId="15" pageNumber="643">
<keyLead id="67679D2CFFE0F8491AA21977FB70F56C" pageId="15" pageNumber="643">1. Pronotum without evident marginal bead from hind angle to, or nearly to, basal impression; at most, lateral bead slightly extended along base beyond hind angle and ended or broadly interrupted by irregular punctation, or margin with a very shallow, usually irregular groove ....................... 2</keyLead>
</paragraph>
<paragraph id="DC293E19FFE0F8491AA2183DFB70F5E9" blockId="15.[707,1194,620,1730]" pageId="15" pageNumber="643">
<keyLead id="67679D2CFFE0F8491AA2183DFB70F5E9" pageId="15" pageNumber="643">
1 
<emphasis id="EEE2E20BFFE0F8491AAE183EFD0DF577" bold="true" box="[719,724,805,827]" pageId="15" pageNumber="643">ʹ</emphasis>
. Pronotum with marginal bead evident from hind angle to, or nearly to, level of basal impression, set off by a finely impressed submarginal groove, rarely with a few small punctures .......................................... 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="676226BCFFE0F8491AA218B1FB71F2D8" pageId="15" pageNumber="643">
<paragraph id="DC293E19FFE0F8491AA218B1FB70F5B9" blockId="15.[707,1194,620,1730]" pageId="15" pageNumber="643">
<keyLead id="67679D2CFFE0F8491AA218B1FB70F5B9" pageId="15" pageNumber="643">2. Striae coarsely or finely crenulate-punctate. Pronotal base with or without scattered punctures .......................................... 3</keyLead>
</paragraph>
<paragraph id="DC293E19FFE0F8491AA218E1FB71F2D8" blockId="15.[707,1194,620,1730]" pageId="15" pageNumber="643">
<keyLead id="67679D2CFFE0F8491AA218E1FB71F2D8" pageId="15" pageNumber="643">
2 
<emphasis id="EEE2E20BFFE0F8491AAE18E1FD0DF25C" bold="true" box="[719,724,1018,1040]" pageId="15" pageNumber="643">ʹ</emphasis>
. Striae not crenulate-punctate, shallowly sulcate with very shallow, irregular punctures more conspicuous near the base. Pronotum without punctures. Large size for genus,&gt;8.0 mm ........... 
<emphasis id="EEE2E20BFFE0F8491BD91F7FFB71F2D8" bold="true" pageId="15" pageNumber="643">
<taxonomicName id="1B96459AFFE0F8491BD91F7FFBFFF2D8" ID-CoL="8MGPM" authority="Will" authorityName="Will" authorityYear="2019" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="643" phylum="Arthropoda" rank="species" species="maximus" status="sp. nov.">
<emphasis id="EEE2E20BFFE0F8491BD91F7FFB70F235" bold="true" box="[952,1193,1124,1145]" italics="true" pageId="15" pageNumber="643">Pediomorphus maximus</emphasis>
Will
</taxonomicName>
, 
<taxonomicNameLabel id="F5D15F70FFE0F8491C521F64FB71F2D8" box="[1075,1192,1151,1172]" pageId="15" pageNumber="643" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="676226BCFFE0F8491AA21F81FB70F37F" pageId="15" pageNumber="643">
<paragraph id="DC293E19FFE0F8491AA21F81FB70F285" blockId="15.[707,1194,620,1730]" pageId="15" pageNumber="643">
<keyLead id="67679D2CFFE0F8491AA21F81FB70F285" pageId="15" pageNumber="643">3. Elytra concolored or with interval 1 paler .............................................. 5</keyLead>
</paragraph>
<paragraph id="DC293E19FFE0F8491AA21FD4FB70F37F" blockId="15.[707,1194,620,1730]" pageId="15" pageNumber="643">
<keyLead id="67679D2CFFE0F8491AA21FD4FB70F37F" pageId="15" pageNumber="643">
3 
<emphasis id="EEE2E20BFFE0F8491AAE1FD5FD0DF2A8" bold="true" box="[719,724,1230,1252]" pageId="15" pageNumber="643">ʹ</emphasis>
. Elytra variably bicolored with a black or infuscated region apparent at least centrally on intervals 2–3 on elytral disc, usually reaching interval 5 (
<figureCitation id="44AD229CFFE0F8491BA51E05FBDEF37F" box="[964,1031,1310,1331]" captionStart="Figs" captionStartId="2.[175,214,1711,1730]" captionTargetBox="[156,1167,936,1693]" captionTargetId="figure-367@2.[154,1169,934,1695]" captionTargetPageId="2" captionText="Figs. 1–2. Pediomorphus species, dorsal habitus. 1) P. planiusculus; 2) P. elongatus. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5402313" httpUri="https://zenodo.org/record/5402313/files/figure.png" pageId="15" pageNumber="643">Figs. 1</figureCitation>
, 
<figureCitation id="44AD229CFFE0F8491C741E05FBF8F37F" box="[1045,1057,1310,1331]" captionStart="Figs" captionStartId="7.[197,236,1640,1659]" captionTargetBox="[177,1183,207,1622]" captionTargetId="figure-0@7.[175,1190,194,1624]" captionTargetPageId="7" captionText="Figs. 7–10. Pediomorphus species, dorsal habitus. 7) P. crenulatus, holotype; 8) P. minor, holotype; 9) P. semilaevis, paratype; 10) P. punctatus, paratype. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5402317" httpUri="https://zenodo.org/record/5402317/files/figure.png" pageId="15" pageNumber="643">7</figureCitation>
) .............. 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="676226BCFFE0F8491AA21E22FB70F03E" pageId="15" pageNumber="643">
<paragraph id="DC293E19FFE0F8491AA21E22FB71F39F" blockId="15.[707,1194,620,1730]" pageId="15" pageNumber="643">
<keyLead id="67679D2CFFE0F8491AA21E22FB71F39F" pageId="15" pageNumber="643">
4. Striae finely punctate, with punctures contiguous or at most with 1 puncture width of separation in basal third of elytra (
<figureCitation id="44AD229CFFE0F8491C341E75FB44F3CF" box="[1109,1181,1390,1411]" captionStart="Figs" captionStartId="12.[175,214,1228,1247]" captionTargetBox="[156,1167,196,1210]" captionTargetId="figure-183@12.[154,1169,194,1212]" captionTargetPageId="12" captionText="Figs. 14–18. Pediomorphus species. P. planiusculus: 14) Labial palp; 15) Base of right elytron. P. crenulatus: 16) Base of right elytron. P. macleayi: 17) Head, showing large antennomere 1. P. robustus: 18) Ventral view of head, arrow indicating subocular carina." figureDoi="http://doi.org/10.5281/zenodo.5402321" httpUri="https://zenodo.org/record/5402321/files/figure.png" pageId="15" pageNumber="643">Fig. 15</figureCitation>
). Pronotal base with or without scattered punctures ................ 
<taxonomicName id="1B96459AFFE0F8491BC31EB8FB71F39F" ID-CoL="75Y5Q" authority="Chaudoir" authorityName="Chaudoir" authorityYear="1878" box="[930,1193,1443,1491]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="643" phylum="Arthropoda" rank="species" species="planiusculus">
<emphasis id="EEE2E20BFFE0F8491BC31EB8FB71F39F" bold="true" box="[930,1193,1443,1491]" pageId="15" pageNumber="643">
<emphasis id="EEE2E20BFFE0F8491BC31EB8FB70F3F4" bold="true" box="[930,1193,1443,1464]" italics="true" pageId="15" pageNumber="643">Pediomorphus planiusculus</emphasis>
Chaudoir
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="DC293E19FFE0F8491AA21EC3FB70F03E" blockId="15.[707,1194,620,1730]" pageId="15" pageNumber="643">
<keyLead id="67679D2CFFE0F8491AA21EC3FB70F03E" pageId="15" pageNumber="643">
4 
<emphasis id="EEE2E20BFFE0F8491AAE1EC3FD0DF3A2" bold="true" box="[719,724,1496,1518]" pageId="15" pageNumber="643">ʹ</emphasis>
. Striae coarsely punctate, with deep punctures 1–2 puncture widths of separation in basal third of elytra (
<figureCitation id="44AD229CFFE0F8491BF21D16FC02F06E" box="[915,987,1549,1570]" captionStart="Figs" captionStartId="12.[175,214,1228,1247]" captionTargetBox="[156,1167,196,1210]" captionTargetId="figure-183@12.[154,1169,194,1212]" captionTargetPageId="12" captionText="Figs. 14–18. Pediomorphus species. P. planiusculus: 14) Labial palp; 15) Base of right elytron. P. crenulatus: 16) Base of right elytron. P. macleayi: 17) Head, showing large antennomere 1. P. robustus: 18) Ventral view of head, arrow indicating subocular carina." figureDoi="http://doi.org/10.5281/zenodo.5402321" httpUri="https://zenodo.org/record/5402321/files/figure.png" pageId="15" pageNumber="643">Fig. 16</figureCitation>
). Pronotal base without punctures. Proepisternum anteromedially impunctate .......... 
<emphasis id="EEE2E20BFFE0F8491BD21D59FB70F03E" bold="true" pageId="15" pageNumber="643">
<taxonomicName id="1B96459AFFE0F8491BD21D59FBF5F03E" ID-CoL="8MGLR" authority="Will" authorityName="Will" authorityYear="2019" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="643" phylum="Arthropoda" rank="species" species="crenulatus" status="sp. nov.">
<emphasis id="EEE2E20BFFE0F8491BD21D59FB70F01B" bold="true" box="[947,1193,1602,1623]" italics="true" pageId="15" pageNumber="643">Pediomorphus crenulatus</emphasis>
Will
</taxonomicName>
, 
<taxonomicNameLabel id="F5D15F70FFE0F8491C561D46FB70F03E" box="[1079,1193,1629,1650]" pageId="15" pageNumber="643" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<caption id="88E96E91FFE0F84918A41D30FDCDF0FA" ID-DOI="http://doi.org/10.5281/zenodo.5402327" ID-Zenodo-Dep="5402327" httpUri="https://zenodo.org/record/5402327/files/figure.png" pageId="15" pageNumber="643" startId="15.[197,236,1579,1598]" targetBox="[185,645,195,1561]" targetPageId="15">
<paragraph id="DC293E19FFE0F84918A41D30FDCDF0FA" blockId="15.[173,660,1579,1718]" pageId="15" pageNumber="643">
<emphasis id="EEE2E20BFFE0F84918A41D30FEEEF072" bold="true" box="[197,311,1579,1598]" pageId="15" pageNumber="643">Figs. 21–32.</emphasis>
<taxonomicName id="1B96459AFFE0F849192D1D30FDCDF072" box="[332,532,1579,1598]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="643" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="EEE2E20BFFE0F849192D1D30FE1EF072" box="[332,455,1579,1598]" italics="true" pageId="15" pageNumber="643">Pediomorphus</emphasis>
species
</taxonomicName>
, aedeagi, left lateral view (A) and ventral view of tip (B). 
<emphasis id="EEE2E20BFFE0F8491A3D1D58FDAEF01A" bold="true" box="[604,631,1603,1622]" pageId="15" pageNumber="643">21)</emphasis>
<taxonomicName id="1B96459AFFE0F8491AE21D58FECEF021" ID-CoL="75Y5Q" authorityName="Chaudoir" authorityYear="1878" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="643" phylum="Arthropoda" rank="species" species="planiusculus">
<emphasis id="EEE2E20BFFE0F8491AE21D58FECEF021" italics="true" pageId="15" pageNumber="643">P. planiusculus</emphasis>
</taxonomicName>
; 
<emphasis id="EEE2E20BFFE0F84919471D40FE9BF022" bold="true" box="[294,322,1627,1646]" pageId="15" pageNumber="643">22)</emphasis>
<taxonomicName id="1B96459AFFE0F849192D1D40FE6FF021" ID-CoL="6TY62" authorityName="Sloane" authorityYear="1898" box="[332,438,1626,1645]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="643" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="EEE2E20BFFE0F849192D1D40FE6FF021" box="[332,438,1626,1645]" italics="true" pageId="15" pageNumber="643">P. elongatus</emphasis>
</taxonomicName>
; 
<emphasis id="EEE2E20BFFE0F84919A41D40FE38F022" bold="true" box="[453,481,1627,1646]" pageId="15" pageNumber="643">23)</emphasis>
<taxonomicName id="1B96459AFFE0F849198A1D40FD96F021" ID-CoL="75XTQ" box="[491,591,1626,1645]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="643" phylum="Arthropoda" rank="species" species="macleayi">
<emphasis id="EEE2E20BFFE0F849198A1D40FD96F021" box="[491,591,1626,1645]" italics="true" pageId="15" pageNumber="643">P. macleayi</emphasis>
</taxonomicName>
; 
<emphasis id="EEE2E20BFFE0F8491A3F1D40FDA3F022" bold="true" box="[606,634,1627,1646]" pageId="15" pageNumber="643">24)</emphasis>
<taxonomicName id="1B96459AFFE0F8491AE21D40FF20F0C9" ID-CoL="75Y62" authorityName="Sloane" authorityYear="1900" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="643" phylum="Arthropoda" rank="species" species="ruficollis">
<emphasis id="EEE2E20BFFE0F8491AE21D40FF20F0C9" italics="true" pageId="15" pageNumber="643">P. ruficollis</emphasis>
</taxonomicName>
; 
<emphasis id="EEE2E20BFFE0F849196F1D68FEF3F0CA" bold="true" box="[270,298,1651,1670]" pageId="15" pageNumber="643">25)</emphasis>
<taxonomicName id="1B96459AFFE0F84919581D68FE70F0C9" ID-CoL="6JJWT" authorityName="Straneo" authorityYear="1960" box="[313,425,1650,1669]" class="Insecta" family="Carabidae" genus="Fouquetius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="643" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="EEE2E20BFFE0F84919581D68FE70F0C9" box="[313,425,1650,1669]" italics="true" pageId="15" pageNumber="643">P. variabilis</emphasis>
</taxonomicName>
; 
<emphasis id="EEE2E20BFFE0F84919DF1D68FE03F0CA" bold="true" box="[446,474,1651,1670]" pageId="15" pageNumber="643">26)</emphasis>
<taxonomicName id="1B96459AFFE0F849198B1D68FDB8F0C9" ID-CoL="8MGLR" authorityName="Will" authorityYear="2019" box="[490,609,1650,1669]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="643" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="EEE2E20BFFE0F849198B1D68FDB8F0C9" box="[490,609,1650,1669]" italics="true" pageId="15" pageNumber="643">P. crenulatus</emphasis>
</taxonomicName>
; 
<emphasis id="EEE2E20BFFE0F8491A161D68FD4AF0CA" bold="true" box="[631,659,1651,1670]" pageId="15" pageNumber="643">27)</emphasis>
<taxonomicName id="1B96459AFFE0F84918CC1D90FF24F0D1" ID-CoL="8M7G9" authorityName="Will" authorityYear="2019" box="[173,253,1675,1693]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="643" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="EEE2E20BFFE0F84918CC1D90FF24F0D1" box="[173,253,1675,1693]" italics="true" pageId="15" pageNumber="643">P. minor</emphasis>
</taxonomicName>
; 
<emphasis id="EEE2E20BFFE0F84919731D90FEF7F0D2" bold="true" box="[274,302,1675,1694]" pageId="15" pageNumber="643">28)</emphasis>
<taxonomicName id="1B96459AFFE0F849195C1D90FE6BF0D1" ID-CoL="8MCWZ" authorityName="Will" authorityYear="2019" box="[317,434,1674,1693]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="643" phylum="Arthropoda" rank="species" species="semilaevis">
<emphasis id="EEE2E20BFFE0F849195C1D90FE6BF0D1" box="[317,434,1674,1693]" italics="true" pageId="15" pageNumber="643">P. semilaevis</emphasis>
</taxonomicName>
; 
<emphasis id="EEE2E20BFFE0F84919A61D90FE3AF0D2" bold="true" box="[455,483,1675,1694]" pageId="15" pageNumber="643">29)</emphasis>
<taxonomicName id="1B96459AFFE0F84919931D90FDBBF0D1" ID-CoL="8MM7P" authorityName="Will" authorityYear="2019" box="[498,610,1675,1693]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="643" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="EEE2E20BFFE0F84919931D90FDBBF0D1" box="[498,610,1675,1693]" italics="true" pageId="15" pageNumber="643">P. punctatus</emphasis>
</taxonomicName>
; 
<emphasis id="EEE2E20BFFE0F8491A161D90FD4AF0D2" bold="true" box="[631,659,1675,1694]" pageId="15" pageNumber="643">30)</emphasis>
<taxonomicName id="1B96459AFFE0F84918CC1DB8FED2F0F9" ID-CoL="8MKD2" authorityName="Will" authorityYear="2019" box="[173,267,1698,1717]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="643" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="EEE2E20BFFE0F84918CC1DB8FED2F0F9" box="[173,267,1698,1717]" italics="true" pageId="15" pageNumber="643">P. robustus</emphasis>
</taxonomicName>
; 
<emphasis id="EEE2E20BFFE0F84919761DB8FEEAF0FA" bold="true" box="[279,307,1699,1718]" pageId="15" pageNumber="643">31)</emphasis>
<taxonomicName id="1B96459AFFE0F849195B1DB8FE50F0F9" ID-CoL="8MB4N" authorityName="Will" authorityYear="2019" box="[314,393,1699,1717]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="643" phylum="Arthropoda" rank="species" species="storeyi">
<emphasis id="EEE2E20BFFE0F849195B1DB8FE50F0F9" box="[314,393,1699,1717]" italics="true" pageId="15" pageNumber="643">P. storeyi</emphasis>
</taxonomicName>
; 
<emphasis id="EEE2E20BFFE0F84919F41DB8FE68F0FA" bold="true" box="[405,433,1699,1718]" pageId="15" pageNumber="643">32)</emphasis>
<taxonomicName id="1B96459AFFE0F84919D91DB8FDD7F0F9" ID-CoL="8LVGJ" authorityName=", Will" authorityYear="2019" box="[440,526,1698,1717]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="643" phylum="Arthropoda" rank="species" species="obtusus">
<emphasis id="EEE2E20BFFE0F84919D91DB8FDD7F0F9" box="[440,526,1698,1717]" italics="true" pageId="15" pageNumber="643">P. obtusus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<keyStep id="676226BCFFE0F8561AA21D63FDA7F76A" lastPageId="16" lastPageNumber="644" pageId="15" pageNumber="643">
<paragraph id="DC293E19FFE0F8491AA21D63FB70F08E" blockId="15.[707,1194,620,1730]" pageId="15" pageNumber="643">
<keyLead id="67679D2CFFE0F8491AA21D63FB70F08E" pageId="15" pageNumber="643">5. Pronotal hind angles nearly right-angled, lateral margins noticeably, though shortly, sinuate just before hind angle ................ 6</keyLead>
</paragraph>
<paragraph id="DC293E19FFFFF85618F61BD9FDA7F76A" blockId="16.[151,639,193,1730]" pageId="16" pageNumber="644">
<keyLead id="67679D2CFFFFF85618F61BD9FDA7F76A" pageId="16" pageNumber="644">
5 
<emphasis id="EEE2E20BFFFFF85618C21BDAFF71F69B" bold="true" box="[163,168,193,215]" pageId="16" pageNumber="644">ʹ</emphasis>
. Pronotal hind angles broadly, obtusely angled, lateral margins at most only very slightly subsinuate just before hind angle ... ....................................................... 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="676226BCFFFFF85618F61A37FDA4F4FA" pageId="16" pageNumber="644">
<paragraph id="DC293E19FFFFF85618F61A37FDA7F7B7" blockId="16.[151,639,193,1730]" pageId="16" pageNumber="644">
<keyLead id="67679D2CFFFFF85618F61A37FDA7F7B7" pageId="16" pageNumber="644">
6. Abdominal ventrites impunctate or with a few scattered, shallow punctulae laterally and across the last ventrite; in some specimens, ventrites irregularly rugose laterally; pronotum and elytra lacking evident microsculpture; small size, 3.0–4.0 mm ............................. ......................... 
<emphasis id="EEE2E20BFFFFF85619EB1AD0FDA7F7B7" bold="true" pageId="16" pageNumber="644">
<taxonomicName id="1B96459AFFFFF85619EB1AD0FDD9F7B7" ID-CoL="8MCWZ" authority="Will" authorityName="Will" authorityYear="2019" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="644" phylum="Arthropoda" rank="species" species="semilaevis" status="sp. nov.">
<emphasis id="EEE2E20BFFFFF85619EB1AD0FDA7F7AC" bold="true" box="[394,638,459,480]" italics="true" pageId="16" pageNumber="644">Pediomorphus semilaevis</emphasis>
Will
</taxonomicName>
, 
<taxonomicNameLabel id="F5D15F70FFFFF8561A6A1AFDFDA7F7B7" box="[523,638,486,507]" pageId="16" pageNumber="644" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="DC293E19FFFFF85618F6191BFDA4F4FA" blockId="16.[151,639,193,1730]" pageId="16" pageNumber="644">
<keyLead id="67679D2CFFFFF85618F6191BFDA4F4FA" pageId="16" pageNumber="644">
6 
<emphasis id="EEE2E20BFFFFF85618C2191BFF71F45A" bold="true" box="[163,168,512,534]" pageId="16" pageNumber="644">ʹ</emphasis>
. Abdominal ventrites punctate throughout, though usually less densely medially; pronotum usually with microscuplture evident basolaterally and elytra with evident transversely stretched microsculpture; small or medium size, but typically larger, 4.0–5.0mm .................. ............. 
<taxonomicName id="1B96459AFFFFF856195519BBFDA4F4FA" ID-CoL="769TQ" authority="(Straneo)" baseAuthorityName="Straneo" baseAuthorityYear="1960" box="[308,637,672,694]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="644" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="EEE2E20BFFFFF856195519BBFDA4F4FA" bold="true" box="[308,637,672,694]" pageId="16" pageNumber="644">
<emphasis id="EEE2E20BFFFFF856195519BBFDC5F4F9" bold="true" box="[308,540,672,693]" italics="true" pageId="16" pageNumber="644">Pediomorphus variabilis</emphasis>
(Straneo)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="676226BCFFFFF85618F619A0FDA7F208" pageId="16" pageNumber="644">
<paragraph id="DC293E19FFFFF85618F619A0FDA7F53C" blockId="16.[151,639,193,1730]" pageId="16" pageNumber="644">
<keyLead id="67679D2CFFFFF85618F619A0FDA7F53C" pageId="16" pageNumber="644">
7. Pronotum distinctly punctate across entire base. Elytral striae 6–7 deeply, densely punctate throughout their length, similar to other striae, only stria 7 with smaller, less deeply impressed punctures very near base .......................... ......................... 
<emphasis id="EEE2E20BFFFFF85619EA185BFDA7F53C" bold="true" pageId="16" pageNumber="644">
<taxonomicName id="1B96459AFFFFF85619EA185BFE26F53C" ID-CoL="8MM7P" authority="Will" authorityName="Will" authorityYear="2019" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="644" phylum="Arthropoda" rank="species" species="punctatus" status="sp. nov.">
<emphasis id="EEE2E20BFFFFF85619EA185BFDA7F519" bold="true" box="[395,638,832,853]" italics="true" pageId="16" pageNumber="644">Pediomorphus punctatus</emphasis>
Will
</taxonomicName>
, 
<taxonomicNameLabel id="F5D15F70FFFFF8561A6B1840FDA7F53C" box="[522,638,859,880]" pageId="16" pageNumber="644" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="DC293E19FFFFF85618F6186EFDA7F208" blockId="16.[151,639,193,1730]" pageId="16" pageNumber="644">
<keyLead id="67679D2CFFFFF85618F6186EFDA7F208" pageId="16" pageNumber="644">
7 
<emphasis id="EEE2E20BFFFFF85618C2186EFF71F5C7" bold="true" box="[163,168,885,907]" pageId="16" pageNumber="644">ʹ</emphasis>
. Pronotal base not punctate, finely punctulate only in and near basal impressions, or with a few scattered punctures medially and/or laterally. Elytral striae 7 and also frequently 5–6 with smaller, shallower, more widely spaced punctures near base and more shallowly impressed compared with other striae .............................................. 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="676226BCFFFFF85618F61F51FDA7F39F" pageId="16" pageNumber="644">
<paragraph id="DC293E19FFFFF85618F61F51FDA4F2B2" blockId="16.[151,639,193,1730]" pageId="16" pageNumber="644">
<keyLead id="67679D2CFFFFF85618F61F51FDA4F2B2" pageId="16" pageNumber="644">
8. Small size, about 4.0 mm; head infuscated darker than pronotum and elytra; outer elytral striae very shallowly punctate and impressed, stria 7 not impressed in basal third, all striae impunctate in the apical third .................. ............................ 
<emphasis id="EEE2E20BFFFFF85619CA1FD5FDA4F2B2" bold="true" pageId="16" pageNumber="644">
<taxonomicName id="1B96459AFFFFF85619CA1FD5FE25F2B2" ID-CoL="8M7G9" authority="Will" authorityName="Will" authorityYear="2019" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="644" phylum="Arthropoda" rank="species" species="minor" status="sp. nov.">
<emphasis id="EEE2E20BFFFFF85619CA1FD5FDA7F2AF" bold="true" box="[427,638,1230,1251]" italics="true" pageId="16" pageNumber="644">Pediomorphus minor</emphasis>
Will
</taxonomicName>
, 
<taxonomicNameLabel id="F5D15F70FFFFF8561A691FF2FDA4F2B2" box="[520,637,1257,1278]" pageId="16" pageNumber="644" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="DC293E19FFFFF85618F61E1FFDA7F39F" blockId="16.[151,639,193,1730]" pageId="16" pageNumber="644">
<keyLead id="67679D2CFFFFF85618F61E1FFDA7F39F" pageId="16" pageNumber="644">
8 
<emphasis id="EEE2E20BFFFFF85618C21E18FF71F355" bold="true" box="[163,168,1283,1305]" pageId="16" pageNumber="644">ʹ</emphasis>
. Medium size,&gt; 
<quantity id="1B6E93FCFFFFF85619001E1FFE73F355" box="[353,426,1284,1305]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="16" pageNumber="644" unit="mm" value="4.5">4.5 mm</quantity>
; head concolorous with pronotum and elytra; all striae on elytral disc deeply, finely punctate and impressed, stria 7 impressed in basal third, although shallower than other striae, all striae punctate and crenulate except in apical fifth or less ................ ............................. 
<emphasis id="EEE2E20BFFFFF85619C51EB8FDA7F39F" bold="true" pageId="16" pageNumber="644">
<taxonomicName id="1B96459AFFFFF85619C51EB8FDD8F39F" ID-CoL="8LVGJ" authority="Will" authorityName=", Will" authorityYear="2019" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="644" phylum="Arthropoda" rank="species" species="obtusus" status="sp. nov.">
<emphasis id="EEE2E20BFFFFF85619C51EB8FDA7F3F4" bold="true" box="[420,638,1443,1464]" italics="true" pageId="16" pageNumber="644">Pediomorphus obtusus</emphasis>
Will
</taxonomicName>
, 
<taxonomicNameLabel id="F5D15F70FFFFF8561A6D1EA5FDA7F39F" box="[524,638,1470,1491]" pageId="16" pageNumber="644" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="676226BCFFFFF85618F61EC3FDA7F08E" pageId="16" pageNumber="644">
<paragraph id="DC293E19FFFFF85618F61EC3FDA7F014" blockId="16.[151,639,193,1730]" pageId="16" pageNumber="644">
<keyLead id="67679D2CFFFFF85618F61EC3FDA7F014" pageId="16" pageNumber="644">
9. First antennomere elongate, typical form for genus (
<figureCitation id="44AD229CFFFFF856192A1EE8FE44F044" box="[331,413,1523,1544]" captionStart="Figs" captionStartId="12.[175,214,1228,1247]" captionTargetBox="[156,1167,196,1210]" captionTargetId="figure-183@12.[154,1169,194,1212]" captionTargetPageId="12" captionText="Figs. 14–18. Pediomorphus species. P. planiusculus: 14) Labial palp; 15) Base of right elytron. P. crenulatus: 16) Base of right elytron. P. macleayi: 17) Head, showing large antennomere 1. P. robustus: 18) Ventral view of head, arrow indicating subocular carina." figureDoi="http://doi.org/10.5281/zenodo.5402321" httpUri="https://zenodo.org/record/5402321/files/figure.png" pageId="16" pageNumber="644">Fig. 18</figureCitation>
). Frontal foveae not impressed, shallow, short or linear. Pronotal base with punctures at least in or near basal impressions ...................................... 10
</keyLead>
</paragraph>
<paragraph id="DC293E19FFFFF85618F61D46FDA7F08E" blockId="16.[151,639,193,1730]" pageId="16" pageNumber="644">
<keyLead id="67679D2CFFFFF85618F61D46FDA7F08E" pageId="16" pageNumber="644">
9 
<emphasis id="EEE2E20BFFFFF85618C21D46FF71F03F" bold="true" box="[163,168,1629,1651]" pageId="16" pageNumber="644">ʹ</emphasis>
. First antennomere relatively short and notably swollen (
<figureCitation id="44AD229CFFFFF85619461D63FEB1F0C1" box="[295,360,1656,1677]" captionStart="Figs" captionStartId="12.[175,214,1228,1247]" captionTargetBox="[156,1167,196,1210]" captionTargetId="figure-183@12.[154,1169,194,1212]" captionTargetPageId="12" captionText="Figs. 14–18. Pediomorphus species. P. planiusculus: 14) Labial palp; 15) Base of right elytron. P. crenulatus: 16) Base of right elytron. P. macleayi: 17) Head, showing large antennomere 1. P. robustus: 18) Ventral view of head, arrow indicating subocular carina." figureDoi="http://doi.org/10.5281/zenodo.5402321" httpUri="https://zenodo.org/record/5402321/files/figure.png" pageId="16" pageNumber="644">Fig. 17</figureCitation>
). Frontal foveae impressed and punctate. Pronotal base without punctures.... ................. 
<taxonomicName id="1B96459AFFFFF856192E1DB7FDA7F08E" ID-CoL="75XTQ" authority="Sloane" authorityName="Sloane" authorityYear="1900" box="[335,638,1708,1730]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="644" phylum="Arthropoda" rank="species" species="macleayi">
<emphasis id="EEE2E20BFFFFF856192E1DB7FDA7F08E" bold="true" box="[335,638,1708,1730]" pageId="16" pageNumber="644">
<emphasis id="EEE2E20BFFFFF856192E1DB7FDEFF08D" bold="true" box="[335,566,1708,1729]" italics="true" pageId="16" pageNumber="644">Pediomorphus macleayi</emphasis>
Sloane
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="676226BCFFFFF8561ACC1BD9FB4AF7DC" pageId="16" pageNumber="644">
<paragraph id="DC293E19FFFFF8561ACC1BD9FB4AF740" blockId="16.[685,1172,194,905]" pageId="16" pageNumber="644">
<keyLead id="67679D2CFFFFF8561ACC1BD9FB4AF740" pageId="16" pageNumber="644">10. Entirely concolorous rufous-brown or with a very slightly infuscated head and paler elytral interval 1. Pronotal form variable ......... 11</keyLead>
</paragraph>
<paragraph id="DC293E19FFFFF8561ACC1A0AFB4AF7DC" blockId="16.[685,1172,194,905]" pageId="16" pageNumber="644">
<keyLead id="67679D2CFFFFF8561ACC1A0AFB4AF7DC" pageId="16" pageNumber="644">
10 
<emphasis id="EEE2E20BFFFFF8561AA41A0AFD13F76B" bold="true" box="[709,714,273,295]" pageId="16" pageNumber="644">ʹ</emphasis>
. Head and elytra piceous, contrasting with rufous-brown pronotum (
<figureCitation id="44AD229CFFFFF8561B821A37FBF8F70D" box="[995,1057,300,321]" captionStart="Figs" captionStartId="2.[175,214,1711,1730]" captionTargetBox="[156,1167,936,1693]" captionTargetId="figure-367@2.[154,1169,934,1695]" captionTargetPageId="2" captionText="Figs. 1–2. Pediomorphus species, dorsal habitus. 1) P. planiusculus; 2) P. elongatus. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5402313" httpUri="https://zenodo.org/record/5402313/files/figure.png" pageId="16" pageNumber="644">Fig. 2</figureCitation>
). Pronotum slightly wider than long, hind angles very broadly obtuse ...................................... .............. 
<taxonomicName id="1B96459AFFFFF8561B391A60FB4AF7DC" ID-CoL="75Y62" authority="Sloane" authorityName="Sloane" authorityYear="1900" box="[856,1171,379,400]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="644" phylum="Arthropoda" rank="species" species="ruficollis">
<emphasis id="EEE2E20BFFFFF8561B391A60FB4AF7DC" bold="true" box="[856,1171,379,400]" pageId="16" pageNumber="644">
<emphasis id="EEE2E20BFFFFF8561B391A60FB90F7DC" bold="true" box="[856,1097,379,400]" italics="true" pageId="16" pageNumber="644">Pediomorphus ruficollis</emphasis>
Sloane
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="676226BCFFFFF8561ACC1A8DFB4DF429" pageId="16" pageNumber="644">
<paragraph id="DC293E19FFFFF8561ACC1A8DFB4DF7B7" blockId="16.[685,1172,194,905]" pageId="16" pageNumber="644">
<keyLead id="67679D2CFFFFF8561ACC1A8DFB4DF7B7" pageId="16" pageNumber="644">
11. Abdominal sterna impunctate. Body form narrow-elongate. Pronotal anterior angles very slightly produced .................................... .............. 
<taxonomicName id="1B96459AFFFFF8561B391AFEFB4DF7B7" ID-CoL="6TY62" authority="Sloane" authorityName="Sloane" authorityYear="1898" box="[856,1172,485,507]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="644" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="EEE2E20BFFFFF8561B391AFEFB4DF7B7" bold="true" box="[856,1172,485,507]" pageId="16" pageNumber="644">
<emphasis id="EEE2E20BFFFFF8561B391AFEFB93F7B6" bold="true" box="[856,1098,485,506]" italics="true" pageId="16" pageNumber="644">Pediomorphus elongatus</emphasis>
Sloane
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="DC293E19FFFFF8561ACC191BFB4DF429" blockId="16.[685,1172,194,905]" pageId="16" pageNumber="644">
<keyLead id="67679D2CFFFFF8561ACC191BFB4DF429" pageId="16" pageNumber="644">
11 
<emphasis id="EEE2E20BFFFFF8561AA4191BFD13F45A" bold="true" box="[709,714,512,534]" pageId="16" pageNumber="644">ʹ</emphasis>
. Abdominal sterna densely and finely punctate or irregularly shallowly punctate laterally. Body form broad. Pronotal anterior angles prominently produced ............... 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="676226BCFFFFF8561ACC1970FB4AF5C5" pageId="16" pageNumber="644">
<paragraph id="DC293E19FFFFF8561ACC1970FB4DF4A6" blockId="16.[685,1172,194,905]" pageId="16" pageNumber="644">
<keyLead id="67679D2CFFFFF8561ACC1970FB4DF4A6" pageId="16" pageNumber="644">
12. Region in and around pronotal basal impressions with distinct, dense punctures; elytral microsculpture not visible at 50X magnification ..................... 
<emphasis id="EEE2E20BFFFFF8561BCC19A1FB4DF4A6" bold="true" pageId="16" pageNumber="644">
<taxonomicName id="1B96459AFFFFF8561BCC19A1FBCCF4A6" ID-CoL="8MKD2" authority="Will" authorityName="Will" authorityYear="2019" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="644" phylum="Arthropoda" rank="species" species="robustus" status="sp. nov.">
<emphasis id="EEE2E20BFFFFF8561BCC19A1FB4DF483" bold="true" box="[941,1172,698,719]" italics="true" pageId="16" pageNumber="644">Pediomorphus robustus</emphasis>
Will
</taxonomicName>
, 
<taxonomicNameLabel id="F5D15F70FFFFF8561C4119CEFB4DF4A6" box="[1056,1172,725,746]" pageId="16" pageNumber="644" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="DC293E19FFFFF8561ACC19EBFB4AF5C5" blockId="16.[685,1172,194,905]" pageId="16" pageNumber="644">
<keyLead id="67679D2CFFFFF8561ACC19EBFB4AF5C5" pageId="16" pageNumber="644">
12 
<emphasis id="EEE2E20BFFFFF8561AA419F4FD13F549" bold="true" box="[709,714,751,773]" pageId="16" pageNumber="644">ʹ</emphasis>
. Region around pronotal basal impressions with a few scattered, very shallow punctulae; elytra with perceptible microsculpture that is transversely stretched and somewhat iridescent ..................... 
<emphasis id="EEE2E20BFFFFF8561BDD1842FB4AF5C5" bold="true" pageId="16" pageNumber="644">
<taxonomicName id="1B96459AFFFFF8561BDD1842FBCBF5C5" ID-CoL="8MB4N" authority="Will" authorityName="Will" authorityYear="2019" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="644" phylum="Arthropoda" rank="species" species="storeyi" status="sp. nov.">
<emphasis id="EEE2E20BFFFFF8561BDD1842FB4DF522" bold="true" box="[956,1172,857,878]" italics="true" pageId="16" pageNumber="644">Pediomorphus storeyi</emphasis>
Will
</taxonomicName>
, 
<taxonomicNameLabel id="F5D15F70FFFFF8561C7F186FFB4AF5C5" box="[1054,1171,884,905]" pageId="16" pageNumber="644" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>