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<document id="F28DD1B452874844FBE9912188257962" ID-CLB-Dataset="133200" ID-DOI="10.1649/0010-065X-73.3.629" ID-GBIF-Dataset="99bf6930-d483-4789-886e-2aaf5b44d9b1" ID-ISSN="1938-4394" ID-Zenodo-Dep="5402311" ID-ZooBank="urn:lsid:zoobank.org:pub:48CFB8E9-AA28-413D-98CB-1841B65ED236" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630336494997" checkinUser="valdenar" docAuthor="Will, Kipling" docDate="2019" docId="543F8F0FFFE6F84F18A81FC2FBEDF08E" docLanguage="en" docName="TheColeopBull.73.3.629-645.pdf" docOrigin="The Coleopterists Bulletin 73 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-73.3.629" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Pediomorphus punctatus Will 2019, new species" docType="treatment" docUuid="4ADE4AF0-A5D8-4CD6-8526-C7B770028546" docUuidSource="ZooBank" docVersion="7" lastPageNumber="637" masterDocId="A806F777FFEFF84618611B1BFFD9F64C" masterDocTitle="Taxonomic Review of the Australian Genus Pediomorphus Chaudoir, 1878 (Coleoptera: Carabidae: Abacetini)" masterLastPageNumber="645" masterPageNumber="629" pageNumber="637" updateTime="1699242009079" updateUser="ExternalLinkService">
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<mods:title id="B076DDD09439606515BD3737F29B444F">Taxonomic Review of the Australian Genus Pediomorphus Chaudoir, 1878 (Coleoptera: Carabidae: Abacetini)</mods:title>
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<mods:namePart id="211BCBE1A40C5F03C1FFB01E3404ED21">Will, Kipling</mods:namePart>
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<mods:title id="D5C565980BA346D79CAD3AAC587B5532">The Coleopterists Bulletin</mods:title>
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<mods:date id="E1F0EB32B540B3AA948C0882A7C0DBBB">2019</mods:date>
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<paragraph id="DC293E19FFE6F84F18A81FC2FDAFF2A2" blockId="9.[185,646,1241,1342]" box="[201,630,1241,1262]" pageId="9" pageNumber="637">
<emphasis id="EEE2E20BFFE6F84F18A81FC2FDAFF2A2" bold="true" box="[201,630,1241,1262]" pageId="9" pageNumber="637">
<taxonomicName id="1B96459AFFE6F84F18A81FC2FE2DF2A2" ID-CoL="8MM7P" authority="Will" authorityName="Will" authorityYear="2019" box="[201,500,1241,1262]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="637" phylum="Arthropoda" rank="species" species="punctatus" status="sp. nov.">
<emphasis id="EEE2E20BFFE6F84F18A81FC2FE18F2A2" bold="true" box="[201,449,1241,1262]" italics="true" pageId="9" pageNumber="637">Pediomorphus punctatus</emphasis>
Will
</taxonomicName>
,
<taxonomicNameLabel id="F5D15F70FFE6F84F1A601FC2FDAFF2A2" box="[513,630,1241,1262]" pageId="9" pageNumber="637" rank="species">new species</taxonomicNameLabel>
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Zoobank.org/
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urn:lsid:zoobank.org:act:
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(
<figureCitation id="44AD229CFFE6F84F19331E32FE7DF372" box="[338,420,1321,1342]" captionStart="Figs" captionStartId="7.[197,236,1640,1659]" captionTargetBox="[177,1183,207,1622]" captionTargetId="figure-0@7.[175,1190,194,1624]" captionTargetPageId="7" captionText="Figs. 710. Pediomorphus species, dorsal habitus. 7) P. crenulatus, holotype; 8) P. minor, holotype; 9) P. semilaevis, paratype; 10) P. punctatus, paratype. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5402317" httpUri="https://zenodo.org/record/5402317/files/figure.png" pageId="9" pageNumber="637">Figs. 10</figureCitation>
,
<figureCitation id="44AD229CFFE6F84F19CE1E32FE10F372" box="[431,457,1321,1342]" captionStart="Fig" captionStartId="13.[197,228,1161,1180]" captionTargetBox="[176,1188,195,1144]" captionTargetId="figure-237@13.[175,1190,194,1146]" captionTargetPageId="13" captionText="Fig. 19. Localities of Pediomorphus species based on material examined and type localities." figureDoi="http://doi.org/10.5281/zenodo.5402323" httpUri="https://zenodo.org/record/5402323/files/figure.png" pageId="9" pageNumber="637">19</figureCitation>
,
<figureCitation id="44AD229CFFE6F84F19B41E32FE37F372" box="[469,494,1321,1342]" captionStart="Figs" captionStartId="15.[197,236,1579,1598]" captionTargetBox="[185,645,195,1561]" captionTargetId="figure-359@15.[184,647,194,1563]" captionTargetPageId="15" captionText="Figs. 2132. Pediomorphus species, aedeagi, left lateral view (A) and ventral view of tip (B). 21) P. planiusculus; 22) P. elongatus; 23) P. macleayi; 24) P. ruficollis; 25) P. variabilis; 26) P. crenulatus; 27) P. minor; 28) P. semilaevis; 29) P. punctatus; 30) P. robustus; 31) P. storeyi; 32) P. obtusus." figureDoi="http://doi.org/10.5281/zenodo.5402327" httpUri="https://zenodo.org/record/5402327/files/figure.png" pageId="9" pageNumber="637">29</figureCitation>
)
</paragraph>
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<paragraph id="DC293E19FFE6F84F18A41E4FFE83F071" blockId="9.[173,660,1363,1730]" pageId="9" pageNumber="637">
<emphasis id="EEE2E20BFFE6F84F18A41E4FFE8BF325" bold="true" box="[197,338,1364,1385]" pageId="9" pageNumber="637">Type Material.</emphasis>
<materialsCitation id="6CFE3444FFE6F84F19371E4FFDDAF3F4" ID-GBIF-Occurrence="3897123330" collectingDate="1967-12-01" collectingDateMax="1967-12-20" collectingDateMin="1967-12-01" collectingMethod="light trap" collectionCode="ANIC" collectorName="W. J. M. Vestjens &amp; U. C. Berkeley" latitude="-14.31" location="Tindal" longLatPrecision="773" longitude="132.22" pageId="9" pageNumber="637" specimenCode="25-067867" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="032D80BBFFE6F84F19371E4FFE0AF325" box="[342,467,1364,1385]" pageId="9" pageNumber="637" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="CA90F590FFE6F84F19B81E4FFDD2F325" box="[473,523,1364,1385]" pageId="9" pageNumber="637" type="male">Male</specimenCount>
.//“
<location id="D94968C2FFE6F84F1A461E4FFDBDF325" LSID="urn:lsid:plazi:treatment:543F8F0FFFE6F84F18A81FC2FBEDF08E:D94968C2FFE6F84F1A461E4FFDBDF325" box="[551,612,1364,1385]" latitude="-14.31" longLatPrecision="773" longitude="132.22" name="Tindal" pageId="9" pageNumber="637">Tindal</location>
, N.T.
<geoCoordinate id="B9A258DEFFE6F84F18CC1E75FF34F3CF" box="[173,237,1390,1411]" degrees="14.31" direction="south" orientation="latitude" pageId="9" pageNumber="637" precision="555" value="-14.31">14.31S</geoCoordinate>
<geoCoordinate id="B9A258DEFFE6F84F18971E75FE9DF3CF" box="[246,324,1390,1411]" degrees="132.22" direction="east" orientation="longitude" pageId="9" pageNumber="637" precision="555" value="132.22">132.22E</geoCoordinate>
,
<date id="A82818D9FFE6F84F19301E75FE3CF3CF" box="[337,485,1390,1411]" pageId="9" pageNumber="637" value="1967-12-01" valueMax="1967-12-20" valueMin="1967-12-01">
<collectingDate id="B86CE131FFE6F84F19301E75FE3CF3CF" box="[337,485,1390,1411]" pageId="9" pageNumber="637" value="1967-12-01" valueMax="1967-12-20" valueMin="1967-12-01">1-20 Dec. 1967</collectingDate>
</date>
<collectingMethod id="05D7460EFFE6F84F198F1E75FD9CF3CF" box="[494,581,1390,1411]" pageId="9" pageNumber="637">light trap</collectingMethod>
,
<collectorName id="71635BCFFFE6F84F1A301E75FF21F3D2" pageId="9" pageNumber="637">W.J.M. Vestjens</collectorName>
”// “
<collectorName id="71635BCFFFE6F84F19471E92FE6BF3D2" box="[294,434,1417,1438]" pageId="9" pageNumber="637">U.C. Berkeley</collectorName>
EMEC705,232”// [deposited
<collectionCode id="BA87A6DCFFE6F84F18971EB8FEE9F3F4" box="[246,304,1443,1464]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="9" pageNumber="637">ANIC</collectionCode>
, Reg. No.
<specimenCode id="8C309662FFE6F84F19F61EB8FE27F3F4" box="[407,510,1443,1464]" pageId="9" pageNumber="637">25-067867</specimenCode>
]
</materialsCitation>
.
<materialsCitation id="6CFE3444FFE6F84F1A6F1EB8FD59F3A1" ID-GBIF-Occurrence="3897123333" collectingDate="1967-12-01" collectingDateMax="1967-12-20" collectingDateMin="1967-12-01" collectionCode="ANIC" collectorName="W. J. M. Vestjens &amp; U. C. Berkeley" latitude="-14.31" longLatPrecision="773" longitude="132.22" pageId="9" pageNumber="637" specimenCode="EMEC705233, EMEC705237" specimenCount="5" stateProvince="Northern Territory" typeStatus="paratype">
<typeStatus id="032D80BBFFE6F84F1A6F1EB8FD48F3F4" box="[526,657,1443,1464]" pageId="9" pageNumber="637" type="paratype">PARATYPES</typeStatus>
:
<collectingRegion id="1E52F0FBFFE6F84F18CC1EA5FE69F39F" box="[173,432,1470,1491]" country="Australia" name="Northern Territory" pageId="9" pageNumber="637">NORTHERN TERRITORY</collectingRegion>
:
<specimenCount id="CA90F590FFE6F84F19D61EA5FDFFF39F" box="[439,550,1470,1491]" pageId="9" pageNumber="637" type="generic">5 specimens</specimenCount>
- same data as holotype,
<specimenCode id="8C309662FFE6F84F197C1EC3FE7BF3A1" box="[285,418,1496,1517]" collectionCode="EMEC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="9" pageNumber="637" type="University or college">EMEC705233</specimenCode>
-
<specimenCode id="8C309662FFE6F84F19C81EC3FDE9F3A1" box="[425,560,1496,1517]" collectionCode="EMEC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="9" pageNumber="637" type="University or college">EMEC705237</specimenCode>
, [
<collectionCode id="BA87A6DCFFE6F84F1A5D1EC3FDA2F3A1" box="[572,635,1496,1517]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="9" pageNumber="637">ANIC</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="6CFE3444FFE6F84F1AE61EC3FE8FF071" ID-GBIF-Occurrence="3897123318" collectingDate="2014-04-29" collectorName="M Langer" latitude="-14.950556" location="Roper Hwy" longLatPrecision="21" longitude="133.54973" pageId="9" pageNumber="637" specimenCount="2" stateProvince="NORTHERN TERRITORY" typeStatus="paratype">
<specimenCount id="CA90F590FFE6F84F1AE61EC3FED2F044" pageId="9" pageNumber="637" type="generic">2 specimens</specimenCount>
-
<locationDeviation id="CD9B273BFFE6F84F19441EE8FDCDF044" box="[293,532,1523,1544]" location="Roper Bar" pageId="9" pageNumber="637">
<quantity id="1B6E93FCFFE6F84F19441EE8FEB6F044" box="[293,367,1523,1544]" metricMagnitude="5" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="637" unit="km" value="150.0">150 km</quantity>
W of Roper Bar
</locationDeviation>
,
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,
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, at light,
<date id="A82818D9FFE6F84F19911D16FD8BF06E" box="[496,594,1549,1570]" pageId="9" pageNumber="637" value="2014-04-29">
<collectingDate id="B86CE131FFE6F84F19911D16FD8BF06E" box="[496,594,1549,1570]" pageId="9" pageNumber="637" value="2014-04-29">
29.iv.
<quantity id="1B6E93FCFFE6F84F1A411D16FD8BF06E" box="[544,594,1549,1570]" metricMagnitude="6" metricUnit="m" metricValue="2.014117" pageId="9" pageNumber="637" unit="m" value="2014117.0">2014</quantity>
</collectingDate>
</date>
, 117m,
<collectorName id="71635BCFFFE6F84F18CC1D33FED2F071" box="[173,267,1576,1597]" pageId="9" pageNumber="637">M Langer</collectorName>
, [cWR]
</materialsCitation>
.
</paragraph>
<paragraph id="DC293E19FFE6F84F18A41D58FE89F08E" blockId="9.[173,660,1363,1730]" pageId="9" pageNumber="637">
<emphasis id="EEE2E20BFFE6F84F18A41D58FDBBF014" bold="true" box="[197,610,1603,1624]" pageId="9" pageNumber="637">Type Locality and Distribution Range.</emphasis>
<collectingMunicipality id="3C4DA463FFE6F84F1A0F1D58FD4AF014" box="[622,659,1603,1624]" pageId="9" pageNumber="637">The</collectingMunicipality>
type locality is as on the
<typeStatus id="032D80BBFFE6F84F19F41D46FE31F03E" box="[405,488,1629,1650]" pageId="9" pageNumber="637" type="holotype">holotype</typeStatus>
label,
<emphasis id="EEE2E20BFFE6F84F1A491D46FDE7F03E" box="[552,574,1629,1650]" italics="true" pageId="9" pageNumber="637">i.e</emphasis>
.,
<location id="D94968C2FFE6F84F1A311D46FD49F03E" LSID="urn:lsid:plazi:treatment:543F8F0FFFE6F84F18A81FC2FBEDF08E:D94968C2FFE6F84F1A311D46FD49F03E" box="[592,656,1629,1650]" latitude="-13.516666" longLatPrecision="1290" longitude="132.36667" municipality="The" name="Tindal" pageId="9" pageNumber="637" stateProvince="Northern Territory">Tindal</location>
,
<collectingRegion id="1E52F0FBFFE6F84F18CC1D63FEBAF0C1" box="[173,355,1656,1677]" country="Australia" name="Northern Territory" pageId="9" pageNumber="637">Northern Territory</collectingRegion>
in the vicinity of
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<geoCoordinate id="B9A258DEFFE6F84F18CC1D89FECDF0EB" box="[173,276,1682,1704]" degrees="132" direction="east" minutes="22" orientation="longitude" pageId="9" pageNumber="637" precision="925" value="132.36667">132°22 E</geoCoordinate>
(
<figureCitation id="44AD229CFFE6F84F19461D89FEADF0EB" box="[295,372,1682,1703]" captionStart="Fig" captionStartId="13.[197,228,1161,1180]" captionTargetBox="[176,1188,195,1144]" captionTargetId="figure-237@13.[175,1190,194,1146]" captionTargetPageId="13" captionText="Fig. 19. Localities of Pediomorphus species based on material examined and type localities." figureDoi="http://doi.org/10.5281/zenodo.5402323" httpUri="https://zenodo.org/record/5402323/files/figure.png" pageId="9" pageNumber="637">Fig. 19</figureCitation>
), which is at approximately
<quantity id="1B6E93FCFFE6F84F18CC1DB6FF33F08E" box="[173,234,1709,1730]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="637" unit="m" value="130.0">
<elevation id="57BBD92AFFE6F84F18CC1DB6FF33F08E" box="[173,234,1709,1730]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="637" unit="m" value="130.0">130 m</elevation>
</quantity>
elevation.
</paragraph>
</subSubSection>
<subSubSection id="948C6D92FFE6F84F1ABA1BD9FCE1F70D" pageId="9" pageNumber="637" type="diagnosis">
<paragraph id="DC293E19FFE6F84F1ABA1BD9FCE1F70D" blockId="9.[707,1194,193,1730]" pageId="9" pageNumber="637">
<emphasis id="EEE2E20BFFE6F84F1ABA1BD9FC9BF69B" bold="true" box="[731,834,194,215]" pageId="9" pageNumber="637">Diagnosis.</emphasis>
<taxonomicName id="1B96459AFFE6F84F1B291BDAFBEDF69B" authorityName="Will" authorityYear="2019" box="[840,1076,193,215]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="637" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="EEE2E20BFFE6F84F1B291BDAFBEDF69B" box="[840,1076,193,215]" italics="true" pageId="9" pageNumber="637">Pediomorphus punctatus</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="1B96459AFFE6F84F1AA21BC7FCE0F6BD" authorityName="Straneo" authorityYear="1960" box="[707,825,219,241]" class="Insecta" family="Carabidae" genus="Fouquetius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="637" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="EEE2E20BFFE6F84F1AA21BC7FCE0F6BD" box="[707,825,219,241]" italics="true" pageId="9" pageNumber="637">P. variabilis</emphasis>
</taxonomicName>
and
<taxonomicName id="1B96459AFFE6F84F1B0E1BC7FC0AF6BD" authorityName=", Will" authorityYear="2019" box="[879,979,219,241]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="637" phylum="Arthropoda" rank="species" species="obtusus">
<emphasis id="EEE2E20BFFE6F84F1B0E1BC7FC0AF6BD" box="[879,979,219,241]" italics="true" pageId="9" pageNumber="637">P. obtusus</emphasis>
</taxonomicName>
but with much more extensive punctation across the base of the pronotum, which also has more obtuse and less prominent hind angles.
</paragraph>
</subSubSection>
<subSubSection id="948C6D92FFE6F84F1ABA1A5DFB97F01A" pageId="9" pageNumber="637" type="description">
<paragraph id="DC293E19FFE6F84F1ABA1A5DFB97F01A" blockId="9.[707,1194,193,1730]" pageId="9" pageNumber="637">
<emphasis id="EEE2E20BFFE6F84F1ABA1A5DFC86F717" bold="true" box="[731,863,326,347]" pageId="9" pageNumber="637">Description.</emphasis>
Description.
<emphasis id="EEE2E20BFFE6F84F1B9D1A5DFBE8F717" bold="true" box="[1020,1073,326,347]" pageId="9" pageNumber="637">Size:</emphasis>
sbl = 4.9 [4.55.1] mm; greatest width across elytra 1.8 = [1.61.8] mm.
<emphasis id="EEE2E20BFFE6F84F1B341A67FC4EF7DD" bold="true" box="[853,919,380,401]" pageId="9" pageNumber="637">Color:</emphasis>
Dorsal and ventral surfaces castaneous to brunneous; legs, mouthparts, and antennae concolorous or slightly paler. Dorsally distinctly and ventrally slightly shiny to moderately dull. Dorsally with spectral iridescence, venter, elytral epipleura, and legs with or without slight spectral iridescent.
<emphasis id="EEE2E20BFFE6F84F1BE71904FC1AF478" bold="true" box="[902,963,543,564]" pageId="9" pageNumber="637">Head:</emphasis>
Dorsal microsculpture hardly evident in mesh sculpticells at 50X magnification. Clypeal-ocular impressions short, divergent, well-delimited laterally, not clearly delimited medially. Ocular ratio = 1.50 [1.491.56]. Eyes relatively large, somewhat prominent, with small posterior orbital area, 1 subocular carina long, sharply defined, sometimes 1 or 2 additional, smaller carinae present. Labrum with anterior margin slightly convex. Median tooth of mentum broad, flat across apex, sometimes with very slight emargination. Antennae long, antennomeres 911 surpassing pronotal base.
<emphasis id="EEE2E20BFFE6F84F1BA1187EFBCBF536" bold="true" box="[960,1042,869,890]" pageId="9" pageNumber="637">Thorax:</emphasis>
Pronotum dorsally with distinct punctures along base in and around basal impressions and medially, laterally usually also punctate, but more sparsely and often not reaching lateral margin, microsculpture not evident at 50X magnification. Pronotum widest slightly anteriad of middle, lateral margins converging to base, not or only very slightly sinuate. Lateral marginal bead narrow and ending just at hind angles or extending slightly and obscurely along base; basal margin without marginal bead; anterior angles not prominent, broadly rounded; hind angles obtuse, not denticulate; basal impressions linear, broad, shallowly impressed, reaching basal margin. Seta at hind angle in marginal bead. Elytra parallel-sided, dorsally depressed, epipleura in dorsal view very slightly sinuate at level of umbilicate marginal puncture 7. Elytral striae finely, shallowly crenulate-punctate. Elytral microsculpture not evident at 50X magnification or scarcely visible as transversely stretch lines. Intervals flat. Elytral plica short, small. Prosternal process rounded, deeply margined. Pro-, meso- and metasterna coarsely punctate.
<emphasis id="EEE2E20BFFE6F84F1B411ECEFC50F3A6" bold="true" box="[800,905,1493,1514]" pageId="9" pageNumber="637">Abdomen:</emphasis>
Abdominal ventrites densely, finely punctate.
<emphasis id="EEE2E20BFFE6F84F1B381EEBFC19F049" bold="true" box="[857,960,1520,1541]" pageId="9" pageNumber="637">Aedeagus:</emphasis>
Medium lobe long, with thin apical portion in lateral view, tip slightly asymmetrically roundly in ventral view, no spines or spinules visible on endophallus in repose.
</paragraph>
</subSubSection>
<subSubSection id="948C6D92FFE6F84F1ABA1D47FBEDF08E" pageId="9" pageNumber="637" type="etymology">
<paragraph id="DC293E19FFE6F84F1ABA1D47FBEDF08E" blockId="9.[707,1194,193,1730]" pageId="9" pageNumber="637">
<emphasis id="EEE2E20BFFE6F84F1ABA1D47FC94F03D" bold="true" box="[731,845,1628,1649]" pageId="9" pageNumber="637">Etymology.</emphasis>
The specific epithet is the Latin masculine nominative
<taxonomicName id="1B96459AFFE6F84F1BFF1D6CFC23F0C0" authorityName="Will" authorityYear="2019" box="[926,1018,1655,1676]" class="Insecta" family="Carabidae" genus="Pediomorphus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="637" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="EEE2E20BFFE6F84F1BFF1D6CFC23F0C0" box="[926,1018,1655,1676]" italics="true" pageId="9" pageNumber="637">punctatus</emphasis>
</taxonomicName>
, which is a reference to the numerous, small punctures across the base of the pronotum of these beetles.
</paragraph>
</subSubSection>
</treatment>
</document>