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<document id="7CA5795F85FB1360437C5D935E949030" ID-CLB-Dataset="61597" ID-DOI="10.11646/phytotaxa.548.2.5" ID-GBIF-Dataset="87c7ff87-12aa-4cc1-811c-3e14ff532ec1" ID-ISSN="1179-3163" ID-Zenodo-Dep="6597796" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1653989760487" checkinUser="plazi" docAuthor="Toprak, Zeynep &amp; Yildiz, Kemal" docDate="2022" docId="5437D461FFB53660A7E47757FA0D81DF" docLanguage="en" docName="phytotaxa.548.2.5.pdf" docOrigin="Phytotaxa 548 (2)" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Silene (sect. Sordidae) K. Yildiz &amp; Toprak 2022, sect. nov." docType="treatment" docVersion="6" lastPageNumber="211" masterDocId="A80EAC19FFB9366CA76C7463FFA7865A" masterDocTitle="Conflicting signals in phylogenetic relationships and taxonomy of Anatolian endemic Silene sordida (Caryophyllaceae)" masterLastPageNumber="222" masterPageNumber="199" pageNumber="211" updateTime="1699338365221" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="21203B9A8616FF589F4EAB4CC7D87E2B">Conflicting signals in phylogenetic relationships and taxonomy of Anatolian endemic Silene sordida (Caryophyllaceae)</mods:title>
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<mods:namePart id="EC3D033EC61B88432178975CCAF7D337">Toprak, Zeynep</mods:namePart>
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<mods:affiliation id="AE5E065FB955E4BCB9624E551BBE31E3">University of Dicle, Faculty of Sciences, Department of Molecular Biology and Genetic, Diyarbakır-Turkey &amp; University of Gothenburg, Faculty of Sciences, Department of Biological and Environmental Sciences, Gothenburg-Sweden &amp; ilgimd @ gmail. com; https: // orcid. org / 0000 - 0002 - 2103 - 4141</mods:affiliation>
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<mods:namePart id="BE5CF8E3A07D64777613795643EF8B8A">Yildiz, Kemal</mods:namePart>
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<mods:affiliation id="10CF9DD1A6A41027CA140EC4E4CB3D83">University of Celal Bayar, Faculty of Sciences, Department of Biology, Manisa-Turkey &amp; kemalyil @ gmail. com; https: // orcid. org / 0000 - 0003 - 4753 - 5005</mods:affiliation>
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<mods:date id="996CD0285B166A5F30CA71644C4D7677">2022</mods:date>
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<subSubSection id="948436FCFFB53660A7E47757FD4D8514" box="[136,746,820,847]" pageId="12" pageNumber="211" type="nomenclature">
<paragraph id="DC216577FFB53660A7E47757FD4D8514" blockId="12.[136,746,820,847]" box="[136,746,820,847]" pageId="12" pageNumber="211">
<heading id="8769D21BFFB53660A7E47757FD4D8514" box="[136,746,820,847]" fontSize="11" level="2" pageId="12" pageNumber="211" reason="3">
<taxonomicName id="1B9E1EF4FFB53660A7E47757FDD08514" authority="K. Yildiz &amp; Toprak" authorityName="K. Yildiz &amp; Toprak" authorityYear="2022" box="[136,631,820,847]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="12" pageNumber="211" phylum="Tracheophyta" rank="section" section="Sordidae" status="sect. nov.">
<emphasis id="EEEAB965FFB53660A7E47757FF778515" bold="true" box="[136,208,820,847]" italics="true" pageId="12" pageNumber="211">Silene</emphasis>
section
<emphasis id="EEEAB965FFB53660A6417757FE338515" bold="true" box="[301,404,820,847]" italics="true" pageId="12" pageNumber="211">Sordidae</emphasis>
K. Yıldız &amp; Toprak
</taxonomicName>
,
<emphasis id="EEEAB965FFB53660A5EE7756FD4D8514" box="[642,746,821,846]" italics="true" pageId="12" pageNumber="211">
<taxonomicNameLabel id="F5D9041EFFB53660A5EE7756FD4D8514" box="[642,746,821,846]" pageId="12" pageNumber="211" rank="section">sect. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="948436FCFFB53660A7E4773FFCE3831C" pageId="12" pageNumber="211" type="materials_examined">
<paragraph id="DC216577FFB53660A7E4773FFD8D852B" blockId="12.[136,554,860,882]" box="[136,554,860,882]" pageId="12" pageNumber="211">
<typeStatus id="0325DBD5FFB53660A7E4773FFF1D852B" box="[136,186,860,881]" pageId="12" pageNumber="211">Type</typeStatus>
:—
<taxonomicName id="1B9E1EF4FFB53660A7BA773FFE328528" authority="Hub." authorityName="Huber-Morath &amp; Reese" authorityYear="1943" box="[214,405,860,882]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="12" pageNumber="211" phylum="Tracheophyta" rank="species" species="sordida">
<emphasis id="EEEAB965FFB53660A7BA773FFEF8852B" box="[214,351,860,881]" italics="true" pageId="12" pageNumber="211">Silene sordida</emphasis>
Hub.
</taxonomicName>
-Mor. &amp; Reese.
</paragraph>
<paragraph id="DC216577FFB53660A7E477C3FC70827C" blockId="12.[136,1452,928,1351]" pageId="12" pageNumber="211">
Diagnosis:—Stem up to
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tall, basal leaves 2542
<emphasis id="EEEAB965FFB53660A5B277C2FD5685E0" box="[734,753,929,954]" italics="true" pageId="12" pageNumber="211">×</emphasis>
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, inflorescence very lax dichasium, cyme, calyx teeth thin and tall, 2.54.0 mm, petals pink to whitish, coronal scales
<quantity id="1B66C892FFB53660A43177A7FC1C8585" box="[861,955,964,991]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="12" pageNumber="211" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
long, obliquely bidentate acute, anthophore
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long, slightly rough, glabrous, seed globose to subglobose, surface flat, rounded at back, testa cells margin undulating, without tubercles, hilum prominent, pollen pore number 2833.
</paragraph>
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Description:—Annual, stem erect, glandular pubescent. Basal leaves spathulate to oblanceolate, cauline leaves ellipsoid to lanceolate; inflorescence a lax dichasial cyme; calyx nerves inconspicuous, crisped unicellular and sparsely glandular pubescent, inflated in fruit; alar flowers erect, distinctly pedicellate, petals limb white to pinkish, rectangular, limb apex approximately
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emarginate, coronal scales
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long; capsule ovoidspheroid, included in the calyx; anthophore
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glabrous; seeds globose to subglobose, hilum prominent. Pore number of pollen grains 2833.
</paragraph>
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Distribution:—Southwest
<collectingCountry id="A48925E7FFB53660A68F716BFDE08378" box="[483,583,1288,1314]" name="Turkey" pageId="12" pageNumber="211">Anatolia</collectingCountry>
.
</paragraph>
<paragraph id="DC216577FFB53660A7D1714EFCE3831C" blockId="12.[136,1452,928,1351]" box="[189,836,1324,1351]" pageId="12" pageNumber="211">Key to the sections:—A proposed diagnostic key follows:</paragraph>
</subSubSection>
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<key id="A80F3246FFB53660A7E47112FA0C804C" pageId="12" pageNumber="211">
<keyStep id="676A7DD2FFB53660A7E47112FA0C83F8" pageId="12" pageNumber="211">
<paragraph id="DC216577FFB53660A7E47112FA0D83DF" blockId="12.[136,1451,1392,1558]" box="[136,1450,1392,1414]" pageId="12" pageNumber="211">
<keyLead id="676FC642FFB53660A7E47112FA0D83DF" box="[136,1450,1392,1414]" pageId="12" pageNumber="211">
1. Inflorescence a monochasium with occasional dichasial branching at base .............................................................
<emphasis id="EEEAB965FFB53660A2617113FA0D83DF" bold="true" box="[1293,1450,1392,1413]" italics="true" pageId="12" pageNumber="211">Dipterospermae</emphasis>
</keyLead>
</paragraph>
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<keyLead id="676FC642FFB53660A7E471EEFA0C83F8" box="[136,1451,1421,1443]" pageId="12" pageNumber="211">1. Inflorescence very lax or asymmetric or compound dichasium.........................................................................................................2</keyLead>
</paragraph>
</keyStep>
<keyStep id="676A7DD2FFB53660A7E471C9FA0C8386" pageId="12" pageNumber="211">
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<keyLead id="676FC642FFB53660A7E471C9FA0C83E5" box="[136,1451,1450,1472]" pageId="12" pageNumber="211">
2. Anthophore pubescent ...................................................................................................................................................
<taxonomicName id="1B9E1EF4FFB53660A24F71C9FA0C83E5" box="[1315,1451,1450,1471]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="12" pageNumber="207" phylum="Tracheophyta" rank="section" section="Cryptoneurae">
<emphasis id="EEEAB965FFB53660A24F71C9FA0C83E5" bold="true" box="[1315,1451,1450,1471]" italics="true" pageId="12" pageNumber="211">Cryptoneurae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="DC216577FFB53660A7E471A4FA0C8386" blockId="12.[136,1451,1392,1558]" box="[136,1451,1479,1501]" pageId="12" pageNumber="211">
<keyLead id="676FC642FFB53660A7E471A4FA0C8386" box="[136,1451,1479,1501]" pageId="12" pageNumber="211">2. Anthophore glabrous ..........................................................................................................................................................................3</keyLead>
</paragraph>
</keyStep>
<keyStep id="676A7DD2FFB53660A7E47187FA0C804C" pageId="12" pageNumber="211">
<paragraph id="DC216577FFB53660A7E47187FA0D83A3" blockId="12.[136,1451,1392,1558]" box="[136,1450,1508,1530]" pageId="12" pageNumber="211">
<keyLead id="676FC642FFB53660A7E47187FA0D83A3" box="[136,1450,1508,1530]" pageId="12" pageNumber="211">
3. Basal leaves rhombic-spathulate .............................................................................................................................................
<taxonomicName id="1B9E1EF4FFB53660A2317187FA0D83A3" ID-CoL="3564" box="[1373,1450,1508,1529]" class="Magnoliopsida" family="Caryophyllaceae" genus="Atocion" kingdom="Plantae" order="Caryophyllales" pageId="12" pageNumber="211" phylum="Tracheophyta" rank="genus">
<emphasis id="EEEAB965FFB53660A2317187FA0D83A3" bold="true" box="[1373,1450,1508,1529]" italics="true" pageId="12" pageNumber="211">Atocion</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="DC216577FFB53660A7E47262FA0C804C" blockId="12.[136,1451,1392,1558]" box="[136,1451,1536,1558]" pageId="12" pageNumber="211">
<keyLead id="676FC642FFB53660A7E47262FA0C804C" box="[136,1451,1536,1558]" pageId="12" pageNumber="211">
3. Basal leaves spathulate to oblanceolate.................................................................................................................................
<taxonomicName id="1B9E1EF4FFB53660A23F7263FA0C804C" authorityName="K. Yildiz &amp; Toprak" authorityYear="2022" box="[1363,1451,1536,1558]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="12" pageNumber="211" phylum="Tracheophyta" rank="section" section="Sordidae">
<emphasis id="EEEAB965FFB53660A23F7263FA0C804C" bold="true" box="[1363,1451,1536,1558]" italics="true" pageId="12" pageNumber="211">Sordidae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
<paragraph id="DC216577FFB53660A7D17226FBA78005" blockId="12.[189,1024,1604,1631]" box="[189,1024,1604,1631]" pageId="12" pageNumber="211">
Key to the species of sect.
<taxonomicName id="1B9E1EF4FFB53660A6857227FDF78005" authorityName="K. Yildiz &amp; Toprak" authorityYear="2022" box="[489,592,1604,1631]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="12" pageNumber="211" phylum="Tracheophyta" rank="section" section="Sordidae">
<emphasis id="EEEAB965FFB53660A6857227FDF78005" bold="true" box="[489,592,1604,1631]" italics="true" pageId="12" pageNumber="211">Sordidae</emphasis>
</taxonomicName>
:—A proposed diagnostic key follows:
</paragraph>
<key id="A80F3246FFB53660A7E472EAFA0D81DF" pageId="12" pageNumber="211">
<keyStep id="676A7DD2FFB53660A7E472EAFA0C80E1" pageId="12" pageNumber="211">
<paragraph id="DC216577FFB53660A7E472EAFA0C80C5" blockId="12.[136,1451,1673,1926]" box="[136,1451,1673,1695]" pageId="12" pageNumber="211">
<keyLead id="676FC642FFB53660A7E472EAFA0C80C5" box="[136,1451,1673,1695]" pageId="12" pageNumber="211">
1. Calyx
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; anthophore
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.................................................................................................................................
<taxonomicName id="1B9E1EF4FFB53660A25772EAFA0C80C5" ID-CoL="4XCD4" authorityName="Barbey" authorityYear="1886" box="[1339,1451,1673,1695]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="12" pageNumber="211" phylum="Tracheophyta" rank="species" species="insularis">
<emphasis id="EEEAB965FFB53660A25772EAFA0C80C5" bold="true" box="[1339,1451,1673,1695]" italics="true" pageId="12" pageNumber="211">S. insularis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
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<keyLead id="676FC642FFB53660A7E472C5FA0C80E1" box="[136,1451,1702,1724]" pageId="12" pageNumber="211">
1. Calyx more than
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; anthophore more than
<quantity id="1B66C892FFB53660A5CD72C5FD7F80E6" box="[673,728,1702,1724]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="211" unit="mm" value="2.0">2 mm</quantity>
......................................................................................................................2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="676A7DD2FFB53660A7E472A0FA0C8148" pageId="12" pageNumber="211">
<paragraph id="DC216577FFB53660A7E472A0FA0D8082" blockId="12.[136,1451,1673,1926]" box="[136,1450,1730,1753]" pageId="12" pageNumber="211">
<keyLead id="676FC642FFB53660A7E472A0FA0D8082" box="[136,1450,1730,1753]" pageId="12" pageNumber="211">
2. Anthophore glabrous, petal limb apex recessed, hilum region of seed prominently protruding.........................................
<taxonomicName id="1B9E1EF4FFB53660A22472A0FA0D8082" ID-CoL="4XD84" authorityName="Huber-Morath &amp; Reese" authorityYear="1943" box="[1352,1450,1730,1752]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="12" pageNumber="211" phylum="Tracheophyta" rank="species" species="sordida">
<emphasis id="EEEAB965FFB53660A22472A0FA0D8082" bold="true" box="[1352,1450,1730,1752]" italics="true" pageId="12" pageNumber="211">S. sordida</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="DC216577FFB53660A7E47283FA0C8148" blockId="12.[136,1451,1673,1926]" pageId="12" pageNumber="211">
<keyLead id="676FC642FFB53660A7E47283FA0C8148" pageId="12" pageNumber="211">2. Anthophore hairy, petal limb apex not recessed, rhomboid, rounded or emarginate, hilum region of seed circular or rectangular.... ............................................................................................................................................................................................................3</keyLead>
</paragraph>
</keyStep>
<keyStep id="676A7DD2FFB53660A7E4737AFA0C8111" pageId="12" pageNumber="211">
<paragraph id="DC216577FFB53660A7E4737AFA0C8175" blockId="12.[136,1451,1673,1926]" box="[136,1451,1817,1839]" pageId="12" pageNumber="211">
<keyLead id="676FC642FFB53660A7E4737AFA0C8175" box="[136,1451,1817,1839]" pageId="12" pageNumber="211">
3. Petal limb apex rhomboid; hilum region of seed circular ..................................................................................................
<taxonomicName id="1B9E1EF4FFB53660A253737AFA0C8175" ID-CoL="4XC4P" box="[1343,1451,1817,1839]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="12" pageNumber="211" phylum="Tracheophyta" rank="species" species="ertekinii">
<emphasis id="EEEAB965FFB53660A253737AFA0C8175" bold="true" box="[1343,1451,1817,1839]" italics="true" pageId="12" pageNumber="211">S. ertekinii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="DC216577FFB53660A7E47355FA0C8111" blockId="12.[136,1451,1673,1926]" box="[136,1451,1846,1868]" pageId="12" pageNumber="211">
<keyLead id="676FC642FFB53660A7E47355FA0C8111" box="[136,1451,1846,1868]" pageId="12" pageNumber="211">3. Petal limb apex rounded or emarginate; hilum region of seed rectangular or circular ......................................................................4</keyLead>
</paragraph>
</keyStep>
<keyStep id="676A7DD2FFB53660A7E47330FA0D81DF" pageId="12" pageNumber="211">
<paragraph id="DC216577FFB53660A7E47330FA0C8132" blockId="12.[136,1451,1673,1926]" box="[136,1451,1875,1897]" pageId="12" pageNumber="211">
<keyLead id="676FC642FFB53660A7E47330FA0C8132" box="[136,1451,1875,1897]" pageId="12" pageNumber="211">
4. Anthophore at least
<quantity id="1B66C892FFB53660A6F97330FE688132" box="[405,463,1875,1896]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="211" unit="mm" value="4.0">4 mm</quantity>
; hilum region of seed rectangular with two opposite sides twisted ..................................
<taxonomicName id="1B9E1EF4FFB53660A2757330FA0C8132" ID-CoL="4XBYG" authorityName="Stapf" authorityYear="1886" box="[1305,1451,1875,1896]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="12" pageNumber="211" phylum="Tracheophyta" rank="species" species="cryptoneura">
<emphasis id="EEEAB965FFB53660A2757330FA0C8132" bold="true" box="[1305,1451,1875,1896]" italics="true" pageId="12" pageNumber="211">S. cryptoneura</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="DC216577FFB53660A7E47313FA0D81DF" blockId="12.[136,1451,1673,1926]" box="[136,1450,1903,1926]" pageId="12" pageNumber="211">
<keyLead id="676FC642FFB53660A7E47313FA0D81DF" box="[136,1450,1903,1926]" pageId="12" pageNumber="211">
4. Anthophore less than
<quantity id="1B66C892FFB53660A6C97313FE7881DF" box="[421,479,1904,1925]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="211" unit="mm" value="4.0">4 mm</quantity>
; hilum region of seed more or less circular, not twisted..................................................
<taxonomicName id="1B9E1EF4FFB53660A2707313FA0D81DF" ID-CoL="4XD45" authorityName="Pampanini" authorityYear="1927" box="[1308,1450,1903,1925]" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" kingdom="Plantae" order="Caryophyllales" pageId="12" pageNumber="211" phylum="Tracheophyta" rank="species" species="salamandra">
<emphasis id="EEEAB965FFB53660A2707313FA0D81DF" bold="true" box="[1308,1450,1903,1925]" italics="true" pageId="12" pageNumber="211">S. salamandra</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>