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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.509.9506" ID-GBIF-Dataset="66954afc-cfb8-414b-8682-210fe363f991" ID-PMC="PMC4493346" ID-Pensoft-Pub="1313-2970-509-123" ID-PubMed="26175606" ID-ZBK="2664655ECBF6456BBC3DFE805E29D745" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-509-123" ModsDocOrigin="ZooKeys 509" ModsDocTitle="The genus Omphreus in Bosnia and Herzegovina and Montenegro, with two new subspecies of O.morio (Coleoptera, Carabidae, Omphreini)" checkinTime="1451244225243" checkinUser="pensoft" docAuthor="Curcic, Srecko, Sciaky, Riccardo, Antic, Dragan &amp; Vesovic, Nikola" docDate="2015" docId="5336D87B354069C007ED64A0E9CE96B5" docLanguage="en" docName="ZooKeys 509: 123-139" docOrigin="ZooKeys 509" docSource="http://dx.doi.org/10.3897/zookeys.509.9506" docTitle="Omphreus (Omphreus) morio subsp. sandeli Curcic &amp; Sciaky, ssp. n." docType="treatment" docUuid="D7777A72-0E69-4CAC-84BC-1EAE851BD480" docUuidSource="ZooBank" docVersion="5" lastPageNumber="127" masterDocId="FFB2FFED4E115B1FFFA4042CFFE98E09" masterDocTitle="The genus Omphreus in Bosnia and Herzegovina and Montenegro, with two new subspecies of O. morio (Coleoptera, Carabidae, Omphreini)" masterLastPageNumber="139" masterPageNumber="123" pageNumber="127" updateTime="1668160421974" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>The genus Omphreus in Bosnia and Herzegovina and Montenegro, with two new subspecies of O. morio (Coleoptera, Carabidae, Omphreini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Curcic, Srecko</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sciaky, Riccardo</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Antic, Dragan</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Vesovic, Nikola</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>509</mods:number>
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<mods:start>123</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.509.9506</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.509.9506</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-509-123</mods:identifier>
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<treatment ID-GBIF-Taxon="152060423" LSID="urn:lsid:zoobank.org:act:D7777A72-0E69-4CAC-84BC-1EAE851BD480" httpUri="http://treatment.plazi.org/id/5336D87B354069C007ED64A0E9CE96B5" lastPageId="5" lastPageNumber="127" pageId="4" pageNumber="127">
<subSubSection pageId="4" pageNumber="127" type="multiple">
<paragraph pageId="4" pageNumber="127">Taxon classification Animalia Coleoptera Carabidae</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="127" type="nomenclature">
<paragraph pageId="4" pageNumber="127">
<taxonomicName LSID="http://zoobank.org/D7777A72-0E69-4CAC-84BC-1EAE851BD480" authority="Curcic &amp; Sciaky" class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="sandeli">
Omphreus (Omphreus) morio sandeli
<normalizedToken originalValue="Ćurčić">Curcic</normalizedToken>
&amp; Sciaky
</taxonomicName>
<taxonomicNameLabel pageId="4" pageNumber="127">ssp. n.</taxonomicNameLabel>
Figures 4-8
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="127" type="material examined">
<paragraph pageId="4" pageNumber="127">Material examined.</paragraph>
<paragraph pageId="4" pageNumber="127">
Holotype male labeled as follows: &quot;eastern Bosnia and Herzegovina, southern slope of Mt. Zelengora,
<normalizedToken originalValue="Čemerno">Cemerno</normalizedToken>
, 1,450 m a.s.l., near Gacko, 22.
<normalizedToken originalValue="VI">VI-</normalizedToken>
14.VII.2012, from pitfall traps, leg. F. Sandel&quot; (white label, printed) / Holotypus
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="sandeli">Omphreus (Omphreus) morio sandeli</taxonomicName>
ssp. n. S.
<normalizedToken originalValue="Ćurčić">Curcic</normalizedToken>
&amp; R. Sciaky det. 2014&quot; (red label, printed) (IZFB). Paratypes: four males and one female, same data as for holotype (CRS); one male labeled as follows: &quot;eastern Bosnia and Herzegovina, southern slope of Mt. Zelengora,
<normalizedToken originalValue="Čemerno">Cemerno</normalizedToken>
, 1,600 m a.s.l., near
<normalizedToken originalValue="Foča">Foca</normalizedToken>
, 07-24.VIII.2011, from pitfall traps, leg. F. Sandel&quot; (CRS); four males and two females labeled as follows: &quot;eastern Bosnia and Herzegovina, Mt.
<normalizedToken originalValue="Maglić">Maglic</normalizedToken>
,
<normalizedToken originalValue="Tjentište">Tjentiste</normalizedToken>
, 1,450 m a.s.l., near
<normalizedToken originalValue="Foča">Foca</normalizedToken>
, 23.
<normalizedToken originalValue="VI">VI-</normalizedToken>
13.VII.2012, from pitfall traps, leg. F. Sandel&quot; (CRS); two males and one female labeled as follows: &quot;eastern Bosnia and Herzegovina, northern slope of Mt. Zelengora,
<normalizedToken originalValue="Tjentište">Tjentiste</normalizedToken>
, 1,450 m a.s.l., near
<normalizedToken originalValue="Foča">Foca</normalizedToken>
, 23.
<normalizedToken originalValue="VI">VI-</normalizedToken>
13.VII.2012, from pitfall traps, leg. F. Sandel&quot; (CRS). All paratypes are labeled with white, printed locality labels and with red printed labels &quot;Paratypus
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="sandeli">Omphreus (Omphreus) morio sandeli</taxonomicName>
ssp. n. S.
<normalizedToken originalValue="Ćurčić">Curcic</normalizedToken>
&amp; R. Sciaky det. 2014&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="127" type="description">
<paragraph pageId="4" pageNumber="127">Description.</paragraph>
<paragraph pageId="4" pageNumber="127">Size large: TL: R 16.88-19.73 mm (M 18.49 mm). Body elongate; elytra ovoid (Figure 4). Body color black, mouthparts, apical antennomeres, and tarsi black-brownish. Tegument shiny, except slightly matt elytra.</paragraph>
<caption pageId="4" pageNumber="127">
<paragraph pageId="4" pageNumber="127">
Figures 4-8.
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="sandeli">Omphreus (Omphreus) morio sandeli</taxonomicName>
ssp. n. from eastern Bosnia and Herzegovina, southern slope of Mt. Zelengora,
<normalizedToken originalValue="Čemerno">Cemerno</normalizedToken>
, 1,450 m a.s.l., near Gacko. 4 holotype male, habitus (dorsal view) 5 holotype male, aedeagus (dorsal view) 6 holotype male, aedeagus (lateral view) 7 holotype male, abdominal sternite IX (urite) 8 paratype female, genitalia. Scale bars 5 mm (4) and 1 mm (5-8).
</paragraph>
</caption>
<paragraph pageId="4" pageNumber="127">Head rounded, somewhat elongated [HW/HL: M 0.96 (R 0.91-1.05)], narrower (HW/PW: M 0.69) than pronotum (Figure 4). Head below eye level somewhat constricted. Labrum broad, medially rounded, carrying four setae. Epistome large, concave anteriorly, with two setae. Both vertex and occiput wrinkled. Frontal foveae deepened and long. Gula bisetose. Mandibles elongated, sickle-shaped, broadened basally. Labial palpomere 1 short, without setae. The labial palpomeres 2 and 3 longer. Both labial and maxillar palpomere 3 broadened distally and densely pubescent. AL: M 10.41 mm (R 9.25-11.16 mm). Antennae pubescent from antennomere 4. AL/TL: M 0.56 (R 0.51-0.58). Antennomere 1 club-like, sharply widened distally, with a few long setae distally, somewhat shorter than the following three antennomeres combined. Antennomere 2 slightly shorter than antennomere 3.</paragraph>
<paragraph pageId="4" pageNumber="127">Pronotum sub-campaniform, elongate, PW/PL: M 0.91 (R 0.885-0.94). Fore angles somewhat prominent, rounded, hind angles obtuse, well rounded (Figure 4). Lateral margins well developed, thickened, arcuate anteriorly, then sinuate and narrowing posteriorly. Anterior pronotal margin somewhat concave, base strongly concave. Pronotum widest between its fore fourth and third. Lateral furrows narrow and shallow, with four anterior setae, one median seta, and one posterior seta each. Median furrow long and deep. Basal foveae deep and long, slightly shorter than one half of pronotum length. Pronotal disc somewhat convex proximally.</paragraph>
<paragraph pageId="4" pageNumber="127">Elytra ovoid, relatively wide, rounded laterally, EW/EL: M 0.58 (R 0.54-0.61) (Figure 4). EL/EW: M 1.73. EW: M 6.26 mm in males, while 6.36 mm in females. Elytral striae shallow, weakly punctate, points somewhat deeper basally. Elytral intervals flattened. Scutellum large, triangular. Scutellar stria present, but without scutellar puncture. Elytra widest around at the middle. A setiferous puncture in interval 7 basally (close to stria 6), 4-6 setiferous punctures in interval 7 medially (close to stria 7), and two setiferous punctures situated on stria 7 apically on each elytron. The punctures deep and large. Shoulders rounded. Umbilicate series regular, with the setae densely distributed. Elytral disc somewhat convex distally.</paragraph>
<paragraph pageId="4" pageNumber="127">Protarsomeres 1 and 2 widened in males. Metacoxae long and rounded. Tarsal claws elongate, glabrous, smooth (Figure 4).</paragraph>
<paragraph pageId="4" pageNumber="127">Aedeagus long, median lobe somewhat widened sub-apically in dorsal view (Figure 5), while curved and strongly widened sub-apically in lateral view (Figure 6), with a straight long acute triangular apex (Figures 5 and 6). Parameres wide, the right being much broader (Figure 5). Basal bulb wide and short (Figures 5 and 6).</paragraph>
<paragraph pageId="4" pageNumber="127">Male abdominal sternite IX (urite) large, sub-triangular (Figure 7).</paragraph>
<paragraph pageId="4" pageNumber="127">Both gonocoxites and gonosubcoxites IX as presented in Figure 8. Gonocoxites IX wide and elongated, somewhat curved, gradually narrowing distally, each with a rounded apex, basally joined with massive rounded gonosubcoxites IX.</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="127" type="variability">
<paragraph pageId="4" pageNumber="127">Variability.</paragraph>
<paragraph pageId="4" pageNumber="127">
It was noticed that the specimens from the population from the southern slope of Mt. Zelengora are of somewhat larger size and more elongate elytra (EW/EL: M 0.56) compared with the specimens belonging to the populations both from Mt.
<normalizedToken originalValue="Maglić">Maglic</normalizedToken>
and the northern slope of Mt. Zelengora (EW/EL: M 0.59). Other characteristics, including the structure of aedeagus, do not show any significant differences among the three analyzed populations of the taxon.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="127" type="differential diagnosis and remarks">
<paragraph pageId="4" pageNumber="127">Differential diagnosis and remarks.</paragraph>
<paragraph pageId="4" pageNumber="127">
The new subspecies is compared here with the morphologically nearest subspecies of
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="species" species="morio" subGenus="Omphreus">Omphreus (Omphreus) morio</taxonomicName>
Dejean, 1828. These are
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="beckianus">Omphreus (Omphreus) morio beckianus</taxonomicName>
Ganglbauer, 1888,
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="serbicus">Omphreus (Omphreus) morio serbicus</taxonomicName>
Winkler, 1933, and
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="durmitorensis">Omphreus (Omphreus) morio durmitorensis</taxonomicName>
ssp. n. (the first antennomere being club-like distally, somewhat shorter than the following three antennomeres combined in all the subspecies mentioned) (
<bibRefCitation author="Ganglbauer, L" journalOrPublisher="Societas entomologica" pageId="12" pageNumber="135" title="Omphreusmoriovar. nov. Beckianus." url="https://archive.org/details/societasentomolo02rh" volume="3" year="1888">Ganglbauer 1888</bibRefCitation>
,
<bibRefCitation author="Winkler, A" journalOrPublisher="Koleopterologische Rundschau" pageId="12" pageNumber="135" pagination="115 - 120" title="Revision der Omphreus-Arten (Carab.)." url="http://www.landesmuseum.at/pdf_frei_remote/KOR_19_1933_0115-0120.pdf" volume="19" year="1933">Winkler 1933</bibRefCitation>
).
</paragraph>
<paragraph pageId="4" pageNumber="127">
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="sandeli">Omphreus (Omphreus) morio sandeli</taxonomicName>
ssp. n. differs from
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="beckianus">Omphreus (Omphreus) morio beckianus</taxonomicName>
in the shape of the elytra (distinctly rounded laterally vs. slightly rounded laterally), the elytra length/width ratio (M 1.73 vs. around 1.66), and the body length (R 16.88-19.73 mm vs. 16-18 mm) (
<bibRefCitation author="Ganglbauer, L" journalOrPublisher="Societas entomologica" pageId="12" pageNumber="135" title="Omphreusmoriovar. nov. Beckianus." url="https://archive.org/details/societasentomolo02rh" volume="3" year="1888">Ganglbauer 1888</bibRefCitation>
,
<bibRefCitation author="Winkler, A" journalOrPublisher="Koleopterologische Rundschau" pageId="12" pageNumber="135" pagination="115 - 120" title="Revision der Omphreus-Arten (Carab.)." url="http://www.landesmuseum.at/pdf_frei_remote/KOR_19_1933_0115-0120.pdf" volume="19" year="1933">Winkler 1933</bibRefCitation>
). The former taxon is distributed both on Mt. Zelengora and Mt.
<normalizedToken originalValue="Maglić">Maglic</normalizedToken>
, while the latter one inhabits Mt.
<normalizedToken originalValue="Visočica">Visocica</normalizedToken>
(the type locality) (
<bibRefCitation author="Ganglbauer, L" journalOrPublisher="Societas entomologica" pageId="12" pageNumber="135" title="Omphreusmoriovar. nov. Beckianus." url="https://archive.org/details/societasentomolo02rh" volume="3" year="1888">Ganglbauer 1888</bibRefCitation>
). In the literature
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="beckianus">Omphreus (Omphreus) morio beckianus</taxonomicName>
is reported to have a wide distribution in Bosnia and Herzegovina, Montenegro and Serbia (from central and southern Bosnia to border with Montenegro, southwestern Serbia and northern Herzegovina, including the high-altitude area of the surroundings of Sarajevo, Mts. Ivan,
<normalizedToken originalValue="Bjelašnica">Bjelasnica</normalizedToken>
, Volujak and Durmitor and the Lim River Valley) (
<bibRefCitation author="Apfelbeck, V" journalOrPublisher="Glasnik Zemaljskog muzeja Bosne i Hercegovine" pageId="11" pageNumber="134" pagination="209 - 223" title="Rezultati koleopteroloskog ispitivanja iz alpijske regije juzno-bosanskih planina." url="http://www.bosniafacts.info/downloads/elibrary/viewcategory/15-glasnik-zemaljskog-muzeja-bosne-i-hercegovine-1889-2009" volume="2" year="1894">Apfelbeck 1894</bibRefCitation>
,
<bibRefCitation author="Apfelbeck, V" journalOrPublisher="R. Friedlaender und Sohn, Berlin" pageId="12" pageNumber="135" title="Die Kaeferfauna der Balkanhalbinsel, mit Beruecksichtigung Klein-Asiens und der Insel Kreta. Erster Band: Familienreihe Caraboidea" year="1904">1904</bibRefCitation>
,
<bibRefCitation author="Wohlberedt-Triebes, O" journalOrPublisher="naturhistorischen Hofmuseums" pageId="12" pageNumber="135" pagination="237 - 262" title="Zur Fauna des Sandschak Novipazar (Mollusken und Kaefer). Annalen des k. k." url="http://www.jstor.org/discover/10.2307/41768064?sid=21105995911413&amp;uid=3738928&amp;uid=2&amp;uid=67&amp;uid=62&amp;uid=19205&amp;uid=3&amp;uid=19204" volume="23" year="1909">Wohlberedt-Triebes 1909</bibRefCitation>
,
<bibRefCitation author="Drovenik, B" editor="Nonveiller, G" journalOrPublisher="The Montenegrin Academy of Sciences and Arts, Titograd" pageId="12" pageNumber="135" pagination="185 - 227" title="Cicindelidae in Carabidae (Insecta, Coleoptera)." volumeTitle="The Fauna of Durmitor" year="1984">Drovenik 1984</bibRefCitation>
). This appears to us very doubtful since
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="genus">Omphreus</taxonomicName>
taxa (species or subspecies) mostly inhabit single mountain ranges (
<bibRefCitation author="Winkler, A" journalOrPublisher="Koleopterologische Rundschau" pageId="12" pageNumber="135" pagination="115 - 120" title="Revision der Omphreus-Arten (Carab.)." url="http://www.landesmuseum.at/pdf_frei_remote/KOR_19_1933_0115-0120.pdf" volume="19" year="1933">Winkler 1933</bibRefCitation>
). The specimens from Mt. Durmitor belong to a new subspecies described herein -
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="durmitorensis">Omphreus (Omphreus) morio durmitorensis</taxonomicName>
ssp. n.; Mt. Volujak is relatively close to Mt. Zelengora and Mt.
<normalizedToken originalValue="Maglić">Maglic</normalizedToken>
and there is a possibility that the population from this mountain might actually belong to
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="sandeli">Omphreus (Omphreus) morio sandeli</taxonomicName>
ssp. n., but this has to be proven. The specimens from all known sites should be compared in the future (including the structure of the male genitalia) in order to conclude whether they belong to a single taxon (subspecies) or to different taxa. Additionally, a detailed study of all
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="species" species="morio" subGenus="Omphreus">Omphreus (Omphreus) morio</taxonomicName>
subspecies (with comparisons of numerous morphological characteristics) would be needed in order to define their real taxonomic status (subspecies or species).
</paragraph>
<paragraph pageId="4" pageNumber="127">
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="sandeli">Omphreus (Omphreus) morio sandeli</taxonomicName>
ssp. n. differs from
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="serbicus">Omphreus (Omphreus) morio serbicus</taxonomicName>
in the shape of the elytra (elongate vs. short), the elytra length/width ratio (M 1.73 vs. around 1.50), and the body length (R 16.88-19.73 mm vs. 15-16 mm) (
<bibRefCitation author="Winkler, A" journalOrPublisher="Koleopterologische Rundschau" pageId="12" pageNumber="135" pagination="115 - 120" title="Revision der Omphreus-Arten (Carab.)." url="http://www.landesmuseum.at/pdf_frei_remote/KOR_19_1933_0115-0120.pdf" volume="19" year="1933">Winkler 1933</bibRefCitation>
).
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="sandeli">Omphreus (Omphreus) morio sandeli</taxonomicName>
ssp. n. is found in eastern Bosnia and Herzegovina (Mt. Zelengora and Mt.
<normalizedToken originalValue="Maglić">Maglic</normalizedToken>
), while
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="serbicus">Omphreus (Omphreus) morio serbicus</taxonomicName>
lives in southwestern Serbia (Mt. Murtenica and Mt. Zlatibor) (
<bibRefCitation author="Winkler, A" journalOrPublisher="Koleopterologische Rundschau" pageId="12" pageNumber="135" pagination="115 - 120" title="Revision der Omphreus-Arten (Carab.)." url="http://www.landesmuseum.at/pdf_frei_remote/KOR_19_1933_0115-0120.pdf" volume="19" year="1933">Winkler 1933</bibRefCitation>
).
</paragraph>
<paragraph pageId="4" pageNumber="127">
Eventually,
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="sandeli">Omphreus (Omphreus) morio sandeli</taxonomicName>
ssp. n. differs from
<taxonomicName class="Insecta" family="Carabidae" genus="Omphreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphreus (Omphreus) morio" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="subSpecies" species="morio" subGenus="Omphreus" subSpecies="durmitorensis">Omphreus (Omphreus) morio durmitorensis</taxonomicName>
ssp. n. in the antennal length (M 10.41 mm vs. 10.38 mm), the shape of the hind pronotal angles (more rounded vs. less rounded), maximum width of the pronotum (between its fore fourth and third vs. in front of the fore third), the shape of the elytra (somewhat widened vs. more elongate), the elytral width/length ratio (M 0.58 vs. 0.56), the elytra length/width ratio (M 1.73 vs. 1.79), the elytral width (M 6.26 mm in males, 6.36 mm in females vs. 5.94 mm in males, 6.12 mm in females), maximum width of the elytra (around at the middle vs. slightly below the middle), the shape of the shoulders (rounded vs. obtusely rounded), the form of the median lobe (somewhat widened sub-apically in dorsal view, while curved and strongly widened sub-apically in lateral view, with a straight long acute triangular apex vs. strongly widened sub-apically in dorsal view, while arcuate, moderately widened and with a shallow concavity in the sub-apical part in lateral view, with a straight short rounded triangular apex), the shape of the basal bulb (wide and short vs. narrow and elongated), the shape of the male abdominal sternite IX (urite) (less elongate vs. more elongate), the form of the apex of the gonocoxites IX (rounded vs. pointed), and the total body length (M 18.49 mm vs. 18.13 mm).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="127" type="etymology">
<paragraph pageId="4" pageNumber="127">Etymology.</paragraph>
<paragraph pageId="4" pageNumber="127">This new subspecies is named after Franco Sandel, friend of the second author and excellent collector, who collected the whole type series of this new subspecies allowing us to freely study the material.</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="127" type="distribution">
<paragraph pageId="4" pageNumber="127">Distribution.</paragraph>
<paragraph pageId="4" pageNumber="127">
So far known only from the type locality (Mt. Zelengora) and the nearby Mt.
<normalizedToken originalValue="Maglić">Maglic</normalizedToken>
, eastern Bosnia and Herzegovina.
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="128" pageId="4" pageNumber="127" type="habitat">
<paragraph pageId="4" pageNumber="127">Habitat.</paragraph>
<paragraph lastPageId="5" lastPageNumber="128" pageId="4" pageNumber="127">
The subspecies prefers high-altitude habitats (1,450-1,600 m a.s.l.) on Mt. Zelengora (at both the northern and southern slopes) and Mt.
<normalizedToken originalValue="Maglić">Maglic</normalizedToken>
in eastern Bosnia and Herzegovina. Type series of the new subspecies was collected by pitfall
<pageBreakToken pageId="5" pageNumber="128" start="start">traps</pageBreakToken>
filled with alcoholic vinegar, placed at different sites in beech forests by the alpine meadows on both mountains, and on a pass and near the first hairpin bend after the pass on the southern slope of Mt. Zelengora.
</paragraph>
</subSubSection>
</treatment>
</document>