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<document id="BFAC5EBB3D0E1726DD7EF1D8C867F8A1" ID="10.11646/zootaxa.4137.4.7" ID-DOI="10.11646/zootaxa.4137.4.7" ID-GBIF-Dataset="7e39308a-d9a3-45f0-9b7d-d015d1666762" ID-ISSN="1175-5326" ID-Zenodo-Dep="271953" ID-ZooBank="F5BB50FE-28CF-4225-8E82-B47BEFCE6465" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1468394896375" checkinUser="plazi" docAuthor="Häckel, Martin &amp; Azadbakhsh, Saeed" docDate="2016" docId="5432D01CFF8EB448FF68BF76FA7117A6" docLanguage="en" docName="zootaxa.4137.4.7.pdf" docOrigin="Zootaxa 4137 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Microcosmodes cruciatus" docType="key" docVersion="7" lastPageNumber="559" masterDocId="A80BA864FF88B44EFFFFBB44FFED121C" masterDocTitle="Two new species of Microcosmodes Strand (Coleoptera: Carabidae: Panagaenini) from Oman and Iran" masterLastPageNumber="560" masterPageNumber="553" pageNumber="559" updateTime="1698364666443" updateUser="plazi">
<mods:mods id="B90D84330CDBC71FE68A19337ECA21E2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4E2B0FDDDADA989E55EDD00D20FD23EE">
<mods:title id="61C49FBB88748E056207ED5802C46E4B">Two new species of Microcosmodes Strand (Coleoptera: Carabidae: Panagaenini) from Oman and Iran</mods:title>
</mods:titleInfo>
<mods:name id="9890C7F34B028BCDD51C32379E72D1DE" type="personal">
<mods:role id="1782FE7BE2C23E034A647809AFC0B0CE">
<mods:roleTerm id="F95C1544E66D9A8EEAB9172FE0208341">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1C80695AEDEF39071F82621BADB373E8">Häckel, Martin</mods:namePart>
</mods:name>
<mods:name id="4F4789AF6E114F152BAC307271459164" type="personal">
<mods:role id="AE49FC80C6193EA3FBD65E59CB04D409">
<mods:roleTerm id="3D5DF68457AD4968C800D44948D30179">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="613F4B5980832B7B525C23E4CC6A4A1A">Azadbakhsh, Saeed</mods:namePart>
</mods:name>
<mods:typeOfResource id="E5279AE5F4AC1509A81D06B712AB5673">text</mods:typeOfResource>
<mods:relatedItem id="A4B3C3954779B95C4A5813E4C0CC5CE1" type="host">
<mods:titleInfo id="C7F4EDB1684CEFBC8E55D2208754C23A">
<mods:title id="BC8F183677E7B0F5D5EFE3763E06BA4D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="BC28EBE8A1EA1270F9CB68C45722574E">
<mods:date id="BD99BD9295A93E38E098E2E4ABB379A7">2016</mods:date>
<mods:detail id="23AA149BD08B4D56C7A4C7F06DCC1BC0" type="volume">
<mods:number id="5944BF6D2B88585B3259BB1A00DC87A3">4137</mods:number>
</mods:detail>
<mods:detail id="D05E7B5D0494080DCF96D5FA93851EA0" type="issue">
<mods:number id="86177A04EBF06E7BE4DCBA6399A57D56">4</mods:number>
</mods:detail>
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<mods:start id="51AFFF2AA757C9EB875A096D09E76316">553</mods:start>
<mods:end id="3671CC2D1B3016E083FE3C6D37B621DD">560</mods:end>
</mods:extent>
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<mods:classification id="69453060ED5A571F4E55618911C52032">journal article</mods:classification>
<mods:identifier id="951D99239FDDBF448829E7377EC2A341" type="DOI">10.11646/zootaxa.4137.4.7</mods:identifier>
<mods:identifier id="25DCD6F86D8DC63CEC2B1D381930989B" type="GBIF-Dataset">7e39308a-d9a3-45f0-9b7d-d015d1666762</mods:identifier>
<mods:identifier id="5F1B3C76C509DD6DDE3C8CF5064A30A4" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="28F143366D023456EF7ABBF3D75EB969" type="Zenodo-Dep">271953</mods:identifier>
<mods:identifier id="146B0EF9CB0BA82EF959C1A6F9912092" type="ZooBank">F5BB50FE-28CF-4225-8E82-B47BEFCE6465</mods:identifier>
</mods:mods>
<treatment id="5432D01CFF8EB448FF68BF76FA7117A6" ID-DOI="http://doi.org/10.5281/zenodo.6067597" ID-GBIF-Taxon="123195305" ID-Zenodo-Dep="6067597" LSID="urn:lsid:plazi:treatment:5432D01CFF8EB448FF68BF76FA7117A6" httpUri="http://treatment.plazi.org/id/5432D01CFF8EB448FF68BF76FA7117A6" lastPageNumber="559" pageId="6" pageNumber="559">
<subSubSection id="94813281FF8EB448FF68BF76FD6F1671" pageId="6" pageNumber="559" type="nomenclature">
<paragraph id="DC24610AFF8EB448FF68BF76FDB51650" blockId="6.[151,642,1074,1133]" box="[151,600,1074,1100]" pageId="6" pageNumber="559">
<heading id="876CD666FF8EB448FF68BF76FDB51650" bold="true" box="[151,600,1074,1100]" fontSize="11" level="1" pageId="6" pageNumber="559" reason="1">
<emphasis id="EEEFBD18FF8EB448FF68BF76FDB51650" bold="true" box="[151,600,1074,1100]" pageId="6" pageNumber="559">
Key to species of
<taxonomicName id="1B9B1A89FF8EB448FE92BF76FDEB1650" ID-CoL="42TZZ" box="[365,518,1074,1100]" class="Insecta" family="Carabidae" genus="Microcosmodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="559" phylum="Arthropoda" rank="species" species="cruciatus">
<emphasis id="EEEFBD18FF8EB448FE92BF76FDEB1650" bold="true" box="[365,518,1074,1100]" italics="true" pageId="6" pageNumber="559">M. cruciatus</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
<paragraph id="DC24610AFF8EB448FF68BF11FD6F1671" blockId="6.[151,642,1074,1133]" box="[151,642,1108,1133]" pageId="6" pageNumber="559">
(except continental Africa and
<collectingCountry id="A48C219AFF8EB448FE0DBF10FD961671" box="[498,635,1108,1133]" name="Madagascar" pageId="6" pageNumber="559">Madagascar</collectingCountry>
)
</paragraph>
</subSubSection>
<subSubSection id="94813281FF8EB448FF68BFD8FA7117A6" pageId="6" pageNumber="559" type="key">
<keyStep id="676F79AFFF8EB448FF68BFD8FA7117A6" pageId="6" pageNumber="559">
<paragraph id="DC24610AFF8EB448FF68BFD8FA71173B" blockId="6.[151,1436,1180,1466]" pageId="6" pageNumber="559">
<keyLead id="676AC23FFF8EB448FF68BFD8FA71173B" pageId="6" pageNumber="559">
1 Body ovoid, pronotum with anterior margin markedly narrower than base, pronotal base slightly narrower than elytral base. Elytron always with two orange maculae, both humeral and apical macula extensive and fully developed, with intervals colored; humeral macula semicircular, laterally widened and covering entire basal third of elytron to humerus; apical macula nearly circular, reaching part of elytral margin. Black reminder of elytra forms pattern of Maltese cross (
<figureCitation id="44A07D8FFF8EB448FB35BFB0FAEF1716" box="[1226,1282,1268,1290]" captionStart="FIGURE 1" captionStartId="2.[151,250,1412,1434]" captionTargetBox="[224,1367,331,1373]" captionTargetId="figure@2.[199,1387,331,1387]" captionTargetPageId="2" captionText="FIGURE 1 Microcosmodes arabicus sp. nov., HT (male): a. dorsal habitus; b. aedeagus, lateral view; c. aedeagus, dorsal view to apex." httpUri="https://zenodo.org/record/271954/files/figure.png" pageId="6" pageNumber="559">Fig. 1</figureCitation>
,
<figureCitation id="44A07D8FFF8EB448FAF0BFB0FAA41716" box="[1295,1353,1268,1290]" captionStart="FIGURES 4 – 11" captionStartId="5.[151,264,1212,1235]" captionTargetBox="[165,1421,193,1189]" captionTargetId="figure@5.[149,1433,193,1189]" captionTargetPageId="5" captionText="FIGURES 4 – 11. Microcosmodes cruciatus group, dorsal habitus (Scale bar = 5 mm): FIg. 4. Microcosmodes arabicus sp. nov., HT (male). FIg. 5. Microcosmodes cruciatus (Dejean, 1831), HT (male, MNHN). FIg. 6. Microcosmodes persicus sp. nov., HT (male). FIg. 7. Microcosmodes persicus sp. nov., PT, female. FIg. 8. Microcosmodes persicus sp. nov., PT, female. FIg. 9. Microcosmodes persicus sp. nov., PT, female. FIg. 10. Microcosmodes flavopilosus (Laferté-Sénectere, 1851), male, Pakistan (Sind). FIg. 11. Microcosmodes laetus (Dejean, 1831), male, Sierra Leone (det. by Alluaud, MNHN)." httpUri="https://zenodo.org/record/271957/files/figure.png" pageId="6" pageNumber="559">Fig. 4</figureCitation>
).
<collectingCountry id="A48C219AFF8EB448FAA4BFB0FA741716" box="[1371,1433,1268,1290]" name="Oman" pageId="6" pageNumber="559">Oman</collectingCountry>
,
<collectingCountry id="A48C219AFF8EB448FF1EBE56FEC9173B" box="[225,292,1298,1319]" name="Yemen" pageId="6" pageNumber="559">Yemen</collectingCountry>
..............................................................................
<taxonomicName id="1B9B1A89FF8EB448FB2ABE56FAA4173B" ID-CoL="73G4D" box="[1237,1353,1297,1319]" class="Insecta" family="Carabidae" genus="Microcosmodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="559" phylum="Arthropoda" rank="species" species="arabicus" status="sp. nov.">
<emphasis id="EEEFBD18FF8EB448FB2ABE56FAA4173B" box="[1237,1353,1297,1319]" italics="true" pageId="6" pageNumber="559">M. arabicus</emphasis>
</taxonomicName>
<emphasis id="EEEFBD18FF8EB448FAB0BE55FA71173B" bold="true" box="[1359,1436,1297,1319]" pageId="6" pageNumber="559">
<taxonomicNameLabel id="F5DC0063FF8EB448FAB0BE55FA71173B" box="[1359,1436,1297,1319]" pageId="6" pageNumber="559" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="DC24610AFF8EB448FF68BE6BFA7117A6" blockId="6.[151,1436,1180,1466]" pageId="6" pageNumber="559">
<keyLead id="676AC23FFF8EB448FF68BE6BFA7117A6" pageId="6" pageNumber="559">
- Body subparallel, pronotum with anterior margin slightly narrower than base, pronotal base markedly narrower than elytral base. Elytron rarely with two orange maculae, more often maculae absent (
<figureCitation id="44A07D8FFF8EB448FC3ABE08FBEC177E" box="[965,1025,1356,1378]" captionStart="FIGURE 2" captionStartId="3.[151,250,1209,1231]" captionTargetBox="[224,1368,187,1174]" captionTargetId="figure@3.[204,1384,187,1188]" captionTargetPageId="3" captionText="FIGURE 2 Microcosmodes persicus sp. nov., HT (male): a. dorsal habitus; b. aedeagus, lateral view; c. aedeagus, dorsal view to apex." httpUri="https://zenodo.org/record/271955/files/figure.png" pageId="6" pageNumber="559">Fig. 2</figureCitation>
,
<figureCitation id="44A07D8FFF8EB448FBF1BE08FB8B177E" box="[1038,1126,1356,1378]" captionStart="FIGURES 4 – 11" captionStartId="5.[151,264,1212,1235]" captionTargetBox="[165,1421,193,1189]" captionTargetId="figure@5.[149,1433,193,1189]" captionTargetPageId="5" captionText="FIGURES 4 – 11. Microcosmodes cruciatus group, dorsal habitus (Scale bar = 5 mm): FIg. 4. Microcosmodes arabicus sp. nov., HT (male). FIg. 5. Microcosmodes cruciatus (Dejean, 1831), HT (male, MNHN). FIg. 6. Microcosmodes persicus sp. nov., HT (male). FIg. 7. Microcosmodes persicus sp. nov., PT, female. FIg. 8. Microcosmodes persicus sp. nov., PT, female. FIg. 9. Microcosmodes persicus sp. nov., PT, female. FIg. 10. Microcosmodes flavopilosus (Laferté-Sénectere, 1851), male, Pakistan (Sind). FIg. 11. Microcosmodes laetus (Dejean, 1831), male, Sierra Leone (det. by Alluaud, MNHN)." httpUri="https://zenodo.org/record/271957/files/figure.png" pageId="6" pageNumber="559">Figs 6–9</figureCitation>
); if maculae present then much reduced, often to minor humeral residua (
<figureCitation id="44A07D8FFF8EB448FD8FBE2DFD411763" box="[624,684,1385,1407]" captionStart="FIGURES 4 – 11" captionStartId="5.[151,264,1212,1235]" captionTargetBox="[165,1421,193,1189]" captionTargetId="figure@5.[149,1433,193,1189]" captionTargetPageId="5" captionText="FIGURES 4 – 11. Microcosmodes cruciatus group, dorsal habitus (Scale bar = 5 mm): FIg. 4. Microcosmodes arabicus sp. nov., HT (male). FIg. 5. Microcosmodes cruciatus (Dejean, 1831), HT (male, MNHN). FIg. 6. Microcosmodes persicus sp. nov., HT (male). FIg. 7. Microcosmodes persicus sp. nov., PT, female. FIg. 8. Microcosmodes persicus sp. nov., PT, female. FIg. 9. Microcosmodes persicus sp. nov., PT, female. FIg. 10. Microcosmodes flavopilosus (Laferté-Sénectere, 1851), male, Pakistan (Sind). FIg. 11. Microcosmodes laetus (Dejean, 1831), male, Sierra Leone (det. by Alluaud, MNHN)." httpUri="https://zenodo.org/record/271957/files/figure.png" pageId="6" pageNumber="559">Fig. 7</figureCitation>
); if both maculae are present then humeral reaches only a small part of elytral margin, never close to humerus, and being more transverse than semicircular; apical macula never reaching elytral margin (
<figureCitation id="44A07D8FFF8EB448FF17BEE0FECE17A6" box="[232,291,1444,1466]" captionStart="FIGURE 2" captionStartId="3.[151,250,1209,1231]" captionTargetBox="[224,1368,187,1174]" captionTargetId="figure@3.[204,1384,187,1188]" captionTargetPageId="3" captionText="FIGURE 2 Microcosmodes persicus sp. nov., HT (male): a. dorsal habitus; b. aedeagus, lateral view; c. aedeagus, dorsal view to apex." httpUri="https://zenodo.org/record/271955/files/figure.png" pageId="6" pageNumber="559">Fig. 2</figureCitation>
,
<figureCitation id="44A07D8FFF8EB448FED2BEE0FE8B17A6" box="[301,358,1444,1466]" captionStart="FIGURES 4 – 11" captionStartId="5.[151,264,1212,1235]" captionTargetBox="[165,1421,193,1189]" captionTargetId="figure@5.[149,1433,193,1189]" captionTargetPageId="5" captionText="FIGURES 4 – 11. Microcosmodes cruciatus group, dorsal habitus (Scale bar = 5 mm): FIg. 4. Microcosmodes arabicus sp. nov., HT (male). FIg. 5. Microcosmodes cruciatus (Dejean, 1831), HT (male, MNHN). FIg. 6. Microcosmodes persicus sp. nov., HT (male). FIg. 7. Microcosmodes persicus sp. nov., PT, female. FIg. 8. Microcosmodes persicus sp. nov., PT, female. FIg. 9. Microcosmodes persicus sp. nov., PT, female. FIg. 10. Microcosmodes flavopilosus (Laferté-Sénectere, 1851), male, Pakistan (Sind). FIg. 11. Microcosmodes laetus (Dejean, 1831), male, Sierra Leone (det. by Alluaud, MNHN)." httpUri="https://zenodo.org/record/271957/files/figure.png" pageId="6" pageNumber="559">Fig. 8</figureCitation>
).
<collectingCountry id="A48C219AFF8EB448FE86BEE0FE4D17A5" box="[377,416,1444,1465]" name="Iran" pageId="6" pageNumber="559">Iran</collectingCountry>
....................................................................
<taxonomicName id="1B9B1A89FF8EB448FB26BEE0FAA517A5" box="[1241,1352,1444,1465]" class="Insecta" family="Carabidae" genus="Microcosmodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="559" phylum="Arthropoda" rank="species" species="perscius" status="sp. nov.">
<emphasis id="EEEFBD18FF8EB448FB26BEE0FAA517A5" box="[1241,1352,1444,1465]" italics="true" pageId="6" pageNumber="559">M. perscius</emphasis>
</taxonomicName>
<emphasis id="EEEFBD18FF8EB448FAB0BEE0FA7117A6" bold="true" box="[1359,1436,1444,1466]" pageId="6" pageNumber="559">
<taxonomicNameLabel id="F5DC0063FF8EB448FAB0BEE0FA7117A6" box="[1359,1436,1444,1466]" pageId="6" pageNumber="559" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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