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<document id="2F922C970934F7471AB1A5B22A249493" ID-CLB-Dataset="34707" ID-DOI="http://dx.doi.org/10.3897/zookeys.271.4062" ID-GBIF-Dataset="c93c3a93-d2e1-4c85-8ece-bf3aa1a5109d" ID-PMC="PMC3652427" ID-Pensoft-Pub="1313-2970-271-1" ID-PubMed="23717185" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-271-1" ModsDocOrigin="ZooKeys 271" ModsDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" checkinTime="1451247655320" checkinUser="pensoft" docAuthor="Caterino, Michael S. &amp; Tishechkin, Alexey K." docDate="2013" docId="54079E4AC32D11A676B15C9003D3F12D" docLanguage="en" docName="ZooKeys 271: 1-401" docOrigin="ZooKeys 271" docSource="http://dx.doi.org/10.3897/zookeys.271.4062" docTitle="Operclipygus mirabilis Wenzel &amp; Dybas 1941, comb. n." docType="treatment" docVersion="5" lastPageNumber="40" masterDocId="4403FF80FFE7FF95FFDA5D0E29259B2F" masterDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" masterLastPageNumber="401" masterPageNumber="1" pageNumber="36" updateTime="1732934305285" updateUser="ExternalLinkService">
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<mods:title id="F82F49BB328E027CCE2A2AA2CB750688">A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
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<mods:namePart id="6D457D8B666C3AC492F57771F9813DC2">Caterino, Michael S.</mods:namePart>
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<mods:namePart id="879708718670A23E1B6F7A3DAE11F12D">Tishechkin, Alexey K.</mods:namePart>
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<mods:identifier id="2F85CAB4427D0DF39DB1F85490396153" type="DOI">http://dx.doi.org/10.3897/zookeys.271.4062</mods:identifier>
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<treatment id="54079E4AC32D11A676B15C9003D3F12D" ID-GBIF-Taxon="152040962" LSID="urn:lsid:plazi:treatment:54079E4AC32D11A676B15C9003D3F12D" httpUri="http://treatment.plazi.org/id/54079E4AC32D11A676B15C9003D3F12D" lastPageId="39" lastPageNumber="40" pageId="35" pageNumber="36" scope_family="Histeridae" scope_order="Coleoptera" scope_tribe="Exosternini">
<subSubSection id="45E2596A08F470DF90EAE213762E3A93" pageId="35" pageNumber="36" type="nomenclature">
<paragraph id="2450D2173E19BFD15E1B36D765593CB2" pageId="35" pageNumber="36">
<taxonomicName id="84D66E8F080F3AF4FA108513E12E448C" ID-CoL="6SNJY" LSID="http://species-id.net/wiki/Operclipygus_mirabilis" authority="Wenzel &amp; Dybas, 1941" authorityName="Wenzel &amp; Dybas" authorityYear="1941" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus mirabilis" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="mirabilis">Operclipygus mirabilis (Wenzel &amp; Dybas, 1941)</taxonomicName>
<taxonomicNameLabel id="977572782B7C321E12A093CE7F80EE45" pageId="35" pageNumber="36">comb. n.</taxonomicNameLabel>
Figs 8A9A10
<normalizedToken id="92BA4C2912BD98ED9B16941E6059B530" originalValue="A–C">A-C</normalizedToken>
, FMap 2
</paragraph>
</subSubSection>
<subSubSection id="B5B32B7BA1A72CC11DB5CB3E1F7538B2" pageId="35" pageNumber="36" type="reference_group">
<paragraph id="637028CDFFCDF0FFD83E6C33B949F6AA" pageId="35" pageNumber="36">
<taxonomicName id="D62BE5B226A0EC3A47BB718BD80BD830" class="Insecta" family="Histeridae" genus="Pseudister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudister mirabilis" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="mirabilis">Pseudister mirabilis</taxonomicName>
Wenzel &amp; Dybas, 1941: 454.
</paragraph>
</subSubSection>
<subSubSection id="0AE1814F53DB41C379E85B86339D8C59" pageId="35" pageNumber="36" type="type locality">
<paragraph id="345BA3CA8E7C6B7FB93FA6BACCCF0654" pageId="35" pageNumber="36">Type locality.</paragraph>
<paragraph id="831C3647D3F0869D1DA71C5674DF5D0B" pageId="35" pageNumber="36">
COLOMBIA: Cundinamarca: Puerto Salgar [
<geoCoordinate id="53775D3D27D6DE21A594DD5E24590BE9" direction="north" orientation="latitude" precision="925" value="5.4666667">5°28'N</geoCoordinate>
,
<geoCoordinate id="608093219F17BC1E4CA0974A3C5051F1" direction="west" orientation="longitude" precision="925" value="-74.65">74°39'W</geoCoordinate>
].
</paragraph>
</subSubSection>
<subSubSection id="5301D05B02EA5EF28A0E2552CE26CED6" pageId="35" pageNumber="36" type="type material">
<paragraph id="7D7DF27B7F65E672CA2402F7FC2DDF1F" pageId="35" pageNumber="36">Type material.</paragraph>
<paragraph id="F3CE3D6A9DA391D81646815F01EBFFE0" pageId="35" pageNumber="36">
Holotype male:
<normalizedToken id="3126E6297286F393D97F73AA082F0515" originalValue="“P’to">&quot;P'to</normalizedToken>
. Salgar, Cund Colomb, VII:31:38&quot; / &quot;Coll. by C.H.Seevers&quot; / &quot;Collection R. L. Wenzel&quot; / &quot;Type
<taxonomicName id="145AE7C8503F02269D17811542A14548" class="Insecta" family="Histeridae" genus="Pseudister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudister mirabilis" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="mirabilis">Pseudister mirabilis</taxonomicName>
Wenzel &amp; Dybas&quot; (FMNH).
</paragraph>
</subSubSection>
<subSubSection id="5EE0DC66FB5ABD8A6715BCB387AF2B23" pageId="36" pageNumber="37" type="other material">
<paragraph id="FC576213F46E1CA738BFB66B044A1A88" pageId="36" pageNumber="37">
<pageBreakToken id="48857E2DEB224BE578E666F0ECA5F21B" pageId="36" pageNumber="37" start="start">Other</pageBreakToken>
material.
</paragraph>
<paragraph id="32536F1260048DE19032AAA865875DDB" pageId="36" pageNumber="37">
PANAMA,
<normalizedToken id="CFA2521CB67F66FF127DCA95AE4D4881" originalValue="Panamá">Panama</normalizedToken>
: 1: Barro Colorado Isl.,
<geoCoordinate id="DF800839C8CBC572FB14E4777D9D4ABB" direction="north" orientation="latitude" precision="925" value="9.183333">09°11'N</geoCoordinate>
,
<geoCoordinate id="8296CC45E0D7FA2D6A4E59B1B2F484D0" direction="west" orientation="longitude" precision="925" value="-79.85">79°51'W</geoCoordinate>
, 8.vii.1994, D. Banks, FIT (SEMC), 1: external refuse deposit
<taxonomicName id="82A993106D61C17A659BBB70BFC508B7" class="Insecta" family="Formicidae" genus="Atta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atta colombica" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="colombica">Atta colombica</taxonomicName>
(
<normalizedToken id="7696D7DD6E87093B4534FF47267B2E60" originalValue="Guérin-Méneville">Guerin-Meneville</normalizedToken>
), 11.ii.1976 (FMNH);
<normalizedToken id="1710AE3CF806381079B8B5454BBA939E" originalValue="Colón">Colon</normalizedToken>
: 1: Parque Nac. San Lorenzo, Achiote,
<geoCoordinate id="86DB8FA2AC38A4F1A7A799D148730894" direction="north" orientation="latitude" precision="925" value="9.2">9°12'N</geoCoordinate>
,
<geoCoordinate id="77B5B8BA82C0E86036FBBDEC2E3B5B10" direction="west" orientation="longitude" precision="925" value="-79.98333">79°59'W</geoCoordinate>
, 10m, 8-22.v.2007, FIT, A. Mercado (FMNH); 1: 50m [from forest edge, as for all A. Mercado specimens; pers. comm.], 9-23.v.2007(GBFM); 1: San Lorenzo Forest, STRI crane site,
<geoCoordinate id="9DD4EE418BCDBA95236578E8D87E1D64" direction="north" orientation="latitude" precision="925" value="9.283334">9°17'N</geoCoordinate>
,
<geoCoordinate id="6EE5F34ED2F3FBB9F32B7352A20595B8" direction="west" orientation="longitude" precision="925" value="-79.96667">79°58'W</geoCoordinate>
, 25-26.v.2003, FIT, A.K. Tishechkin (GBFM), 1: ex. refuse pile
<taxonomicName id="BA6127AB1873F2197262C6CF3A305685" class="Insecta" family="Formicidae" genus="Atta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atta colombica" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="colombica">Atta colombica</taxonomicName>
, 25.x.2003 (AKTC), 1: FIT, 11-12.v.2004 (LSAM).
</paragraph>
</subSubSection>
<subSubSection id="4200372BB6FFEBD50AB3137E425FC9C8" lastPageId="39" lastPageNumber="40" pageId="36" pageNumber="37" type="description">
<paragraph id="257B999360F59B1775076EE7E1370A65" pageId="36" pageNumber="37">Diagnostic description.</paragraph>
<paragraph id="ADBF99E80DCBCE483A05882FDA6F1875" lastPageId="39" lastPageNumber="40" pageId="36" pageNumber="37">
Length: 1.90-1.97 mm, width: 1.50-1.68 mm; Body rufo-brunneous, sides rounded, moderately convex; lower part of frons and epistoma
<pageBreakToken id="E02EEB47BF95F13EB7090EE2943235E3" pageId="37" pageNumber="38" start="start">weakly</pageBreakToken>
depressed at middle, frontal stria outwardly arcuate, faintly sinuate at middle, rarely interrupted at sides; supraorbital stria absent; labrum rather narrow, about 1.5
<normalizedToken id="85B095FE6650FA31EE8B0EF2644907AA" originalValue="×">x</normalizedToken>
as wide as long, shallowly emarginate at apex; left mandible untoothed, right mandible with very weak basal incisor tooth; pronotum with strong plicae in basal half, in front of 3rd elytral stria, lateral submarginal pronotal stria complete in female, in male ending one-third behind anterior corner in anterolateral pronotal depression; anterior submarginal stria detached, with ends recurved about one-sixth pronotal length; anterior marginal pronotal stra narrowly interrupted behind head; male with depression in anterolateral corner with central tubercle bearing median pronotal gland opening at its apex; female with median pair of gland openings just laterad ends of recurved anterior submarginal stria, about 8 diameters from anterior margin; pronotal disk with few coarse punctures intermingled with fine ground punctures toward sides; elytra with two complete epipleural striae, outer subhumeral stria briefly interrupted at midpoint, otherwise complete, inner subhumeral stria present, obsolete at anterior and posterior ends, all other dorsal elytral striae complete, 5th and sutural striae connected by an angulate basal arch; few coarse punctures present near elytral apex; prosternum with carinal striae complete, sinuate, close at midpoint, diverging slightly to front, connected by narrow anterior arch, keel with faint secondary carinal striae laterad primary striae; prosternal keel weakly produced posteriorly; anterior mesoventral margin broadly emarginate in continuous arc from corner to corner, marginal mesoventral stria complete, mesometaventral stria arched forward to middle of mesoventral disk, crenulate, extend
<pageBreakToken id="04ED0C967848856F5CADEA71CE3D650E" pageId="38" pageNumber="39" start="start">ing</pageBreakToken>
posterad to middle of each metacoxa; 1st abdominal ventrite with two lateral striae, outer stria abbreviated posteriorly; propygidium with sparse fine ground punctation and coarser punctures scattered, separated by about 2
<normalizedToken id="B9088486126B82260822834D416EBA64" originalValue="×">x</normalizedToken>
their diameters; pygidium with similar ground punctation, and very few coarser punctures, particularly near anterolateral corners; pygidial marginal stria absent. Male genitalia (Figs 10
<normalizedToken id="0E6F98859FB04D9AB0F379E225BFF271" originalValue="A–C">A-C</normalizedToken>
, F): accessory sclerites present; T8 with deep, rather narrow basal emargination with well sclerotized basal edge, basal membrane attachment line not intersecting basal emargination, apical emargination shallow, ventrolateral apodemes not meeting at midline; S8 narrowing weakly to apex, with apical guides developed only at apex, ventral halves meeting along most of midline but with median area markedly desclerotized; T9 with apices simple, pointed inward; T10 with halves fully separate; base of S9 widened gradually in basal two-thirds, lacking apical emargination, apical flanges separate, apicolateral flanges
<pageBreakToken id="AAEC431153FD4A36ED968F06A87A05F3" pageId="39" pageNumber="40" start="start">well</pageBreakToken>
developed; tegmen widest basad midpoint, more strongly narrowed to apex, apex strongly bent ventrad, medioventral process
<normalizedToken id="F64BF8ED552E686A90E2F6ED7B1AF222" originalValue="‘U’">'U'</normalizedToken>
-shaped, projecting beneath about one-fourth from base; median lobe about one-third tegmen length, proximal apodemes uniform; basal piece about one-third tegmen length.
</paragraph>
</subSubSection>
<subSubSection id="FEA788A6ED1C423032606CA8BE0A5EF1" pageId="39" pageNumber="40" type="remarks">
<paragraph id="88950B226F8902A5D7CDB5BA59C31E25" pageId="39" pageNumber="40">Remarks.</paragraph>
<paragraph id="A848193AFFB7FA0A6C25F9C509262052" pageId="39" pageNumber="40">
The nominate species in this group is the most easily recognized, at least based on males. The distinctive depression bearing the median pronotal gland opening on a small tubercle (Fig. 9A) identifies them instantly. Males of the following species,
<taxonomicName id="614A9A625AB99A9CB0C351A012C20A64" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus pustulifer" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="pustulifer">Operclipygus pustulifer</taxonomicName>
, have a smaller tubercle and no surrounding depression (Fig. 9B). Both of these species have the 5th and sutural elytral striae complete and connected by a basal arch, and also have the inner subhumeral stria more or less complete. The females are more difficult to identify, and are extremely similar to those of
<taxonomicName id="4EA96E7CD4419606B4E019E29516827E" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus pustulifer" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="pustulifer">Operclipygus pustulifer</taxonomicName>
. Those of
<taxonomicName id="D23352551796971D6E8E78AF52DA15AB" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus mirabilis" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mirabilis">Operclipygus mirabilis</taxonomicName>
, however,have more conspicuously coarse punctures along the sides of the pronotum and the pygidium.
</paragraph>
<paragraph id="D47125FB508026B8C6DF2BD871C51710" pageId="39" pageNumber="40">
The holotype was reportedly collected in a 'rubbish heap of
<taxonomicName id="AA4AB1EAA11FA88E4E515C4CB0C405C4" class="Insecta" family="Formicidae" genus="Atta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atta" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">Atta</taxonomicName>
sp.' (
<bibRefCitation id="F55F480309C59D370E5E73E0DAA7462D" author="Wenzel, RL" journalOrPublisher="Fieldiana, Zoology" pageId="377" pageNumber="378" pagination="433 - 472" title="New and little known neotropical Histeridae (Coleoptera)." volume="22" year="1941">Wenzel and Dybas 1941</bibRefCitation>
), and several recent records confirm this association, specifically with
<taxonomicName id="9422DD6027B196352D4732687A3D3BFD" class="Insecta" family="Formicidae" genus="Atta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atta colombica" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="colombica">Atta colombica</taxonomicName>
.
</paragraph>
<caption id="F69179F76FD3891759157F2714E911BB" pageId="39" pageNumber="40">
<paragraph id="6CB61B16CA716EB12EC7B39470AABAE5" pageId="39" pageNumber="40">
Figure 8.
<taxonomicName id="43CF65ADB4CE5B6212BF5C6CAA96FDA1" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus mirabilis" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mirabilis">Operclipygus mirabilis</taxonomicName>
group. A Dorsal habitus of
<taxonomicName id="C72FC56D4DB97D762F56AE2022C79107" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus mirabilis" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mirabilis">Operclipygus mirabilis</taxonomicName>
B Dorsal habitus of
<taxonomicName id="79D6D5C4F21DAE938EAA299482C49CFB" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus mutuca" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mutuca">Operclipygus mutuca</taxonomicName>
C Dorsal habitus of
<taxonomicName id="1DB1319D4E7C854584A82E316350289F" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus carinistrius" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="carinistrius">Operclipygus carinistrius</taxonomicName>
(lectotype) D Dorsal habitus of
<taxonomicName id="5DA107040A5D1390DE3A764C56BD4456" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus parensis" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="parensis">Operclipygus parensis</taxonomicName>
E Dorsal habitus of
<taxonomicName id="2F220B9072B7BFA78AB5BAE15CA7FC5A" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus schlingeri" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="schlingeri">Operclipygus schlingeri</taxonomicName>
F Frons of
<taxonomicName id="40B5AEB1C844E3A4B476FB9F042A60EB" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus schlingeri" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="schlingeri">Operclipygus schlingeri</taxonomicName>
.
</paragraph>
</caption>
<caption id="09F5A6BF16D621677DB72850C185310D" pageId="39" pageNumber="40">
<paragraph id="526835ECBA909779D693288D3C3BF4DA" pageId="39" pageNumber="40">
Figure 9.
<taxonomicName id="43F7C405F821640D213F7EC79A9E946E" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus mirabilis" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mirabilis">Operclipygus mirabilis</taxonomicName>
group. A Pronotum of
<taxonomicName id="5E2CAA074CEB2311997C03AA0FD3B85F" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus mirabilis" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mirabilis">Operclipygus mirabilis</taxonomicName>
B Pronotum of
<taxonomicName id="DF134C845C235454B43676CC314D80D1" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus pustulifer" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="pustulifer">Operclipygus pustulifer</taxonomicName>
C Pronotum of
<taxonomicName id="76DBE1E39DDDD6C1EC5EE1B7D8B26013" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus plaumanni" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="plaumanni">Operclipygus plaumanni</taxonomicName>
D Pronotum of
<taxonomicName id="5B2381723517E5AAB1EF0A40973F25DE" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus sinuatus" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="sinuatus">Operclipygus sinuatus</taxonomicName>
.
</paragraph>
</caption>
<caption id="0EFF584891AF452C7030713AFF9C881E" pageId="39" pageNumber="40">
<paragraph id="74B320FFC3518AA5E0289A754B1DB651" pageId="39" pageNumber="40">
Map 2. Records of the
<taxonomicName id="FE1C3B40A259AE3B1C3F9B181F86C47A" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus mirabilis" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mirabilis">Operclipygus mirabilis</taxonomicName>
group.
</paragraph>
</caption>
<caption id="10D1002633A7527E2A1D2B4A7A34FAAD" pageId="39" pageNumber="40">
<paragraph id="F2FA3070EC696F16EFB5B83205C15625" pageId="39" pageNumber="40">
Figure 10. Male genitalia of
<taxonomicName id="D5D4998BBADA4B8DC88C33485F20C8FA" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus mirabilis" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mirabilis">Operclipygus mirabilis</taxonomicName>
group. A T8 of
<taxonomicName id="D2F14E72F8F1F72430289BFD0F21055F" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus mirabilis" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mirabilis">Operclipygus mirabilis</taxonomicName>
B S8 of
<taxonomicName id="F1261158AE06AAC436C280B63D77E17F" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus mirabilis" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mirabilis">Operclipygus mirabilis</taxonomicName>
C S9 of
<taxonomicName id="8630CB32C46DC1FDB51EF41B6227A0E5" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus mirabilis" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mirabilis">Operclipygus mirabilis</taxonomicName>
D S9 of
<taxonomicName id="8A0D2E0F1803382274A3B3B77A71B07C" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus sinuatus" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="sinuatus">Operclipygus sinuatus</taxonomicName>
E S9 of
<taxonomicName id="B109DF4382AFD66AC09C7CF5862DA226" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus schlingeri" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="schlingeri">Operclipygus schlingeri</taxonomicName>
F Aedeagus, dorsal and lateral views, of
<taxonomicName id="617AC66A4BD7803615B6CD64529674E2" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus mirabilis" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mirabilis">Operclipygus mirabilis</taxonomicName>
G Aedeagus, dorsal and lateral views, of
<taxonomicName id="DA3B5E3D9FC94A5CE4AC8B5AD7917685" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus sinuatus" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="sinuatus">Operclipygus sinuatus</taxonomicName>
H Aedeagus, dorsal and lateral views, of
<taxonomicName id="1ECD2436F41B4A939D2D53E4AC624C0F" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus schlingeri" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="schlingeri">Operclipygus schlingeri</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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