treatments-xml/data/54/06/C2/5406C22A0A3B2140C032FC2729DF47A2.xml

<document id="AAF426FA6D340B7EA1D65C9601A12A15" ID-CLB-Dataset="26370" ID-DOI="10.1206/00030090-426.1.1" ID-GBIF-Dataset="ed64cd3a-2478-45f1-a0e6-ae0384f6c740" ID-Handle="http://digitallibrary.amnh.org/handle/2246/6925" ID-Zenodo-Dep="3259018" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1561701537349" checkinUser="plazi" docAuthor="Magalhaes, Ivan L. F. &amp; Ramírez, Martín J." docDate="2019" docId="5406C22A0A3B2140C032FC2729DF47A2" docLanguage="en" docName="BulAmeMusNatHis.2019.426.1-89.pdf" docOrigin="Bulletin of the American Museum of Natural History 426" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.9:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="9" docTitle="Kukulcania brignolii Magalhaes &amp; Ramírez 2019" docType="treatment" docVersion="10" lastPageNumber="102" masterDocId="A83FBA520A5B2126C365FF9F2A4C4147" masterDocTitle="The Crevice Weaver Spider Genus Kukulcania (Araneae: Filistatidae)" masterLastPageNumber="153" masterPageNumber="1" pageNumber="96" updateTime="1698737499883" updateUser="ExternalLinkService">
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<mods:title id="6634F81A76DD9A4937409CAD8CE83186">The Crevice Weaver Spider Genus Kukulcania (Araneae: Filistatidae)</mods:title>
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<mods:namePart id="BEE42086BDBC41BA5858D7220EB81A61">Ramírez, Martín J.</mods:namePart>
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<subSubSection id="94B520B70A3B2146C032FC272E2F42B6" pageId="96" pageNumber="96" type="nomenclature">
<paragraph id="DC10733C0A3B2146C032FC272E2F42B6" blockId="96.[822,1123,952,1009]" pageId="96" pageNumber="96">
<taxonomicName id="1BAF08BF0A3B2146C032FC27299242B6" ID-CoL="3RFGY" authority="(Alayon, 1981)" authorityName="Magalhaes &amp; Ramírez" authorityYear="2019" baseAuthorityName="Alayon" baseAuthorityYear="1981" class="Arachnida" family="Filistatidae" genus="Kukulcania" kingdom="Animalia" order="Araneae" pageId="96" pageNumber="101" phylum="Arthropoda" rank="species" species="brignolii" status="comb. nov.">
<emphasis id="EEDBAF2E0A3B2146C032FC27299242B6" bold="true" pageId="96" pageNumber="96">
<heading id="8758C4500A3B2146C032FC272E0C4297" bold="true" box="[855,1088,952,976]" centered="true" fontSize="10" level="1" pageId="96" pageNumber="96" reason="4">
<emphasis id="EEDBAF2E0A3B2146C032FC272E0C4297" bold="true" box="[855,1088,952,976]" italics="true" pageId="96" pageNumber="96">Kukulcania brignolii</emphasis>
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<heading id="8758C4500A3B2146C053FC46299242B6" box="[822,990,985,1009]" centered="true" fontSize="10" level="3" pageId="96" pageNumber="96" reason="4">
(
<bibRefCitation id="B83E0ECD0A3B2146C058FC46299A42B6" author="Alayon G., G." box="[829,982,985,1009]" journalOrPublisher="Poeyana" pageId="96" pageNumber="96" pagination="1 - 3" part="229" refId="ref85600" refString="Alayon G., G. 1981. Nueva especie de Filistata Latreille, 1810 (Araneae: Filistatidae), de Mexico. Poeyana 229: 1 - 3." title="Nueva especie de Filistata Latreille, 1810 (Araneae: Filistatidae), de Mexico." type="journal article" year="1981">Alayón, 1981</bibRefCitation>
)
</heading>
</emphasis>
</taxonomicName>
, comb. nov.
</paragraph>
</subSubSection>
<subSubSection id="94B520B70A3B2146C06BFB942EC74564" box="[782,1163,1035,1059]" pageId="96" pageNumber="96" type="description">
<paragraph id="DC10733C0A3B2146C06BFB942EC74564" blockId="96.[782,1163,1035,1059]" box="[782,1163,1035,1059]" pageId="96" pageNumber="96">
<figureCitation id="44946FB90A3B2146C06BFB9429CD4564" box="[782,897,1035,1059]" captionStart="FIGURE 2" captionStartId="5.[108,190,1567,1588]" captionTargetBox="[109,1213,248,1538]" captionTargetId="figure@5.[109,1213,248,1538]" captionTargetPageId="5" captionText="FIGURE 2. Kukulcania spp., living specimens. A. K . hurca (Chamberlin and Ivie, 1942), male from California, San Diego Co., Anza-Borrego Desert State Park (SDSU). Photo by M. Hedin. B. Kukulcania sp. from California. Photo by M. Hedin. C. K . arizonica (Chamberlin and Ivie, 1935), male (MACN-Ar 33844). D. K . tequila, sp. nov., male from Mexico, Colima, Cuauhtémoc. Photo by S. Longhorn.. E. K .geophila (Chamberlin and Ivie, 1935), female from California (CAS 9058471). Photo by K.-I. Ueda. F. Kukulcania sp., probably K . brignolii (Alayón, 1981), comb. nov., female with eggsac from Mexico, Puebla, Reserva de la Biosfera Tehuacán-Cuicatlán. Photo by F. Vol. G–I. Kukulcania tractans (O.Pickard-Cambridge, 1896) from Mexico, Guerrero. Photos by S. Longhorn. G. Webs in stone wall in Xochipala. H. Female guarding eggsac in Venta Vieja (=Venta del Zopilote). I. (Subadult?) female from Amula." httpUri="https://zenodo.org/record/3259022/files/figure.png" pageId="96" pageNumber="96">Figures 2F</figureCitation>
, 
<figureCitation id="44946FB90A3B2146C0EEFB9429E64564" box="[907,938,1035,1059]" captionStart="FIGURE 4" captionStartId="8.[133,215,1640,1661]" captionTargetBox="[132,1236,248,1612]" captionTargetId="figure@8.[132,1236,248,1612]" captionTargetPageId="8" captionText="FIGURE 4. Distribution map of A, Kukulcania tractans (O. Pickard-Cambridge, 1896) and K . santosi. (white dots represent literature records from Brescovit and Santos, 2013), B, K . benita, sp. nov., K . brignolii (Alayón, 1981), comb. nov., K . cochimi, sp. nov., K . geophila (Chamberlin and Ivie, 1935), K . gertschi, sp. nov., and K . tequila, and C, K . arizonica (Chamberlin and Ivie, 1935), K . bajacali, sp. nov., K . chingona, sp. nov., and K . mexicana, sp. nov." httpUri="https://zenodo.org/record/3259026/files/figure.png" pageId="96" pageNumber="96">4B</figureCitation>
, 
<figureCitation id="44946FB90A3B2146C0D3FB94299B4564" box="[950,983,1035,1059]" captionStart="FIGURE 5" captionStartId="9.[108,195,1620,1641]" captionTargetBox="[111,1206,248,1592]" captionTargetId="figure@9.[111,1206,248,1592]" captionTargetPageId="9" captionText="FIGURE 5. W.J. Gertsch’s unpublished drawings and sketches of Kukulcania spp., AMNH Library Archives DR 191, reproduced with permission from the AMNH library. A. K . tractans (O. Pickard-Cambridge, 1896), male right bulb. B. K . geophila (Chamberlin and Ivie, 1935), female habitus, dorsal. C. K . utahana (Chamberlin and Ivie, 1935), male bulb. D. K . brignolii (Alayón, 1981), male bulb. E. K . santosi, spermathecae. F. K . hurca (Chamberlin and Ivie, 1942), spermathecae. G. K . cochimi, spermathecae. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/3259028/files/figure.png" pageId="96" pageNumber="96">5D</figureCitation>
, 
<figureCitation id="44946FB90A3B2146C086FB942E494564" box="[995,1029,1035,1059]" captionStart="FIGURE 8" captionStartId="15.[109,196,1640,1661]" captionTargetBox="[188,1113,248,1610]" captionTargetId="figure@15.[109,1213,248,1612]" captionTargetPageId="15" captionText="FIGURE 8. Kukulcania spp., female spermathecae, ventral. A. K . brignolii (Alayón, 1981), comb. nov. (AMNH IFM-1524). B. K . mexicana, paratype (CNAN-Ar DNA0043). C. K . santosi, paratype (AMNH IFM-1528). Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/3259034/files/figure.png" pageId="96" pageNumber="96">8A</figureCitation>
, 
<figureCitation id="44946FB90A3B2146C774FB942E714564" box="[1041,1085,1035,1059]" captionStart="FIGURE 31" captionStartId="40.[131,218,1577,1598]" captionTargetBox="[131,1235,248,1549]" captionTargetId="figure@40.[131,1235,248,1549]" captionTargetPageId="40" captionText="FIGURE 31. Kukulcania spp., female spermathecae, ventral view except where noted. A. Kukulcania hurca (Chamberlin and Ivie, 1942) (AMNH IFM-1402). B. K . brignolii (Alayón, 1981), comb. nov. (AMNH IFM- 1406). C. K . mexicana (AMNH IFM-1405). D. K . santosi (AMNH IFM-1403), dorsal. E. K . tractans (O. Pickard-Cambridge, 1896) (AMNH IFM-1416). F. K . tequila, paratype (AMNH IFM-1666). G. Same, anterior. H. K . geophila (Chamberlin and Ivie, 1935) (AMNH IFM-1391)." httpUri="https://zenodo.org/record/3259080/files/figure.png" pageId="96" pageNumber="96">31B</figureCitation>
, 
<figureCitation id="44946FB90A3B2146C72DFB942EC74564" box="[1096,1163,1035,1059]" captionStart-0="FIGURE 57" captionStart-1="FIGURE 58" captionStart-2="FIGURE 59" captionStart-3="FIGURE 60" captionStartId-0="98.[132,219,1644,1665]" captionStartId-1="99.[108,195,1183,1204]" captionStartId-2="100.[131,218,1130,1151]" captionStartId-3="101.[108,192,1577,1598]" captionTargetBox-0="[132,1233,248,1602]" captionTargetBox-1="[111,1213,248,1149]" captionTargetBox-2="[132,1217,248,1087]" captionTargetBox-3="[109,1213,248,1549]" captionTargetId-0="figure@98.[131,1235,248,1612]" captionTargetId-1="figure@99.[109,1213,248,1155]" captionTargetId-2="figure@100.[132,1236,248,1101]" captionTargetId-3="figure@101.[109,1213,248,1549]" captionTargetPageId-0="98" captionTargetPageId-1="99" captionTargetPageId-2="100" captionTargetPageId-3="101" captionText-0="FIGURE 57. Kukulcania brignolii (Alayón, 1981), comb. nov., male holotype from Mexico, Oaxaca, Monte Albán (MCVR), habitus. A. Dorsal. B. Lateral. C. Ventral. D. Subanterior. E. Labels associated to the holotype. Scale bars = 1 mm." captionText-1="FIGURE 58. Kukulcania brignolii (Alayón, 1981), comb. nov., male holotype from Mexico, Oaxaca, Monte Albán (MCVR), leg I and genitalia. A. Left leg I, prolateral. B. Left bulb, prolateral. C. Right bulb, prolateral, mirrored. D. Left palp, prolateral. E. Left bulb, dorsal. F. Left bulb, retrolateral. Scale bars = 1 mm (A–B), 0.1 mm (C–F)." captionText-2="FIGURE 59. Kukulcania brignolii (Alayón, 1981), comb. nov., female from Mexico, Oaxaca, Monte Albán (AMNH IFM-1637), habitus. A. Dorsal. B. Lateral. C. Ventral. Scale bars = 1 mm." captionText-3="FIGURE 60. Kukulcania brignolii (Alayón, 1981), comb. nov., female spermathecae, ventral. A. Oaxaca, Monte Albán (AMNH IFM-1637). B. Puebla, Tehuacán (AMNH-1524). C. Puebla, Tehuacán (AMNH-1525). D. Oaxaca, carretera Ixtlán-Oaxaca (CNAN-Ar 6635). E. Puebla, 12 miles N Acatlán (AMNH IFM-1569). F. Oaxaca, 7.4 miles N of Telixtlahuaca (CAS 9057632). G. Puebla, 62 km west of of Orizaba (AMNH IFM-1573). H. Sierra Mixteca(?) (MfN). Abbreviations: GPS, glandular portion of spermathecae apex; IF, interpulmonary fold; MPS, membranous portion of spermathecae apex; SB, sclerotized bars; UE, uterus externus." httpUri-0="https://zenodo.org/record/3259134/files/figure.png" httpUri-1="https://zenodo.org/record/3259136/files/figure.png" httpUri-2="https://zenodo.org/record/3259138/files/figure.png" httpUri-3="https://zenodo.org/record/3259140/files/figure.png" pageId="96" pageNumber="96">57–60</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="94B520B70A3B2146C1A1FBDE292F45D9" pageId="96" pageNumber="96" type="reference_group">
<paragraph id="DC10733C0A3B2146C1A1FBDE292F45D9" blockId="96.[708,1231,1089,1182]" pageId="96" pageNumber="96">
<taxonomicName id="1BAF08BF0A3B2146C1A1FBDE2E6B451E" ID-CoL="933VZ" authority="Alayon, 1981: 2" authorityName="Alayon" authorityPageNumber="2" authorityYear="1981" box="[708,1063,1089,1113]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="96" pageNumber="96" phylum="Arthropoda" rank="species" species="brignolii">
<emphasis id="EEDBAF2E0A3B2146C1A1FBDE2934451E" box="[708,888,1089,1113]" italics="true" pageId="96" pageNumber="96">Filistata brignolii</emphasis>
<bibRefCitation id="B83E0ECD0A3B2146C01AFBDE2E6B451E" author="Alayon G., G." box="[895,1063,1089,1113]" journalOrPublisher="Poeyana" pageId="96" pageNumber="96" pagination="1 - 3" part="229" refId="ref85600" refString="Alayon G., G. 1981. Nueva especie de Filistata Latreille, 1810 (Araneae: Filistatidae), de Mexico. Poeyana 229: 1 - 3." title="Nueva especie de Filistata Latreille, 1810 (Araneae: Filistatidae), de Mexico." type="journal article" year="1981">Alayón, 1981: 2</bibRefCitation>
</taxonomicName>
, figs. 1–2. Male holotype from Mexico, Oaxaca (MCVR), examined.
</paragraph>
</subSubSection>
<subSubSection id="94B520B70A3B2146C181FB2529604472" pageId="96" pageNumber="96" type="materials_examined">
<paragraph id="DC10733C0A3B2146C181FB2529604472" blockId="96.[708,1236,1210,1765]" pageId="96" pageNumber="96">
EXAMINED TYPE MATERIAL: 
<materialsCitation id="6CC779610A3B2146C776FB2529604472" ID-GBIF-Occurrence="2270855588" collectingDate="1973-08-07" collectionCode="MCVR" collectorName="P. Brignoli" country="MEX- ICO" latitude="17.0438" location="Monte Alban" longLatPrecision="5" longitude="-96.76811" municipality="Oaxaca" pageId="96" pageNumber="96" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="0314CD9E0A3B2146C776FB252EC04595" box="[1043,1164,1210,1234]" pageId="96" pageNumber="96" type="holotype">HOLOTYPE</typeStatus>
: 
<emphasis id="EEDBAF2E0A3B2146C7F0FB2528BC45B4" bold="true" pageId="96" pageNumber="96">
<collectingCountry id="A4B833AC0A3B2146C7F0FB2528BC45B4" pageId="96" pageNumber="96">MEX- ICO</collectingCountry>
</emphasis>
. 
<emphasis id="EEDBAF2E0A3B2146C075FB44293D45B4" bold="true" box="[784,881,1243,1267]" pageId="96" pageNumber="96">
<collectingMunicipality id="3C74E9460A3B2146C075FB44292145B4" box="[784,877,1243,1267]" pageId="96" pageNumber="96">Oaxaca</collectingMunicipality>
:
</emphasis>
<emphasis id="EEDBAF2E0A3B2146C0ECFB442E7C45B4" box="[905,1072,1243,1267]" italics="true" pageId="96" pageNumber="96">
<location id="D97025E70A3B2146C0ECFB442E7C45B4" LSID="urn:lsid:plazi:treatment:5406C22A0A3B2140C032FC2729DF47A2:D97025E70A3B2146C0ECFB442E7C45B4" box="[905,1072,1243,1267]" country="MEX- ICO" latitude="17.0438" longLatPrecision="5" longitude="-96.76811" municipality="Oaxaca" name="Monte Alban" pageId="96" pageNumber="96">Monte Albán</location>
</emphasis>
[
<geoCoordinate id="B99B15FB0A3B2146C72AFB442E9C45B4" box="[1103,1232,1243,1267]" degrees="17.0438" direction="north" orientation="latitude" pageId="96" pageNumber="96" precision="5" value="17.0438">N17.0438°</geoCoordinate>
, 
<geoCoordinate id="B99B15FB0A3B2146C1A1FB6329144453" box="[708,856,1276,1300]" degrees="96.76811" direction="west" orientation="longitude" pageId="96" pageNumber="96" precision="1" value="-96.76811">W96.76811°</geoCoordinate>
], 
<collectorName id="715A16EA0A3B2146C01DFB632E4E4453" box="[888,1026,1276,1300]" pageId="96" pageNumber="96">P. Brignoli</collectorName>
, 
<date id="A81155FC0A3B2146C77EFB632EDA4453" box="[1051,1174,1276,1300]" pageId="96" pageNumber="96" value="1973-08-07">
<collectingDate id="B855AC140A3B2146C77EFB632EDA4453" box="[1051,1174,1276,1300]" pageId="96" pageNumber="96" value="1973-08-07">7.viii.1973</collectingDate>
</date>
, 
<specimenCount id="CAA9B8B50A3B2146C7CAFB632E984456" box="[1199,1236,1274,1300]" pageId="96" pageNumber="96" type="male">1♂</specimenCount>
(
<collectionCode id="BABEEBF90A3B2146C1A9FA82296D4472" box="[716,801,1309,1333]" collectionName="MCVR" pageId="96" pageNumber="96">MCVR</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="94B520B70A3B2147C181FAA1283A4030" lastPageId="97" lastPageNumber="97" pageId="96" pageNumber="96" type="discussion">
<paragraph id="DC10733C0A3B2147C181FAA1283A4030" blockId="96.[708,1236,1210,1765]" lastBlockId="97.[108,637,252,1764]" lastPageId="97" lastPageNumber="97" pageId="96" pageNumber="96">
REMARKS: 
<bibRefCitation id="B83E0ECD0A3B2146C039FAA129B64411" author="Alayon G., G." box="[860,1018,1342,1366]" journalOrPublisher="Poeyana" pageId="96" pageNumber="96" pagination="1 - 3" part="229" refId="ref85600" refString="Alayon G., G. 1981. Nueva especie de Filistata Latreille, 1810 (Araneae: Filistatidae), de Mexico. Poeyana 229: 1 - 3." title="Nueva especie de Filistata Latreille, 1810 (Araneae: Filistatidae), de Mexico." type="journal article" year="1981">Alayón (1981)</bibRefCitation>
described this species under 
<taxonomicName id="1BAF08BF0A3B2146C05EFAC029DD4430" authorityName="Latreille" authorityYear="1810" box="[827,913,1375,1399]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="96" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="EEDBAF2E0A3B2146C05EFAC029DD4430" box="[827,913,1375,1399]" italics="true" pageId="96" pageNumber="96">Filistata</emphasis>
</taxonomicName>
stating that 
<bibRefCitation id="B83E0ECD0A3B2146C77FFAC02E984430" author="Lehtinen, P. T." box="[1050,1236,1375,1399]" journalOrPublisher="Annales Zoologici Fennici" pageId="96" pageNumber="96" pagination="199 - 468" part="4" refId="ref86818" refString="Lehtinen, P. T. 1967. Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici 4: 199 - 468." title="Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha" type="journal article" year="1967">Lehtinen’s (1967)</bibRefCitation>
decision to erect 
<taxonomicName id="1BAF08BF0A3B2146C01EFA1E29B944DE" authorityName="Lehtinen. The" authorityYear="1967" box="[891,1013,1409,1433]" class="Arachnida" family="Filistatidae" genus="Kukulcania" kingdom="Animalia" order="Araneae" pageId="96" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="EEDBAF2E0A3B2146C01EFA1E29B944DE" box="[891,1013,1409,1433]" italics="true" pageId="96" pageNumber="96">Kukulcania</emphasis>
</taxonomicName>
for New World filistatines was not well justified and that additional studies on the phylogenetic relationships of the family were necessary. He did not explicitly state his reasons to doubt the validity of 
<emphasis id="EEDBAF2E0A3B2146C736F99A2E98475A" box="[1107,1236,1541,1565]" italics="true" pageId="96" pageNumber="96">
<taxonomicName id="1BAF08BF0A3B2146C736F99A2E81475A" authorityName="Lehtinen. The" authorityYear="1967" box="[1107,1229,1541,1565]" class="Arachnida" family="Filistatidae" genus="Kukulcania" kingdom="Animalia" order="Araneae" pageId="96" pageNumber="96" phylum="Arthropoda" rank="genus">Kukulcania</taxonomicName>
.
</emphasis>
The possession of sclerotized bars in the spermathecae of 
<emphasis id="EEDBAF2E0A3B2146C04CF9D629D24727" box="[809,926,1608,1632]" italics="true" pageId="96" pageNumber="96">
K. 
<taxonomicName id="1BAF08BF0A3B2146C023F9D729D24727" authorityName="Alayon" authorityYear="1981" box="[838,926,1608,1632]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="96" pageNumber="96" phylum="Arthropoda" rank="species" species="brignolii">brignolii</taxonomicName>
</emphasis>
is a character so far recorded only in 
<emphasis id="EEDBAF2E0A3B2146C04BF9F629F947C6" box="[814,949,1641,1665]" italics="true" pageId="96" pageNumber="96">
<taxonomicName id="1BAF08BF0A3B2146C04BF9F629FD47C6" authorityName="Lehtinen. The" authorityYear="1967" box="[814,945,1641,1665]" class="Arachnida" family="Filistatidae" genus="Kukulcania" kingdom="Animalia" order="Araneae" pageId="96" pageNumber="96" phylum="Arthropoda" rank="genus">Kukulcania</taxonomicName>
,
</emphasis>
indicating this species belongs here and not in 
<emphasis id="EEDBAF2E0A3B2146C0B2F9152E7847E5" box="[983,1076,1674,1698]" italics="true" pageId="96" pageNumber="96">
<taxonomicName id="1BAF08BF0A3B2146C0B2F9152E6147E5" authorityName="Latreille" authorityYear="1810" box="[983,1069,1674,1698]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="96" pageNumber="96" phylum="Arthropoda" rank="genus">Filistata</taxonomicName>
.
</emphasis>
Hence, a new combination is proposed. With this transfer, all American species once placed in 
<taxonomicName id="1BAF08BF0A3B2146C725F9522ED447A2" authorityName="Latreille" authorityYear="1810" box="[1088,1176,1741,1765]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="96" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="EEDBAF2E0A3B2146C725F9522ED447A2" box="[1088,1176,1741,1765]" italics="true" pageId="96" pageNumber="96">Filistata</emphasis>
</taxonomicName>
have been accommodated in New World genera. In the years before the publication of 
<bibRefCitation id="B83E0ECD0A3A2147C145FE822AFE4011" author="Alayon G., G." journalOrPublisher="Poeyana" pageId="97" pageNumber="97" pagination="1 - 3" part="229" refId="ref85600" refString="Alayon G., G. 1981. Nueva especie de Filistata Latreille, 1810 (Araneae: Filistatidae), de Mexico. Poeyana 229: 1 - 3." title="Nueva especie de Filistata Latreille, 1810 (Araneae: Filistatidae), de Mexico." type="journal article" year="1981">Alayón’s (1981)</bibRefCitation>
work, W.J. Gertsch recognized this species under the manuscript name “ 
<emphasis id="EEDBAF2E0A3A2147C2D5FEC028274030" box="[432,619,351,375]" italics="true" pageId="97" pageNumber="97">
<taxonomicName id="1BAF08BF0A3A2147C2D5FEC028294030" box="[432,613,351,375]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="97" pageNumber="97" phylum="Arthropoda" rank="species" species="tehuana">Filistata tehuana</taxonomicName>
.
</emphasis>
”
</paragraph>
</subSubSection>
<subSubSection id="94B520B70A3A2147C3E9FE1F2B3542EE" pageId="97" pageNumber="97" type="diagnosis">
<paragraph id="DC10733C0A3A2147C3E9FE1F2B3542EE" blockId="97.[108,637,252,1764]" pageId="97" pageNumber="97">
DIAGNOSIS: Males are similar to those of 
<emphasis id="EEDBAF2E0A3A2147C101FE1E2A9940FE" italics="true" pageId="97" pageNumber="97">
K. 
<taxonomicName id="1BAF08BF0A3A2147C309FE3D2A9940FE" authorityName="Magalhaes &amp; Ramírez" authorityYear="2019" box="[108,213,418,441]" class="Arachnida" family="Filistatidae" genus="Kukulcania" kingdom="Animalia" order="Araneae" pageId="97" pageNumber="97" phylum="Arthropoda" rank="species" species="mexicana">mexicana</taxonomicName>
</emphasis>
and 
<emphasis id="EEDBAF2E0A3A2147C270FE3D2BC940FE" box="[277,389,418,441]" italics="true" pageId="97" pageNumber="97">
K. 
<taxonomicName id="1BAF08BF0A3A2147C25FFE3D2BC940FE" authorityName="Magalhaes &amp; Ramírez" authorityYear="2019" box="[314,389,418,441]" class="Arachnida" family="Filistatidae" genus="Kukulcania" kingdom="Animalia" order="Araneae" pageId="97" pageNumber="97" phylum="Arthropoda" rank="species" species="santosi">santosi</taxonomicName>
</emphasis>
in having incrassate, strong setae in the prolateral border of the cymbium. They differ in having a short, uncoiled embolus (long, two coils in 
<emphasis id="EEDBAF2E0A3A2147C2FEFD9A286E435B" box="[411,546,517,540]" italics="true" pageId="97" pageNumber="97">
K. 
<taxonomicName id="1BAF08BF0A3A2147C2DEFD9A286E435B" authorityName="Magalhaes &amp; Ramírez" authorityYear="2019" box="[443,546,517,540]" class="Arachnida" family="Filistatidae" genus="Kukulcania" kingdom="Animalia" order="Araneae" pageId="97" pageNumber="97" phylum="Arthropoda" rank="species" species="mexicana">mexicana</taxonomicName>
</emphasis>
) and by the simple apex of the incrassate setae on the cymbium (fig. 58) (spatulate in 
<emphasis id="EEDBAF2E0A3A2147C29DFDD728204318" box="[504,620,584,607]" italics="true" pageId="97" pageNumber="97">
K. 
<taxonomicName id="1BAF08BF0A3A2147C145FDD728204318" authorityName="Magalhaes &amp; Ramírez" authorityYear="2019" box="[544,620,584,607]" class="Arachnida" family="Filistatidae" genus="Kukulcania" kingdom="Animalia" order="Araneae" pageId="97" pageNumber="97" phylum="Arthropoda" rank="species" species="santosi">santosi</taxonomicName>
</emphasis>
). Females are similar to 
<emphasis id="EEDBAF2E0A3A2147C21DFDF6284943C7" box="[376,517,617,640]" italics="true" pageId="97" pageNumber="97">
K. 
<taxonomicName id="1BAF08BF0A3A2147C2F9FDF6284943C7" authorityName="Magalhaes &amp; Ramírez" authorityYear="2019" box="[412,517,617,640]" class="Arachnida" family="Filistatidae" genus="Kukulcania" kingdom="Animalia" order="Araneae" pageId="97" pageNumber="97" phylum="Arthropoda" rank="species" species="mexicana">mexicana</taxonomicName>
</emphasis>
in having long, curved sclerotized bars that do not taper posteriorly, and spermathecae that project anteriorly to the margin of the uterus externus. 
<emphasis id="EEDBAF2E0A3A2147C101FD532A8A4243" italics="true" pageId="97" pageNumber="97">
K. 
<taxonomicName id="1BAF08BF0A3A2147C309FD732A8A4243" authorityName="Alayon" authorityYear="1981" box="[108,198,748,772]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="97" pageNumber="97" phylum="Arthropoda" rank="species" species="brignolii">brignolii</taxonomicName>
</emphasis>
might be distinguished by the smaller glandular portion of the spermathecae apex, which is placed ventrally to the membranous portion (larger, ectally placed glandular portion in 
<emphasis id="EEDBAF2E0A3A2147C3EBFCEE2B5742CF" box="[142,283,881,904]" italics="true" pageId="97" pageNumber="97">
K. 
<taxonomicName id="1BAF08BF0A3A2147C3D7FCEE2B5742CF" authorityName="Magalhaes &amp; Ramírez" authorityYear="2019" box="[178,283,881,904]" class="Arachnida" family="Filistatidae" genus="Kukulcania" kingdom="Animalia" order="Araneae" pageId="97" pageNumber="97" phylum="Arthropoda" rank="species" species="mexicana">mexicana</taxonomicName>
</emphasis>
), and by the usually less bent sclerotized bars (fig. 60).
</paragraph>
</subSubSection>
<subSubSection id="94B520B70A3A2147C3E9FC2C29C447E5" pageId="97" pageNumber="97" type="description">
<paragraph id="DC10733C0A3A2147C3E9FC2C2E0540BC" blockId="97.[108,637,252,1764]" lastBlockId="97.[684,1214,252,1764]" pageId="97" pageNumber="97">
DESCRIPTION: Male holotype from Monte Albán, Oaxaca, Mexico (MCVR). Coloration yellowish cream. Carapace finely stippled with light brown. Abdomen dorsum light brown around cardiac area and in posterior end. Clypeus short. Sternum oval, with two pairs of sigillae. Total length 7.45. Carapace length 3.36, width 2.81, clypeus length 0.27. Eye diameters and interdistances: AME‰.15; PME‰.214; 
<taxonomicName id="1BAF08BF0A3A2147C2D5FB252B924594" authorityName="Pascoe" authorityYear="1865" box="[432,478,1210,1235]" class="Insecta" family="Cerambycidae" genus="Ale" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="97" pageNumber="97" phylum="Arthropoda" rank="genus">ALE</taxonomicName>
‰.249; PLE 0.235; AME–AME 0.074; PME–PME 0.233. Sternum length 1.54, width 1.62. Palp: femur length 4.33, height 0.54; tibia length 4.43, height 0.64. Leg I: femur (fe) 7.21; patella (pa) 1.43; tibia (ti) 7.29; metatarsus (mt) 8.37; tarsus (ta) 4.11. II: fe 5.68; pa 1.32; ti 5.11; mt 5.84; ta 2.5. III: fe 4.67; pa 1.11; ti 4.36; mt 5.67; ta 3.64. IV: fe 6.39; pa 1.4; ti 6.2; mt 7.72; ta 3.42. Abdomen: length 4.33, width 2.1. Palp macrosetae long, in several rows along femur ventral face. Leg macrosetae: fe I 1d, 15p, 4v, 8r; ti I 4p, 7v; mt I 1p, 14v, 1r; ta I 7v; fe II 4d, 6p, 5v, 4r; ti II 2p, 6v; mt II 3p, 12v, 3r; ta II 9v; fe III 5d, 5p, 12v, 1r; ti III 1d, 3p, 4v, 3r; mt III 1d, 2p, 16v, 2r; ta III 11v; fe IV 5d, 3p, 6v, 3r; ti IV 1p, 2v, 2r; mt IV 2p, 15v, 2r; ta IV 11v. Palp: cymbium long, with prolateral edge slightly protruding and bearing a forelock of incrassate setae; bulb short, piriform; sperm duct with three tightly packed coils; embolus short, pointing dorsally, with a small keel near the base. State of the specimen: well preserved but very fragile, left leg I broken from the base of the femur, left leg II loose, left palp dissected, missing some of the prolateral thick setae.
</paragraph>
<paragraph id="DC10733C0A3A2147C1A9FD9A29C447E5" blockId="97.[684,1214,252,1764]" pageId="97" pageNumber="97">
Female from Monte Albán, Oaxaca, Mexico (AMNH IFM-1637). Coloration dark yellowish brown. Carapace stippled with dark brown. Sternum and first femora and tibiae hirsute, with long setae. Legs brown, except for yellowish-brown metatarsi and tarsi, coxae with lighter, longitudinal markings. Abdomen dorsum brown. Anterior margin of the carapace unmodified. Sternum oval, with two pairs of sigillae. Total length 10.29. Carapace length 4.55, width 3.34, clypeus length 0.64. Eye diameters and interdistances: AME 0.142; PME 0.252; 
<taxonomicName id="1BAF08BF0A3A2147C02DFC0D293042ED" authorityName="Pascoe" authorityYear="1865" box="[840,892,914,938]" class="Insecta" family="Cerambycidae" genus="Ale" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="97" pageNumber="103" phylum="Arthropoda" rank="genus">ALE</taxonomicName>
0.302; PLE 0.327; AME– AME 0.114; PME–PME 0.28. Sternum length 1.98, width 2.01. Palp: femur length 2.56, height 0.89; tibia length 1.4, height 0.75. Leg I: femur (fe) 5.05; patella (pa) 1.72; tibia (ti) 4.9; metatarsus (mt) 4.34; tarsus (ta) 2.1. II: fe 4.06; pa 1.65; ti 3.45; mt 3.16; ta 1.76. III: fe 3.21; pa 1.37; ti 2.62; mt 2.81; ta 1.58. IV: fe 4.55; pa 1.5; ti 3.96; mt 3.68; ta 1.73. Abdomen: length 5.88, width 3.59. Palp macrosetae on ventral surface of tibia and tarsus. Leg macrosetae present on ventral surfaces of tibiae, metatarsi and tarsi; femora with 2–4 dorsal macrosetae. Calamistrum with three rows with 7–10 setae each. Interpulmonary fold large, rounded, covering the spermathecae dorsally. Sclerotized lateral bars present, elongate, slightly curved, positioned posteriorly to receptacles; membranous portion of the spermathecae apex subrounded; glandular portion positioned ventrally to the membranous portion. State of the specimen: good, many setae missing, right legs I and II missing from tibia, genitalia dissected.
</paragraph>
</subSubSection>
<subSubSection id="94B520B70A3A2140C1A9F9342B2D4385" lastPageId="102" lastPageNumber="102" pageId="97" pageNumber="97" type="discussion">
<paragraph id="DC10733C0A3A2140C1A9F9342B2D4385" blockId="97.[684,1214,252,1764]" lastBlockId="102.[132,661,252,1751]" lastPageId="102" lastPageNumber="102" pageId="97" pageNumber="97">
INTRASPECIFIC VARIATION: Males (
<emphasis id="EEDBAF2E0A3A2147C700F9332ED84784" box="[1125,1172,1708,1731]" italics="true" pageId="97" pageNumber="97">N =</emphasis>
2): total length 7.45–8.34 (7.9), carapace length 3.36–3.71 (3.54), femur I length 7.21–7.46 (7.34), tibia I length 6.98–7.29 (7.14), femur/ carapace ratio 2.01–2.15 (2.08). Females (
<emphasis id="EEDBAF2E0A3D2140C102FEA028D94011" box="[615,661,319,342]" italics="true" pageId="102" pageNumber="102">N =</emphasis>
5): total length 10.29–12.77 (12.14), carapace length 4.55–5.6 (5.08), femur I length 5.05– 6.34 (5.85), tibia I length 4.9–6.08 (5.6), femur/ carapace ratio 1.11–1.26 (1.15). The shape of the female genitalia is variable, especially the shape of the sclerotized bars (fig. 60), and some specimens are somewhat intermediate with the morphology found in 
<emphasis id="EEDBAF2E0A3D2140C298FDD728C24318" box="[509,654,584,607]" italics="true" pageId="102" pageNumber="102">
K. 
<taxonomicName id="1BAF08BF0A3D2140C144FDD728C24318" authority="Magalhaes &amp; Ramírez, 2019" authorityName="Magalhaes &amp; Ramírez" authorityYear="2019" box="[545,654,584,607]" class="Arachnida" family="Filistatidae" genus="Kukulcania" kingdom="Animalia" order="Araneae" pageId="102" pageNumber="102" phylum="Arthropoda" rank="species" species="mexicana" status="sp. nov.">mexicana</taxonomicName>
</emphasis>
, 
<taxonomicNameLabel id="F5E812550A3D2140C3E1FDF72A904338" box="[132,220,616,639]" pageId="102" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
, although these have been collected in areas with males or females with typical morphology (fig. 60H).
</paragraph>
</subSubSection>
<caption id="88D023B40A392144C3E1F9F32B0547FA" httpUri="https://zenodo.org/record/3259134/files/figure.png" pageId="98" pageNumber="98" startId="98.[132,219,1644,1665]" targetBox="[132,1233,248,1602]" targetPageId="98">
<paragraph id="DC10733C0A392144C3E1F9F32B0547FA" blockId="98.[132,1237,1642,1725]" pageId="98" pageNumber="98">
FIGURE 57. 
<taxonomicName id="1BAF08BF0A392144C26FF9F4283047C4" authority="(Alayon, 1981)" authorityName="Magalhaes &amp; Ramírez" authorityYear="2019" baseAuthorityName="Alayon" baseAuthorityYear="1981" box="[266,636,1642,1667]" class="Arachnida" family="Filistatidae" genus="Kukulcania" kingdom="Animalia" order="Araneae" pageId="98" pageNumber="98" phylum="Arthropoda" rank="species" species="brignolii" status="comb. nov.">
<emphasis id="EEDBAF2E0A392144C26FF9F42B3047C4" box="[266,380,1643,1667]" italics="true" pageId="98" pageNumber="98">Kukulcania</emphasis>
<emphasis id="EEDBAF2E0A392144C2E0F9F42B9547C4" box="[389,473,1643,1667]" italics="true" pageId="98" pageNumber="98">brignolii</emphasis>
(
<bibRefCitation id="B83E0ECD0A392144C28DF9F5283947C4" author="Alayon G., G." box="[488,629,1642,1667]" journalOrPublisher="Poeyana" pageId="98" pageNumber="98" pagination="1 - 3" part="229" refId="ref85600" refString="Alayon G., G. 1981. Nueva especie de Filistata Latreille, 1810 (Araneae: Filistatidae), de Mexico. Poeyana 229: 1 - 3." title="Nueva especie de Filistata Latreille, 1810 (Araneae: Filistatidae), de Mexico." type="journal article" year="1981">Alayón, 1981</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="F5E812550A392144C1ECF9F528B147C5" box="[649,765,1642,1666]" pageId="98" pageNumber="98" rank="species">comb. nov.</taxonomicNameLabel>
, male holotype from Mexico, Oaxaca, Monte Albán (MCVR), habitus. 
<emphasis id="EEDBAF2E0A392144C2E4F9172BD547E7" bold="true" box="[385,409,1672,1696]" pageId="98" pageNumber="98">A.</emphasis>
Dorsal. 
<emphasis id="EEDBAF2E0A392144C288F917284F47E7" bold="true" box="[493,515,1672,1696]" pageId="98" pageNumber="98">B.</emphasis>
Lateral. 
<emphasis id="EEDBAF2E0A392144C13FF917283D47E7" bold="true" box="[602,625,1672,1696]" pageId="98" pageNumber="98">C.</emphasis>
Ventral. 
<emphasis id="EEDBAF2E0A392144C1A9F91728A847E7" bold="true" box="[716,740,1672,1696]" pageId="98" pageNumber="98">D.</emphasis>
Subanterior. 
<emphasis id="EEDBAF2E0A392144C00FF917293347E7" bold="true" box="[874,895,1672,1696]" pageId="98" pageNumber="98">E.</emphasis>
Labels associated to the holotype. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="88D023B40A382145C309FB002AAC4449" httpUri="https://zenodo.org/record/3259136/files/figure.png" pageId="99" pageNumber="99" startId="99.[108,195,1183,1204]" targetBox="[111,1213,248,1149]" targetPageId="99">
<paragraph id="DC10733C0A382145C309FB002AAC4449" blockId="99.[108,1212,1181,1294]" pageId="99" pageNumber="99">
FIGURE 58. 
<taxonomicName id="1BAF08BF0A382145C397FB02282845F1" authority="(Alayon, 1981)" authorityName="Magalhaes &amp; Ramírez" authorityYear="2019" baseAuthorityName="Alayon" baseAuthorityYear="1981" box="[242,612,1181,1206]" class="Arachnida" family="Filistatidae" genus="Kukulcania" kingdom="Animalia" order="Araneae" pageId="99" pageNumber="99" phylum="Arthropoda" rank="species" species="brignolii" status="comb. nov.">
<emphasis id="EEDBAF2E0A382145C397FB022B2845F2" box="[242,356,1181,1205]" italics="true" pageId="99" pageNumber="99">Kukulcania</emphasis>
<emphasis id="EEDBAF2E0A382145C208FB022B8D45F2" box="[365,449,1181,1205]" italics="true" pageId="99" pageNumber="99">brignolii</emphasis>
(
<bibRefCitation id="B83E0ECD0A382145C2B5FB02281145F1" author="Alayon G., G." box="[464,605,1181,1206]" journalOrPublisher="Poeyana" pageId="99" pageNumber="99" pagination="1 - 3" part="229" refId="ref85600" refString="Alayon G., G. 1981. Nueva especie de Filistata Latreille, 1810 (Araneae: Filistatidae), de Mexico. Poeyana 229: 1 - 3." title="Nueva especie de Filistata Latreille, 1810 (Araneae: Filistatidae), de Mexico." type="journal article" year="1981">Alayón, 1981</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="F5E812550A382145C114FB0228A945F2" box="[625,741,1181,1205]" pageId="99" pageNumber="99" rank="species">comb. nov.</taxonomicNameLabel>
, male holotype from Mexico, Oaxaca, Monte Albán (MCVR), leg I and genitalia. 
<emphasis id="EEDBAF2E0A382145C2BFFB242BBE4594" bold="true" box="[474,498,1211,1235]" pageId="99" pageNumber="99">A.</emphasis>
Left leg I, prolateral. 
<emphasis id="EEDBAF2E0A382145C1ABFB2428A84594" bold="true" box="[718,740,1211,1235]" pageId="99" pageNumber="99">B.</emphasis>
Left bulb, prolateral. 
<emphasis id="EEDBAF2E0A382145C0A5FB24299B4594" bold="true" box="[960,983,1211,1235]" pageId="99" pageNumber="99">C.</emphasis>
Right bulb, prolateral, mirrored. 
<emphasis id="EEDBAF2E0A382145C3B1FB472AA145B7" bold="true" box="[212,237,1240,1264]" pageId="99" pageNumber="99">D.</emphasis>
Left palp, prolateral. 
<emphasis id="EEDBAF2E0A382145C2A2FB472B9045B7" bold="true" box="[455,476,1240,1264]" pageId="99" pageNumber="99">E.</emphasis>
Left bulb, dorsal. 
<emphasis id="EEDBAF2E0A382145C1F1FB4728E845B7" bold="true" box="[660,676,1240,1264]" pageId="99" pageNumber="99">F.</emphasis>
Left bulb, retrolateral. Scale bars = 1 mm (A–B), 0.1 mm (C–F).
</paragraph>
</caption>
<caption id="88D023B40A3F2142C3E6FBF5298045DA" httpUri="https://zenodo.org/record/3259138/files/figure.png" pageId="100" pageNumber="100" startId="100.[131,218,1130,1151]" targetBox="[132,1217,248,1087]" targetPageId="100">
<paragraph id="DC10733C0A3F2142C3E6FBF5298045DA" blockId="100.[131,1235,1128,1182]" pageId="100" pageNumber="100">
FIGURE 59. 
<taxonomicName id="1BAF08BF0A3F2142C26FFBF728CC45C7" authority="(Alayon, 1981)" authorityName="Magalhaes &amp; Ramírez" authorityYear="2019" baseAuthorityName="Alayon" baseAuthorityYear="1981" box="[266,640,1128,1152]" class="Arachnida" family="Filistatidae" genus="Kukulcania" kingdom="Animalia" order="Araneae" pageId="100" pageNumber="100" phylum="Arthropoda" rank="species" species="brignolii" status="comb. nov.">
<emphasis id="EEDBAF2E0A3F2142C26FFBF72B3145C7" box="[266,381,1128,1152]" italics="true" pageId="100" pageNumber="100">Kukulcania</emphasis>
<emphasis id="EEDBAF2E0A3F2142C2E3FBF72B9745C7" box="[390,475,1128,1152]" italics="true" pageId="100" pageNumber="100">brignolii</emphasis>
(
<bibRefCitation id="B83E0ECD0A3F2142C28FFBF7283445C7" author="Alayon G., G." box="[490,632,1128,1152]" journalOrPublisher="Poeyana" pageId="100" pageNumber="100" pagination="1 - 3" part="229" refId="ref85600" refString="Alayon G., G. 1981. Nueva especie de Filistata Latreille, 1810 (Araneae: Filistatidae), de Mexico. Poeyana 229: 1 - 3." title="Nueva especie de Filistata Latreille, 1810 (Araneae: Filistatidae), de Mexico." type="journal article" year="1981">Alayón, 1981</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="F5E812550A3F2142C1E8FBF7294D4538" box="[653,769,1128,1152]" pageId="100" pageNumber="100" rank="species">comb. nov.</taxonomicNameLabel>
, female from Mexico, Oaxaca, Monte Albán (AMNH IFM-1637), habitus. 
<emphasis id="EEDBAF2E0A3F2142C2D6FB1A2B8745DA" bold="true" box="[435,459,1157,1181]" pageId="100" pageNumber="100">A.</emphasis>
Dorsal. 
<emphasis id="EEDBAF2E0A3F2142C147FB19287445D9" bold="true" box="[546,568,1158,1182]" pageId="100" pageNumber="100">B.</emphasis>
Lateral. 
<emphasis id="EEDBAF2E0A3F2142C1F7FB1A28E645DA" bold="true" box="[658,682,1157,1181]" pageId="100" pageNumber="100">C.</emphasis>
Ventral. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="88D023B40A3E2143C309F9B6298B4795" httpUri="https://zenodo.org/record/3259140/files/figure.png" pageId="101" pageNumber="101" startId="101.[108,192,1577,1598]" targetBox="[109,1213,248,1549]" targetPageId="101">
<paragraph id="DC10733C0A3E2143C309F9B6298B4795" blockId="101.[108,1212,1575,1746]" pageId="101" pageNumber="101">
FIGURE 60. 
<taxonomicName id="1BAF08BF0A3E2143C38EF9B8281D4707" authority="(Alayon, 1981)" authorityName="Magalhaes &amp; Ramírez" authorityYear="2019" baseAuthorityName="Alayon" baseAuthorityYear="1981" box="[235,593,1575,1600]" class="Arachnida" family="Filistatidae" genus="Kukulcania" kingdom="Animalia" order="Araneae" pageId="101" pageNumber="101" phylum="Arthropoda" rank="species" species="brignolii" status="comb. nov.">
<emphasis id="EEDBAF2E0A3E2143C38EF9B82B164778" box="[235,346,1575,1599]" italics="true" pageId="101" pageNumber="101">Kukulcania</emphasis>
<emphasis id="EEDBAF2E0A3E2143C207F9B82BFF4778" box="[354,435,1575,1599]" italics="true" pageId="101" pageNumber="101">brignolii</emphasis>
(
<bibRefCitation id="B83E0ECD0A3E2143C2A7F9B828054707" author="Alayon G., G." box="[450,585,1575,1600]" journalOrPublisher="Poeyana" pageId="101" pageNumber="101" pagination="1 - 3" part="229" refId="ref85600" refString="Alayon G., G. 1981. Nueva especie de Filistata Latreille, 1810 (Araneae: Filistatidae), de Mexico. Poeyana 229: 1 - 3." title="Nueva especie de Filistata Latreille, 1810 (Araneae: Filistatidae), de Mexico." type="journal article" year="1981">Alayón, 1981</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="F5E812550A3E2143C139F9B828804778" box="[604,716,1575,1599]" pageId="101" pageNumber="101" rank="species">comb. nov.</taxonomicNameLabel>
, female spermathecae, ventral. 
<emphasis id="EEDBAF2E0A3E2143C760F9B82E514778" bold="true" box="[1029,1053,1575,1599]" pageId="101" pageNumber="101">A.</emphasis>
Oaxaca, Monte Albán (AMNH IFM-1637). 
<emphasis id="EEDBAF2E0A3E2143C2E1F9DA2BD6471A" bold="true" box="[388,410,1605,1629]" pageId="101" pageNumber="101">B.</emphasis>
Puebla, Tehuacán (AMNH-1524). 
<emphasis id="EEDBAF2E0A3E2143C192F9DA2942471A" bold="true" box="[759,782,1605,1629]" pageId="101" pageNumber="101">C.</emphasis>
Puebla, Tehuacán (AMNH-1525). 
<emphasis id="EEDBAF2E0A3E2143C70EF9DA2EC8471A" bold="true" box="[1131,1156,1605,1629]" pageId="101" pageNumber="101">D.</emphasis>
Oaxaca, carretera Ixtlán-Oaxaca (CNAN-Ar 6635). 
<emphasis id="EEDBAF2E0A3E2143C12FF9FD2812473D" bold="true" box="[586,606,1634,1658]" pageId="101" pageNumber="101">E.</emphasis>
Puebla, 12 miles N Acatlán (AMNH IFM-1569). 
<emphasis id="EEDBAF2E0A3E2143C731F9FD2E2A473D" bold="true" box="[1108,1126,1634,1658]" pageId="101" pageNumber="101">F.</emphasis>
Oaxaca, 7.4 miles N of Telixtlahuaca (CAS 9057632). 
<emphasis id="EEDBAF2E0A3E2143C143F9E0287347D0" bold="true" box="[550,575,1663,1687]" pageId="101" pageNumber="101">G.</emphasis>
Puebla, 62 km west of of Orizaba (AMNH IFM-1573). 
<emphasis id="EEDBAF2E0A3E2143C707F91F2E3047DF" bold="true" box="[1122,1148,1664,1688]" pageId="101" pageNumber="101">H.</emphasis>
Sierra Mixteca(?) (MfN). Abbreviations: 
<emphasis id="EEDBAF2E0A3E2143C2DBF9022BBE47F2" bold="true" box="[446,498,1693,1717]" pageId="101" pageNumber="101">GPS,</emphasis>
glandular portion of spermathecae apex; 
<emphasis id="EEDBAF2E0A3E2143C0EBF90229EA47F2" bold="true" box="[910,934,1693,1717]" pageId="101" pageNumber="101">IF,</emphasis>
interpulmonary fold; 
<emphasis id="EEDBAF2E0A3E2143C7E1F9022EF047F2" bold="true" box="[1156,1212,1693,1717]" pageId="101" pageNumber="101">MPS,</emphasis>
membranous portion of spermathecae apex; 
<emphasis id="EEDBAF2E0A3E2143C14EF92528014795" bold="true" box="[555,589,1722,1746]" pageId="101" pageNumber="101">SB,</emphasis>
sclerotized bars; 
<emphasis id="EEDBAF2E0A3E2143C19CF925296D4795" bold="true" box="[761,801,1722,1746]" pageId="101" pageNumber="101">UE,</emphasis>
uterus externus.
</paragraph>
</caption>
<subSubSection id="94B520B70A3D2140C3C1FD542B9E4262" pageId="102" pageNumber="102" type="biology_ecology">
<paragraph id="DC10733C0A3D2140C3C1FD542B9E4262" blockId="102.[132,661,252,1751]" pageId="102" pageNumber="102">NATURAL HISTORY: Label data indicate this species has been collected under stones, on rocky slopes and in pine-oak forests.</paragraph>
</subSubSection>
<subSubSection id="94B520B70A3D2140C3C1FCB128604220" pageId="102" pageNumber="102" type="distribution">
<paragraph id="DC10733C0A3D2140C3C1FCB128604220" blockId="102.[132,661,252,1751]" pageId="102" pageNumber="102">DISTRIBUTION: Mexico, in Estado de México, Oaxaca, Puebla and Veracruz (fig. 4B).</paragraph>
</subSubSection>
<subSubSection id="94B520B70A3D2140C3C1FCF229DF47A2" pageId="102" pageNumber="102" type="materials_examined">
<paragraph id="DC10733C0A3D2140C3C1FCF229DF47A2" blockId="102.[132,661,252,1751]" lastBlockId="102.[708,1237,249,1765]" pageId="102" pageNumber="102">
ADDITIONAL MATERIAL EXAMINED: 
<emphasis id="EEDBAF2E0A3D2140C157FCF328C342C3" bold="true" box="[562,655,876,900]" pageId="102" pageNumber="102">MEXICO</emphasis>
. 
<emphasis id="EEDBAF2E0A3D2140C3E1FC152B1942E6" bold="true" box="[132,341,905,930]" pageId="102" pageNumber="102">Estado de México:</emphasis>
<emphasis id="EEDBAF2E0A3D2140C23AFC152A9C42F8" italics="true" pageId="102" pageNumber="102">Pyramids of San Juán Teotihuacán</emphasis>
[N19.69227°, W98.8435°], Bentzien, 15. ix.1977, 3♀ 3 imm. (UCB). 
<emphasis id="EEDBAF2E0A3D2140C2A5FC5B285B429B" bold="true" box="[448,535,964,988]" pageId="102" pageNumber="102">Oaxaca:</emphasis>
no further data [N17.0438°, W96.76811°], R. Greenfield, vvi.1938, 2♀ (USNM); [N17.05942°, W96.72162°, 1534m], C. and P. Vaurie, 17.vii.1955, 1♀ (AMNH); [N17.05942°, W96.72162°], 6.ii.1939, 1♀ 1 imm. (AMNH); R.V. Chamberlin, 6.ii.1939, 2♀ (AMNH); 
<emphasis id="EEDBAF2E0A3D2140C3E1FBEA2B0245CB" box="[132,334,1140,1165]" italics="true" pageId="102" pageNumber="102">10 miles E Oaxaca</emphasis>
[N17.00505°, W96.54714°], J. Reddell and J. Fish, 15.viii.1967, 3♀ 4 imm. (AMNH); 
<emphasis id="EEDBAF2E0A3D2140C267FB302BB84580" box="[258,500,1199,1223]" italics="true" pageId="102" pageNumber="102">10 miles S Tomellín</emphasis>
[N17.62498°, W96.91975°], J. Reddell, J. Fish and T. Evans, 14. viii.1967, 1♀ 2 imm. (AMNH); 
<emphasis id="EEDBAF2E0A3D2140C2ABFB7528D84445" box="[462,660,1258,1282]" italics="true" pageId="102" pageNumber="102">28 km S Nochixtlán</emphasis>
[N17.2261°, W96.87526°], S.A. Stockwell, 11. viii.1988, 1♀ 1 imm. (AMNH); 
<emphasis id="EEDBAF2E0A3D2140C2B5FABA28D9447B" box="[464,661,1316,1341]" italics="true" pageId="102" pageNumber="102">3.7 km N El Moral,</emphasis>
under stones (N17.50223°, W96.93485°, 2050m), L. Prendini, O. Francke, E. Gonzaléz and J. Ponce, 23.vii.2002, 5♀ (AMNH); 
<emphasis id="EEDBAF2E0A3D2140C2C4FAE2281144D3" box="[417,605,1404,1428]" italics="true" pageId="102" pageNumber="102">5 miles NE Mitla,</emphasis>
near “El Crucero” ruins [N16.9379°, W96.28138°, 2073– 2195m], M.R. Bogert, G. Sluder and N. Bucknall, 27.viii.1963, 1♀ 1 imm. (AMNH); 
<emphasis id="EEDBAF2E0A3D2140C28AFA4A2A8C474D" italics="true" pageId="102" pageNumber="102">5.8 miles N Teotitlán,</emphasis>
off Mex-131 [N17.03016°, W96.52073°], L.R. Erickson and M.E. Soleglad, 31.vii.1973, 2♀ (AMNH); 
<emphasis id="EEDBAF2E0A3D2140C394F9B228444702" box="[241,520,1581,1605]" italics="true" pageId="102" pageNumber="102">7.4 miles N of Telixtlahuaca</emphasis>
[N17.34993°, W96.89013°, 1800m], V.F. Lee, 16.xi.1987, 1♀ (CAS 9057632); 
<emphasis id="EEDBAF2E0A3D2140C25DF9F8284D4738" box="[312,513,1639,1663]" italics="true" pageId="102" pageNumber="102">8 km NE El Punto</emphasis>
[N17.30739°, W96.55186°, 2134m], S.A. Stockwell, 16.viii.1988, 1♀ (AMNH); 
<emphasis id="EEDBAF2E0A3D2140C240F93D28D947FD" box="[293,661,1698,1722]" italics="true" pageId="102" pageNumber="102">9 km N de San Lorenzo Mixtepec</emphasis>
(N16.17521°, W96.1983°, 1925m), O. Francke et al., 23.vi.2006, 1♀ (CNAN-Ar 8181); 
<emphasis id="EEDBAF2E0A3D2140C74FFF632E984054" box="[1066,1236,251,275]" italics="true" pageId="102" pageNumber="102">Disto. De Ejutla,</emphasis>
1.5 km W San Andres Zabache, on rocky slope [N16.59905°, W96.85799°, 1539m], C.M. Bogert, 15.viii.1970, 1♀ (AMNH); 
<emphasis id="EEDBAF2E0A3D2140C09FFECA2E2D402B" box="[1018,1121,341,365]" italics="true" pageId="102" pageNumber="102">El Catrín</emphasis>
(N17.21°, W96.57°), J. and W. Ivie, 3.ix.1964, 2♀ 3 imm. (AMNH); 
<emphasis id="EEDBAF2E0A3D2140C056FE0F292E40EF" box="[819,866,400,424]" italics="true" pageId="102" pageNumber="102">Etla,</emphasis>
2 miles N of El Moral, in pine-oak forest [N17.47762°, W96.93724°], C.M. Bogert, 10.ix.1966, 1♀ (AMNH); 
<emphasis id="EEDBAF2E0A3D2140C087FE532E0C40A3" box="[994,1088,460,484]" italics="true" pageId="102" pageNumber="102">Guelatao</emphasis>
[N17.31763°, W96.49114°], M.R. Bogert, viii.1962, 1♀ (AMNH); 
<emphasis id="EEDBAF2E0A3D2140C1A1FD972E744367" box="[708,1080,520,544]" italics="true" pageId="102" pageNumber="102">Km 171 carretera Ixtlán-Oaxaca</emphasis>
(N17.29725°, W96.54295°, 2006m), H. Montaña, 16.iii.2008, 1♀ (CNAN-Ar 6635); 
<emphasis id="EEDBAF2E0A3D2140C0CDFDDB2E73431C" box="[936,1087,579,603]" italics="true" pageId="102" pageNumber="102">Monte Albán</emphasis>
[N17.0438°, W96.76811°], 11.xii.1948, 2♀ 1 imm. (AMNH); E.S. Ross, 11.xii.1948, 1♀ (AMNH IFM-1637); 
<emphasis id="EEDBAF2E0A3D2140C7D1FDE029F043F3" italics="true" pageId="102" pageNumber="102">NE slope of Cerro San Felipe</emphasis>
[N17.09253°, W96.74474°, 2499–2743m], M.R. Bogert, 6.viii.1961, 1♀ (AMNH); 
<emphasis id="EEDBAF2E0A3D2140C055FD4729D243B7" box="[816,926,728,752]" italics="true" pageId="102" pageNumber="102">Nochixtlán</emphasis>
(N17.27°, W97.14°), J. and W. Ivie, 4.ix.1963, 2♀ (AMNH); 
<emphasis id="EEDBAF2E0A3D2140C092FD692E09424A" box="[1015,1093,758,781]" italics="true" pageId="102" pageNumber="102">Oaxaca</emphasis>
[N17.05942°, W96.72162°], 1♂ (AMNH); B. Malkin, 13–20. ix.1947, 1♀ (AMNH); 
<emphasis id="EEDBAF2E0A3D2140C0DCFCAD2E1C420E" box="[953,1104,818,841]" italics="true" pageId="102" pageNumber="102">Sierra Mixteca</emphasis>
[N17.0438°, W96.76811°], C.A. Purpus, 1♀ 3 imm. (ZMB IFM- 0826). 
<emphasis id="EEDBAF2E0A3D2140C074FCF2292842C2" bold="true" box="[785,868,877,901]" pageId="102" pageNumber="102">Puebla:</emphasis>
<emphasis id="EEDBAF2E0A3D2140C00BFCF22E7342C2" box="[878,1087,877,901]" italics="true" pageId="102" pageNumber="102">12 miles N Acatlán</emphasis>
[N18.38001°, W98.05422°], L.I. and A.M. Davis, 3.vii.1947, 1♀ 1 imm. (AMNH IFM-1569); 
<emphasis id="EEDBAF2E0A3D2140C093FC362E984286" box="[1014,1236,937,961]" italics="true" pageId="102" pageNumber="102">5 miles N Tehuacán</emphasis>
(N19.35°, W97.24°), J. and W. Ivie, 3.viii.1966, 2♀ 2 imm. (AMNH); 
<emphasis id="EEDBAF2E0A3D2140C0ECFC7B2E0142BB" box="[905,1101,996,1020]" italics="true" pageId="102" pageNumber="102">62 km W Orizaba,</emphasis>
under rocks [N18.72052°, W97.68412°], G. Dingerkus et al., 11.viii.1975, 2♀ (AMNH IFM-1573); 
<emphasis id="EEDBAF2E0A3D2140C73BFBBF297C4511" italics="true" pageId="102" pageNumber="102">7 miles NE Tehuacán,</emphasis>
ground, dry rocky slope [N19.35°, W97.24°], W. Peck, 7.iv.1967, 1♀ (CAS 9060623); 
<emphasis id="EEDBAF2E0A3D2140C1A1FBE6290745D6" box="[708,843,1145,1169]" italics="true" pageId="102" pageNumber="102">Cacaloapán,</emphasis>
0.5 mile west on highway 50 [N18.57893°, W97.58413°], R.M. Schick and D. Schroeder, 25.vii.1965, 1♀ (AMNH); 
<emphasis id="EEDBAF2E0A3D2140C715FB2A2E98458A" box="[1136,1236,1205,1229]" italics="true" pageId="102" pageNumber="102">Tehuacán</emphasis>
[N18.46586°, W97.40005°], D.M. Bogert and H.E. Vokes, 8.xi.1939, missing spermathecae, 1♀ 1 imm. (AMNH IFM-1525); H. Wagner, 17–24.x.1944, 10♀ 1 imm. (AMNH IFM-1406); [N18.46651°, W97.40038°], W.J. Gertsch and V. Roth, 24.vii.1956, 1♂ 1♀ 2 imm. (AMNH); 
<emphasis id="EEDBAF2E0A3D2140C0BFFAF82E094438" box="[986,1093,1383,1407]" italics="true" pageId="102" pageNumber="102">Tlacotepec</emphasis>
[N18.68163°, W97.64832°], V. Roth and W. Gertsch, 24.vii.1956, 7♀ 8 imm. (AMNH); 
<emphasis id="EEDBAF2E0A3D2140C0CBFA3C2E9844FC" box="[942,1236,1443,1467]" italics="true" pageId="102" pageNumber="102">toll road 21 km N Tehuacán,</emphasis>
under rocks (N18.64475°, W97.45636°, 2249m), W.G. Villegas, O. Francke and R. Paredes, 31. xi.2004, 2♀ (AMNH). 
<emphasis id="EEDBAF2E0A3D2140C0ACFA622E7F4752" bold="true" box="[969,1075,1533,1557]" pageId="102" pageNumber="102">Veracruz:</emphasis>
<emphasis id="EEDBAF2E0A3D2140C725FA6229534775" italics="true" pageId="102" pageNumber="102">15.5 miles W Orizaba,</emphasis>
off Mex-150 [N18.71566°, W97.30777°], L.R. Erickson and M.E. Soleglad, 30.vii.1973, 1♀ (AMNH); 
<emphasis id="EEDBAF2E0A3D2140C05DF9C92E7E4729" box="[824,1074,1622,1646]" italics="true" pageId="102" pageNumber="102">2 miles NE Acultzingo,</emphasis>
J.A. Beatty, 4. vii.1963, 1♀ 5 imm. (AMNH); 
<emphasis id="EEDBAF2E0A3D2140C717F9EB2E9947CC" box="[1138,1237,1652,1675]" italics="true" pageId="102" pageNumber="102">Veracruz</emphasis>
[N19.17377°, W96.13422°], O’Rourke, iii.1968, 2♀ (AMNH). 
<emphasis id="EEDBAF2E0A3D2140C051F9302E4E4780" bold="true" box="[820,1026,1711,1735]" pageId="102" pageNumber="102">Uncertain locality:</emphasis>
label illegible, C.A. Purpus, 1♀ (ZMB).
</paragraph>
</subSubSection>
</treatment>
</document>