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<document id="0BFC18E4DDD330A95E59F6AF94B48E97" ID-CLB-Dataset="28554" ID-DOI="10.5852/ejt.2019.494" ID-GBIF-Dataset="5600128b-7356-4530-bf52-32f74f7fa9af" ID-Zenodo-Dep="2561924" ID-ZooBank="60C94F41-D9CB-471A-BAB7-2D558F2D3357" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1550010391328" checkinUser="plazi" docAuthor="Peñafiel-Vinueza, Ana Danitza &amp; Rafael, Violeta" docDate="2019" docId="53544669FF97FFA5EE44FD2DFD88D17C" docLanguage="en" docName="ejt-494_penafil-vinueza_rafael.pdf.imf" docOrigin="European Journal of Taxonomy 494" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Drosophila warmi Peñafiel-Vinueza &amp; Rafael 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="7" masterDocId="AF6D3E11FF95FFA3EC3FFF90FFF9D241" masterDocTitle="Five new species of the Drosophila tripunctata group (Diptera: Drosophilidae) from Podocarpus National Park, Ecuador" masterLastPageNumber="18" masterPageNumber="1" originalUpdateDomain="Boston" originalUpdateTime="1549887238809" originalUpdateUser="plazi" pageNumber="3" updateTime="1698698774925" updateUser="ExternalLinkService">
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<mods:title id="D4FD985946308591B9B9168BBE4FB199">Five new species of the Drosophila tripunctata group (Diptera: Drosophilidae) from Podocarpus National Park, Ecuador</mods:title>
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<mods:name id="93A82B71102B85332D0A035EB0217432" type="personal">
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<mods:namePart id="E45B1AE88980CEA530964C0E23BFDD91">Peñafiel-Vinueza, Ana Danitza</mods:namePart>
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<mods:namePart id="738D0D787BBD8C9A942380BB722B1994">Rafael, Violeta</mods:namePart>
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<mods:title id="DDA6FCE08E977248BAD140963204ECE0">European Journal of Taxonomy</mods:title>
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<mods:date id="40C81E070C0117DDD42805C42CB8D5B3">2019</mods:date>
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<mods:number id="42BBE8F6880C0DD1C2E5A3D252D38933">2019-02-01</mods:number>
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<treatment id="53544669FF97FFA5EE44FD2DFD88D17C" ID-DOI="http://doi.org/10.5281/zenodo.5940640" ID-GBIF-Taxon="152541876" ID-Zenodo-Dep="5940640" LSID="urn:lsid:plazi:treatment:53544669FF97FFA5EE44FD2DFD88D17C" httpUri="http://treatment.plazi.org/id/53544669FF97FFA5EE44FD2DFD88D17C" lastPageId="6" lastPageNumber="7" pageId="2" pageNumber="3">
<subSubSection id="93E7A4F4FF97FFA1EE44FD2DFB39D0BA" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="DB42F77FFF97FFA1EE44FD2DFB39D0BA" blockId="2.[371,1216,701,798]" pageId="2" pageNumber="3">
<heading id="800A4013FF97FFA1EE44FD2DFC40D099" box="[635,953,701,728]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
<taxonomicName id="1CFD8CFCFF97FFA1EE44FD2DFCAED096" ID-CoL="9HNBH" authority="Peñafiel-Vinueza &amp; Rafael, 2019" authorityName="Peñafiel-Vinueza &amp; Rafael" authorityYear="2019" box="[635,855,701,728]" class="Insecta" family="Drosophilidae" genus="Drosophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="warmi" status="sp. nov.">
<emphasis id="E9892B6DFF97FFA1EE44FD2DFCAED096" bold="true" box="[635,855,701,728]" italics="true" pageId="2" pageNumber="3">Drosophila warmi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2BA9616FF97FFA1EF61FD2EFC40D099" box="[862,953,702,728]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
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<uri id="AF6CFB7DFF97FFA1ED4CFD70FB39D0BA" box="[371,1216,736,763]" pageId="2" pageNumber="3">
urn:lsid:zoobank.org:act:
<uuid id="AF5BCDAAFF97FFA1EEACFD70FB39D0BA" box="[659,1216,736,763]" pageId="2" pageNumber="3">F64D7E97-D209-4D70-B011-493B438BFE59</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="93E7A4F4FF97FFA1EED8FC94FCB4D15F" box="[743,845,772,798]" pageId="2" pageNumber="3" type="description">
<paragraph id="DB42F77FFF97FFA1EED8FC94FCB4D15F" blockId="2.[371,1216,701,798]" box="[743,845,772,798]" pageId="2" pageNumber="3">
<figureCitation id="43C6EBFAFF97FFA1EED8FC94FCB4D15F" box="[743,845,772,798]" captionStart-0="Fig. 1" captionStart-1="Fig. 2" captionStart-2="Fig. 3" captionStartId-0="3.[189,232,1964,1990]" captionStartId-1="4.[189,232,1928,1954]" captionStartId-2="5.[189,232,1894,1920]" captionTargetBox-0="[317,1269,830,1935]" captionTargetBox-1="[360,1228,516,1907]" captionTargetBox-2="[275,1313,651,1860]" captionTargetId-0="figure@3.[317,1269,830,1935]" captionTargetId-1="figure@4.[360,1228,516,1907]" captionTargetId-2="figure@5.[275,1313,651,1860]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Fig. 1. Drosophila warmi sp. nov. Holotype, ♀ (QCAZ-I 3344). A. Abdomen and wing, dorsal view. B. Ovipositor. C. Spermatheca." captionText-1="Fig. 2. Drosophila warmi sp. nov. Allotype, ♂ (QCAZ-I 3345). A. Abdomen, dorsal view. B. Epandrium, cerci, surstylus, decasternum. C. Hypandrium, gonopods and paraphyses in ventral view. DF. Aedeagus in ventral, lateral and dorsal view, respectively." captionText-2="Fig. 3. Drosophila warmi sp. nov. AC. Allotype, ♂ (QCAZ-I 3345), aedeagus in ventro-lateral to lateral view, respectively; arrows show the two lateral sclerotized projections. D. Holotype, ♀ (QCAZ-I 3344), spermatheca; arrow shows the centrodistal spines. E. Paratype, ♀ (QCAZ-I 33XX), spermatheca; arrow shows the centrodistal spines." httpUri-0="https://zenodo.org/record/2561926/files/figure.png" httpUri-1="https://zenodo.org/record/2561928/files/figure.png" httpUri-2="https://zenodo.org/record/2561930/files/figure.png" pageId="2" pageNumber="3">Figs 13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="93E7A4F4FF97FFA1EC82FCD5FD8BD6E6" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="DB42F77FFF97FFA1EC82FCD5FECFD11E" blockId="2.[189,310,837,863]" box="[189,310,837,863]" pageId="2" pageNumber="3">
<heading id="800A4013FF97FFA1EC82FCD5FECFD11E" bold="true" box="[189,310,837,863]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="E9892B6DFF97FFA1EC82FCD5FECFD11E" bold="true" box="[189,310,837,863]" pageId="2" pageNumber="3">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DB42F77FFF97FFA1EC82FCE4FD8BD6E6" blockId="2.[189,1399,884,1191]" pageId="2" pageNumber="3">
Aristae with six dorsal and three ventral branches, plus terminal fork. Prominent vibrissa. Thorax yellowish brown. Wings transparent yellow, bM-Cu slightly clouded. Male abdomen yellow with dorsal midline, 2
<superScript id="2C885A37FF97FFA1ED08FC28FEB1D187" attach="left" box="[311,328,952,966]" fontSize="6" pageId="2" pageNumber="3">nd</superScript>
to 4
<superScript id="2C885A37FF97FFA1EDBDFC28FE76D187" attach="left" box="[386,399,952,966]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
tergites with thin pigmented bands, 5
<superScript id="2C885A37FF97FFA1EF64FC28FC91D187" attach="left" box="[859,872,952,966]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
tergite without pigmentation and 6
<superScript id="2C885A37FF97FFA1E92CFC28FAD9D187" attach="left" box="[1299,1312,952,966]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
with a median spot. Female abdomen yellow with dorsal midline, 1
<superScript id="2C885A37FF97FFA1EFBAFC4CFC69D1AB" attach="left" box="[901,912,988,1002]" fontSize="6" pageId="2" pageNumber="3">st</superScript>
tergite yellow, 2
<superScript id="2C885A37FF97FFA1E86AFC4CFB9FD1AB" attach="left" box="[1109,1126,988,1002]" fontSize="6" pageId="2" pageNumber="3">nd</superScript>
to 4
<superScript id="2C885A37FF97FFA1E8A4FC4CFB51D1AB" attach="left" box="[1179,1192,988,1002]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
tergites with thin pigmented bands, 5
<superScript id="2C885A37FF97FFA1ED99FC6FFE4AD64C" attach="left" box="[422,435,1023,1037]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
and 6
<superScript id="2C885A37FF97FFA1EDC0FC6FFDF5D64C" attach="left" box="[511,524,1023,1037]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
with a median spot between both tergites. Cerci not fused to epandrium. Hypandrium shield-shaped. Gonopod bearing one thick bristle. Paraphyses fused to gonopod bearing one small bristle. Aedeagus tubular and bifid, with two lateral sclerotized and serrated projections, and two ventral membranous enlargements covered in bright studs. Spermatheca balloon-shaped, with a distal dimple covered in short spines.
</paragraph>
</subSubSection>
<subSubSection id="93E7A4F4FF97FFA1EC82FB5FFA8ED759" box="[189,1399,1231,1304]" pageId="2" pageNumber="3" type="etymology">
<paragraph id="DB42F77FFF97FFA1EC82FB5FFEBDD6A8" blockId="2.[189,324,1231,1257]" box="[189,324,1231,1257]" pageId="2" pageNumber="3">
<heading id="800A4013FF97FFA1EC82FB5FFEBDD6A8" bold="true" box="[189,324,1231,1257]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="E9892B6DFF97FFA1EC82FB5FFEBDD6A8" bold="true" box="[189,324,1231,1257]" pageId="2" pageNumber="3">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="DB42F77FFF97FFA1EC82FB6EFA8ED759" blockId="2.[189,1399,1277,1304]" box="[189,1399,1277,1304]" pageId="2" pageNumber="3">
In the Kichwa language,
<taxonomicName id="1CFD8CFCFF97FFA1EDD8FB6EFDCBD756" authorityName="Peñafiel-Vinueza &amp; Rafael" authorityYear="2019" box="[487,562,1278,1303]" class="Insecta" family="Drosophilidae" genus="Drosophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="warmi">
<emphasis id="E9892B6DFF97FFA1EDD8FB6EFDCBD756" box="[487,562,1278,1303]" italics="true" pageId="2" pageNumber="3">warmi</emphasis>
</taxonomicName>
refers to woman. The species name is in honor of Ecuadorian women.
</paragraph>
</subSubSection>
<subSubSection id="93E7A4F4FF97FFA1EC82FAAEFA88D565" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="DB42F77FFF97FFA1EC82FAAEFE53D719" blockId="2.[189,426,1342,1368]" box="[189,426,1342,1368]" pageId="2" pageNumber="3">
<heading id="800A4013FF97FFA1EC82FAAEFE53D719" bold="true" box="[189,426,1342,1368]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="E9892B6DFF97FFA1EC82FAAEFE53D719" bold="true" box="[189,426,1342,1368]" pageId="2" pageNumber="3">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DB42F77FFF97FFA1ECD5FAFDFEA5D7C6" blockId="2.[189,1399,1389,1521]" box="[234,348,1389,1415]" pageId="2" pageNumber="3">
<typeStatus id="044649DDFF97FFA1ECD5FAFDFEA5D7C6" box="[234,348,1389,1415]" pageId="2" pageNumber="3" type="holotype">
<emphasis id="E9892B6DFF97FFA1ECD5FAFDFEA5D7C6" bold="true" box="[234,348,1389,1415]" pageId="2" pageNumber="3">Holotype</emphasis>
</typeStatus>
</paragraph>
<paragraph id="DB42F77FFF97FFA1EC82FA01FE5CD7B0" blockId="2.[189,1399,1389,1521]" pageId="2" pageNumber="3">
<materialsCitation id="6B95FD22FF97FFA1EC82FA01FE5CD7B0" ID-GBIF-Occurrence="1993784576" collectingDate="2015-04-23" collectionCode="QCAZ" collectorName="A. Peñafiel" country="Ecuador" determinerName="V. Rafael" elevation="1000" latitude="-4.116611" location="Bombuscaro" longLatPrecision="1" longitude="-78.968025" pageId="2" pageNumber="3" specimenCode="QCAZ-I 3344" specimenCount="1" specimenCount-female="1" stateProvince="Zamora Chinchipe" typeStatus="holotype">
<collectingCountry id="A3EAB7EFFF97FFA1EC82FA01FEB2D7EA" box="[189,331,1425,1451]" name="Ecuador" pageId="2" pageNumber="3">ECUADOR</collectingCountry>
<specimenCount id="CDFB3CF6FF97FFA1ED61FA01FE8DD7EA" box="[350,372,1425,1451]" pageId="2" pageNumber="3" type="female"></specimenCount>
(dissected, terminalia in microvial, dry mounted);
<collectingRegion id="1939399DFF97FFA1EF8AFA01FB77D7EB" box="[949,1166,1424,1451]" country="Ecuador" name="Zamora-Chinchipe" pageId="2" pageNumber="3">Zamora Chinchipe</collectingRegion>
Prov.,
<location id="DE22A1A4FF97FFA1E8E4FA00FA8BD7EB" LSID="urn:lsid:plazi:treatment:53544669FF97FFA5EE44FD2DFD88D17C:DE22A1A4FF97FFA1E8E4FA00FA8BD7EB" box="[1243,1394,1424,1450]" country="Ecuador" latitude="-4.116611" longLatPrecision="1" longitude="-78.968025" name="Bombuscaro" pageId="2" pageNumber="3" stateProvince="Zamora Chinchipe">Bombuscaro</location>
,
<elevation id="50D0104CFF97FFA1EC82FA24FEA1D78F" box="[189,344,1460,1486]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="3" unit="m" value="1000.0">
<quantity id="1C055A9AFF97FFA1EC82FA24FEEED78F" box="[189,279,1460,1486]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="3" unit="m" value="1000.0">1000 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="BEC991B8FF97FFA1ED58FA22FDF4D78F" box="[359,525,1458,1486]" direction="south" orientation="latitude" pageId="2" pageNumber="3" precision="1" value="-4.116611">04°0659.8″ S</geoCoordinate>
,
<geoCoordinate id="BEC991B8FF97FFA1EE24FA22FD37D78F" box="[539,718,1458,1486]" direction="west" orientation="longitude" pageId="2" pageNumber="3" precision="1" value="-78.968025">78°5804.9″ W</geoCoordinate>
;
<collectingDate id="BF072857FF97FFA1EEE4FA24FC80D78F" box="[731,889,1459,1486]" pageId="2" pageNumber="3" value="2015-04-23">
<date id="AF43D1BFFF97FFA1EEE4FA24FC80D78F" box="[731,889,1459,1486]" pageId="2" pageNumber="3" value="2015-04-23">23 Apr. 2015</date>
</collectingDate>
;
<collectorName id="760892A9FF97FFA1EFBBFA24FBF1D78F" box="[900,1032,1460,1486]" pageId="2" pageNumber="3">A. Peñafiel</collectorName>
leg.;
<collectorName id="760892A9FF97FFA1E874FA24FB37D78F" box="[1099,1230,1460,1486]" pageId="2" pageNumber="3">A. Peñafiel</collectorName>
and
<determinerName id="2E9E1BBBFF97FFA1E936FA24FA8ED78F" box="[1289,1399,1460,1486]" pageId="2" pageNumber="3">V. Rafael</determinerName>
det.;
<specimenCode id="8B5B5F04FF97FFA1ECC7FA47FE59D7B0" box="[248,416,1495,1521]" pageId="2" pageNumber="3">QCAZ-I 3344</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="DB42F77FFF97FFA1ECD5F988FEADD473" blockId="2.[189,1398,1560,1657]" box="[234,340,1560,1586]" pageId="2" pageNumber="3">
<typeStatus id="044649DDFF97FFA1ECD5F988FEADD473" box="[234,340,1560,1586]" pageId="2" pageNumber="3" type="allotype">
<emphasis id="E9892B6DFF97FFA1ECD5F988FEADD473" bold="true" box="[234,340,1560,1586]" pageId="2" pageNumber="3">Allotype</emphasis>
</typeStatus>
</paragraph>
<paragraph id="DB42F77FFF97FFA1EC82F9ACFD5CD438" blockId="2.[189,1398,1560,1657]" pageId="2" pageNumber="3">
<materialsCitation id="6B95FD22FF97FFA1EC82F9ACFD5CD438" ID-GBIF-Occurrence="1994756077" collectingDate="2015-04-23" collectionCode="QCAZ" collectorName="A. Peñafiel" country="Ecuador" determinerName="V. Rafael" elevation="1000" latitude="-4.116611" location="Bombuscaro" longLatPrecision="1" longitude="-78.968025" pageId="2" pageNumber="3" specimenCode="QCAZ-I 3345" specimenCount="1" specimenCount-male="1" stateProvince="Zamora Chinchipe" typeStatus="allotype">
<collectingCountry id="A3EAB7EFFF97FFA1EC82F9ACFEB2D417" box="[189,331,1596,1622]" name="Ecuador" pageId="2" pageNumber="3">ECUADOR</collectingCountry>
• ♂ (dissected, terminalia in microvial, dry mounted, descendant from isofemale line F
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); same data as for holotype; QCAZ-I 3345.
</materialsCitation>
</paragraph>
<paragraph id="DB42F77FFF97FFA1ECD5F930FE9ED4FB" blockId="2.[189,1399,1696,1834]" box="[234,359,1696,1722]" pageId="2" pageNumber="3">
<emphasis id="E9892B6DFF97FFA1ECD5F930FE9ED4FB" bold="true" box="[234,359,1696,1722]" pageId="2" pageNumber="3">
<typeStatus id="044649DDFF97FFA1ECD5F930FE9ED4FB" box="[234,359,1696,1722]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DB42F77FFF97FFA1EC82F953FA88D565" blockId="2.[189,1399,1696,1834]" pageId="2" pageNumber="3">
<materialsCitation id="6B95FD22FF97FFA1EC82F953FC06D540" ID-GBIF-Occurrence="1993784574" collectingDate="2015-04-23" collectionCode="QCAZ" collectorName="A. Peñafiel" country="Ecuador" determinerName="V. Rafael" elevation="1000" latitude="-4.116611" location="Bombuscaro" longLatPrecision="1" longitude="-78.968025" pageId="2" pageNumber="3" specimenCode="QCAZ-I 3344" specimenCount="8" specimenCount-female="8" stateProvince="Zamora Chinchipe" typeStatus="paratype">
<collectingCountry id="A3EAB7EFFF97FFA1EC82F953FEB2D49C" box="[189,331,1731,1757]" name="Ecuador" pageId="2" pageNumber="3">ECUADOR</collectingCountry>
<specimenCount id="CDFB3CF6FF97FFA1ED5CF953FE5CD49F" box="[355,421,1731,1758]" pageId="2" pageNumber="3" type="female">8 ♀♀</specimenCount>
(dissected, terminalia in microvial, dry mounted, from among 24 descendants from isofemale line F
<subScript id="4779F53AFF97FFA1EDBFF968FE70D547" attach="left" box="[384,393,1784,1798]" fontSize="6" pageId="2" pageNumber="3">1</subScript>
); same data as for holotype; QCAZ-I 3347 to 3354
</materialsCitation>
<materialsCitation id="6B95FD22FF97FFA1E821F977FA94D565" ID-GBIF-Occurrence="1993784573" country="Ecuador" pageId="2" pageNumber="3" specimenCount="1" typeStatus="paratype">
1 ♂ (dissected, terminalia in microvial, dry mounted, descendant from isofemale line F
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); same data as for holotype; QCAZ-I 3346
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="93E7A4F4FF97FFA5EC82F8DBFEF7D07B" lastPageId="6" lastPageNumber="7" pageId="2" pageNumber="3" type="description">
<paragraph id="DB42F77FFF97FFA1EC82F8DBFEB7D524" blockId="2.[189,334,1867,1893]" box="[189,334,1867,1893]" pageId="2" pageNumber="3">
<heading id="800A4013FF97FFA1EC82F8DBFEB7D524" bold="true" box="[189,334,1867,1893]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="E9892B6DFF97FFA1EC82F8DBFEB7D524" bold="true" box="[189,334,1867,1893]" pageId="2" pageNumber="3">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DB42F77FFF97FFA1ECD5F8EAFAE6D59A" blockId="2.[189,1398,1914,2011]" pageId="2" pageNumber="3">
<emphasis id="E9892B6DFF97FFA1ECD5F8EAFEBCD5D5" bold="true" box="[234,325,1914,1940]" pageId="2" pageNumber="3">Female</emphasis>
The female was chosen as the
<typeStatus id="044649DDFF97FFA1EE2DF80DFD80D5F6" box="[530,633,1949,1975]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
for this species because both males are in poor condition.
<typeStatus id="044649DDFF97FFA1E936F80DFA8FD5F6" box="[1289,1398,1949,1975]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
external morphology: total length (body + wings)
<quantity id="1C055A9AFF97FFA1EF36F850FCA9D59A" box="[777,848,1984,2011]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="3" unit="mm" value="4.0">4 mm</quantity>
, body length
<quantity id="1C055A9AFF97FFA1EFD1F850FBCCD59A" box="[1006,1077,1984,2011]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="3" unit="mm" value="3.0">3 mm</quantity>
. Body color yellow.
</paragraph>
<paragraph id="DB42F77FFF96FFA0EC82FE9CFE1DD3B8" blockId="3.[189,1399,267,505]" pageId="3" pageNumber="4">
HEAD. Aristae with six dorsal and three ventral branches, plus terminal fork and fine hairs. Head with orbital plate yellowish brown, frontal length
<quantity id="1C055A9AFF96FFA0EEF5FEBFFCCFD308" box="[714,822,303,329]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="4" unit="mm" value="0.27">0.27 mm</quantity>
, frontal index = 0.61, top to bottom width ratio = 0.18. Medial vertical seta closer to lateral vertical seta and slightly towards outer edge of orbital plate, vt index = 0.19; or1or3 ratio = 1, or2or1 ratio = 0.91. Ocellar triangle brown, ocellus yellow; frontal triangle yellowish brown. Frontal vitta yellowish brown. Gena and postgena yellow, cheek index = 9.5. Vibrissa prominent, vibrissa index = 0.27. Carina prominent and not sulcate. Proboscis yellow. Eyes scarlet, eye index = 4.02.
</paragraph>
<paragraph id="DB42F77FFF96FFA0EC82FDB0FAB2D0C1" blockId="3.[189,1398,544,641]" pageId="3" pageNumber="4">
THORAX. Yellowish brown, thorax length
<quantity id="1C055A9AFF96FFA0EEA2FDB0FCF1D07B" box="[669,776,544,570]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" pageId="3" pageNumber="4" unit="mm" value="0.94">0.94 mm</quantity>
, acrostichal hairs in seven rows between two anterior dorsocentral setae, h index = 1.37, dc index = 0.82. Scutellum yellow; basal scutellar setae divergent, scut index = 1.12. Medial katepisternal seta same size as anterior, sterno index = 0.51. Legs yellow.
</paragraph>
<paragraph id="DB42F77FFF96FFA0EC82FD38FDE9D148" blockId="3.[189,1399,680,777]" pageId="3" pageNumber="4">
WINGS. Transparent yellow, bM-Cu slightly clouded (
<figureCitation id="43C6EBFAFF96FFA0EF66FD38FC47D083" box="[857,958,680,706]" captionStart="Fig. 1" captionStartId="3.[189,232,1964,1990]" captionTargetBox="[317,1269,830,1935]" captionTargetId="figure@3.[317,1269,830,1935]" captionTargetPageId="3" captionText="Fig. 1. Drosophila warmi sp. nov. Holotype, ♀ (QCAZ-I 3344). A. Abdomen and wing, dorsal view. B. Ovipositor. C. Spermatheca." httpUri="https://zenodo.org/record/2561926/files/figure.png" pageId="3" pageNumber="4">Fig. 1A</figureCitation>
). Alar length
<quantity id="1C055A9AFF96FFA0E84BFD38FB1ED083" box="[1140,1255,680,707]" metricMagnitude="-3" metricUnit="m" metricValue="2.51" pageId="3" pageNumber="4" unit="mm" value="2.51">2.51 mm</quantity>
, alar width
<quantity id="1C055A9AFF96FFA0EC82FD5BFED2D0A7" box="[189,299,715,742]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="4" unit="mm" value="1.1">1.10 mm</quantity>
. Alar indices: alar = 2.26; C = 4.21; a c = 1.68; hb = 0.47; 4c = 0.6;
<date id="AF43D1BFFF96FFA0E84BFD5BFB68D0A4" box="[1140,1169,715,741]" pageId="3" pageNumber="4">4v</date>
= 1.42;
<date id="AF43D1BFFF96FFA0E8C5FD5BFAE1D0A7" box="[1274,1304,715,742]" pageId="3" pageNumber="4">5x</date>
= 1.08; M = 0.38 and prox. x = 0.58.
</paragraph>
<caption id="8F82A7F7FF96FFA0EC82F83CFDD4D5A8" httpUri="https://zenodo.org/record/2561926/files/figure.png" pageId="3" pageNumber="4" startId="3.[189,232,1964,1990]" targetBox="[317,1269,830,1935]" targetPageId="3">
<paragraph id="DB42F77FFF96FFA0EC82F83CFDD4D5A8" blockId="3.[189,1399,1964,2025]" pageId="3" pageNumber="4">
<emphasis id="E9892B6DFF96FFA0EC82F83CFEF4D587" bold="true" box="[189,269,1964,1990]" pageId="3" pageNumber="4">Fig. 1.</emphasis>
<taxonomicName id="1CFD8CFCFF96FFA0ED29F83CFE16D587" authority="Peñafiel-Vinueza &amp; Rafael, 2019" authorityName="Peñafiel-Vinueza &amp; Rafael" authorityYear="2019" box="[278,495,1964,1990]" class="Insecta" family="Drosophilidae" genus="Drosophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="warmi" status="sp. nov.">
<emphasis id="E9892B6DFF96FFA0ED29F83CFE16D587" box="[278,495,1964,1990]" italics="true" pageId="3" pageNumber="4">Drosophila warmi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2BA9616FF96FFA0EDC7F83CFDACD587" box="[504,597,1964,1990]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
Holotype, ♀ (QCAZ-I 3344).
<emphasis id="E9892B6DFF96FFA0EFF8F83CFC25D587" bold="true" box="[967,988,1964,1990]" pageId="3" pageNumber="4">A</emphasis>
. Abdomen and wing, dorsal view.
<emphasis id="E9892B6DFF96FFA0EC82F85FFF28D5A8" bold="true" box="[189,209,1999,2025]" pageId="3" pageNumber="4">B</emphasis>
. Ovipositor.
<emphasis id="E9892B6DFF96FFA0ED55F85FFE86D5A8" bold="true" box="[362,383,1999,2025]" pageId="3" pageNumber="4">C</emphasis>
. Spermatheca.
</paragraph>
</caption>
<paragraph id="DB42F77FFF91FFA7EC82FE9CFC68D308" blockId="4.[189,1398,266,329]" pageId="4" pageNumber="5">
ABDOMEN. Yellow with dorsal midline, 1
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tergite yellow, 2
<superScript id="2C885A37FF91FFA7EF4CFE9AFC7DD359" attach="left" box="[883,900,266,280]" fontSize="6" pageId="4" pageNumber="5">nd</superScript>
to 4
<superScript id="2C885A37FF91FFA7EF83FE9AFC30D359" attach="left" box="[956,969,266,280]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
tergites with thin pigmented bands, 5
<superScript id="2C885A37FF91FFA7ECF3FEBDFF20D37A" attach="left" box="[204,217,301,315]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
and 6
<superScript id="2C885A37FF91FFA7ED1EFEBDFED7D37A" attach="left" box="[289,302,301,315]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
with a median spot between both tergites (
<figureCitation id="43C6EBFAFF91FFA7EF1AFEBEFC7BD308" box="[805,898,302,329]" captionStart="Fig. 1" captionStartId="3.[189,232,1964,1990]" captionTargetBox="[317,1269,830,1935]" captionTargetId="figure@3.[317,1269,830,1935]" captionTargetPageId="3" captionText="Fig. 1. Drosophila warmi sp. nov. Holotype, ♀ (QCAZ-I 3344). A. Abdomen and wing, dorsal view. B. Ovipositor. C. Spermatheca." httpUri="https://zenodo.org/record/2561926/files/figure.png" pageId="4" pageNumber="5">Fig. 1A</figureCitation>
).
</paragraph>
<paragraph id="DB42F77FFF91FFA7EC82FEE0FDBCD391" blockId="4.[189,1398,368,464]" pageId="4" pageNumber="5">
TERMINALIA. Ovipositor sclerotized, slipper-shaped, with 15 marginal and five discal teeth, one long bristle and three fine hairs (
<figureCitation id="43C6EBFAFF91FFA7EE2FFE03FD96D3EC" box="[528,623,403,429]" captionStart="Fig. 1" captionStartId="3.[189,232,1964,1990]" captionTargetBox="[317,1269,830,1935]" captionTargetId="figure@3.[317,1269,830,1935]" captionTargetPageId="3" captionText="Fig. 1. Drosophila warmi sp. nov. Holotype, ♀ (QCAZ-I 3344). A. Abdomen and wing, dorsal view. B. Ovipositor. C. Spermatheca." httpUri="https://zenodo.org/record/2561926/files/figure.png" pageId="4" pageNumber="5">Fig. 1B</figureCitation>
). Spermatheca sclerotized, balloon-shaped, with a distal dimple covered in short spines (
<figureCitation id="43C6EBFAFF91FFA7EDE4FE26FDCFD391" box="[475,566,438,464]" captionStart="Fig. 1" captionStartId="3.[189,232,1964,1990]" captionTargetBox="[317,1269,830,1935]" captionTargetId="figure@3.[317,1269,830,1935]" captionTargetPageId="3" captionText="Fig. 1. Drosophila warmi sp. nov. Holotype, ♀ (QCAZ-I 3344). A. Abdomen and wing, dorsal view. B. Ovipositor. C. Spermatheca." httpUri="https://zenodo.org/record/2561926/files/figure.png" pageId="4" pageNumber="5">Fig. 1C</figureCitation>
).
</paragraph>
<caption id="8F82A7F7FF91FFA7EC82F818FD1FD5A8" httpUri="https://zenodo.org/record/2561928/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,232,1928,1954]" targetBox="[360,1228,516,1907]" targetPageId="4">
<paragraph id="DB42F77FFF91FFA7EC82F818FD1FD5A8" blockId="4.[189,1399,1927,2025]" pageId="4" pageNumber="5">
<emphasis id="E9892B6DFF91FFA7EC82F818FEF0D5E3" bold="true" box="[189,265,1928,1954]" pageId="4" pageNumber="5">Fig. 2.</emphasis>
<taxonomicName id="1CFD8CFCFF91FFA7ED31F818FE1DD5E3" authority="Peñafiel-Vinueza &amp; Rafael, 2019" authorityName="Peñafiel-Vinueza &amp; Rafael" authorityYear="2019" box="[270,484,1928,1954]" class="Insecta" family="Drosophilidae" genus="Drosophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="warmi" status="sp. nov.">
<emphasis id="E9892B6DFF91FFA7ED31F818FE1DD5E3" box="[270,484,1928,1954]" italics="true" pageId="4" pageNumber="5">Drosophila warmi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2BA9616FF91FFA7EDD6F819FDBBD5E2" box="[489,578,1929,1955]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
Allotype, ♂ (QCAZ-I 3345).
<emphasis id="E9892B6DFF91FFA7EFA6F818FC57D5E3" bold="true" box="[921,942,1928,1954]" pageId="4" pageNumber="5">A</emphasis>
. Abdomen, dorsal view.
<emphasis id="E9892B6DFF91FFA7E8F3F819FB19D5E2" bold="true" box="[1228,1248,1929,1955]" pageId="4" pageNumber="5">B</emphasis>
. Epandrium, cerci, surstylus, decasternum.
<emphasis id="E9892B6DFF91FFA7EE26F83BFDD7D584" bold="true" box="[537,558,1963,1989]" pageId="4" pageNumber="5">C</emphasis>
. Hypandrium, gonopods and paraphyses in ventral view.
<emphasis id="E9892B6DFF91FFA7E8FDF83CFB0ED587" bold="true" box="[1218,1271,1964,1990]" pageId="4" pageNumber="5">DF</emphasis>
. Aedeagus in ventral, lateral and dorsal view, respectively.
</paragraph>
</caption>
<paragraph id="DB42F77FFF90FFA6ECD5FE9BFCCCD367" blockId="5.[189,1399,267,435]" box="[234,821,267,294]" pageId="5" pageNumber="6">
<emphasis id="E9892B6DFF90FFA6ECD5FE9BFDFAD367" bold="true" box="[234,515,267,294]" pageId="5" pageNumber="6">
Variation in
<typeStatus id="044649DDFF90FFA6EDB8FE9CFDFAD367" box="[391,515,268,294]" pageId="5" pageNumber="6" type="paratype">paratypes</typeStatus>
</emphasis>
(dry mounted specimens)
</paragraph>
<paragraph id="DB42F77FFF90FFA6EC82FEBEFCFCD3F2" blockId="5.[189,1399,267,435]" pageId="5" pageNumber="6">
Head: frontal length
<quantity id="1C055A9AFF90FFA6ED8BFEBEFD9AD308" box="[436,611,302,329]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" metricValueMax="3.2" metricValueMin="2.6" pageId="5" pageNumber="6" unit="mm" value="0.29" valueMax="0.32" valueMin="0.26">0.260.32 mm</quantity>
, frontal index = 0.630.72, top to bottom width ratio = 0.130.22; vt index = 0.140.22, or1or3 ratio = 0.841.33, or2or1 ratio = 0.671.27; cheek index = 8.7112.4; vibrissa index = 0.140.39; eye index = 2.23.87. Thorax: h index = 0.702.2, dc index = 0.740.82; scut index = 1.121.47; sterno index = 0.510.66.
</paragraph>
<paragraph id="DB42F77FFF90FFA6ECD5FE4AFED3D3B5" blockId="5.[189,1399,474,606]" box="[234,298,474,500]" pageId="5" pageNumber="6">
<emphasis id="E9892B6DFF90FFA6ECD5FE4AFED3D3B5" bold="true" box="[234,298,474,500]" pageId="5" pageNumber="6">Male</emphasis>
</paragraph>
<paragraph id="DB42F77FFF90FFA6EC82FE6DFD8FD01C" blockId="5.[189,1399,474,606]" pageId="5" pageNumber="6">
Reared from an isofemale line. Only two males emerged and both died crushed into the culture media. Morphologically with same characteristics as female except for abdomen, which is yellow and with a faint tiny spot on 6
<superScript id="2C885A37FF90FFA6EDA4FDD1FE51D00E" attach="left" box="[411,424,577,591]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
tergite (
<figureCitation id="43C6EBFAFF90FFA6EE35FDD3FD91D01C" box="[522,616,579,605]" captionStart="Fig. 2" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[360,1228,516,1907]" captionTargetId="figure@4.[360,1228,516,1907]" captionTargetPageId="4" captionText="Fig. 2. Drosophila warmi sp. nov. Allotype, ♂ (QCAZ-I 3345). A. Abdomen, dorsal view. B. Epandrium, cerci, surstylus, decasternum. C. Hypandrium, gonopods and paraphyses in ventral view. DF. Aedeagus in ventral, lateral and dorsal view, respectively." httpUri="https://zenodo.org/record/2561928/files/figure.png" pageId="5" pageNumber="6">Fig. 2A</figureCitation>
).
</paragraph>
<caption id="8F82A7F7FF90FFA6EC82F8F6FD9FD5AB" httpUri="https://zenodo.org/record/2561930/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1894,1920]" targetBox="[275,1313,651,1860]" targetPageId="5">
<paragraph id="DB42F77FFF90FFA6EC82F8F6FD9FD5AB" blockId="5.[189,1399,1893,2027]" pageId="5" pageNumber="6">
<emphasis id="E9892B6DFF90FFA6EC82F8F6FEF7D5C1" bold="true" box="[189,270,1894,1920]" pageId="5" pageNumber="6">Fig. 3.</emphasis>
<taxonomicName id="1CFD8CFCFF90FFA6ED25F8F6FE0CD5C1" authority="Peñafiel-Vinueza &amp; Rafael, 2019" authorityName="Peñafiel-Vinueza &amp; Rafael" authorityYear="2019" box="[282,501,1894,1920]" class="Insecta" family="Drosophilidae" genus="Drosophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="warmi" status="sp. nov.">
<emphasis id="E9892B6DFF90FFA6ED25F8F6FE0CD5C1" box="[282,501,1894,1920]" italics="true" pageId="5" pageNumber="6">Drosophila warmi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2BA9616FF90FFA6EE3FF8F7FDA7D5C0" box="[512,606,1895,1921]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="E9892B6DFF90FFA6EE56F8F6FD5BD5C1" bold="true" box="[617,674,1894,1920]" pageId="5" pageNumber="6">AC</emphasis>
. Allotype, ♂ (QCAZ-I 3345), aedeagus in ventro-lateral to lateral view, respectively; arrows show the two lateral sclerotized projections.
<emphasis id="E9892B6DFF90FFA6E871F81AFB9AD5E5" bold="true" box="[1102,1123,1930,1956]" pageId="5" pageNumber="6">D</emphasis>
. Holotype, ♀ (QCAZ-I 3344), spermatheca; arrow shows the centrodistal spines.
<emphasis id="E9892B6DFF90FFA6EF66F83DFC94D586" bold="true" box="[857,877,1965,1991]" pageId="5" pageNumber="6">E</emphasis>
. Paratype, ♀ (QCAZ-I 33XX), spermatheca; arrow shows the centrodistal spines.
</paragraph>
</caption>
<paragraph id="DB42F77FFF93FFA5EC82FE9CFB5FD3CE" blockId="6.[189,1398,267,399]" pageId="6" pageNumber="7">
MALE TERMINALIA. Epandrium dorsally microtrichose, ventral lobe with 9 bristles on right and 6 on left side. Cerci not fused to epandrium, dorsally microtrichose. Surstylus with 9 primary teeth and 13 marginal bristles on right side and 14 on left (
<figureCitation id="43C6EBFAFF93FFA5EECFFEC2FCB6D32D" box="[752,847,338,364]" captionStart="Fig. 2" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[360,1228,516,1907]" captionTargetId="figure@4.[360,1228,516,1907]" captionTargetPageId="4" captionText="Fig. 2. Drosophila warmi sp. nov. Allotype, ♂ (QCAZ-I 3345). A. Abdomen, dorsal view. B. Epandrium, cerci, surstylus, decasternum. C. Hypandrium, gonopods and paraphyses in ventral view. DF. Aedeagus in ventral, lateral and dorsal view, respectively." httpUri="https://zenodo.org/record/2561928/files/figure.png" pageId="6" pageNumber="7">Fig. 2B</figureCitation>
). Hypandrium shield-shaped, with sclerotized edge. Gonopod U-shaped, with some bright studs, bearing one thick bristle (
<figureCitation id="43C6EBFAFF93FFA5E803FEE5FB60D3CE" box="[1084,1177,373,399]" captionStart="Fig. 2" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[360,1228,516,1907]" captionTargetId="figure@4.[360,1228,516,1907]" captionTargetPageId="4" captionText="Fig. 2. Drosophila warmi sp. nov. Allotype, ♂ (QCAZ-I 3345). A. Abdomen, dorsal view. B. Epandrium, cerci, surstylus, decasternum. C. Hypandrium, gonopods and paraphyses in ventral view. DF. Aedeagus in ventral, lateral and dorsal view, respectively." httpUri="https://zenodo.org/record/2561928/files/figure.png" pageId="6" pageNumber="7">Fig. 2C</figureCitation>
).
</paragraph>
<paragraph id="DB42F77FFF93FFA5EC82FE27FEF7D07B" blockId="6.[189,1399,438,570]" pageId="6" pageNumber="7">
AEDEAGUS. Tubular and bifid, with two lateral sclerotized and serrated projections, below these projections with two ventral, membranous enlargements covered in bright studs. Aedeagal apodeme shorter than aedeagus. Ventral rod slightly developed (
<figureCitation id="43C6EBFAFF93FFA5EE94FE6DFCD3D056" box="[683,810,509,535]" captionStart="Fig. 2" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[360,1228,516,1907]" captionTargetId="figure@4.[360,1228,516,1907]" captionTargetPageId="4" captionText="Fig. 2. Drosophila warmi sp. nov. Allotype, ♂ (QCAZ-I 3345). A. Abdomen, dorsal view. B. Epandrium, cerci, surstylus, decasternum. C. Hypandrium, gonopods and paraphyses in ventral view. DF. Aedeagus in ventral, lateral and dorsal view, respectively." httpUri="https://zenodo.org/record/2561928/files/figure.png" pageId="6" pageNumber="7">Fig. 2DF</figureCitation>
). Paraphyses fused to gonopod, bearing one small bristle.
</paragraph>
</subSubSection>
<subSubSection id="93E7A4F4FF93FFA5EC82FDF1FD88D17C" pageId="6" pageNumber="7" type="distribution">
<paragraph id="DB42F77FFF93FFA5EC82FDF1FEAFD03A" blockId="6.[189,342,609,635]" box="[189,342,609,635]" pageId="6" pageNumber="7">
<heading id="800A4013FF93FFA5EC82FDF1FEAFD03A" bold="true" box="[189,342,609,635]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="E9892B6DFF93FFA5EC82FDF1FEAFD03A" bold="true" box="[189,342,609,635]" pageId="6" pageNumber="7">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DB42F77FFF93FFA5EC82FD00FDA0D0EB" blockId="6.[189,601,656,682]" box="[189,601,656,682]" pageId="6" pageNumber="7">
Known only from the
<typeStatus id="044649DDFF93FFA5EDFDFD00FE0CD0EB" box="[450,501,656,682]" pageId="6" pageNumber="7">type</typeStatus>
locality.
</paragraph>
<paragraph id="DB42F77FFF93FFA5EC82FD41FDDBD0AA" blockId="6.[189,546,721,748]" box="[189,546,721,748]" pageId="6" pageNumber="7">
<heading id="800A4013FF93FFA5EC82FD41FDDBD0AA" bold="true" box="[189,546,721,748]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="E9892B6DFF93FFA5EC82FD41FDDBD0AA" bold="true" box="[189,546,721,748]" pageId="6" pageNumber="7">Relationship to other species</emphasis>
</heading>
</paragraph>
<paragraph id="DB42F77FFF93FFA5EC82FC90FD88D17C" blockId="6.[189,1399,768,830]" pageId="6" pageNumber="7">
This species belongs to subgroup II of the
<taxonomicName id="1CFD8CFCFF93FFA5EEFEFC91FCBAD15B" authorityName="Loew" authorityYear="1862" box="[705,835,769,794]" class="Insecta" family="Drosophilidae" genus="Drosophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tripunctata">
<emphasis id="E9892B6DFF93FFA5EEFEFC91FCBAD15B" box="[705,835,769,794]" italics="true" pageId="6" pageNumber="7">tripunctata</emphasis>
</taxonomicName>
group. The most similar species is
<taxonomicName id="1CFD8CFCFF93FFA5E8CDFC90FD94D17C" authority="Bachli, Vilela &amp; Ratcov, 2000" authorityName="Bachli, Vilela &amp; Ratcov" authorityYear="2000" class="Insecta" family="Drosophilidae" genus="Drosophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="cuaso">
<emphasis id="E9892B6DFF93FFA5E8CDFC90FEF8D17C" italics="true" pageId="6" pageNumber="7">Drosophila cuaso</emphasis>
Bächli,
<bibRefCitation id="BF6C8A8EFF93FFA5ED5CFCB3FD94D17C" author="Vilela C. &amp; Bachli G." box="[355,621,803,830]" pageId="6" pageNumber="7" pagination="49" refId="ref7943" refString="Vilela C. &amp; Bachli G. 2000. Five new species of Neotropical Drosophila. Bulletin de la Societe entomologique suisse 73: 49 ‾ 65." type="journal article" year="2000">Vilela &amp; Ratcov, 2000</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>