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<document id="257C372B2A54307DC842A7E228401BCD" ID-CLB-Dataset="299942" ID-DOI="10.11646/zootaxa.5481.1.5" ID-GBIF-Dataset="0423a610-a41f-4de4-9d39-289e7ec7168f" ID-ISSN="1175-5326" ID-Zenodo-Dep="12742641" ID-ZooBank="987E3EFF-7867-4186-A26A-79359DF762C8" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1721033360048" checkinUser="plazi" docAuthor="Wimolsuthikul, Chaipat, Yamane, Seiki, Samung, Yudthana &amp; Jaitrong, Weeyawat" docDate="2024" docId="542287ABFFBD412B2CD641E849DCF9AA" docLanguage="en" docName="zootaxa.5481.1.5.pdf" docOrigin="Zootaxa 5481 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5481.1.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Carebara pygmaea" docType="treatment" docVersion="4" lastPageNumber="81" masterDocId="A81BFFD3FFBC41292C4144224C40FF9D" masterDocTitle="A revision of the Carebara pygmaea (Emery, 1887) species group (Hymenoptera: Formicidae) in Thailand" masterLastPageNumber="100" masterPageNumber="79" pageNumber="80" updateTime="1732492712269" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3CA450B5277420276053A0F0E213A8CB">A revision of the Carebara pygmaea (Emery, 1887) species group (Hymenoptera: Formicidae) in Thailand</mods:title>
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<mods:namePart id="0A09EA7A4FD4FFD97DC3B1469B3339AC">Wimolsuthikul, Chaipat</mods:namePart>
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<mods:namePart id="0162E50EBDCA42FABB320B295AD8A8D4">Yamane, Seiki</mods:namePart>
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<mods:affiliation id="79E504AB4FB1429C0015770E7DAFA7BC">Haruyama-chô 1054 - 1, Kagoshima-shi, 899 - 2704 Japan.</mods:affiliation>
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<mods:namePart id="2A5AE6B61E75FFA8F61A8A56A72D2054">Samung, Yudthana</mods:namePart>
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<mods:affiliation id="37A024D19E978AE0F032529E6B47617C">Khlong &amp; Department of Medical Entomology, Faculty of Tropical Medicine, Mahidol University, Bangkok 10400, Thailand.</mods:affiliation>
<mods:nameIdentifier id="64990A12EF0CA99AF30940E560A07F8C" type="email">yudthana.sam@mahidol.ac.th</mods:nameIdentifier>
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<mods:namePart id="0B8E4357F6BC2746FB67B55549CBD541">Jaitrong, Weeyawat</mods:namePart>
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<mods:affiliation id="3DFFE9B0C4828B92ABD809B12BB03988">Office of Natural Science Research, National Science Museum, 39, Moo &amp; Office of Natural Science Research, National Science Museum, 39, Moo &amp; Khlong Luang, Pathum Thani, 12120 Thailand.</mods:affiliation>
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<treatment id="542287ABFFBD412B2CD641E849DCF9AA" ID-DOI="http://doi.org/10.5281/zenodo.12751219" ID-GBIF-Taxon="235587386" ID-Zenodo-Dep="12751219" LSID="urn:lsid:plazi:treatment:542287ABFFBD412B2CD641E849DCF9AA" httpUri="http://treatment.plazi.org/id/542287ABFFBD412B2CD641E849DCF9AA" lastPageId="2" lastPageNumber="81" pageId="1" pageNumber="80" scope_family="Formicidae" scope_order="Hymenoptera">
<subSubSection id="94916536FFBD41282CD641E84E59FA78" box="[151,537,1482,1509]" pageId="1" pageNumber="80" type="nomenclature">
<paragraph id="DC3436BDFFBD41282CD641E84E59FA78" blockId="1.[151,537,1482,1509]" box="[151,537,1482,1509]" pageId="1" pageNumber="80">
<heading id="877C81D1FFBD41282CD641E84E59FA78" bold="true" box="[151,537,1482,1509]" fontSize="11" level="1" pageId="1" pageNumber="80" reason="1">
<emphasis id="EEFFEAAFFFBD41282CD641E84E59FA78" bold="true" box="[151,537,1482,1509]" pageId="1" pageNumber="80">
<taxonomicName id="1B8B4D3EFFBD41282CD641E84D32FA78" ID-CoL="CLGD8" authority="(Emery, 1887)" baseAuthorityName="Emery" baseAuthorityYear="1887" box="[151,370,1482,1509]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="80" phylum="Arthropoda" rank="species" species="pygmaea">
<emphasis id="EEFFEAAFFFBD41282CD641E84D32FA78" bold="true" box="[151,370,1482,1509]" italics="true" pageId="1" pageNumber="80">Carebara pygmaea</emphasis>
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species group
</emphasis>
</heading>
</paragraph>
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<subSubSection id="94916536FFBD412B2CD642304FA3FE30" lastPageId="2" lastPageNumber="81" pageId="1" pageNumber="80" type="diagnosis">
<paragraph id="DC3436BDFFBD41282CD64230492EF824" blockId="1.[151,1437,1554,1977]" pageId="1" pageNumber="80">
<emphasis id="EEFFEAAFFFBD41282CD642304D52F9B1" bold="true" box="[151,274,1554,1580]" pageId="1" pageNumber="80">Diagnosis.</emphasis>
Dimorphic.
<emphasis id="EEFFEAAFFFBD41282DDE42304DB0F9B1" bold="true" box="[415,496,1554,1580]" pageId="1" pageNumber="80">Major.</emphasis>
Frons with longitudinal striation; posterior portion of head in full-face view usually with transverse striation (except for
<taxonomicName id="1B8B4D3EFFBD41282E7A42154E96F9CC" authorityName="Matsuura &amp; Hosoishi" authorityYear="2024" box="[571,726,1591,1617]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="80" phylum="Arthropoda" rank="species" species="bokorensis">
<emphasis id="EEFFEAAFFFBD41282E7A42154E96F9CC" box="[571,726,1591,1617]" italics="true" pageId="1" pageNumber="80">C. bokorensis</emphasis>
</taxonomicName>
lacking this striation); vertex without a pair of horns; frontal carinae with three pairs of erect hairs; mandible short, subtriangular, masticatory margin with five teeth (including large apical tooth); promesonotum in profile view distinctly convex to form a dome; promesonotal suture indistinct; metanotal groove present; propodeum distinctly lower than promesonotum; propodeal spine short, usually shorter than width at base; petiole in profile, with long anterior peduncle; postpetiole clearly shorter than petiole; gastral tergite I clearly larger than remaining segments combined.
<emphasis id="EEFFEAAFFFBD41282F5B42C84F2BF899" bold="true" box="[794,875,1770,1796]" pageId="1" pageNumber="80">Minor.</emphasis>
Head small, smooth and shiny; mandible relatively short, subtriangular, masticatory margin with five teeth (including large apical tooth); eye very small, present as a dot; promesonotum in profile view convex, smooth and shiny; promesonotal suture absent; metanotal groove distinct; propodeal spine absent; petiole with long anterior peduncle; postpetiole clearly shorter than petiole; gastral tergite I clearly larger than remaining segments combined.
<emphasis id="EEFFEAAFFFBD41282F7343584FB2F808" bold="true" box="[818,1010,1914,1941]" pageId="1" pageNumber="80">Ergatoid queen.</emphasis>
Morphological character is similar to major worker in having large head but mesosoma is slightly larger than in major worker and ocelli are present.
</paragraph>
<paragraph id="DC3436BDFFBE412B2C8644B54FA9FEDC" blockId="2.[151,1437,151,501]" pageId="2" pageNumber="81">
We placed
<taxonomicName id="1B8B4D3EFFBE412B2D0444B54D9FFF2C" authorityName="Matsuura &amp; Hosoishi" authorityYear="2024" box="[325,479,151,177]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="bokorensis">
<emphasis id="EEFFEAAFFFBE412B2D0444B54D9FFF2C" box="[325,479,151,177]" italics="true" pageId="2" pageNumber="81">C. bokorensis</emphasis>
</taxonomicName>
under the
<taxonomicName id="1B8B4D3EFFBE412B2E1A44B54EA1FF2D" baseAuthorityName="Emery" baseAuthorityYear="1887" box="[603,737,151,177]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="pygmaea">
<emphasis id="EEFFEAAFFFBE412B2E1A44B54EA1FF2D" box="[603,737,151,177]" italics="true" pageId="2" pageNumber="81">C. pygmaea</emphasis>
</taxonomicName>
species group because almost all characters of the minor and the major workers agreed well with the characteristics of the species group. The body sculptures and the condition of the hairs can be used to distinguish the species within the species group. We have tried to extract DNA using dry specimens but were unsuccessful using the PCR process. In future work, DNA barcoding should help to distinguish members of the
<taxonomicName id="1B8B4D3EFFBE412B2D0D45054D92FEDD" baseAuthorityName="Emery" baseAuthorityYear="1887" box="[332,466,295,321]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="pygmaea">
<emphasis id="EEFFEAAFFFBE412B2D0D45054D92FEDD" box="[332,466,295,321]" italics="true" pageId="2" pageNumber="81">C. pygmaea</emphasis>
</taxonomicName>
species group when obtaining fresh specimens.
</paragraph>
<paragraph id="DC3436BDFFBE412B2C8645684FA3FE30" blockId="2.[151,1437,151,501]" pageId="2" pageNumber="81">
<emphasis id="EEFFEAAFFFBE412B2C8645684F95FEF9" bold="true" box="[199,981,330,357]" pageId="2" pageNumber="81">
Currently, valid names included in the species group of
<collectingCountry id="A49C762DFFBE412B2F2045684F90FEF9" box="[865,976,330,356]" name="Thailand" pageId="2" pageNumber="81">Thailand</collectingCountry>
.
</emphasis>
<taxonomicName id="1B8B4D3EFFBE412B2F9C456948E1FEF8" authorityName="Wimolsuthikul &amp; Yamane &amp; Samung &amp; Jaitrong" authorityYear="2024" box="[989,1185,331,357]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="adisaki" status="sp. nov.">
<emphasis id="EEFFEAAFFFBE412B2F9C456948E1FEF8" box="[989,1185,331,357]" italics="true" pageId="2" pageNumber="81">Carebara adisaki</emphasis>
</taxonomicName>
<emphasis id="EEFFEAAFFFBE412B28E945694942FEF8" bold="true" box="[1192,1282,331,357]" pageId="2" pageNumber="81">
<taxonomicNameLabel id="F5CC57D4FFBE412B28E945694942FEF8" box="[1192,1282,331,357]" pageId="2" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<taxonomicName id="1B8B4D3EFFBE412B295045694933FEF9" authorityName="Wimolsuthikul &amp; Yamane &amp; Samung &amp; Jaitrong" authorityYear="2024" box="[1297,1395,331,357]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="parva" status="sp. nov.">
<emphasis id="EEFFEAAFFFBE412B295045694933FEF9" box="[1297,1395,331,357]" italics="true" pageId="2" pageNumber="81">C. parva</emphasis>
</taxonomicName>
<emphasis id="EEFFEAAFFFBE412B293A45694C88FE14" bold="true" pageId="2" pageNumber="81">
<taxonomicNameLabel id="F5CC57D4FFBE412B293A45694C88FE14" pageId="2" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<taxonomicName id="1B8B4D3EFFBE412B2C98454D4F58FE14" authority="Jaitrong, Pitaktunsakul &amp; Jantarit, 2021" authorityName="Jaitrong, Pitaktunsakul &amp; Jantarit" authorityYear="2021" box="[217,792,367,393]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="panhai">
<emphasis id="EEFFEAAFFFBE412B2C98454D4D09FE14" box="[217,329,367,393]" italics="true" pageId="2" pageNumber="81">C. panhai</emphasis>
<bibRefCitation id="B81A4B4CFFBE412B2D10454D4F58FE14" author="Jaitrong, W. &amp; Pitaktunsakul, P. &amp; Jantarit, S." box="[337,792,367,393]" pageId="2" pageNumber="81" refId="ref12424" refString="Jaitrong, W., Pitaktunsakul, P. &amp; Jantarit, S. (2021) A new species of the genus Carebara westwood, 1840 (Hymenoptera: Formicidae: Myrmecinae) inhabits a cave in Thailand. Far Eastern Entomologist, 425, 7 - 20. [http: // zoobank. org / References / 191 E 3 FC 3 - 9 FFE- 4 DA 8 - B 887 - E 6028469 CDEE] https: // doi. org / 10.25221 / fee. 425.2" type="url" year="2021">Jaitrong, Pitaktunsakul &amp; Jantarit, 2021</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="1B8B4D3EFFBE412B2F64454D4948FE14" authority="Matsuura &amp; Hosoishi, 2024" authorityName="Matsuura &amp; Hosoishi" authorityYear="2024" box="[805,1288,367,393]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="bokorensis">
<emphasis id="EEFFEAAFFFBE412B2F64454D4FFFFE14" box="[805,959,367,393]" italics="true" pageId="2" pageNumber="81">C. bokorensis</emphasis>
<bibRefCitation id="B81A4B4CFFBE412B2F89454D4948FE14" author="Matsuura, K. &amp; Hosoishi, S. &amp; Sokh, H." box="[968,1288,367,393]" pageId="2" pageNumber="81" pagination="1 - 9" refId="ref12643" refString="Matsuura, K., Hosoishi, S. &amp; Sokh, H. (2024) A remarkable new species from the pygmaea species group of the genus Carebara Westwood, 1840 (Hymenoptera: Formicidae: Myrmicinae) from Cambodia. Far Eastern Entomologist, 496, 1 - 9. https: // doi. org / 10.25221 / fee. 496.1" type="journal article" year="2024">Matsuura &amp; Hosoishi, 2024</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="1B8B4D3EFFBE412B2954454D4D7CFE30" authority="(Emery, 1887)" baseAuthorityName="Emery" baseAuthorityYear="1887" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="pygmaea">
<emphasis id="EEFFEAAFFFBE412B2954454D49DCFE15" box="[1301,1436,367,393]" italics="true" pageId="2" pageNumber="81">C. pygmaea</emphasis>
(
<bibRefCitation id="B81A4B4CFFBE412B2CDE45B14D73FE30" author="Emery, C." box="[159,307,403,429]" pageId="2" pageNumber="81" pagination="465 - 473" refId="ref11909" refString="Emery, C. (1887) Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte terza. Formiche della regione Indo-Malese e dell'Australia (continuazione e fine). Annali del Museo civico di storia naturale di Genova, 25 [= (2) (5)], 465 - 473." type="journal article" year="1887">Emery, 1887</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="1B8B4D3EFFBE412B2D0645B14E0DFE30" authority="(Smith, 1860)" baseAuthorityName="F. Smith" baseAuthorityYear="1860" box="[327,589,403,429]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="rubra">
<emphasis id="EEFFEAAFFFBE412B2D0645B14DE8FE30" box="[327,424,403,429]" italics="true" pageId="2" pageNumber="81">C. rubra</emphasis>
(Smith, 1860)
</taxonomicName>
, and
<taxonomicName id="1B8B4D3EFFBE412B2EC945B14F9FFE30" authority="(Smith, 1860)" baseAuthorityName="F. Smith" baseAuthorityYear="1860" box="[648,991,403,429]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="transversalis">
<emphasis id="EEFFEAAFFFBE412B2EC945B14F7AFE30" box="[648,826,403,429]" italics="true" pageId="2" pageNumber="81">C. transversalis</emphasis>
(Smith, 1860)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="94916536FFBE412B2C8645944D5DFE68" pageId="2" pageNumber="81" type="distribution">
<paragraph id="DC3436BDFFBE412B2C8645944D5DFE68" blockId="2.[151,1437,151,501]" pageId="2" pageNumber="81">
<emphasis id="EEFFEAAFFFBE412B2C8645944D21FE4D" bold="true" box="[199,353,438,464]" pageId="2" pageNumber="81">Distribution.</emphasis>
<collectingCountry id="A49C762DFFBE412B2D2845954D9BFE4C" box="[361,475,439,465]" name="Sri Lanka" pageId="2" pageNumber="81">Sri Lanka</collectingCountry>
,
<collectingCountry id="A49C762DFFBE412B2DA645954E60FE4C" box="[487,544,439,465]" name="Laos" pageId="2" pageNumber="81">Laos</collectingCountry>
,
<collectingCountry id="A49C762DFFBE412B2E6C45954EE3FE4C" box="[557,675,439,465]" name="Cambodia" pageId="2" pageNumber="81">Cambodia</collectingCountry>
,
<collectingCountry id="A49C762DFFBE412B2EEE45954F55FE4C" box="[687,789,439,465]" name="Thailand" pageId="2" pageNumber="81">Thailand</collectingCountry>
,
<collectingCountry id="A49C762DFFBE412B2F6345954FC9FE4C" box="[802,905,439,465]" name="Malaysia" pageId="2" pageNumber="81">Malaysia</collectingCountry>
(Borneo),
<collectingCountry id="A49C762DFFBE412B284445954832FE4C" box="[1029,1138,439,465]" name="Indonesia" pageId="2" pageNumber="81">Indonesia</collectingCountry>
(
<collectingCountry id="A49C762DFFBE412B28C345954898FE4C" box="[1154,1240,439,465]" name="Indonesia" pageId="2" pageNumber="81">Ternate</collectingCountry>
to
<collectingRegion id="1E4FF85FFFBE412B28BF45954920FE4C" box="[1278,1376,439,465]" country="Indonesia" name="Sumatera Utara" pageId="2" pageNumber="81">Sumatra</collectingRegion>
), and
<collectingCountry id="A49C762DFFBE412B2CD645F94D59FE68" box="[151,281,475,501]" name="Philippines" pageId="2" pageNumber="81">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="94916536FFBE412B2CD6466149DCFB87" pageId="2" pageNumber="81" type="key">
<paragraph id="DC3436BDFFBE412B2CD64661482FFDC0" blockId="2.[151,1135,578,605]" box="[151,1135,578,605]" pageId="2" pageNumber="81">
<emphasis id="EEFFEAAFFFBE412B2CD64661482FFDC0" bold="true" box="[151,1135,578,605]" pageId="2" pageNumber="81">
Key to Thai species of
<taxonomicName id="1B8B4D3EFFBE412B2DF146604ED5FDC0" baseAuthorityName="Emery" baseAuthorityYear="1887" box="[432,661,578,605]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="pygmaea">
<emphasis id="EEFFEAAFFFBE412B2DF146604ED5FDC0" bold="true" box="[432,661,578,605]" italics="true" pageId="2" pageNumber="81">Carebara pygmaea</emphasis>
</taxonomicName>
species group based on major worker
</emphasis>
</paragraph>
<keyStep id="677F2E18FFBE412B2CD646AA49DCFD68" pageId="2" pageNumber="81">
<paragraph id="DC3436BDFFBE412B2CD646AA49DDFD26" blockId="2.[151,1437,648,1051]" pageId="2" pageNumber="81">
<keyLead id="677A9588FFBE412B2CD646AA49DDFD26" pageId="2" pageNumber="81">
1 In dorsal view, posterior half of head and entire gastral tergite I without both erect and decumbent hairs (
<figureCitation id="44B02A38FFBE412B288F46AA4968FD00" box="[1230,1320,648,670]" captionStart="FIGURE 10" captionStartId="16.[151,250,1891,1915]" captionTargetBox="[195,1393,561,1868]" captionTargetId="figure-190@16.[195,1393,561,1868]" captionTargetPageId="16" captionText="FIGURE 10. Carebara pygmaea, non-type major worker (THNHM-I-2013-05969). A, Body in profile view; B, head in fullface view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742667" httpUri="https://zenodo.org/record/12742667/files/figure.png" pageId="2" pageNumber="81">Fig. 10A</figureCitation>
)....................................................................................................
<taxonomicName id="1B8B4D3EFFBE412B296B468749DDFD26" baseAuthorityName="Emery" baseAuthorityYear="1887" box="[1322,1437,677,699]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="pygmaea">
<emphasis id="EEFFEAAFFFBE412B296B4687497AFD27" box="[1322,1338,677,698]" italics="true" pageId="2" pageNumber="81">C</emphasis>
.
<emphasis id="EEFFEAAFFFBE412B2907468449DDFD26" box="[1350,1437,678,699]" italics="true" pageId="2" pageNumber="81">pygmaea</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="DC3436BDFFBE412B2CD646E149DCFD68" blockId="2.[151,1437,648,1051]" pageId="2" pageNumber="81">
<keyLead id="677A9588FFBE412B2CD646E149DCFD68" pageId="2" pageNumber="81">
- In dorsal view, posterior half of head and entire gastral tergite I with dense or sparse erect or decumbent hairs (
<figureCitation id="44B02A38FFBE412B295946E0492EFD4A" box="[1304,1390,706,728]" captionStart="FIGURE 1" captionStartId="3.[151,250,1708,1732]" captionTargetBox="[215,1373,489,1685]" captionTargetId="figure-149@3.[215,1373,489,1685]" captionTargetPageId="3" captionText="FIGURE 1. Carebara adisaki Wimolsuthikul &amp; Jaitrong, sp. nov., holotype major worker (THNHM-I-27491). A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742645" httpUri="https://zenodo.org/record/12742645/files/figure.png" pageId="2" pageNumber="81">Figs. 1A</figureCitation>
,
<figureCitation id="44B02A38FFBE412B293846E049D8FD4A" box="[1401,1432,706,727]" captionStart="FIGURE 3" captionStartId="6.[151,250,1976,2000]" captionTargetBox="[157,1429,509,1946]" captionTargetId="figure-104@6.[157,1429,509,1946]" captionTargetPageId="6" captionText="FIGURE 3. Carebara bokorensis, non-type major worker (THNHM-I-00028081).A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742649" httpUri="https://zenodo.org/record/12742649/files/figure.png" pageId="2" pageNumber="81">3A</figureCitation>
,
<figureCitation id="44B02A38FFBE412B2CA346C24D41FD6B" box="[226,257,736,758]" captionStart="FIGURE 6" captionStartId="9.[151,250,1981,2005]" captionTargetBox="[277,1311,812,1957]" captionTargetId="figure-317@9.[277,1311,812,1957]" captionTargetPageId="9" captionText="FIGURE 6. Carebara panhai, holotype major worker (THNHM-I-14191). A, Body in profile view; B, head in full-face view; 3C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742655" httpUri="https://zenodo.org/record/12742655/files/figure.png" pageId="2" pageNumber="81">6A</figureCitation>
,
<figureCitation id="44B02A38FFBE412B2D4A46C24D6CFD68" box="[267,300,736,757]" captionStart="FIGURE 8" captionStartId="12.[151,250,1945,1969]" captionTargetBox="[222,1365,705,1921]" captionTargetId="figure-234@12.[222,1365,705,1921]" captionTargetPageId="12" captionText="FIGURE 8. Carebara parva Wimolsuthikul &amp; Jaitrong, sp. nov., holotype major worker (THNHM-I-14199).A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742661" httpUri="https://zenodo.org/record/12742661/files/figure.png" pageId="2" pageNumber="81">8A</figureCitation>
)............................................................................................. 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="677F2E18FFBE412B2CD646DF49DCFCF7" pageId="2" pageNumber="81">
<paragraph id="DC3436BDFFBE412B2CD646DF49DCFCB2" blockId="2.[151,1437,648,1051]" pageId="2" pageNumber="81">
<keyLead id="677A9588FFBE412B2CD646DF49DCFCB2" pageId="2" pageNumber="81">
2 Posterior one-third of head in full-face view with dense foveae, without transverse striation (
<figureCitation id="44B02A38FFBE412B281946DF48E2FC8F" box="[1112,1186,765,787]" captionStart="FIGURE 1" captionStartId="3.[151,250,1708,1732]" captionTargetBox="[215,1373,489,1685]" captionTargetId="figure-149@3.[215,1373,489,1685]" captionTargetPageId="3" captionText="FIGURE 1. Carebara adisaki Wimolsuthikul &amp; Jaitrong, sp. nov., holotype major worker (THNHM-I-27491). A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742645" httpUri="https://zenodo.org/record/12742645/files/figure.png" pageId="2" pageNumber="81">Fig. 1B</figureCitation>
); gastral tergite I punctate (
<figureCitation id="44B02A38FFBE412B2CA847384D72FCB2" box="[233,306,794,816]" captionStart="FIGURE 3" captionStartId="6.[151,250,1976,2000]" captionTargetBox="[157,1429,509,1946]" captionTargetId="figure-104@6.[157,1429,509,1946]" captionTargetPageId="6" captionText="FIGURE 3. Carebara bokorensis, non-type major worker (THNHM-I-00028081).A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742649" httpUri="https://zenodo.org/record/12742649/files/figure.png" pageId="2" pageNumber="81">Fig. 3C</figureCitation>
)..................................................................................
<taxonomicName id="1B8B4D3EFFBE412B2958473949DCFCB2" authorityName="Matsuura &amp; Hosoishi" authorityYear="2024" box="[1305,1436,794,816]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="bokorensis">
<emphasis id="EEFFEAAFFFBE412B2958473949DCFCB2" box="[1305,1436,794,816]" italics="true" pageId="2" pageNumber="81">C. bokorensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="DC3436BDFFBE412B2CD6471A49DCFCF7" blockId="2.[151,1437,648,1051]" pageId="2" pageNumber="81">
<keyLead id="677A9588FFBE412B2CD6471A49DCFCF7" pageId="2" pageNumber="81">
- Posterior one-third of head in full-face view with transverse striation (
<figureCitation id="44B02A38FFBE412B2FC9471A4F98FCD0" box="[904,984,824,846]" captionStart="FIGURE 4" captionStartId="7.[151,250,1642,1666]" captionTargetBox="[151,1436,181,1618]" captionTargetId="figure-17@7.[151,1436,181,1618]" captionTargetPageId="7" captionText="FIGURE 4. Carebara bokorensis, ergatoid queen (THNHM-I-00028220). A, Body in profile view; B, head in full-face view; 2C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742651" httpUri="https://zenodo.org/record/12742651/files/figure.png" pageId="2" pageNumber="81">Figs 4B</figureCitation>
,
<figureCitation id="44B02A38FFBE412B2FA3471A4840FCD3" box="[994,1024,824,846]" captionStart="FIGURE 6" captionStartId="9.[151,250,1981,2005]" captionTargetBox="[277,1311,812,1957]" captionTargetId="figure-317@9.[277,1311,812,1957]" captionTargetPageId="9" captionText="FIGURE 6. Carebara panhai, holotype major worker (THNHM-I-14191). A, Body in profile view; B, head in full-face view; 3C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742655" httpUri="https://zenodo.org/record/12742655/files/figure.png" pageId="2" pageNumber="81">6B</figureCitation>
,
<figureCitation id="44B02A38FFBE412B284A471A4869FCD0" box="[1035,1065,824,845]" captionStart="FIGURE 8" captionStartId="12.[151,250,1945,1969]" captionTargetBox="[222,1365,705,1921]" captionTargetId="figure-234@12.[222,1365,705,1921]" captionTargetPageId="12" captionText="FIGURE 8. Carebara parva Wimolsuthikul &amp; Jaitrong, sp. nov., holotype major worker (THNHM-I-14199).A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742661" httpUri="https://zenodo.org/record/12742661/files/figure.png" pageId="2" pageNumber="81">8B</figureCitation>
); gastral tergite I smooth and shiny, at most weakly shagreened (
<figureCitation id="44B02A38FFBE412B2D9A47774E6EFCF7" box="[475,558,853,875]" captionStart="FIGURE 1" captionStartId="3.[151,250,1708,1732]" captionTargetBox="[215,1373,489,1685]" captionTargetId="figure-149@3.[215,1373,489,1685]" captionTargetPageId="3" captionText="FIGURE 1. Carebara adisaki Wimolsuthikul &amp; Jaitrong, sp. nov., holotype major worker (THNHM-I-27491). A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742645" httpUri="https://zenodo.org/record/12742645/files/figure.png" pageId="2" pageNumber="81">Figs. 1C</figureCitation>
,
<figureCitation id="44B02A38FFBE412B2E7847774E17FCF6" box="[569,599,853,875]" captionStart="FIGURE 6" captionStartId="9.[151,250,1981,2005]" captionTargetBox="[277,1311,812,1957]" captionTargetId="figure-317@9.[277,1311,812,1957]" captionTargetPageId="9" captionText="FIGURE 6. Carebara panhai, holotype major worker (THNHM-I-14191). A, Body in profile view; B, head in full-face view; 3C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742655" httpUri="https://zenodo.org/record/12742655/files/figure.png" pageId="2" pageNumber="81">6C</figureCitation>
,
<figureCitation id="44B02A38FFBE412B2E2047774EC0FCF7" box="[609,640,853,874]" captionStart="FIGURE 8" captionStartId="12.[151,250,1945,1969]" captionTargetBox="[222,1365,705,1921]" captionTargetId="figure-234@12.[222,1365,705,1921]" captionTargetPageId="12" captionText="FIGURE 8. Carebara parva Wimolsuthikul &amp; Jaitrong, sp. nov., holotype major worker (THNHM-I-14199).A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742661" httpUri="https://zenodo.org/record/12742661/files/figure.png" pageId="2" pageNumber="81">8C</figureCitation>
)................................................................. 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="677F2E18FFBE412B2CD6475049DCFC38" pageId="2" pageNumber="81">
<paragraph id="DC3436BDFFBE412B2CD6475049DDFC1A" blockId="2.[151,1437,648,1051]" box="[151,1437,882,904]" pageId="2" pageNumber="81">
<keyLead id="677A9588FFBE412B2CD6475049DDFC1A" box="[151,1437,882,904]" pageId="2" pageNumber="81">
3 Mesosomal dorsum with erect hairs (
<figureCitation id="44B02A38FFBE412B2E0747504ED1FC15" box="[582,657,882,904]" captionStart="FIGURE 6" captionStartId="9.[151,250,1981,2005]" captionTargetBox="[277,1311,812,1957]" captionTargetId="figure-317@9.[277,1311,812,1957]" captionTargetPageId="9" captionText="FIGURE 6. Carebara panhai, holotype major worker (THNHM-I-14191). A, Body in profile view; B, head in full-face view; 3C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742655" httpUri="https://zenodo.org/record/12742655/files/figure.png" pageId="2" pageNumber="81">Fig. 6A</figureCitation>
)........................................................
<taxonomicName id="1B8B4D3EFFBE412B297F475149DDFC1A" authorityName="Jaitrong, Pitaktunsakul &amp; Jantarit" authorityYear="2021" box="[1342,1437,882,904]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="panhai">
<emphasis id="EEFFEAAFFFBE412B297F475149DDFC1A" box="[1342,1437,882,904]" italics="true" pageId="2" pageNumber="81">C. panhai</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="DC3436BDFFBE412B2CD647B249DCFC38" blockId="2.[151,1437,648,1051]" box="[151,1436,912,934]" pageId="2" pageNumber="81">
<keyLead id="677A9588FFBE412B2CD647B249DCFC38" box="[151,1436,912,934]" pageId="2" pageNumber="81">
- Mesosomal dorsum without erect hairs (
<figureCitation id="44B02A38FFBE412B2E2547B24EF9FC38" box="[612,697,912,934]" captionStart="FIGURE 1" captionStartId="3.[151,250,1708,1732]" captionTargetBox="[215,1373,489,1685]" captionTargetId="figure-149@3.[215,1373,489,1685]" captionTargetPageId="3" captionText="FIGURE 1. Carebara adisaki Wimolsuthikul &amp; Jaitrong, sp. nov., holotype major worker (THNHM-I-27491). A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742645" httpUri="https://zenodo.org/record/12742645/files/figure.png" pageId="2" pageNumber="81">Figs. 1A</figureCitation>
,
<figureCitation id="44B02A38FFBE412B2E8547B24EA4FC38" box="[708,740,912,933]" captionStart="FIGURE 8" captionStartId="12.[151,250,1945,1969]" captionTargetBox="[222,1365,705,1921]" captionTargetId="figure-234@12.[222,1365,705,1921]" captionTargetPageId="12" captionText="FIGURE 8. Carebara parva Wimolsuthikul &amp; Jaitrong, sp. nov., holotype major worker (THNHM-I-14199).A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742661" httpUri="https://zenodo.org/record/12742661/files/figure.png" pageId="2" pageNumber="81">8A</figureCitation>
)........................................................ 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="677F2E18FFBE412B2CD6478F49DCFB87" pageId="2" pageNumber="81">
<paragraph id="DC3436BDFFBE412B2CD6478F49DCFC42" blockId="2.[151,1437,648,1051]" pageId="2" pageNumber="81">
<keyLead id="677A9588FFBE412B2CD6478F49DCFC42" pageId="2" pageNumber="81">
4 Propodeal spines shorter than diameter of propodeal spiracle; first gastral tergite with erect hairs only along its posterior margin (
<figureCitation id="44B02A38FFBE412B2CA847E84D75FC42" box="[233,309,970,992]" captionStart="FIGURE 1" captionStartId="3.[151,250,1708,1732]" captionTargetBox="[215,1373,489,1685]" captionTargetId="figure-149@3.[215,1373,489,1685]" captionTargetPageId="3" captionText="FIGURE 1. Carebara adisaki Wimolsuthikul &amp; Jaitrong, sp. nov., holotype major worker (THNHM-I-27491). A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742645" httpUri="https://zenodo.org/record/12742645/files/figure.png" pageId="2" pageNumber="81">Fig. 1A</figureCitation>
)..............................................................................
<taxonomicName id="1B8B4D3EFFBE412B28A947E94909FC42" authorityName="Wimolsuthikul &amp; Yamane &amp; Samung &amp; Jaitrong" authorityYear="2024" box="[1256,1353,970,992]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="adisaki" status="sp. nov.">
<emphasis id="EEFFEAAFFFBE412B28A947E94909FC42" box="[1256,1353,970,992]" italics="true" pageId="2" pageNumber="81">C. adisaki</emphasis>
</taxonomicName>
<emphasis id="EEFFEAAFFFBE412B291147E849DCFC42" bold="true" box="[1360,1436,970,991]" pageId="2" pageNumber="81">
<taxonomicNameLabel id="F5CC57D4FFBE412B291147E849DCFC42" box="[1360,1436,970,991]" pageId="2" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="DC3436BDFFBE412B2CD647CA49DCFB87" blockId="2.[151,1437,648,1051]" pageId="2" pageNumber="81">
<keyLead id="677A9588FFBE412B2CD647CA49DCFB87" pageId="2" pageNumber="81">
- Propodeal spine almost as long as or slightly longer than diameter of propodeal spiracle; entire first gastral tergite covered with erect hairs (
<figureCitation id="44B02A38FFBE412B2D1340274DE0FB87" box="[338,416,1029,1051]" captionStart="FIGURE 8" captionStartId="12.[151,250,1945,1969]" captionTargetBox="[222,1365,705,1921]" captionTargetId="figure-234@12.[222,1365,705,1921]" captionTargetPageId="12" captionText="FIGURE 8. Carebara parva Wimolsuthikul &amp; Jaitrong, sp. nov., holotype major worker (THNHM-I-14199).A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742661" httpUri="https://zenodo.org/record/12742661/files/figure.png" pageId="2" pageNumber="81">Fig. 8A</figureCitation>
)......................................................................
<taxonomicName id="1B8B4D3EFFBE412B28B74027490AFB86" authorityName="Wimolsuthikul &amp; Yamane &amp; Samung &amp; Jaitrong" authorityYear="2024" box="[1270,1354,1029,1051]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="parva" status="sp. nov.">
<emphasis id="EEFFEAAFFFBE412B28B74027490AFB86" box="[1270,1354,1029,1051]" italics="true" pageId="2" pageNumber="81">C. parva</emphasis>
</taxonomicName>
<emphasis id="EEFFEAAFFFBE412B2911402749DCFB87" bold="true" box="[1360,1436,1029,1050]" pageId="2" pageNumber="81">
<taxonomicNameLabel id="F5CC57D4FFBE412B2911402749DCFB87" box="[1360,1436,1029,1050]" pageId="2" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
<subSubSection id="94916536FFBE412B2CD64042482FFBE7" box="[151,1135,1119,1146]" pageId="2" pageNumber="81" type="multiple">
<paragraph id="DC3436BDFFBE412B2CD64042482FFBE7" blockId="2.[151,1135,1119,1146]" box="[151,1135,1119,1146]" pageId="2" pageNumber="81">
<emphasis id="EEFFEAAFFFBE412B2CD64042482FFBE7" bold="true" box="[151,1135,1119,1146]" pageId="2" pageNumber="81">
Key to Thai species of
<taxonomicName id="1B8B4D3EFFBE412B2DF1407D4ED5FBE7" baseAuthorityName="Emery" baseAuthorityYear="1887" box="[432,661,1119,1146]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="pygmaea">
<emphasis id="EEFFEAAFFFBE412B2DF1407D4ED5FBE7" bold="true" box="[432,661,1119,1146]" italics="true" pageId="2" pageNumber="81">Carebara pygmaea</emphasis>
</taxonomicName>
species group based on minor worker
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="94916536FFBE412B2CD6408749DCF9AA" pageId="2" pageNumber="81" type="key">
<keyStep id="677F2E18FFBE412B2CD6408749DCFB4A" pageId="2" pageNumber="81">
<paragraph id="DC3436BDFFBE412B2CD6408749DCFB27" blockId="2.[151,1437,1189,1592]" box="[151,1436,1189,1211]" pageId="2" pageNumber="81">
<keyLead id="677A9588FFBE412B2CD6408749DCFB27" box="[151,1436,1189,1211]" pageId="2" pageNumber="81">
1 Propodeal dorsum entirely punctate (
<figureCitation id="44B02A38FFBE412B2E0440874EEEFB27" box="[581,686,1189,1211]" captionStart="FIGURE 5" captionStartId="8.[151,250,1728,1752]" captionTargetBox="[206,1381,525,1705]" captionTargetId="figure-159@8.[206,1381,525,1705]" captionTargetPageId="8" captionText="FIGURE 5. Carebara bokorensis, non-type minor worker (THNHM-I-00028204).A, Body in profile view; B, head in full-face view; 2C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742653" httpUri="https://zenodo.org/record/12742653/files/figure.png" pageId="2" pageNumber="81">Fig. 5A, C</figureCitation>
)...................................................
<taxonomicName id="1B8B4D3EFFBE412B2958408749DCFB27" authorityName="Matsuura &amp; Hosoishi" authorityYear="2024" box="[1305,1436,1189,1210]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="bokorensis">
<emphasis id="EEFFEAAFFFBE412B2958408749DCFB27" box="[1305,1436,1189,1210]" italics="true" pageId="2" pageNumber="81">C. bokorensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="DC3436BDFFBE412B2CD640E149DCFB4A" blockId="2.[151,1437,1189,1592]" box="[151,1436,1218,1240]" pageId="2" pageNumber="81">
<keyLead id="677A9588FFBE412B2CD640E149DCFB4A" box="[151,1436,1218,1240]" pageId="2" pageNumber="81">
- Propodeal dorsum smooth and shiny (
<figureCitation id="44B02A38FFBE412B2E0F40E04EE3FB4A" box="[590,675,1218,1240]" captionStart="FIGURE 2" captionStartId="4.[151,250,1936,1960]" captionTargetBox="[151,1437,525,1913]" captionTargetId="figure-180@4.[151,1437,525,1913]" captionTargetPageId="4" captionText="FIGURE 2. Carebara adisaki Wimolsuthikul &amp; Jaitrong, sp. nov., paratype minor worker (THNHM-I-27500). A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742647" httpUri="https://zenodo.org/record/12742647/files/figure.png" pageId="2" pageNumber="81">Figs. 2A</figureCitation>
,
<figureCitation id="44B02A38FFBE412B2EEC40E04E8CFB4A" box="[685,716,1218,1239]" captionStart="FIGURE 7" captionStartId="10.[151,250,1970,1994]" captionTargetBox="[230,1357,701,1946]" captionTargetId="figure-190@10.[230,1357,701,1946]" captionTargetPageId="10" captionText="FIGURE 7. Carebara panhai, paratype minor worker (THNHM-I-14193). A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742659" httpUri="https://zenodo.org/record/12742659/files/figure.png" pageId="2" pageNumber="81">7A</figureCitation>
,
<figureCitation id="44B02A38FFBE412B2E9640E04EB6FB4A" box="[727,758,1218,1239]" captionStart="FIGURE 9" captionStartId="13.[151,250,1429,1453]" captionTargetBox="[226,1360,181,1405]" captionTargetId="figure-17@13.[226,1360,181,1405]" captionTargetPageId="13" captionText="FIGURE 9. Carebara parva Wimolsuthikul &amp; Jaitrong, sp. nov., paratype minor worker (THNHM-I-14201).A, Body in profile view; B, head in full-face view; 6C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742665" httpUri="https://zenodo.org/record/12742665/files/figure.png" pageId="2" pageNumber="81">9A</figureCitation>
,
<figureCitation id="44B02A38FFBE412B2F4140E04F69FB4A" box="[768,809,1218,1239]" captionStart="FIGURE 11" captionStartId="17.[151,250,1455,1479]" captionTargetBox="[238,1349,181,1431]" captionTargetId="figure-17@17.[238,1349,181,1431]" captionTargetPageId="17" captionText="FIGURE 11. Carebara pygmaea, non-type minor worker (THNHM-I-2013-05970). A, Body in profile view; B, head in fullface view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742669" httpUri="https://zenodo.org/record/12742669/files/figure.png" pageId="2" pageNumber="81">11A</figureCitation>
).................................................. 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="677F2E18FFBE412B2CD640C249DCFAD0" pageId="2" pageNumber="81">
<paragraph id="DC3436BDFFBE412B2CD640C249DDFA8E" blockId="2.[151,1437,1189,1592]" pageId="2" pageNumber="81">
<keyLead id="677A9588FFBE412B2CD640C249DDFA8E" pageId="2" pageNumber="81">
2 Antennal scape not reaching posterior margin of head (
<figureCitation id="44B02A38FFBE412B2EB340C24F09FB68" box="[754,841,1248,1270]" captionStart="FIGURE 11" captionStartId="17.[151,250,1455,1479]" captionTargetBox="[238,1349,181,1431]" captionTargetId="figure-17@17.[238,1349,181,1431]" captionTargetPageId="17" captionText="FIGURE 11. Carebara pygmaea, non-type minor worker (THNHM-I-2013-05970). A, Body in profile view; B, head in fullface view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742669" httpUri="https://zenodo.org/record/12742669/files/figure.png" pageId="2" pageNumber="81">Fig. 11B</figureCitation>
); hairs on pronotum shorter than diameter of eye (
<figureCitation id="44B02A38FFBE412B296A40C249C3FB68" box="[1323,1411,1248,1270]" captionStart="FIGURE 11" captionStartId="17.[151,250,1455,1479]" captionTargetBox="[238,1349,181,1431]" captionTargetId="figure-17@17.[238,1349,181,1431]" captionTargetPageId="17" captionText="FIGURE 11. Carebara pygmaea, non-type minor worker (THNHM-I-2013-05970). A, Body in profile view; B, head in fullface view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742669" httpUri="https://zenodo.org/record/12742669/files/figure.png" pageId="2" pageNumber="81">Fig. 11A</figureCitation>
)...........................................................................................
<taxonomicName id="1B8B4D3EFFBE412B296B40DF49DDFA8E" baseAuthorityName="Emery" baseAuthorityYear="1887" box="[1322,1437,1277,1299]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="pygmaea">
<emphasis id="EEFFEAAFFFBE412B296B40DF49DDFA8E" box="[1322,1437,1277,1299]" italics="true" pageId="2" pageNumber="81">C. pygmaea</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="DC3436BDFFBE412B2CD6413949DCFAD0" blockId="2.[151,1437,1189,1592]" pageId="2" pageNumber="81">
<keyLead id="677A9588FFBE412B2CD6413949DCFAD0" pageId="2" pageNumber="81">
- Antennal scape reaching or extending beyond posterior margin of head (
<figureCitation id="44B02A38FFBE412B2FF141384847FAB2" box="[944,1031,1306,1328]" captionStart="FIGURE 2" captionStartId="4.[151,250,1936,1960]" captionTargetBox="[151,1437,525,1913]" captionTargetId="figure-180@4.[151,1437,525,1913]" captionTargetPageId="4" captionText="FIGURE 2. Carebara adisaki Wimolsuthikul &amp; Jaitrong, sp. nov., paratype minor worker (THNHM-I-27500). A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742647" httpUri="https://zenodo.org/record/12742647/files/figure.png" pageId="2" pageNumber="81">Figs. 2B</figureCitation>
,
<figureCitation id="44B02A38FFBE412B285241394871FAAD" box="[1043,1073,1307,1328]" captionStart="FIGURE 7" captionStartId="10.[151,250,1970,1994]" captionTargetBox="[230,1357,701,1946]" captionTargetId="figure-190@10.[230,1357,701,1946]" captionTargetPageId="10" captionText="FIGURE 7. Carebara panhai, paratype minor worker (THNHM-I-14193). A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742659" httpUri="https://zenodo.org/record/12742659/files/figure.png" pageId="2" pageNumber="81">7B</figureCitation>
,
<figureCitation id="44B02A38FFBE412B287C4138481CFAB2" box="[1085,1116,1306,1327]" captionStart="FIGURE 9" captionStartId="13.[151,250,1429,1453]" captionTargetBox="[226,1360,181,1405]" captionTargetId="figure-17@13.[226,1360,181,1405]" captionTargetPageId="13" captionText="FIGURE 9. Carebara parva Wimolsuthikul &amp; Jaitrong, sp. nov., paratype minor worker (THNHM-I-14201).A, Body in profile view; B, head in full-face view; 6C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742665" httpUri="https://zenodo.org/record/12742665/files/figure.png" pageId="2" pageNumber="81">9B</figureCitation>
); hairs on pronotum longer than diameter of eye....................................................................................... 3
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<keyStep id="677F2E18FFBE412B2CD6417749DCFA5F" pageId="2" pageNumber="81">
<paragraph id="DC3436BDFFBE412B2CD6417749DDFA1A" blockId="2.[151,1437,1189,1592]" pageId="2" pageNumber="81">
<keyLead id="677A9588FFBE412B2CD6417749DDFA1A" pageId="2" pageNumber="81">
3 Propodeal junction roundly convex (
<figureCitation id="44B02A38FFBE412B2E0741774ED4FAF7" box="[582,660,1365,1387]" captionStart="FIGURE 7" captionStartId="10.[151,250,1970,1994]" captionTargetBox="[230,1357,701,1946]" captionTargetId="figure-190@10.[230,1357,701,1946]" captionTargetPageId="10" captionText="FIGURE 7. Carebara panhai, paratype minor worker (THNHM-I-14193). A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742659" httpUri="https://zenodo.org/record/12742659/files/figure.png" pageId="2" pageNumber="81">Fig. 7A</figureCitation>
); antennal scape extending beyond posterior margin of head (
<figureCitation id="44B02A38FFBE412B28AC4177497AFAF7" box="[1261,1338,1365,1387]" captionStart="FIGURE 7" captionStartId="10.[151,250,1970,1994]" captionTargetBox="[230,1357,701,1946]" captionTargetId="figure-190@10.[230,1357,701,1946]" captionTargetPageId="10" captionText="FIGURE 7. Carebara panhai, paratype minor worker (THNHM-I-14193). A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742659" httpUri="https://zenodo.org/record/12742659/files/figure.png" pageId="2" pageNumber="81">Fig. 7B</figureCitation>
); dorsa of petiole and postpetiole each with 24 erect hairs (
<figureCitation id="44B02A38FFBE412B2EFF41504F4CFA1A" box="[702,780,1394,1416]" captionStart="FIGURE 7" captionStartId="10.[151,250,1970,1994]" captionTargetBox="[230,1357,701,1946]" captionTargetId="figure-190@10.[230,1357,701,1946]" captionTargetPageId="10" captionText="FIGURE 7. Carebara panhai, paratype minor worker (THNHM-I-14193). A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742659" httpUri="https://zenodo.org/record/12742659/files/figure.png" pageId="2" pageNumber="81">Fig. 7A</figureCitation>
)..............................................
<taxonomicName id="1B8B4D3EFFBE412B297F415149DDFA1A" authorityName="Jaitrong, Pitaktunsakul &amp; Jantarit" authorityYear="2021" box="[1342,1437,1394,1416]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="panhai">
<emphasis id="EEFFEAAFFFBE412B297F415149DDFA1A" box="[1342,1437,1394,1416]" italics="true" pageId="2" pageNumber="81">C. panhai</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="DC3436BDFFBE412B2CD641B249DCFA5F" blockId="2.[151,1437,1189,1592]" pageId="2" pageNumber="81">
<keyLead id="677A9588FFBE412B2CD641B249DCFA5F" pageId="2" pageNumber="81">
- Propodeal junction bluntly angulated (
<figureCitation id="44B02A38FFBE412B2E1141B24EE5FA38" box="[592,677,1424,1446]" captionStart="FIGURE 2" captionStartId="4.[151,250,1936,1960]" captionTargetBox="[151,1437,525,1913]" captionTargetId="figure-180@4.[151,1437,525,1913]" captionTargetPageId="4" captionText="FIGURE 2. Carebara adisaki Wimolsuthikul &amp; Jaitrong, sp. nov., paratype minor worker (THNHM-I-27500). A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742647" httpUri="https://zenodo.org/record/12742647/files/figure.png" pageId="2" pageNumber="81">Figs. 2A</figureCitation>
,
<figureCitation id="44B02A38FFBE412B2EEE41B24E8FFA38" box="[687,719,1424,1445]" captionStart="FIGURE 9" captionStartId="13.[151,250,1429,1453]" captionTargetBox="[226,1360,181,1405]" captionTargetId="figure-17@13.[226,1360,181,1405]" captionTargetPageId="13" captionText="FIGURE 9. Carebara parva Wimolsuthikul &amp; Jaitrong, sp. nov., paratype minor worker (THNHM-I-14201).A, Body in profile view; B, head in full-face view; 6C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742665" httpUri="https://zenodo.org/record/12742665/files/figure.png" pageId="2" pageNumber="81">9A</figureCitation>
); antennal scape reaching posterior margin of head (
<figureCitation id="44B02A38FFBE412B288041B24955FA38" box="[1217,1301,1424,1446]" captionStart="FIGURE 2" captionStartId="4.[151,250,1936,1960]" captionTargetBox="[151,1437,525,1913]" captionTargetId="figure-180@4.[151,1437,525,1913]" captionTargetPageId="4" captionText="FIGURE 2. Carebara adisaki Wimolsuthikul &amp; Jaitrong, sp. nov., paratype minor worker (THNHM-I-27500). A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742647" httpUri="https://zenodo.org/record/12742647/files/figure.png" pageId="2" pageNumber="81">Figs. 2B</figureCitation>
,
<figureCitation id="44B02A38FFBE412B295E41B2497EFA38" box="[1311,1342,1424,1445]" captionStart="FIGURE 9" captionStartId="13.[151,250,1429,1453]" captionTargetBox="[226,1360,181,1405]" captionTargetId="figure-17@13.[226,1360,181,1405]" captionTargetPageId="13" captionText="FIGURE 9. Carebara parva Wimolsuthikul &amp; Jaitrong, sp. nov., paratype minor worker (THNHM-I-14201).A, Body in profile view; B, head in full-face view; 6C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742665" httpUri="https://zenodo.org/record/12742665/files/figure.png" pageId="2" pageNumber="81">9B</figureCitation>
); dorsa of petiole and postpetiole without erect hairs (
<figureCitation id="44B02A38FFBE412B2EC0418F4E96FA5F" box="[641,726,1453,1475]" captionStart="FIGURE 2" captionStartId="4.[151,250,1936,1960]" captionTargetBox="[151,1437,525,1913]" captionTargetId="figure-180@4.[151,1437,525,1913]" captionTargetPageId="4" captionText="FIGURE 2. Carebara adisaki Wimolsuthikul &amp; Jaitrong, sp. nov., paratype minor worker (THNHM-I-27500). A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742647" httpUri="https://zenodo.org/record/12742647/files/figure.png" pageId="2" pageNumber="81">Figs. 2A</figureCitation>
,
<figureCitation id="44B02A38FFBE412B2EA1418F4F41FA5F" box="[736,769,1453,1474]" captionStart="FIGURE 9" captionStartId="13.[151,250,1429,1453]" captionTargetBox="[226,1360,181,1405]" captionTargetId="figure-17@13.[226,1360,181,1405]" captionTargetPageId="13" captionText="FIGURE 9. Carebara parva Wimolsuthikul &amp; Jaitrong, sp. nov., paratype minor worker (THNHM-I-14201).A, Body in profile view; B, head in full-face view; 6C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742665" httpUri="https://zenodo.org/record/12742665/files/figure.png" pageId="2" pageNumber="81">9A</figureCitation>
)...................................................... 4
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<paragraph id="DC3436BDFFBE412B2CD641E849DCFA60" blockId="2.[151,1437,1189,1592]" pageId="2" pageNumber="81">
<keyLead id="677A9588FFBE412B2CD641E849DCFA60" pageId="2" pageNumber="81">
4 Peduncle of petiole punctate; PI1 240275; larger species (HW
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in minor worker;
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in major worker)...............................................................................
<taxonomicName id="1B8B4D3EFFBE412B28A941CA4909FA60" authorityName="Wimolsuthikul &amp; Yamane &amp; Samung &amp; Jaitrong" authorityYear="2024" box="[1256,1353,1512,1533]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="adisaki" status="sp. nov.">
<emphasis id="EEFFEAAFFFBE412B28A941CA48B8FA60" box="[1256,1272,1512,1533]" italics="true" pageId="2" pageNumber="81">C</emphasis>
.
<emphasis id="EEFFEAAFFFBE412B294541CA4909FA60" box="[1284,1353,1512,1533]" italics="true" pageId="2" pageNumber="81">adisaki</emphasis>
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<emphasis id="EEFFEAAFFFBE412B291141CA49DCFA60" bold="true" box="[1360,1436,1512,1533]" pageId="2" pageNumber="81">
<taxonomicNameLabel id="F5CC57D4FFBE412B291141CA49DCFA60" box="[1360,1436,1512,1533]" pageId="2" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="DC3436BDFFBE412B2CD6422749DCF9AA" blockId="2.[151,1437,1189,1592]" pageId="2" pageNumber="81">
<keyLead id="677A9588FFBE412B2CD6422749DCF9AA" pageId="2" pageNumber="81">
- Peduncle of petiole smooth and shiny; PI1 160200; smaller species (HW
<quantity id="1B739B58FFBE412B2FF642274803F987" box="[951,1091,1541,1563]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" metricValueMax="4.0" metricValueMin="3.5999999999999996" pageId="2" pageNumber="81" unit="mm" value="0.38" valueMax="0.4" valueMin="0.36">0.360.40 mm</quantity>
in minor worker;
<quantity id="1B739B58FFBE412B28B7422749C2F987" box="[1270,1410,1541,1562]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" metricValueMax="8.299999999999999" metricValueMin="7.9" pageId="2" pageNumber="81" unit="mm" value="0.81" valueMax="0.83" valueMin="0.79">0.790.83 mm</quantity>
in major worker)...........................................................................
<taxonomicName id="1B8B4D3EFFBE412B28B74201490AF9A5" authorityName="Wimolsuthikul &amp; Yamane &amp; Samung &amp; Jaitrong" authorityYear="2024" box="[1270,1354,1571,1592]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="parva" status="sp. nov.">
<emphasis id="EEFFEAAFFFBE412B28B742014946F9A5" box="[1270,1286,1571,1592]" italics="true" pageId="2" pageNumber="81">C</emphasis>
.
<emphasis id="EEFFEAAFFFBE412B29534201490AF9A5" box="[1298,1354,1571,1592]" italics="true" pageId="2" pageNumber="81">parva</emphasis>
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<taxonomicNameLabel id="F5CC57D4FFBE412B2911420049DCF9AA" box="[1360,1436,1570,1591]" pageId="2" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
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