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<document id="7E444D222737257705C528E85DA6BCC1" ID-CLB-Dataset="28554" ID-DOI="10.5852/ejt.2019.494" ID-GBIF-Dataset="5600128b-7356-4530-bf52-32f74f7fa9af" ID-Zenodo-Dep="2561924" ID-ZooBank="60C94F41-D9CB-471A-BAB7-2D558F2D3357" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1550010391328" checkinUser="plazi" docAuthor="Peñafiel-Vinueza, Ana Danitza &amp; Rafael, Violeta" docDate="2019" docId="53544669FF99FFAEEE4DFB67FDB3D59E" docLanguage="en" docName="ejt-494_penafil-vinueza_rafael.pdf.imf" docOrigin="European Journal of Taxonomy 494" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Drosophila ayauma Peñafiel-Vinueza &amp; Rafael 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="14" masterDocId="AF6D3E11FF95FFA3EC3FFF90FFF9D241" masterDocTitle="Five new species of the Drosophila tripunctata group (Diptera: Drosophilidae) from Podocarpus National Park, Ecuador" masterLastPageNumber="18" masterPageNumber="1" originalUpdateDomain="Boston" originalUpdateTime="1549887238809" originalUpdateUser="plazi" pageNumber="13" updateTime="1698698774925" updateUser="ExternalLinkService">
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<mods:title id="7706C3FE4121C682D7832EB7C3BD3946">Five new species of the Drosophila tripunctata group (Diptera: Drosophilidae) from Podocarpus National Park, Ecuador</mods:title>
</mods:titleInfo>
<mods:name id="A1CCBF7C9D9AB3D781A14BFACA515D5A" type="personal">
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<mods:namePart id="BEEFE2DD049B0B7F4B65541FC2BB8F77">Peñafiel-Vinueza, Ana Danitza</mods:namePart>
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<mods:namePart id="A2CA5278ED780F82C227C7705F88C850">Rafael, Violeta</mods:namePart>
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<mods:typeOfResource id="E71111161D17374B46AEAF27717DE3BA">text</mods:typeOfResource>
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<mods:title id="488896A9AF7AF98E423E4F5D77E71329">European Journal of Taxonomy</mods:title>
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<mods:date id="ED85CD21B9FB5FE414A87ABEB7302935">2019</mods:date>
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<mods:detail id="E3DE7B7C37069DA3282DDAA0D10442B4" type="volume">
<mods:number id="DD7DDD9C415F30507D06AB02264D6163">494</mods:number>
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<treatment id="53544669FF99FFAEEE4DFB67FDB3D59E" ID-DOI="http://doi.org/10.5281/zenodo.5940650" ID-GBIF-Taxon="152541868" ID-Zenodo-Dep="5940650" LSID="urn:lsid:plazi:treatment:53544669FF99FFAEEE4DFB67FDB3D59E" httpUri="http://treatment.plazi.org/id/53544669FF99FFAEEE4DFB67FDB3D59E" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<subSubSection id="93E7A4F4FF99FFAFEE4DFB67FB3BD774" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="DB42F77FFF99FFAFEE4DFB67FC3BD753" blockId="12.[370,1218,1271,1368]" box="[626,962,1271,1298]" pageId="12" pageNumber="13">
<heading id="800A4013FF99FFAFEE4DFB67FC3BD753" box="[626,962,1271,1298]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="13" reason="2">
<taxonomicName id="1CFD8CFCFF99FFAFEE4DFB67FC99D753" ID-CoL="9HNB3" authority="Peñafiel-Vinueza &amp; Rafael, 2019" authorityName="Peñafiel-Vinueza &amp; Rafael" authorityYear="2019" box="[626,864,1271,1298]" class="Insecta" family="Drosophilidae" genus="Drosophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ayauma" status="sp. nov.">
<emphasis id="E9892B6DFF99FFAFEE4DFB67FC99D753" bold="true" box="[626,864,1271,1298]" italics="true" pageId="12" pageNumber="13">Drosophila ayauma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2BA9616FF99FFAFEF57FB68FC3BD753" box="[872,962,1272,1298]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="DB42F77FFF99FFAFED4DFA8AFB3BD774" blockId="12.[370,1218,1271,1368]" box="[370,1218,1306,1333]" pageId="12" pageNumber="13">
<uri id="AF6CFB7DFF99FFAFED4DFA8AFB3BD774" box="[370,1218,1306,1333]" pageId="12" pageNumber="13">
urn:lsid:zoobank.org:act:
<uuid id="AF5BCDAAFF99FFAFEEACFA8AFB3BD774" box="[659,1218,1306,1333]" pageId="12" pageNumber="13">5CEBC281-6F5E-41B8-B837-03ABF2237494</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="93E7A4F4FF99FFAFEEC8FAADFCC5D719" box="[759,828,1341,1368]" pageId="12" pageNumber="13" type="description">
<paragraph id="DB42F77FFF99FFAFEEC8FAADFCC5D719" blockId="12.[370,1218,1271,1368]" box="[759,828,1341,1368]" pageId="12" pageNumber="13">
<figureCitation id="43C6EBFAFF99FFAFEEC8FAADFCC5D719" box="[759,828,1341,1368]" captionStart="Fig. 7" captionStartId="14.[189,232,1930,1956]" captionTargetBox="[386,1202,646,1899]" captionTargetId="figure@14.[386,1202,646,1899]" captionTargetPageId="14" captionText="Fig. 7. Drosophila ayauma sp. nov. Holotype, ♂ (QCAZ-I 3336). A. Abdomen. B. Epandrium, cerci, surstylus, decasternum. C. Hypandrium, gonopods and paraphyses in ventral view. DF. Aedeagus in ventral, lateral and dorsal view, respectively." httpUri="https://zenodo.org/record/2561938/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="93E7A4F4FF99FFAFEC82FAEFFDA2D436" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="DB42F77FFF99FFAFEC82FAEFFECFD7D8" blockId="12.[189,310,1407,1433]" box="[189,310,1407,1433]" pageId="12" pageNumber="13">
<heading id="800A4013FF99FFAFEC82FAEFFECFD7D8" bold="true" box="[189,310,1407,1433]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="E9892B6DFF99FFAFEC82FAEFFECFD7D8" bold="true" box="[189,310,1407,1433]" pageId="12" pageNumber="13">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DB42F77FFF99FFAFEC82FA3DFDA2D436" blockId="12.[189,1399,1453,1656]" pageId="12" pageNumber="13">
Body color yellowish brown. Aristae with four dorsal and two ventral branches. Two prominent oral bristles plus terminal fork. Thorax yellowish brown. Abdomen yellow with dorsal midline, 2
<superScript id="2C885A37FF99FFAFE921FA5FFAD6D79C" attach="left" box="[1310,1327,1487,1501]" fontSize="6" pageId="12" pageNumber="13">nd</superScript>
to 5
<superScript id="2C885A37FF99FFAFE956FA5FFA8FD79C" attach="none" box="[1385,1398,1487,1501]" fontSize="6" pageId="12" pageNumber="13">th</superScript>
tergites with triangular pigmentation that thins laterally. Cerci not fused to epandrium. Aedeagus wide, with two dorsal sclerotized projections ending in a point, ventrally with two sclerotized rounded sheets. Hypandrium shield-shaped. Gonopod bearing one long bristle. Paraphyses microtrichose, fused to gonopod, bearing one small bristle.
</paragraph>
</subSubSection>
<subSubSection id="93E7A4F4FF99FFAFEC82F90FFC77D54A" pageId="12" pageNumber="13" type="etymology">
<paragraph id="DB42F77FFF99FFAFEC82F90FFEBDD4F8" blockId="12.[189,324,1695,1721]" box="[189,324,1695,1721]" pageId="12" pageNumber="13">
<heading id="800A4013FF99FFAFEC82F90FFEBDD4F8" bold="true" box="[189,324,1695,1721]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="E9892B6DFF99FFAFEC82F90FFEBDD4F8" bold="true" box="[189,324,1695,1721]" pageId="12" pageNumber="13">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="DB42F77FFF99FFAFEC82F95DFC77D54A" blockId="12.[189,1399,1741,1803]" pageId="12" pageNumber="13">
The name
<taxonomicName id="1CFD8CFCFF99FFAFED7AF95EFE5BD4A6" authorityName="Peñafiel-Vinueza &amp; Rafael" authorityYear="2019" box="[325,418,1742,1767]" class="Insecta" family="Drosophilidae" genus="Drosophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ayauma">
<emphasis id="E9892B6DFF99FFAFED7AF95EFE5BD4A6" box="[325,418,1742,1767]" italics="true" pageId="12" pageNumber="13">ayauma</emphasis>
</taxonomicName>
comes from Kichwa
<emphasis id="E9892B6DFF99FFAFEE89F95EFD19D4A6" box="[694,736,1742,1767]" italics="true" pageId="12" pageNumber="13">aya</emphasis>
= spirit and
<emphasis id="E9892B6DFF99FFAFEFA2F95EFC29D4A6" box="[925,976,1742,1767]" italics="true" pageId="12" pageNumber="13">uma</emphasis>
= head, meaning spirit from the head. The apex of the aedeagus resembles two demon horns.
</paragraph>
</subSubSection>
<subSubSection id="93E7A4F4FF99FFAEEC82F8A2FADCD3CE" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="DB42F77FFF99FFAFEC82F8A2FE53D50D" blockId="12.[189,426,1842,1868]" box="[189,426,1842,1868]" pageId="12" pageNumber="13">
<heading id="800A4013FF99FFAFEC82F8A2FE53D50D" bold="true" box="[189,426,1842,1868]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="E9892B6DFF99FFAFEC82F8A2FE53D50D" bold="true" box="[189,426,1842,1868]" pageId="12" pageNumber="13">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DB42F77FFF99FFAFECD5F8F1FEA5D53A" blockId="12.[189,1399,1889,1986]" box="[234,348,1889,1915]" pageId="12" pageNumber="13">
<typeStatus id="044649DDFF99FFAFECD5F8F1FEA5D53A" box="[234,348,1889,1915]" pageId="12" pageNumber="13" type="holotype">
<emphasis id="E9892B6DFF99FFAFECD5F8F1FEA5D53A" bold="true" box="[234,348,1889,1915]" pageId="12" pageNumber="13">Holotype</emphasis>
</typeStatus>
</paragraph>
<paragraph id="DB42F77FFF99FFAFEC82F814FB06D580" blockId="12.[189,1399,1889,1986]" pageId="12" pageNumber="13">
<materialsCitation id="6B95FD22FF99FFAFEC82F814FB06D580" ID-GBIF-Occurrence="1994756072" collectingDate="2015-11-19" collectorName="A. Penafiel" country="Ecuador" determinerName="V. Rafael" elevation="2800" latitude="-4.116361" location="Cajanuma" longLatPrecision="1" longitude="-79.16997" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Loja" typeStatus="holotype">
<collectingCountry id="A3EAB7EFFF99FFAFEC82F814FEB2D5DF" box="[189,331,1924,1950]" name="Ecuador" pageId="12" pageNumber="13">ECUADOR</collectingCountry>
• ♂ (dissected, terminalia in microvial);
<collectingRegion id="1939399DFF99FFAFEF1EF814FCAED5DF" box="[801,855,1924,1950]" country="Ecuador" name="Loja" pageId="12" pageNumber="13">Loja</collectingRegion>
Prov.,
<location id="DE22A1A4FF99FFAFEF98F814FBD8D5DF" LSID="urn:lsid:plazi:treatment:53544669FF99FFAEEE4DFB67FDB3D59E:DE22A1A4FF99FFAFEF98F814FBD8D5DF" box="[935,1057,1924,1950]" country="Ecuador" latitude="-4.116361" longLatPrecision="1" longitude="-79.16997" name="Cajanuma" pageId="12" pageNumber="13" stateProvince="Loja">Cajanuma</location>
,
<elevation id="50D0104CFF99FFAFE814F814FB38D5DF" box="[1067,1217,1924,1950]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="12" pageNumber="13" unit="m" value="2800.0">
<quantity id="1C055A9AFF99FFAFE814F814FB7BD5DF" box="[1067,1154,1924,1950]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="12" pageNumber="13" unit="m" value="2800.0">2800 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="BEC991B8FF99FFAFE8F2F812FA88D5DF" box="[1229,1393,1922,1950]" direction="south" orientation="latitude" pageId="12" pageNumber="13" precision="1" value="-4.116361">04°0658.9″ S</geoCoordinate>
,
<geoCoordinate id="BEC991B8FF99FFAFEC82F835FE94D583" box="[189,365,1957,1986]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="1" value="-79.16997">79°1011.9″ W</geoCoordinate>
;
<collectingDate id="BF072857FF99FFAFED46F837FDE2D583" box="[377,539,1959,1986]" pageId="12" pageNumber="13" value="2015-11-19">
<date id="AF43D1BFFF99FFAFED46F837FDE2D583" box="[377,539,1959,1986]" pageId="12" pageNumber="13" value="2015-11-19">19 Nov. 2015</date>
</collectingDate>
;
<collectorName id="760892A9FF99FFAFEE19F837FD51D580" box="[550,680,1959,1985]" pageId="12" pageNumber="13">A. Peñafiel</collectorName>
leg.;
<collectorName id="760892A9FF99FFAFEED7F837FC92D580" box="[744,875,1959,1985]" pageId="12" pageNumber="13">A. Peñafiel</collectorName>
and
<determinerName id="2E9E1BBBFF99FFAFEF9CF838FBE9D580" box="[931,1040,1959,1986]" pageId="12" pageNumber="13">V. Rafael</determinerName>
det.; QCAZ-I 3336.
</materialsCitation>
</paragraph>
<paragraph id="DB42F77FFF98FFAEECD5FE9BFE9ED364" blockId="13.[189,1399,267,399]" box="[234,359,267,293]" pageId="13" pageNumber="14">
<typeStatus id="044649DDFF98FFAEECD5FE9BFE9ED364" box="[234,359,267,293]" pageId="13" pageNumber="14" type="paratype">
<emphasis id="E9892B6DFF98FFAEECD5FE9BFE9ED364" bold="true" box="[234,359,267,293]" pageId="13" pageNumber="14">Paratypes</emphasis>
</typeStatus>
</paragraph>
<paragraph id="DB42F77FFF98FFAEEC82FEBEFADCD3CE" blockId="13.[189,1399,267,399]" pageId="13" pageNumber="14">
<materialsCitation id="6B95FD22FF98FFAEEC82FEBEFF01D32A" ID-GBIF-Occurrence="1993784567" collectingDate="2015-11-19" collectorName="A. Penafiel" country="Ecuador" determinerName="V. Rafael" elevation="2800" latitude="-4.116361" location="Cajanuma" longLatPrecision="1" longitude="-79.16997" pageId="13" pageNumber="14" specimenCount="1" stateProvince="Loja" typeStatus="paratype">
<collectingCountry id="A3EAB7EFFF98FFAEEC82FEBEFEB2D309" box="[189,331,302,328]" name="Ecuador" pageId="13" pageNumber="14">ECUADOR</collectingCountry>
• 6 ♂♂ (dissected, terminalia in microvial); same data as for holotype; QCAZ-I 3337 to 3342
</materialsCitation>
<materialsCitation id="6B95FD22FF98FFAEED2EFEC1FAD8D3CE" ID-GBIF-Occurrence="1993784565" collectingDate="2015-09-19" collectorName="A. B. Manzano &amp; A. Penafiel" country="Ecuador" determinerName="V. Rafael" elevation="2548" latitude="-0.5455556" location="Rio Guango" longLatPrecision="1" longitude="-77.95372" pageId="13" pageNumber="14" specimenCount="1" stateProvince="Napo" typeStatus="paratype">
1 ♂ (dissected, terminalia in microvial);
<collectingRegion id="1939399DFF98FFAEEECDFEC2FCCBD32D" box="[754,818,338,364]" country="Ecuador" name="Napo" pageId="13" pageNumber="14">Napo</collectingRegion>
Prov.,
<location id="DE22A1A4FF98FFAEEFBAFEC1FBEED32A" LSID="urn:lsid:plazi:treatment:53544669FF99FFAEEE4DFB67FDB3D59E:DE22A1A4FF98FFAEEFBAFEC1FBEED32A" box="[901,1047,337,363]" country="Ecuador" latitude="-0.5455556" longLatPrecision="1" longitude="-77.95372" name="Rio Guango" pageId="13" pageNumber="14" stateProvince="Napo">Río Guango</location>
,
<elevation id="50D0104CFF98FFAEE81CFEC1FB44D32A" box="[1059,1213,337,364]" metricMagnitude="3" metricUnit="m" metricValue="2.548" pageId="13" pageNumber="14" unit="m" value="2548.0">
<quantity id="1C055A9AFF98FFAEE81CFEC1FB85D32D" box="[1059,1148,337,364]" metricMagnitude="3" metricUnit="m" metricValue="2.548" pageId="13" pageNumber="14" unit="m" value="2548.0">2548 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="BEC991B8FF98FFAEE8F4FEDFFA88D32A" box="[1227,1393,335,363]" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="1" value="-0.5455556">00°3244.0″ S</geoCoordinate>
,
<geoCoordinate id="BEC991B8FF98FFAEEC82FEE3FE97D3CE" box="[189,366,371,399]" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="1" value="-77.95372">77°5713.4″ W</geoCoordinate>
;
<collectingDate id="BF072857FF98FFAEED45FEE5FDE1D3CE" box="[378,536,372,399]" pageId="13" pageNumber="14" value="2015-09-19">
<date id="AF43D1BFFF98FFAEED45FEE5FDE1D3CE" box="[378,536,372,399]" pageId="13" pageNumber="14" value="2015-09-19">19 Sep. 2015</date>
</collectingDate>
;
<collectorName id="760892A9FF98FFAEEE1DFEE5FD37D3CE" box="[546,718,373,399]" pageId="13" pageNumber="14">A.B. Manzano</collectorName>
leg.;
<collectorName id="760892A9FF98FFAEEF31FEE5FC68D3CF" box="[782,913,372,399]" pageId="13" pageNumber="14">A. Peñafiel</collectorName>
and
<determinerName id="2E9E1BBBFF98FFAEEFF6FEE5FBCFD3CF" box="[969,1078,372,399]" pageId="13" pageNumber="14">V. Rafael</determinerName>
det.; QCAZ-I 3343
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="93E7A4F4FF98FFAEEC82FE2DFA8ED487" pageId="13" pageNumber="14" type="description">
<paragraph id="DB42F77FFF98FFAEEC82FE2DFEB7D396" blockId="13.[189,334,445,471]" box="[189,334,445,471]" pageId="13" pageNumber="14">
<heading id="800A4013FF98FFAEEC82FE2DFEB7D396" bold="true" box="[189,334,445,471]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="E9892B6DFF98FFAEEC82FE2DFEB7D396" bold="true" box="[189,334,445,471]" pageId="13" pageNumber="14">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DB42F77FFF98FFAEECD5FE7CFE70D00D" blockId="13.[189,1398,492,588]" pageId="13" pageNumber="14">
<emphasis id="E9892B6DFF98FFAEECD5FE7CFED3D047" bold="true" box="[234,298,492,518]" pageId="13" pageNumber="14">Male</emphasis>
<typeStatus id="044649DDFF98FFAEEC82FD9FFED3D068" box="[189,298,527,553]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
external morphology: total length (body + wings)
<quantity id="1C055A9AFF98FFAEEF47FD9FFC1AD06B" box="[888,995,527,554]" metricMagnitude="-3" metricUnit="m" metricValue="7.43" pageId="13" pageNumber="14" unit="mm" value="7.43">7.43 mm</quantity>
, body length
<quantity id="1C055A9AFF98FFAEE841FD9FFB10D06B" box="[1150,1257,527,554]" metricMagnitude="-3" metricUnit="m" metricValue="4.22" pageId="13" pageNumber="14" unit="mm" value="4.22">4.22 mm</quantity>
. Body color yellowish brown.
</paragraph>
<paragraph id="DB42F77FFF98FFAEEC82FDECFBF9D104" blockId="13.[189,1399,635,838]" pageId="13" pageNumber="14">
HEAD. Aristae with 4 dorsal and 2 ventral branches plus terminal fork and fine hairs. Orbital plate yellowish brown, frontal length
<quantity id="1C055A9AFF98FFAEEE0BFD0FFD66D0F8" box="[564,671,671,697]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="13" pageNumber="14" unit="mm" value="0.37">0.37 mm</quantity>
; frontal index = 1.02; top to bottom width ratio = 1.51. Medial vertical seta closer to lateral vertical seta and slightly towards outer edge of orbital plate; vt index = 0.86; or1or3 ratio = 0.74; or2or1 ratio = 0.43. Ocellar triangle brown, ocellus yellow, frontal vitta yellowish brown. Cheek index = 5.9. Two prominent oral bristles of same size, vibrissa index = 0.78. Carina yellowish brown, not sulcate. Eyes bright red, eye index = 1.66.
</paragraph>
<paragraph id="DB42F77FFF98FFAEEC82FCE5FC43D194" blockId="13.[189,1399,884,981]" pageId="13" pageNumber="14">THORAX. Yellowish brown, acrostichal hairs in 8 rows between two anterior dorsocentral setae that decrease in number towards rear. h index = 1.27; dc index = 0.83. Scut index = 1.09. Medial katepisternal seta same size as previous seta, sterno index = 1.47. Legs yellow.</paragraph>
<paragraph id="DB42F77FFF98FFAEEC82FB94FBF5D600" blockId="13.[189,1399,1028,1090]" pageId="13" pageNumber="14">
WINGS. Brown. Alar length
<quantity id="1C055A9AFF98FFAEEDC1FB94FD90D65F" box="[510,617,1028,1055]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" pageId="13" pageNumber="14" unit="mm" value="3.55">3.55 mm</quantity>
, alar width
<quantity id="1C055A9AFF98FFAEEECEFB94FCA5D65F" box="[753,860,1028,1055]" metricMagnitude="-3" metricUnit="m" metricValue="1.51" pageId="13" pageNumber="14" unit="mm" value="1.51">1.51 mm</quantity>
. Alar indices: alar = 2.26; C = 4.98; ac = 1.78; hb = 0.20; 4c = 0.43;
<date id="AF43D1BFFF98FFAEED85FBB7FE21D600" box="[442,472,1063,1089]" pageId="13" pageNumber="14">4v</date>
= 1.20;
<date id="AF43D1BFFF98FFAEEE06FBB7FDAED603" box="[569,599,1063,1090]" pageId="13" pageNumber="14">5x</date>
= 1.15; M = 0.32 and prox. x = 0.34.
</paragraph>
<paragraph id="DB42F77FFF98FFAEEC82FBE0FDF4D6EC" blockId="13.[189,1399,1134,1197]" pageId="13" pageNumber="14">
ABDOMEN. Yellow with dorsal midline, 1
<superScript id="2C885A37FF98FFAEEEB3FBFFFD6ED63C" attach="left" box="[652,663,1135,1149]" fontSize="6" pageId="13" pageNumber="14">st</superScript>
tergite brown and 2
<superScript id="2C885A37FF98FFAEEF44FBFEFC75D63D" attach="left" box="[891,908,1134,1148]" fontSize="6" pageId="13" pageNumber="14">nd</superScript>
to 5
<superScript id="2C885A37FF98FFAEEF82FBFEFC33D63D" attach="left" box="[957,970,1134,1148]" fontSize="6" pageId="13" pageNumber="14">th</superScript>
tergites with triangular pigmentation that thins laterally (
<figureCitation id="43C6EBFAFF98FFAEED9EFB03FDF9D6EC" box="[417,512,1171,1197]" captionStart="Fig. 7" captionStartId="14.[189,232,1930,1956]" captionTargetBox="[386,1202,646,1899]" captionTargetId="figure@14.[386,1202,646,1899]" captionTargetPageId="14" captionText="Fig. 7. Drosophila ayauma sp. nov. Holotype, ♂ (QCAZ-I 3336). A. Abdomen. B. Epandrium, cerci, surstylus, decasternum. C. Hypandrium, gonopods and paraphyses in ventral view. DF. Aedeagus in ventral, lateral and dorsal view, respectively." httpUri="https://zenodo.org/record/2561938/files/figure.png" pageId="13" pageNumber="14">Fig. 7A</figureCitation>
).
</paragraph>
<paragraph id="DB42F77FFF98FFAEEC82FB4DFA8FD721" blockId="13.[189,1399,1244,1377]" pageId="13" pageNumber="14">
MALE TERMINALIA. Epandrium dorsally microtrichose, with no upper or lower bristles, ventral lobe fused to surstylus by a membrane bearing one long bristle. Cerci not fused to epandrium, ventral side not microtrichose and with several long bristles towards middle. Surstylus with 9 primary teeth on each side, 15 marginal bristles on right and 14 on left (
<figureCitation id="43C6EBFAFF98FFAEEEFEFAD6FCE7D721" box="[705,798,1350,1376]" captionStart="Fig. 7" captionStartId="14.[189,232,1930,1956]" captionTargetBox="[386,1202,646,1899]" captionTargetId="figure@14.[386,1202,646,1899]" captionTargetPageId="14" captionText="Fig. 7. Drosophila ayauma sp. nov. Holotype, ♂ (QCAZ-I 3336). A. Abdomen. B. Epandrium, cerci, surstylus, decasternum. C. Hypandrium, gonopods and paraphyses in ventral view. DF. Aedeagus in ventral, lateral and dorsal view, respectively." httpUri="https://zenodo.org/record/2561938/files/figure.png" pageId="13" pageNumber="14">Fig. 7B</figureCitation>
). Hypandrium sclerotized, shield-shaped (
<figureCitation id="43C6EBFAFF98FFAEE933FAD6FA90D721" box="[1292,1385,1350,1376]" captionStart="Fig. 7" captionStartId="14.[189,232,1930,1956]" captionTargetBox="[386,1202,646,1899]" captionTargetId="figure@14.[386,1202,646,1899]" captionTargetPageId="14" captionText="Fig. 7. Drosophila ayauma sp. nov. Holotype, ♂ (QCAZ-I 3336). A. Abdomen. B. Epandrium, cerci, surstylus, decasternum. C. Hypandrium, gonopods and paraphyses in ventral view. DF. Aedeagus in ventral, lateral and dorsal view, respectively." httpUri="https://zenodo.org/record/2561938/files/figure.png" pageId="13" pageNumber="14">Fig. 7C</figureCitation>
).
</paragraph>
<paragraph id="DB42F77FFF98FFAEEC82FA1FFAF0D453" blockId="13.[189,1399,1423,1555]" pageId="13" pageNumber="14">
AEDEAGUS. Wide, with two dorsal sclerotized projections ending in a point, ventrally with two sclerotized rounded sheets, middle part of aedeagus membranous, covered in bright studs. Ventral rod primitive. Aedeagus apodeme slightly sclerotized and straight. Gonopod microtrichose, bearing one long bristle (
<figureCitation id="43C6EBFAFF98FFAEECF9FA68FEBDD452" box="[198,324,1528,1555]" captionStart="Fig. 7" captionStartId="14.[189,232,1930,1956]" captionTargetBox="[386,1202,646,1899]" captionTargetId="figure@14.[386,1202,646,1899]" captionTargetPageId="14" captionText="Fig. 7. Drosophila ayauma sp. nov. Holotype, ♂ (QCAZ-I 3336). A. Abdomen. B. Epandrium, cerci, surstylus, decasternum. C. Hypandrium, gonopods and paraphyses in ventral view. DF. Aedeagus in ventral, lateral and dorsal view, respectively." httpUri="https://zenodo.org/record/2561938/files/figure.png" pageId="13" pageNumber="14">Fig. 7DF</figureCitation>
). Paraphyses microtrichose, fused to gonopod, bearing one small bristle (
<figureCitation id="43C6EBFAFF98FFAEE8A0FA68FB02D453" box="[1183,1275,1528,1554]" captionStart="Fig. 7" captionStartId="14.[189,232,1930,1956]" captionTargetBox="[386,1202,646,1899]" captionTargetId="figure@14.[386,1202,646,1899]" captionTargetPageId="14" captionText="Fig. 7. Drosophila ayauma sp. nov. Holotype, ♂ (QCAZ-I 3336). A. Abdomen. B. Epandrium, cerci, surstylus, decasternum. C. Hypandrium, gonopods and paraphyses in ventral view. DF. Aedeagus in ventral, lateral and dorsal view, respectively." httpUri="https://zenodo.org/record/2561938/files/figure.png" pageId="13" pageNumber="14">Fig. 7C</figureCitation>
).
</paragraph>
<paragraph id="DB42F77FFF98FFAEECD5F9D1FCCCD41D" blockId="13.[189,1399,1601,1734]" box="[234,821,1601,1628]" pageId="13" pageNumber="14">
<emphasis id="E9892B6DFF98FFAEECD5F9D1FDFAD41D" bold="true" box="[234,515,1601,1628]" pageId="13" pageNumber="14">
Variation in
<typeStatus id="044649DDFF98FFAEEDB8F9D2FDFAD41D" box="[391,515,1602,1628]" pageId="13" pageNumber="14" type="paratype">paratypes</typeStatus>
</emphasis>
(dry mounted specimens)
</paragraph>
<paragraph id="DB42F77FFF98FFAEEC82F9F4FA8ED487" blockId="13.[189,1399,1601,1734]" pageId="13" pageNumber="14">
Abdomen: other specimens show triangular pigmentation that thins laterally and then reaches and covers exterior margin of each tergite. Head: frontal length
<quantity id="1C055A9AFF98FFAEEF0DF918FC1BD4E3" box="[818,994,1672,1698]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" metricValueMax="4.2" metricValueMin="2.2" pageId="13" pageNumber="14" unit="mm" value="0.32" valueMax="0.42" valueMin="0.22">0.220.42 mm</quantity>
, frontal index = 0.530.84, top to bottom width ratio = 1.21.66; cheek index = 57.57; vibrissa index = 0.450.78; eye index = 1.151.64.
</paragraph>
</subSubSection>
<subSubSection id="93E7A4F4FF98FFAEEC82F964FDB3D59E" pageId="13" pageNumber="14" type="distribution">
<paragraph id="DB42F77FFF98FFAEEC82F964FEAFD54F" blockId="13.[189,342,1780,1806]" box="[189,342,1780,1806]" pageId="13" pageNumber="14">
<heading id="800A4013FF98FFAEEC82F964FEAFD54F" bold="true" box="[189,342,1780,1806]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="E9892B6DFF98FFAEEC82F964FEAFD54F" bold="true" box="[189,342,1780,1806]" pageId="13" pageNumber="14">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DB42F77FFF98FFAEEC82F8B3FB25D57D" blockId="13.[189,1244,1826,1853]" box="[189,1244,1826,1853]" pageId="13" pageNumber="14">
Known from the
<typeStatus id="044649DDFF98FFAEEDB9F8B3FE40D57C" box="[390,441,1827,1853]" pageId="13" pageNumber="14">type</typeStatus>
locality and from the montane forest of Río Guango at
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a.s.l.
</paragraph>
<paragraph id="DB42F77FFF98FFAEEC82F8E3FDDBD5CC" blockId="13.[189,546,1907,1933]" box="[189,546,1907,1933]" pageId="13" pageNumber="14">
<heading id="800A4013FF98FFAEEC82F8E3FDDBD5CC" bold="true" box="[189,546,1907,1933]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="E9892B6DFF98FFAEEC82F8E3FDDBD5CC" bold="true" box="[189,546,1907,1933]" pageId="13" pageNumber="14">Relationship to other species</emphasis>
</heading>
</paragraph>
<paragraph id="DB42F77FFF98FFAEEC82F831FDB3D59E" blockId="13.[189,1398,1953,2015]" pageId="13" pageNumber="14">
The general shape of the male terminalia does not suggest any relationship to other described species of the
<taxonomicName id="1CFD8CFCFF98FFAEECD7F855FE0FD59E" authorityName="Loew" authorityYear="1862" box="[232,502,1989,2015]" class="Insecta" family="Drosophilidae" genus="Drosophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tripunctata">
<emphasis id="E9892B6DFF98FFAEECD7F855FE0FD59E" box="[232,502,1989,2015]" italics="true" pageId="13" pageNumber="14">Drosophila tripunctata</emphasis>
</taxonomicName>
group.
</paragraph>
</subSubSection>
</treatment>
</document>