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<document id="F0E3D32D0D76E00EB5178852A6DBD771" ID-CLB-Dataset="22478" ID-DOI="10.5852/ejt.2017.258" ID-GBIF-Dataset="35792d9c-db1a-45f1-b983-f52105f8d004" ID-ISSN="2118-9773" ID-Zenodo-Dep="3776244" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588180985326" checkinUser="carolina" docAuthor="Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn &amp; Alpert, Gary D." docDate="2017" docId="543E8C277C71FA6BFD85FB2DFAD3187D" docLanguage="en" docName="ejt-2017-258.pdf.imf" docOrigin="European Journal of Taxonomy 258" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Nesomyrmex innocens" docType="treatment" docVersion="12" lastPageNumber="27" masterDocId="A807F45F7C69FA71FFBAFFA7FFE21A12" masterDocTitle="Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique" masterLastPageNumber="31" masterPageNumber="1" pageNumber="25" updateTime="1732492056321" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="92D8BCDF73721C5A8B1444AFE849D4A2">Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique</mods:title>
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<mods:namePart id="93782303C7F2C82AECF9D4AD8B1F052E">Garcia, Francisco Hita</mods:namePart>
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<mods:affiliation id="7E4E35C5A54461817D74C840A0DE29ED">Okinawa Institute of Science and Technology, Onna-son, Okinawa, Japan. &amp; urn: lsid: zoobank. org: author: B 7 ADF 56 F- 935 D- 4 BD 8 - ADB 3 - 50 E 96 F 8 BB 463 &amp; Corresponding author: fhitagarcia @ gmail. com</mods:affiliation>
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<mods:namePart id="3CDAFDC91113CB2DECBE0A822F122F20">Mbanyana, Nokuthula</mods:namePart>
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<mods:affiliation id="E0BB7183DD9D1136E30093135CAB755F">Iziko South African Museum, Cape Town, South Africa. &amp; Email: nmbanyana @ iziko. org. za &amp; urn: lsid: zoobank. org: author: 157 A 51 ED- 7 C 85 - 4 CA 3 - 84 E 4 - B 7 BD 1 E 2 A 7 E 8 F</mods:affiliation>
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<mods:namePart id="7CFAA3784B9F99E505DC27349D87FFDC">Audisio, Tracy Lynn</mods:namePart>
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<mods:affiliation id="CA9DA8B979E63E67D33674264FD8729D">Okinawa Institute of Science and Technology, Onna-son, Okinawa, Japan. &amp; Email: tracy @ troglophile. com &amp; urn: lsid: zoobank. org: author: C 821 F 482 - 744 F- 467 A-A 480 - 1 E 353001 A 51 F</mods:affiliation>
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<mods:namePart id="26E7BF0682376D23FD54C5842CDBEE20">Alpert, Gary D.</mods:namePart>
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<mods:affiliation id="144DB3D333A5C695D8E2CCFDD8CA308B">Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, U. S. A. &amp; Email: garydalpert @ gmail. com &amp; urn: lsid: zoobank. org: author: 5 BD 71 BB 5 - 1 EDA- 4 E 05 - 84 E 1 - FE 71 EA 366767</mods:affiliation>
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<treatment id="543E8C277C71FA6BFD85FB2DFAD3187D" ID-DOI="http://doi.org/10.5281/zenodo.3796566" ID-GBIF-Taxon="163876685" ID-Zenodo-Dep="3796566" LSID="urn:lsid:plazi:treatment:543E8C277C71FA6BFD85FB2DFAD3187D" httpUri="http://treatment.plazi.org/id/543E8C277C71FA6BFD85FB2DFAD3187D" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25" scope_family="Formicidae" scope_subFamily="Myrmicinae">
<subSubSection id="948D6EBA7C71FA69FD85FB2DFC161EB6" box="[575,1012,1161,1188]" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="DC283D317C71FA69FD85FB2DFC161EB6" blockId="24.[575,1012,1161,1223]" box="[575,1012,1161,1188]" pageId="24" pageNumber="25">
<heading id="87608A5D7C71FA69FD85FB2DFC161EB6" box="[575,1012,1161,1188]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="25" reason="2">
<taxonomicName id="1B9746B27C71FA69FD85FB2DFC161EB6" ID-CoL="CM7YR" authority="(Forel, 1913)" baseAuthorityName="Forel" baseAuthorityYear="1913" box="[575,1012,1161,1188]" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="innocens">
<emphasis id="EEE3E1237C71FA69FD85FB2DFCB01EB1" bold="true" box="[575,850,1161,1188]" italics="true" pageId="24" pageNumber="25">Nesomyrmex innocens</emphasis>
(
<bibRefCitation id="B80640C07C71FA69FCDBFB2EFC0E1EB6" author="Forel A." box="[865,1004,1161,1188]" pageId="24" pageNumber="25" pagination="306 - 351" refId="ref10286" refString="Forel A. 1913. Formicides du Congo Belge. Recoltes par MM. Bequaert, Luja, etc. Revue de Zoologie Africaine 2: 306 - 351." type="journal article" year="1913">Forel, 1913</bibRefCitation>
)
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</heading>
</paragraph>
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<subSubSection id="948D6EBA7C71FA69FD14FB0AFC671ED5" box="[686,901,1197,1223]" pageId="24" pageNumber="25" type="description">
<paragraph id="DC283D317C71FA69FD14FB0AFC671ED5" blockId="24.[575,1012,1161,1223]" box="[686,901,1197,1223]" pageId="24" pageNumber="25">
<figureCitation id="44AC21B47C71FA69FD14FB0AFCF91ED5" box="[686,795,1197,1223]" captionStart="Fig" captionStartId="13.[189,232,1507,1533]" captionTargetBox="[189,1399,1124,1472]" captionTargetId="figure@13.[189,1399,1124,1472]" captionTargetPageId="13" captionText="Fig. 12. Head, mesosoma, and waist segments in profile. A. Nesomyrmex denticulatus (Mayr, 1901) (CASENT0914923). B. N. innocens (Forel, 1913) (CASENT0906195)." figureDoi="http://doi.org/10.5281/zenodo.3776268" httpUri="https://zenodo.org/record/3776268/files/figure.png" pageId="24" pageNumber="25">Figs 12B</figureCitation>
,
<figureCitation id="44AC21B47C71FA69FC9DFB0AFCBE1ED5" box="[807,860,1197,1223]" captionStart="Fig" captionStartId="13.[189,232,1964,1990]" captionTargetBox="[189,1399,1595,1941]" captionTargetId="figure@13.[189,1399,1595,1941]" captionTargetPageId="13" captionText="Fig. 13. Mesosoma and waist segments in profile. A. Nesomyrmex innocens (Forel, 1913) (CASENT0908995). B. N. stramineus (Arnold, 1948) (CASENT0922011)." figureDoi="http://doi.org/10.5281/zenodo.3776270" httpUri="https://zenodo.org/record/3776270/files/figure.png" pageId="24" pageNumber="25">13A</figureCitation>
,
<figureCitation id="44AC21B47C71FA69FCD2FB0AFC671ED5" box="[872,901,1197,1223]" captionStart="Fig" captionStartId="25.[189,232,1928,1954]" captionTargetBox="[204,1380,272,1902]" captionTargetId="figure@25.[204,1380,272,1902]" captionTargetPageId="25" captionText="Fig. 19. Nesomyrmex innocens (Forel, 1913) (CASENT0906195). A. Body in profile view. B. Body in dorsal view. C. Head in full-face view. D. Map of Africa and Madagascar showing currently known distribution." figureDoi="http://doi.org/10.5281/zenodo.3776282" httpUri="https://zenodo.org/record/3776282/files/figure.png" pageId="24" pageNumber="25">19</figureCitation>
</paragraph>
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<subSubSection id="948D6EBA7C71FA69FF07FB48FBE01F7F" pageId="24" pageNumber="25" type="reference_group">
<paragraph id="DC283D317C71FA69FF07FB48FBF01F1A" blockId="24.[189,1042,1262,1289]" box="[189,1042,1262,1289]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="FC871A1F7C71FA69FF07FB48FBF01F1A" box="[189,1042,1262,1289]" pageId="24" pageNumber="25">
<treatmentCitation id="5D361B207C71FA69FF07FB48FCDB1F1A" author="Forel A." box="[189,825,1262,1289]" page="317" pageId="24" pageNumber="25" year="1913">
<taxonomicName id="1B9746B27C71FA69FF07FB48FCDB1F1A" ID-CoL="CMV5M" authority="Forel, 1913: 317" authorityName="Forel" authorityPageNumber="317" authorityYear="1913" box="[189,825,1262,1289]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="innocens" subGenus="Leptothorax">
<emphasis id="EEE3E1237C71FA69FF07FB48FEB71F1A" box="[189,341,1263,1288]" italics="true" pageId="24" pageNumber="25">Tetramorium</emphasis>
(
<emphasis id="EEE3E1237C71FA69FEDCFB49FE171F1A" box="[358,501,1262,1288]" italics="true" pageId="24" pageNumber="25">Leptothorax</emphasis>
)
<emphasis id="EEE3E1237C71FA69FDBDFB48FD8D1F1A" box="[519,623,1263,1288]" italics="true" pageId="24" pageNumber="25">innocens</emphasis>
<bibRefCitation id="B80640C07C71FA69FDCCFB49FCDB1F1A" author="Forel A." box="[630,825,1262,1288]" pageId="24" pageNumber="25" pagination="306 - 351" refId="ref10286" refString="Forel A. 1913. Formicides du Congo Belge. Recoltes par MM. Bequaert, Luja, etc. Revue de Zoologie Africaine 2: 306 - 351." type="journal article" year="1913">Forel, 1913: 317</bibRefCitation>
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(w.), D.R.
<collectingCountry id="A4807DA17C71FA69FC06FB49FBEC1F1A" box="[956,1038,1262,1288]" name="Democratic Republic of the Congo" pageId="24" pageNumber="25">Congo</collectingCountry>
.
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</paragraph>
<paragraph id="DC283D317C71FA69FF07FA97FD7E1F58" blockId="24.[189,1026,1327,1389]" box="[189,668,1327,1354]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="FC871A1F7C71FA69FF07FA97FD7E1F58" box="[189,668,1327,1354]" pageId="24" pageNumber="25">
<taxonomicName id="1B9746B27C71FA69FF07FA97FE5E1F5B" ID-CoL="3TY28" baseAuthorityName="Forel" baseAuthorityYear="1913" box="[189,444,1328,1354]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="innocens">
<emphasis id="EEE3E1237C71FA69FF07FA97FE5E1F5B" box="[189,444,1328,1354]" italics="true" pageId="24" pageNumber="25">Leptothorax innocens</emphasis>
</taxonomicName>
<treatmentCitation id="5D361B207C71FA69FE63FA88FD7B1F58" author="Forel A." box="[473,665,1327,1354]" page="425" pageId="24" pageNumber="25" year="1916">
<bibRefCitation id="B80640C07C71FA69FE63FA88FD7B1F58" author="Forel A." box="[473,665,1327,1354]" pageId="24" pageNumber="25" pagination="397 - 460" refId="ref10316" refString="Forel A. 1916. Fourmis du Congo et d'autres provenances recoltees par MM. Hermann Kohl, Luja, Mayne, etc. Revue Suisse de Zoologie 24: 397 - 460." type="journal article" year="1916">Forel 1916: 425</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="DC283D317C71FA69FF07FAF3FBE01F7F" blockId="24.[189,1026,1327,1389]" box="[189,1026,1363,1389]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="FC871A1F7C71FA69FF07FAF3FBE01F7F" box="[189,1026,1363,1389]" pageId="24" pageNumber="25">
<taxonomicName id="1B9746B27C71FA69FF07FAF3FE261F7F" ID-CoL="CM7YR" baseAuthorityName="Forel" baseAuthorityYear="1913" box="[189,452,1364,1389]" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="innocens">
<emphasis id="EEE3E1237C71FA69FF07FAF3FE261F7F" box="[189,452,1364,1389]" italics="true" pageId="24" pageNumber="25">Nesomyrmex innocens</emphasis>
</taxonomicName>
<treatmentCitation id="5D361B207C71FA69FE5BFAF4FD4D1F7F" author="Bolton B." box="[481,687,1363,1389]" page="272" pageId="24" pageNumber="25" year="2003">
<bibRefCitation id="B80640C07C71FA69FE5BFAF4FD4D1F7F" author="Bolton B." box="[481,687,1363,1389]" pageId="24" pageNumber="25" pagination="1 - 370" refId="ref9875" refString="Bolton B. 2003. Synopsis and classification of Formicidae. Memoirs of the American Entomological Institute 71: 1 - 370." type="journal article" year="2003">
<collectingRegion id="1E53F3D37C71FA69FE5BFAF4FDD31F7F" box="[481,561,1363,1389]" country="United Kingdom" name="Bolton" pageId="24" pageNumber="25">Bolton</collectingRegion>
2003: 272
</bibRefCitation>
</treatmentCitation>
(see also
<treatmentCitation id="5D361B207C71FA69FC99FAF4FC141F7F" author="Bolton B." box="[803,1014,1363,1389]" page="330" pageId="24" pageNumber="25" year="1982">
<bibRefCitation id="B80640C07C71FA69FC99FAF4FC141F7F" author="Bolton B." box="[803,1014,1363,1389]" pageId="24" pageNumber="25" pagination="245 - 307" refId="ref9811" refString="Bolton B. 1982. A revision of six minor genera of Myrmicinae (Hymenoptera: Formicidae) in the Ethiopian zoogeographical region. Bulletin of the British Museum (Natural History), Entomology 43: 245 - 307." type="journal article" year="1982">
<collectingRegion id="1E53F3D37C71FA69FC99FAF4FC911F7F" box="[803,883,1363,1389]" country="United Kingdom" name="Bolton" pageId="24" pageNumber="25">Bolton</collectingRegion>
1982: 330
</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="948D6EBA7C71FA69FF07FA33FD821CA2" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="DC283D317C71FA69FF07FA33FED41FBC" blockId="24.[189,310,1428,1454]" box="[189,310,1428,1454]" pageId="24" pageNumber="25">
<heading id="87608A5D7C71FA69FF07FA33FED41FBC" bold="true" box="[189,310,1428,1454]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="EEE3E1237C71FA69FF07FA33FED41FBC" bold="true" box="[189,310,1428,1454]" pageId="24" pageNumber="25">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DC283D317C71FA69FF07FA64FD821CA2" blockId="24.[189,1399,1475,1712]" pageId="24" pageNumber="25">
The following character combination distinguishes
<taxonomicName id="1B9746B27C71FA69FC9AFA63FC491FCF" baseAuthorityName="Forel" baseAuthorityYear="1913" box="[800,939,1476,1501]" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="innocens">
<emphasis id="EEE3E1237C71FA69FC9AFA63FC491FCF" box="[800,939,1476,1501]" italics="true" pageId="24" pageNumber="25">N. innocens</emphasis>
</taxonomicName>
from the other members of the group: eyes with 79 ommatidia in longest row; in profile mesosomal dorsum with conspicuously impressed metanotal groove; propodeal spines short and thick, elongate-triangular and only weakly longer than their basal width; in profile petiolar node nodiform, appearing approximately as long as high; in dorsal view petiolar node laterally denticulate; subpetiolar process without a long cuticular flange running back to the postpetiolar junction; dorsum of propodeum with standing hairs; first gastral tergite with standing hairs evenly distributed throughout.
</paragraph>
</subSubSection>
<subSubSection id="948D6EBA7C71FA6BFF07F970FCFB1B9D" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25" type="discussion">
<paragraph id="DC283D317C71FA69FF07F970FE251CE0" blockId="24.[189,455,1751,1778]" box="[189,455,1751,1778]" pageId="24" pageNumber="25">
<heading id="87608A5D7C71FA69FF07F970FE251CE0" bold="true" box="[189,455,1751,1778]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="EEE3E1237C71FA69FF07F970FE251CE0" bold="true" box="[189,455,1751,1778]" pageId="24" pageNumber="25">Diagnostic comments</emphasis>
</heading>
</paragraph>
<paragraph id="DC283D317C71FA6BFF07F8A1FCFB1B9D" blockId="24.[189,1399,1798,2000]" lastBlockId="26.[189,1399,267,399]" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25">
As noted in the description of
<taxonomicName id="1B9746B27C71FA69FDB0F8A0FD5A1D32" baseAuthorityName="Mayr" baseAuthorityYear="1901" box="[522,696,1798,1824]" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="EEE3E1237C71FA69FDB0F8A0FD5A1D32" box="[522,696,1798,1824]" italics="true" pageId="24" pageNumber="25">N.denticulatus</emphasis>
</taxonomicName>
, the latter,
<taxonomicName id="1B9746B27C71FA69FC8BF8A0FC5A1D32" baseAuthorityName="Forel" baseAuthorityYear="1913" box="[817,952,1799,1824]" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="innocens">
<emphasis id="EEE3E1237C71FA69FC8BF8A0FC5A1D32" box="[817,952,1799,1824]" italics="true" pageId="24" pageNumber="25">N.innocens</emphasis>
</taxonomicName>
and
<taxonomicName id="1B9746B27C71FA69FC50F8A0FB681D32" baseAuthorityName="Arnold" baseAuthorityYear="1948" box="[1002,1162,1799,1824]" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="stramineus">
<emphasis id="EEE3E1237C71FA69FC50F8A0FB681D32" box="[1002,1162,1799,1824]" italics="true" pageId="24" pageNumber="25">N. stramineus</emphasis>
</taxonomicName>
are morphologically relatively close.
<taxonomicName id="1B9746B27C71FA69FE32F88DFD761D51" baseAuthorityName="Forel" baseAuthorityYear="1913" box="[392,660,1834,1859]" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="innocens">
<emphasis id="EEE3E1237C71FA69FE32F88DFD761D51" box="[392,660,1834,1859]" italics="true" pageId="24" pageNumber="25">Nesomyrmex innocens</emphasis>
</taxonomicName>
and
<taxonomicName id="1B9746B27C71FA69FD6DF88DFC9D1D51" baseAuthorityName="Arnold" baseAuthorityYear="1948" box="[727,895,1834,1859]" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="stramineus">
<emphasis id="EEE3E1237C71FA69FD6DF88DFC9D1D51" box="[727,895,1834,1859]" italics="true" pageId="24" pageNumber="25">N. stramineus</emphasis>
</taxonomicName>
differ from
<taxonomicName id="1B9746B27C71FA69FBA7F88DFB311D51" baseAuthorityName="Mayr" baseAuthorityYear="1901" box="[1053,1235,1833,1859]" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="EEE3E1237C71FA69FBA7F88DFB311D51" box="[1053,1235,1833,1859]" italics="true" pageId="24" pageNumber="25">N. denticulatus</emphasis>
</taxonomicName>
by generally smaller body size, smaller eyes with less ommatidia, and a subpetiolar process without a long cuticular flange running back to the postpetiolar junction. The separation of
<taxonomicName id="1B9746B27C71FA69FC63F8D7FB861D9B" baseAuthorityName="Forel" baseAuthorityYear="1913" box="[985,1124,1904,1929]" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="innocens">
<emphasis id="EEE3E1237C71FA69FC63F8D7FB861D9B" box="[985,1124,1904,1929]" italics="true" pageId="24" pageNumber="25">N. innocens</emphasis>
</taxonomicName>
and
<taxonomicName id="1B9746B27C71FA69FB1AF8D7FAA71D9B" baseAuthorityName="Arnold" baseAuthorityYear="1948" box="[1184,1349,1904,1929]" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="stramineus">
<emphasis id="EEE3E1237C71FA69FB1AF8D7FAA71D9B" box="[1184,1349,1904,1929]" italics="true" pageId="24" pageNumber="25">N. stramineus</emphasis>
</taxonomicName>
is a bit more difficult, as already mentioned by
<bibRefCitation id="B80640C07C71FA69FD0BF834FCBB1DBF" author="Bolton B." box="[689,857,1939,1965]" pageId="24" pageNumber="25" pagination="245 - 307" refId="ref9811" refString="Bolton B. 1982. A revision of six minor genera of Myrmicinae (Hymenoptera: Formicidae) in the Ethiopian zoogeographical region. Bulletin of the British Museum (Natural History), Entomology 43: 245 - 307." type="journal article" year="1982">
<collectingRegion id="1E53F3D37C71FA69FD0BF834FCE31DBF" box="[689,769,1939,1965]" country="United Kingdom" name="Bolton" pageId="24" pageNumber="25">Bolton</collectingRegion>
(1982)
</bibRefCitation>
.
<taxonomicName id="1B9746B27C71FA69FCD9F833FB8B1DBF" baseAuthorityName="Forel" baseAuthorityYear="1913" box="[867,1129,1940,1965]" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="innocens">
<emphasis id="EEE3E1237C71FA69FCD9F833FB8B1DBF" box="[867,1129,1940,1965]" italics="true" pageId="24" pageNumber="25">Nesomyrmex innocens</emphasis>
</taxonomicName>
has shorter and thicker propodeal spines and a lower and thicker petiolar node compared to
<taxonomicName id="1B9746B27C71FA69FC58F810FB671DC2" baseAuthorityName="Arnold" baseAuthorityYear="1948" box="[994,1157,1975,2000]" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="stramineus">
<emphasis id="EEE3E1237C71FA69FC58F810FB671DC2" box="[994,1157,1975,2000]" italics="true" pageId="24" pageNumber="25">N. stramineus</emphasis>
</taxonomicName>
. It is not clear at the moment whether or not these character states are sufficient to maintain their heterospecificity in the long term.
<bibRefCitation id="B80640C07C73FA6BFEB8FE89FE451B5B" author="Bolton B." box="[258,423,302,329]" pageId="26" pageNumber="27" pagination="245 - 307" refId="ref9811" refString="Bolton B. 1982. A revision of six minor genera of Myrmicinae (Hymenoptera: Formicidae) in the Ethiopian zoogeographical region. Bulletin of the British Museum (Natural History), Entomology 43: 245 - 307." type="journal article" year="1982">
<collectingRegion id="1E53F3D37C73FA6BFEB8FE89FEB01B5A" box="[258,338,302,328]" country="United Kingdom" name="Bolton" pageId="26" pageNumber="27">Bolton</collectingRegion>
(1982)
</bibRefCitation>
had some doubts about this, too, and it is possible that they represent geographical varieties of the same species. However, at present, based on the scarcity of the material, especially of
<taxonomicName id="1B9746B27C73FA6BFF07FED1FEA51B9D" baseAuthorityName="Forel" baseAuthorityYear="1913" box="[189,327,374,399]" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="innocens">
<emphasis id="EEE3E1237C73FA6BFF07FED1FEA51B9D" box="[189,327,374,399]" italics="true" pageId="26" pageNumber="27">N. innocens</emphasis>
</taxonomicName>
, we treat them as two different species.
</paragraph>
</subSubSection>
<caption id="88E86DB97C70FA68FF07F82FFEAE1DFB" ID-DOI="http://doi.org/10.5281/zenodo.3776282" ID-Zenodo-Dep="3776282" httpUri="https://zenodo.org/record/3776282/files/figure.png" pageId="25" pageNumber="26" startId="25.[189,232,1928,1954]" targetBox="[204,1380,272,1902]" targetPageId="25">
<paragraph id="DC283D317C70FA68FF07F82FFEAE1DFB" blockId="25.[189,1399,1928,2025]" pageId="25" pageNumber="26">
<emphasis id="EEE3E1237C70FA68FF07F82FFEFE1DB0" bold="true" box="[189,284,1928,1954]" pageId="25" pageNumber="26">Fig. 19.</emphasis>
<taxonomicName id="1B9746B27C70FA68FE9CF82EFD3A1DB1" authority="(Forel, 1913)" baseAuthorityName="Forel" baseAuthorityYear="1913" box="[294,728,1928,1955]" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="innocens">
<emphasis id="EEE3E1237C70FA68FE9CF82EFDD21DB0" box="[294,560,1929,1954]" italics="true" pageId="25" pageNumber="26">Nesomyrmex innocens</emphasis>
(
<bibRefCitation id="B80640C07C70FA68FDF9F82FFD321DB1" author="Forel A." box="[579,720,1928,1955]" pageId="25" pageNumber="26" pagination="306 - 351" refId="ref10286" refString="Forel A. 1913. Formicides du Congo Belge. Recoltes par MM. Bequaert, Luja, etc. Revue de Zoologie Africaine 2: 306 - 351." type="journal article" year="1913">Forel, 1913</bibRefCitation>
)
</taxonomicName>
(CASENT0906195).
<emphasis id="EEE3E1237C70FA68FC5AF82FFC171DB0" bold="true" box="[992,1013,1928,1954]" pageId="25" pageNumber="26">A</emphasis>
. Body in profile view.
<emphasis id="EEE3E1237C70FA68FAA8F82EFAC41DB1" bold="true" box="[1298,1318,1929,1955]" pageId="25" pageNumber="26">B</emphasis>
. Body in dorsal view.
<emphasis id="EEE3E1237C70FA68FEC8F80CFE651DD7" bold="true" box="[370,391,1963,1989]" pageId="25" pageNumber="26">C</emphasis>
. Head in full-face view.
<emphasis id="EEE3E1237C70FA68FD17F80BFD201DD4" bold="true" box="[685,706,1964,1990]" pageId="25" pageNumber="26">D</emphasis>
. Map of Africa and Madagascar showing currently known distribution.
</paragraph>
</caption>
<subSubSection id="948D6EBA7C73FA6BFF07FE11FBC01BED" box="[189,1058,438,511]" pageId="26" pageNumber="27" type="biology_ecology">
<paragraph id="DC283D317C73FA6BFF07FE11FEFE1BC2" blockId="26.[189,284,438,464]" box="[189,284,438,464]" pageId="26" pageNumber="27">
<heading id="87608A5D7C73FA6BFF07FE11FEFE1BC2" bold="true" box="[189,284,438,464]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="EEE3E1237C73FA6BFF07FE11FEFE1BC2" bold="true" box="[189,284,438,464]" pageId="26" pageNumber="27">Biology</emphasis>
</heading>
</paragraph>
<paragraph id="DC283D317C73FA6BFF07FE42FBC01BED" blockId="26.[189,1058,485,511]" box="[189,1058,485,511]" pageId="26" pageNumber="27">
Based on the limited data available,
<taxonomicName id="1B9746B27C73FA6BFDDDFE41FD131BED" baseAuthorityName="Forel" baseAuthorityYear="1913" box="[615,753,486,511]" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="innocens">
<emphasis id="EEE3E1237C73FA6BFDDDFE41FD131BED" box="[615,753,486,511]" italics="true" pageId="26" pageNumber="27">N. innocens</emphasis>
</taxonomicName>
nests in the stem of trees.
</paragraph>
</subSubSection>
<subSubSection id="948D6EBA7C73FA6BFF07FD81FAD3187D" pageId="26" pageNumber="27" type="distribution">
<paragraph id="DC283D317C73FA6BFF07FD81FE0D1852" blockId="26.[189,495,550,577]" box="[189,495,550,577]" pageId="26" pageNumber="27">
<heading id="87608A5D7C73FA6BFF07FD81FE0D1852" bold="true" box="[189,495,550,577]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="EEE3E1237C73FA6BFF07FD81FE0D1852" bold="true" box="[189,495,550,577]" pageId="26" pageNumber="27">Distribution and biology</emphasis>
</heading>
</paragraph>
<paragraph id="DC283D317C73FA6BFF07FDF2FAD3187D" blockId="26.[189,1329,597,623]" box="[189,1329,597,623]" pageId="26" pageNumber="27">
This species is only known from very few specimens, collected from the D.R.
<collectingCountry id="A4807DA17C73FA6BFBECFDF2FB46187D" box="[1110,1188,597,623]" name="Democratic Republic of the Congo" pageId="26" pageNumber="27">Congo</collectingCountry>
and
<collectingCountry id="A4807DA17C73FA6BFB67FDF2FACF187D" box="[1245,1325,597,623]" name="Kenya" pageId="26" pageNumber="27">Kenya</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>