treatments-xml/data/53/30/87/533087D1CB5AFFA3E473F9A2EE24FCA9.xml

634 lines
91 KiB
XML
Raw Permalink Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="192FDDDE04F6E4D93FFD4C6BC88E8E45" ID-DOI="10.5852/ejt.2025.976.2773" ID-ISSN="2118-9773" ID-Zenodo-Dep="14622251" ID-ZooBank="33C36006-66C0-4B29-9D49-447AAEDB1398" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1736432095769" checkinUser="plazi" docAuthor="Romero-Rincon, Juan, Alvear, Santiago, Martínez-Torres, Daniela &amp; Robles-Piñeros, Jairo" docDate="2025" docId="533087D1CB5AFFA3E473F9A2EE24FCA9" docLanguage="en" docName="EJT.2025.976.33-46.pdf" docOrigin="European Journal of Taxonomy 976 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2773/12679" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Inodesmus mesibovi Romero-Rincon, Alvear, Martínez-Torres &amp; Robles-Piñeros, 2025, sp. nov." docType="treatment" docVersion="3" lastPageNumber="40" masterDocId="AF09FFA9CB58FFA4E61DFF9CEA45FFF2" masterDocTitle="First record of the millipede family Haplodesmidae (Diplopoda: Polydesmida) from Colombia: two new species of Inodesmus Cook, 1896 from the Northern Andes" masterLastPageNumber="46" masterPageNumber="33" pageNumber="35" updateTime="1737635427589" updateUser="valdenar" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="5C58BE13B92F79919C0198D5BFF42E07" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DC2F6C9CE23C3C7D7BCFA9AF0B4A51E0">
<mods:title id="015C2E95302C5D17818BF8EACBBF105F">First record of the millipede family Haplodesmidae (Diplopoda: Polydesmida) from Colombia: two new species of Inodesmus Cook, 1896 from the Northern Andes</mods:title>
</mods:titleInfo>
<mods:name id="31F7BC5E1A21C5E33CDF8BD9B4A0EAFE" type="personal">
<mods:role id="C1BBB5A2D2A5BA89B5AC83CA8055FE2C">
<mods:roleTerm id="D722813232485E122DCBE885C5311ED2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2ACCE8B5DB3B0C36C25FD14CB4E145DE">Romero-Rincon, Juan</mods:namePart>
<mods:nameIdentifier id="1EB91338CAB715FD714D5808277F63B2" type="LSID">4AAEDF28-ABEB-44E7-9549-C3FE2D377383</mods:nameIdentifier>
<mods:affiliation id="0B32FD2D87607F202BF2B1CFB182568E">Museo de Historia Natural, Universidad Pedagógica Nacional, Bogotá, Colombia.</mods:affiliation>
<mods:nameIdentifier id="C7672F621E04CEF40C265EF56A084854" type="email">romerorjuanc@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="641B41EB2850E4CE7514A4387555293C" type="personal">
<mods:role id="C6F3AB2C2B69051DE59365DB5D6E5654">
<mods:roleTerm id="5534765CB67C01B585E90CA6C11ABBE5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="24FEC041CBB97506073F542FE7D8ED16">Alvear, Santiago</mods:namePart>
<mods:nameIdentifier id="3C6943FE484D85343F1FC0DB577ABB6C" type="LSID">B0D52AEC-D281-445A-B3A6-4962BF637A8D</mods:nameIdentifier>
<mods:affiliation id="3DFDD51F323F894E05C003739E6CD1F7">Semillero de Investigación en Diversidad Funcional y Servicios Ecosistémicos - Grupo de Estudios Ambientales, Universidad del Cauca, Popayán, Colombia.</mods:affiliation>
<mods:nameIdentifier id="826A5F5A0D5A3822420CBF5623F7B955" type="email">alexs.alvear@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="0786E7A09A7ABD5CBA92FF87F5EB72D4" type="personal">
<mods:role id="CCEF156EAEEE2EF636D0262939838A59">
<mods:roleTerm id="2ED4FE8723BF622EB89949361B7C17E5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8FC54A8837080C7A1972BE76D9ACAAC9">Martínez-Torres, Daniela</mods:namePart>
<mods:nameIdentifier id="2A9446F21BC5723A24290FF8C68C81EF" type="LSID">3D6A9085-FE64-4647-9A4E-EC1599DD26D4</mods:nameIdentifier>
<mods:affiliation id="35992FA1DEACA219570EB1AEF580C5C7">Grupo de Investigación en Aracnología &amp; Miriapodología (GAM- UN), Universidad Nacional de Colombia, Bogotá, Colombia.</mods:affiliation>
<mods:nameIdentifier id="98F4C2D039096163D78A97DC94E1B1BF" type="email">sdmartinezt@unal.edu.co</mods:nameIdentifier>
</mods:name>
<mods:name id="598D331E64A37ABDC3FC3D78D905BC50" type="personal">
<mods:role id="82FCAD3886251606FD0EC0B5C1C8D609">
<mods:roleTerm id="CA90F20FCD9D1D49B4037B55F25EC923">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5E23F9C9FEFD78158799E37449669C8E">Robles-Piñeros, Jairo</mods:namePart>
<mods:nameIdentifier id="51742EDCAEB51398BD8990F2E58E9E72" type="LSID">20C57158-BF00-4F31-A02C-4F7831E4D66C</mods:nameIdentifier>
<mods:affiliation id="E52B93EB20D7434B32BFFAB0C935639E">Departamento de Biología, Universidad Pedagógica Nacional, Bogotá, Colombia.</mods:affiliation>
<mods:nameIdentifier id="72D012696358031FF0CFDE325DE69A69" type="email">jroblesp@upn.edu.co</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="21474311EE5A096EF3564B72AE9B08E7">text</mods:typeOfResource>
<mods:relatedItem id="A5A53CB0CF98641B8C29FDEA4E116F8E" type="host">
<mods:titleInfo id="3C7658E6286791AD019B032284AF529E">
<mods:title id="B7BC69C68FD6A453763EAE4E8FC4DDF1">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="F6378E79BCE3020CE5CF7FB972192FFC">
<mods:date id="58DAB061B6DCC274074DF61C14195A31">2025</mods:date>
<mods:detail id="D36CB54FAC577FE6305B3719381C4E08" type="pubDate">
<mods:number id="05AA9EC87B54AEF8EA731B6D1585BF54">2025-01-06</mods:number>
</mods:detail>
<mods:detail id="628E2176E7803482C68072A343C3BDBE" type="volume">
<mods:number id="CB6FBB3A75A5FFAA1FF84148FC766875">976</mods:number>
</mods:detail>
<mods:detail id="2D25F182E5BCE2BDEC7E53FDA22FA882" type="issue">
<mods:number id="9D7B2CCC4F35C2B7699FC0DDE79A4F41">1</mods:number>
</mods:detail>
<mods:extent id="2F7B9986F55E7ACB7649D80CE90C4FEF" unit="page">
<mods:start id="356D91B3040D7E57A9F6937B08BD0C53">33</mods:start>
<mods:end id="EF9C55FA86E55904C7BDCD216428D5F3">46</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="B72A480F17E4AA08FFB3623B475BD656">
<mods:url id="8C946B9CF4151CCC8D5AF15C19328A43">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2773/12679</mods:url>
</mods:location>
<mods:classification id="91A4AE1B5F892E60A6091057D9C9DD06">journal article</mods:classification>
<mods:identifier id="5EF8593E83F9448D567FAE457297BA21" type="DOI">10.5852/ejt.2025.976.2773</mods:identifier>
<mods:identifier id="555E1EDD3FB7B86DE30FD540C6099CFD" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="03A9608CACC5FBC67A2B2692A2873F76" type="Zenodo-Dep">14622251</mods:identifier>
<mods:identifier id="79B8B0428248EDB08D1D3B5F7B3F095D" type="ZooBank">33C36006-66C0-4B29-9D49-447AAEDB1398</mods:identifier>
</mods:mods>
<treatment id="533087D1CB5AFFA3E473F9A2EE24FCA9" LSID="urn:lsid:plazi:treatment:533087D1CB5AFFA3E473F9A2EE24FCA9" httpUri="http://treatment.plazi.org/id/533087D1CB5AFFA3E473F9A2EE24FCA9" lastPageId="7" lastPageNumber="40" pageId="2" pageNumber="35">
<subSubSection id="9383654CCB5AFFA6E473F9A2E983F9AB" box="[622,966,1598,1625]" pageId="2" pageNumber="35" type="nomenclature">
<paragraph id="DB2636C7CB5AFFA6E473F9A2E983F9AB" blockId="2.[375,1213,1598,1695]" box="[622,966,1598,1625]" pageId="2" pageNumber="35">
<heading id="806E81ABCB5AFFA6E473F9A2E983F9AB" box="[622,966,1598,1625]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="35" reason="2">
<taxonomicName id="1C994D44CB5AFFA6E473F9A2E921F9AB" authorityName="Romero-Rincon &amp; Alvear &amp; Martínez-Torres &amp; Robles-Piñeros" authorityYear="2025" box="[622,868,1598,1625]" class="Diplopoda" family="Comodesmidae" genus="Inodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="35" phylum="Arthropoda" rank="species" species="mesibovi" status="sp. nov.">
<emphasis id="E9EDEAD5CB5AFFA6E473F9A2E921F9AB" bold="true" box="[622,868,1598,1625]" italics="true" pageId="2" pageNumber="35">Inodesmus mesibovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2DE57AECB5AFFA6E576F9A3E983F9AB" box="[875,966,1599,1625]" pageId="2" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9383654CCB5AFFA6E76AF9FEE919F96D" pageId="2" pageNumber="35" type="description">
<paragraph id="DB2636C7CB5AFFA6E76AF9FEEEF8F98F" blockId="2.[375,1213,1598,1695]" box="[375,1213,1634,1661]" pageId="2" pageNumber="35">
<uri id="AF083AC5CB5AFFA6E76AF9FEEEF8F98F" box="[375,1213,1634,1661]" pageId="2" pageNumber="35">
urn:lsid:zoobank.org:act:
<uuid id="AF3F0C12CB5AFFA6E488F9FEEEF8F98F" box="[661,1213,1634,1661]" pageId="2" pageNumber="35">945A115E-0BE9-484A-9581-FF7E09C59803</uuid>
</uri>
</paragraph>
<paragraph id="DB2636C7CB5AFFA6E4C5F919E919F96D" blockId="2.[375,1213,1598,1695]" box="[728,860,1669,1695]" pageId="2" pageNumber="35">
<figureCitation id="43A22A42CB5AFFA6E4C5F919E905F96D" box="[728,832,1669,1695]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="3.[189,232,1961,1987]" captionStartId-1="4.[189,232,1961,1987]" captionStartId-2="5.[189,232,1785,1811]" captionStartId-3="6.[189,232,1925,1951]" captionTargetBox-0="[227,1202,978,1932]" captionTargetBox-1="[189,1366,1332,1937]" captionTargetBox-2="[188,1397,532,1759]" captionTargetBox-3="[231,1366,973,1898]" captionTargetId-0="figure-222@3.[189,1399,966,1947]" captionTargetId-1="figure-413@4.[189,1399,1332,1937]" captionTargetId-2="figure-243@5.[172,1414,518,1775]" captionTargetId-3="figure-343@6.[189,1399,958,1912]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="Fig. 1. Habitus of Inodesmus mesibovi sp. nov., male paratype (above; MHNUC-MD-276) and female paratype (below; MHNUC-MD-276). Scale bar: 1 mm." captionText-1="Fig. 2. Inodesmus mesibovi sp. nov., male holotype (ICN-MD-2913). A. Ventral view of the gonopods. B. Left gonopod in ventral view. Scale bars: 0.1 mm." captionText-2="Fig. 3. Inodesmus mesibovi sp. nov., male holotype (A, CD, HI; ICN-MD-2913) and male paratype (B, EG; MHN-UPN-MD-168). A. Ventrolateral view of head. B. Dorsolateral view of the anterior part of the body. C. Lateral view of the anterior part of the body. D. Midbody sternite. E. Sculpture of the prozonite. F. Ozopore. G. Lateral view of the posterior part of the body. H. Ventral view of telson. I. Spinnerets. Abbreviations: a = anterior part of prozonite; b = posterior part of prozonite; L = limbus; lo = lobe; o = ozopore; r = ridge; s = spherical knobs; t = tooth-like lobe. Scale bars: A, H = 0.2 mm; B, G = 0.5 mm; C = 1 mm; D = 0.1 mm; EF = 0.02 mm; I = 0.05 mm." captionText-3="Fig. 4. Teratologies inInodesmus mesibovi sp. nov., female paratype (MHN-UPN-MD-177). A. Anterior part of body. B. Middle part of the body. The numbers 6 and 9 correspond to the body ring number. Scale bar: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14622253" figureDoi-1="http://doi.org/10.5281/zenodo.14622257" figureDoi-2="http://doi.org/10.5281/zenodo.14622259" figureDoi-3="http://doi.org/10.5281/zenodo.14622263" httpUri-0="https://zenodo.org/record/14622253/files/figure.png" httpUri-1="https://zenodo.org/record/14622257/files/figure.png" httpUri-2="https://zenodo.org/record/14622259/files/figure.png" httpUri-3="https://zenodo.org/record/14622263/files/figure.png" pageId="2" pageNumber="35">Figs 14</figureCitation>
,
<figureCitation id="43A22A42CB5AFFA6E550F919E919F96D" box="[845,860,1669,1695]" captionStart="Fig" captionStartId="11.[189,232,1151,1177]" captionTargetBox="[182,1410,282,1138]" captionTargetId="figure-243@11.[174,1414,265,1142]" captionTargetPageId="11" captionText="Fig. 8. Distribution map of the known species of Inodesmus Cook, 1896 in Colombia." figureDoi="http://doi.org/10.5281/zenodo.14622271" httpUri="https://zenodo.org/record/14622271/files/figure.png" pageId="2" pageNumber="35">8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="9383654CCB5AFFA6E6A0F952EE12F835" pageId="2" pageNumber="35" type="diagnosis">
<paragraph id="DB2636C7CB5AFFA6E6A0F952EB73F91A" blockId="2.[189,310,1742,1768]" box="[189,310,1742,1768]" pageId="2" pageNumber="35">
<heading id="806E81ABCB5AFFA6E6A0F952EB73F91A" bold="true" box="[189,310,1742,1768]" fontSize="11" level="3" pageId="2" pageNumber="35" reason="3">
<emphasis id="E9EDEAD5CB5AFFA6E6A0F952EB73F91A" bold="true" box="[189,310,1742,1768]" pageId="2" pageNumber="35">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DB2636C7CB5AFFA6E6A0F961EE12F835" blockId="2.[189,1399,1789,1991]" pageId="2" pageNumber="35">
Males with head + 19 rings, females with head + 20 rings. This species differs from its congeners in having a coxal projection on male leg 6. Telopodite (
<figureCitation id="43A22A42CB5AFFA6E523F8BCE9C2F8C8" box="[830,903,1824,1850]" captionStart="Fig" captionStartId="4.[189,232,1961,1987]" captionTargetBox="[189,1366,1332,1937]" captionTargetId="figure-413@4.[189,1399,1332,1937]" captionTargetPageId="4" captionText="Fig. 2. Inodesmus mesibovi sp. nov., male holotype (ICN-MD-2913). A. Ventral view of the gonopods. B. Left gonopod in ventral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14622257" httpUri="https://zenodo.org/record/14622257/files/figure.png" pageId="2" pageNumber="35">Fig. 2</figureCitation>
) with
<emphasis id="E9EDEAD5CB5AFFA6E5C5F8BCE9BCF8C8" bold="true" box="[984,1017,1824,1850]" pageId="2" pageNumber="35">pp</emphasis>
straight;
<emphasis id="E9EDEAD5CB5AFFA6E271F8BCEEC1F8C8" bold="true" box="[1132,1156,1824,1850]" pageId="2" pageNumber="35">at</emphasis>
in transverse plane, large, somewhat fusiform, and slightly bent posteriorly;
<emphasis id="E9EDEAD5CB5AFFA6E541F8DFE938F8AF" bold="true" box="[860,893,1859,1885]" pageId="2" pageNumber="35">dp</emphasis>
directed laterobasally, basal portion of the
<emphasis id="E9EDEAD5CB5AFFA6E6A3F8FAEA9AF872" bold="true" box="[190,223,1894,1920]" pageId="2" pageNumber="35">dp</emphasis>
linguiform;
<emphasis id="E9EDEAD5CB5AFFA6E766F8FAEBF4F872" bold="true" box="[379,433,1894,1920]" pageId="2" pageNumber="35">meb</emphasis>
curving strongly behind
<emphasis id="E9EDEAD5CB5AFFA6E4F4F8FAE965F872" bold="true" box="[745,800,1894,1920]" pageId="2" pageNumber="35">mab</emphasis>
, bifurcated basally, with a long, acuminate axial process, and a hook-shaped process, with another, small process at midlength;
<emphasis id="E9EDEAD5CB5AFFA6E27EF816EEDFF856" bold="true" box="[1123,1178,1930,1956]" pageId="2" pageNumber="35">mab</emphasis>
concave medially, broad distally, curved outward, and with a small subtriangular process distally.
</paragraph>
</subSubSection>
<subSubSection id="9383654CCB5BFFA7E6A0FE95EE15FE86" pageId="3" pageNumber="36" type="etymology">
<paragraph id="DB2636C7CB5BFFA7E6A0FE95EB01FED1" blockId="3.[189,324,265,291]" box="[189,324,265,291]" pageId="3" pageNumber="36">
<heading id="806E81ABCB5BFFA7E6A0FE95EB01FED1" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="3" pageNumber="36" reason="3">
<emphasis id="E9EDEAD5CB5BFFA7E6A0FE95EB01FED1" bold="true" box="[189,324,265,291]" pageId="3" pageNumber="36">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="DB2636C7CB5BFFA7E6A0FEABEE15FE86" blockId="3.[189,1398,311,373]" pageId="3" pageNumber="36">
The specific epithet,
<emphasis id="E9EDEAD5CB5BFFA7E7A8FEABE861FEA3" box="[437,548,311,337]" italics="true" pageId="3" pageNumber="36">mesobovi</emphasis>
(noun, genetive case), is a patronym, honoring Dr. Robert Mesibov for his valuable contributions to the knowledge of the Australian millipede fauna.
</paragraph>
</subSubSection>
<subSubSection id="9383654CCB5BFFA0E6A0FE07EB5EFCE3" lastPageId="4" lastPageNumber="37" pageId="3" pageNumber="36" type="materials_examined">
<paragraph id="DB2636C7CB5BFFA7E6A0FE07EBEFFE47" blockId="3.[189,426,411,437]" box="[189,426,411,437]" pageId="3" pageNumber="36">
<heading id="806E81ABCB5BFFA7E6A0FE07EBEFFE47" bold="true" box="[189,426,411,437]" fontSize="11" level="3" pageId="3" pageNumber="36" reason="3">
<emphasis id="E9EDEAD5CB5BFFA7E6A0FE07EBEFFE47" bold="true" box="[189,426,411,437]" pageId="3" pageNumber="36">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DB2636C7CB5BFFA7E6F7FE56E9A8FDBC" blockId="3.[189,1398,458,590]" pageId="3" pageNumber="36">
<materialsCitation id="6BF13C9ACB5BFFA7E6F7FE56E9A8FDBC" collectedFrom="under a decaying fallen log" collectingDate="2023-01-12" collectingMethod="daytime hand collection" collectorName="J. Romero-Rincon &amp; J. Veloza" country="Colombia" elevation="2253" latitude="4.617133" location="Parque Natural Chicaque" longLatPrecision="1" longitude="-74.313835" municipality="San Antonio del Tequendama" pageId="3" pageNumber="36" specimenCode="ICN-MD-2913" specimenCount="1" specimenCount-male="1" stateProvince="Cundinamarca" typeStatus="holotype">
<emphasis id="E9EDEAD5CB5BFFA7E6F7FE56EB19FE16" bold="true" box="[234,348,458,484]" pageId="3" pageNumber="36">
<typeStatus id="04228865CB5BFFA7E6F7FE56EB19FE16" box="[234,348,458,484]" pageId="3" pageNumber="36" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="A38E7657CB5BFFA7E6A0FE72EB1DFDFA" box="[189,344,494,520]" name="Colombia" pageId="3" pageNumber="36">COLOMBIA</collectingCountry>
<specimenCount id="CD9FFD4ECB5BFFA7E76EFE72EBC9FDFA" box="[371,396,494,520]" count="1" pageId="3" pageNumber="36" type="male"></specimenCount>
;
<collectingRegion id="195DF825CB5BFFA7E784FE71E802FDF5" box="[409,583,493,519]" country="Colombia" name="Cundinamarca" pageId="3" pageNumber="36">Cundinamarca</collectingRegion>
,
<collectingMunicipality id="3B42ACBDCB5BFFA7E449FE72E9F1FDF5" box="[596,948,493,520]" pageId="3" pageNumber="36">San Antonio del Tequendama</collectingMunicipality>
,
<location id="DE46601CCB5BFFA7E5DDFE72EEA9FDF5" LSID="urn:lsid:plazi:treatment:533087D1CB5AFFA3E473F9A2EE24FCA9:DE46601CCB5BFFA7E5DDFE72EEA9FDF5" box="[960,1260,493,520]" country="Colombia" latitude="4.617133" longLatPrecision="1" longitude="-74.313835" municipality="San Antonio del Tequendama" name="Parque Natural Chicaque" pageId="3" pageNumber="36" stateProvince="Cundinamarca">Parque Natural Chicaque</location>
,
<location id="DE46601CCB5BFFA7E2E4FE71EF37FDF5" LSID="urn:lsid:plazi:treatment:533087D1CB5AFFA3E473F9A2EE24FCA9:DE46601CCB5BFFA7E2E4FE71EF37FDF5" box="[1273,1394,493,519]" country="Colombia" latitude="4.617133" longLatPrecision="1" longitude="-74.313835" municipality="San Antonio del Tequendama" name="oak forest" pageId="3" pageNumber="36" stateProvince="Cundinamarca">oak forest</location>
; [
<geoCoordinate id="BEAD5000CB5BFFA7E6D9FD93EB21FDD9" box="[196,356,527,555]" degrees="04" direction="north" minutes="37.028" orientation="latitude" pageId="3" pageNumber="36" precision="1" value="4.617133">04º37.028 N</geoCoordinate>
,
<geoCoordinate id="BEAD5000CB5BFFA7E76DFD93E85DFDD9" box="[368,536,527,555]" degrees="74" direction="west" minutes="18.830" orientation="longitude" pageId="3" pageNumber="36" precision="1" value="-74.313835">74°18.830 W</geoCoordinate>
];
<quantity id="1C619B22CB5BFFA7E431FD8CE88AFDD9" box="[556,719,528,555]" metricMagnitude="3" metricUnit="m" metricValue="2.2535" metricValueMax="2.275" metricValueMin="2.232" pageId="3" pageNumber="36" unit="m" value="2253.5" valueMax="2275.0" valueMin="2232.0">22322275 m</quantity>
a.s.l.;
<date id="AF271007CB5BFFA7E501FD8DE9F0FDD9" box="[796,949,529,555]" pageId="3" pageNumber="36" value="2023-01-12">
<collectingDate id="BF63E9EFCB5BFFA7E501FD8DE9F0FDD9" box="[796,949,529,555]" pageId="3" pageNumber="36" value="2023-01-12">12 Jan. 2023</collectingDate>
</date>
;
<collectorName id="766C5311CB5BFFA7E5DCFD8DEEDCFDD9" box="[961,1177,529,555]" pageId="3" pageNumber="36">J. Romero-Rincon</collectorName>
and
<collectorName id="766C5311CB5BFFA7E2CFFD8DEF79FDD9" box="[1234,1340,529,555]" pageId="3" pageNumber="36">J. Veloza</collectorName>
leg.;
<collectedFrom id="6885753ACB5BFFA7E6A0FDA8E845FDBC" box="[189,512,564,590]" pageId="3" pageNumber="36">under a decaying fallen log</collectedFrom>
;
<collectingMethod id="02D84ED0CB5BFFA7E411FDA8E96FFDBC" box="[524,810,564,590]" pageId="3" pageNumber="36">daytime hand collection</collectingMethod>
;
<specimenCode id="8B3F9EBCCB5BFFA7E528FDA8E9ACFDBC" box="[821,1001,564,590]" pageId="3" pageNumber="36">ICN-MD-2913</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="DB2636C7CB5BFFA0E6F7FDE9EB5EFCE3" blockId="3.[189,1398,629,902]" lastBlockId="4.[189,1401,265,785]" lastPageId="4" lastPageNumber="37" pageId="3" pageNumber="36">
<materialsCitation id="6BF13C9ACB5BFFA7E6F7FDE9EF5CFD40" collectedFrom="under a decaying fallen log" collectingDate="2023-01-12" collectingMethod="daytime hand collection" collectorName="J. Romero-Rincon &amp; J. Veloza" country="Colombia" elevation="2253" latitude="4.617133" location="COLOMBIA" longLatPrecision="1" longitude="-74.313835" municipality="San Antonio del Tequendama" pageId="3" pageNumber="36" specimenCode="MHN-UPN-MD-168" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Cundinamarca" typeStatus="paratype">
<emphasis id="E9EDEAD5CB5BFFA7E6F7FDE9EB22FD7D" bold="true" box="[234,359,629,655]" pageId="3" pageNumber="36">
<typeStatus id="04228865CB5BFFA7E6F7FDE9EB22FD7D" box="[234,359,629,655]" pageId="3" pageNumber="36" type="paratype">Paratypes</typeStatus>
</emphasis>
(11 ♂♂, 15 ♀♀, 17 immatures)
<collectingCountry id="A38E7657CB5BFFA7E6A0FD05EB1DFD41" box="[189,344,665,691]" name="Colombia" pageId="3" pageNumber="36">COLOMBIA</collectingCountry>
<specimenCount id="CD9FFD4ECB5BFFA7E79CFD05EBF0FD41" box="[385,437,665,691]" count="1" pageId="3" pageNumber="36" type="male">1 ♂</specimenCount>
(fragmented),
<specimenCount id="CD9FFD4ECB5BFFA7E469FD05E8ECFD41" box="[628,681,665,691]" count="1" pageId="3" pageNumber="36" type="female">1 ♀</specimenCount>
; same data as for holotype;
<specimenCode id="8B3F9EBCCB5BFFA7E23FFD04EF5CFD40" box="[1058,1305,664,690]" pageId="3" pageNumber="36">MHN-UPN-MD-168</specimenCode>
</materialsCitation>
<materialsCitation id="6BF13C9ACB5BFFA7E35FFD05E9E6FD27" collectedFrom="under a decaying fallen log" collectingDate="2023-01-12" collectingMethod="daytime hand collection" collectorName="J. Romero-Rincon &amp; J. Veloza" country="Colombia" elevation="2253" latitude="4.617133" location="Parque Natural Chicaque" longLatPrecision="1" longitude="-74.313835" municipality="San Antonio del Tequendama" pageId="3" pageNumber="36" specimenCode="MHN-UPN-MD-176" specimenCount="1" specimenCount-male="1" stateProvince="Cundinamarca">
<specimenCount id="CD9FFD4ECB5BFFA7E35FFD05EF33FD41" box="[1346,1398,665,691]" count="1" pageId="3" pageNumber="36" type="male">1 ♂</specimenCount>
(fragmented); same data as for holotype;
<specimenCode id="8B3F9EBCCB5BFFA7E4B6FD27E9E6FD27" box="[683,931,699,725]" pageId="3" pageNumber="36">MHN-UPN-MD-176</specimenCode>
</materialsCitation>
<materialsCitation id="6BF13C9ACB5BFFA7E5A2FD20EBDCFD0B" collectedFrom="under a decaying fallen log" collectingDate="2023-01-12" collectingMethod="daytime hand collection" collectorName="J. Romero-Rincon &amp; J. Veloza" country="Colombia" elevation="2253" latitude="4.617133" location="Parque Natural Chicaque" longLatPrecision="1" longitude="-74.313835" municipality="San Antonio del Tequendama" pageId="3" pageNumber="36" specimenCode="MHNUC-MD-276" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Cundinamarca">
<specimenCount id="CD9FFD4ECB5BFFA7E5A2FD20E9AAFD24" box="[959,1007,700,726]" count="1" pageId="3" pageNumber="36" type="male">1 ♂</specimenCount>
,
<specimenCount id="CD9FFD4ECB5BFFA7E5E0FD20EE69FD24" box="[1021,1068,700,726]" count="1" pageId="3" pageNumber="36" type="female">1 ♀</specimenCount>
; same data as for holotype;
<specimenCode id="8B3F9EBCCB5BFFA7E6A0FD43EBDCFD0B" box="[189,409,735,761]" pageId="3" pageNumber="36">MHNUC-MD-276</specimenCode>
</materialsCitation>
<materialsCitation id="6BF13C9ACB5BFFA7E7A5FD43EF53FD0B" box="[440,1302,735,761]" collectedFrom="under a decaying fallen log" collectingDate="2023-01-12" collectingMethod="daytime hand collection" collectorName="J. Romero-Rincon &amp; J. Veloza" country="Colombia" elevation="2253" latitude="4.617133" location="Parque Natural Chicaque" longLatPrecision="1" longitude="-74.313835" municipality="San Antonio del Tequendama" pageId="3" pageNumber="36" specimenCode="MHN-UPN-MD-177" specimenCount="1" specimenCount-female="1" stateProvince="Cundinamarca">
<specimenCount id="CD9FFD4ECB5BFFA7E7A5FD43EBADFD0B" box="[440,488,735,761]" count="1" pageId="3" pageNumber="36" type="female">1 ♀</specimenCount>
; same data as for holotype; with teratologies;
<specimenCode id="8B3F9EBCCB5BFFA7E202FD43EF53FD0B" box="[1055,1302,735,761]" pageId="3" pageNumber="36">MHN-UPN-MD-177</specimenCode>
</materialsCitation>
<materialsCitation id="6BF13C9ACB5BFFA7E35DFD43E9AAFCEE" collectedFrom="under a decaying fallen log" collectingDate="2023-01-12" collectingMethod="daytime hand collection" collectorName="J. Romero-Rincon &amp; J. Veloza" country="Colombia" elevation="2253" latitude="4.617133" location="Parque Natural Chicaque" longLatPrecision="1" longitude="-74.313835" municipality="San Antonio del Tequendama" pageId="3" pageNumber="36" specimenCode="MHN-UPN-MD-178" specimenCount="11" specimenCount-female="3" specimenCount-immature="7" specimenCount-male="1" stateProvince="Cundinamarca">
<specimenCount id="CD9FFD4ECB5BFFA7E35DFD43EF34FD0B" box="[1344,1393,735,761]" count="1" pageId="3" pageNumber="36" type="male">1 ♂</specimenCount>
,
<specimenCount id="CD9FFD4ECB5BFFA7E6A0FC9EEB47FCEF" box="[189,258,770,797]" count="3" pageId="3" pageNumber="36" type="female">3 ♀♀</specimenCount>
,
<specimenCount id="CD9FFD4ECB5BFFA7E70CFC9FEBEDFCEE" box="[273,424,770,797]" count="7" pageId="3" pageNumber="36" type="immature">7 immatures</specimenCount>
; same data as for holotype;
<specimenCode id="8B3F9EBCCB5BFFA7E4E5FC9EE9AAFCEE" box="[760,1007,770,796]" pageId="3" pageNumber="36">MHN-UPN-MD-178</specimenCode>
</materialsCitation>
<materialsCitation id="6BF13C9ACB5BFFA7E211FC9EEBA6FCCD" collectedFrom="under a decaying fallen log" collectingDate="2023-01-12" collectingMethod="daytime hand collection" collectorName="J. Romero-Rincon &amp; J. Veloza" country="Colombia" elevation="2253" latitude="4.617133" location="Parque Natural Chicaque" longLatPrecision="1" longitude="-74.313835" municipality="San Antonio del Tequendama" pageId="3" pageNumber="36" specimenCode="ICN-MD-3130" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Cundinamarca">
<specimenCount id="CD9FFD4ECB5BFFA7E211FC9EEE14FCEF" box="[1036,1105,770,797]" count="2" pageId="3" pageNumber="36" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CD9FFD4ECB5BFFA7E242FC9EEEE1FCEF" box="[1119,1188,770,797]" count="2" pageId="3" pageNumber="36" type="female">2 ♀♀</specimenCount>
; same data as for holotype;
<specimenCode id="8B3F9EBCCB5BFFA7E72EFCB9EBA6FCCD" box="[307,483,805,831]" pageId="3" pageNumber="36">ICN-MD-3130</specimenCode>
</materialsCitation>
<materialsCitation id="6BF13C9ACB5BFFA7E7E1FCB9EECAFC91" collectedFrom="under decaying log" collectingDate="2022-09-26" collectingDateMax="2022-09-29" collectingDateMin="2022-09-26" collectingMethod="daytime hand collection" collectorName="J. Romero-Rincon &amp; L. Poveda" country="Colombia" elevation="2253" latitude="4.617133" location="Parque Natural Chicaque" longLatPrecision="1" longitude="-74.313835" municipality="San Antonio del Tequendama" pageId="3" pageNumber="36" specimenCode="MHN-UPN-MD-92" specimenCount="1" specimenCount-male="1" stateProvince="Cundinamarca">
<specimenCount id="CD9FFD4ECB5BFFA7E7E1FCB9E86EFCB2" box="[508,555,805,832]" count="1" pageId="3" pageNumber="36" type="male">1 ♂</specimenCount>
; same locality as for holotype;
<date id="AF271007CB5BFFA7E584FCB9EE27FCCD" box="[921,1122,805,831]" pageId="3" pageNumber="36" value="2022-09-26" valueMax="2022-09-29" valueMin="2022-09-26">
<collectingDate id="BF63E9EFCB5BFFA7E584FCB9EE27FCCD" box="[921,1122,805,831]" pageId="3" pageNumber="36" value="2022-09-26" valueMax="2022-09-29" valueMin="2022-09-26">2629 Sep. 2022</collectingDate>
</date>
;
<collectorName id="766C5311CB5BFFA7E273FCBAEF00FCCD" box="[1134,1349,805,832]" pageId="3" pageNumber="36">J. Romero-Rincon</collectorName>
and
<collectorName id="766C5311CB5BFFA7E6A0FCD5EB73FC90" box="[189,310,840,867]" pageId="3" pageNumber="36">L. Poveda</collectorName>
leg.;
<collectedFrom id="6885753ACB5BFFA7E763FCD4E822FC90" box="[382,615,840,866]" pageId="3" pageNumber="36">under decaying log</collectedFrom>
;
<collectingMethod id="02D84ED0CB5BFFA7E46BFCD4E9DCFC90" box="[630,921,840,866]" pageId="3" pageNumber="36">daytime hand collection</collectingMethod>
;
<specimenCode id="8B3F9EBCCB5BFFA7E5BAFCD5EECAFC91" box="[935,1167,841,867]" pageId="3" pageNumber="36">MHN-UPN-MD-92</specimenCode>
</materialsCitation>
<materialsCitation id="6BF13C9ACB5BFFA7E2B3FCD5E931FC74" collectedFrom="under decaying log" collectingDate="2022-09-26" collectingDateMax="2022-09-29" collectingDateMin="2022-09-26" collectingMethod="daytime hand collection" collectorName="J. Romero-Rincon &amp; L. Poveda" country="Colombia" elevation="2253" latitude="4.617133" location="Parque Natural Chicaque" longLatPrecision="1" longitude="-74.313835" municipality="San Antonio del Tequendama" pageId="3" pageNumber="36" specimenCode="MHN-UPN-94" specimenCount="2" specimenCount-female="1" specimenCount-immature="1" stateProvince="Cundinamarca">
<specimenCount id="CD9FFD4ECB5BFFA7E2B3FCD5EE99FC91" box="[1198,1244,841,867]" count="1" pageId="3" pageNumber="36" type="female">1 ♀</specimenCount>
,
<specimenCount id="CD9FFD4ECB5BFFA7E2F0FCD5EF33FC90" box="[1261,1398,840,867]" count="1" pageId="3" pageNumber="36" type="immature">1 immature</specimenCount>
(fragmented); same data as for preceding;
<specimenCode id="8B3F9EBCCB5BFFA7E4D9FCF0E931FC74" box="[708,884,876,902]" pageId="3" pageNumber="36">MHN-UPN-94</specimenCode>
</materialsCitation>
<materialsCitation id="6BF13C9ACB5BFFA0E589FCF0E875FEB4" collectedFrom="inside a decaying fallen log" collectingDate="2022-05-29" collectingDateMax="2022-09-29" collectingDateMin="2022-09-26" collectingMethod="daytime hand collection" collectionCode="R" collectorName="J. Romero-Rincon &amp; R. De La" country="Colombia" elevation="2253" lastPageId="4" lastPageNumber="37" latitude="4.617133" location="Parque Natural Chicaque" longLatPrecision="1" longitude="-74.313835" municipality="San Antonio del Tequendama" pageId="3" pageNumber="36" specimenCode="MHN-UPN-MD-96" specimenCount="1" specimenCount-female="1" stateProvince="Cundinamarca">
<specimenCount id="CD9FFD4ECB5BFFA7E589FCF0E980FC74" box="[916,965,876,902]" count="1" pageId="3" pageNumber="36" type="female">1 ♀</specimenCount>
; same locality as for preceding;
<date id="AF271007CB5BFFA0E344FCF0EB79FED1" lastPageId="4" lastPageNumber="37" pageId="3" pageNumber="36" value="2022-05-29">
<collectingDate id="BF63E9EFCB5BFFA0E344FCF0EB79FED1" lastPageId="4" lastPageNumber="37" pageId="3" pageNumber="36" value="2022-05-29">29 May 2022</collectingDate>
</date>
;
<collectorName id="766C5311CB5CFFA0E750FE95E86DFED1" box="[333,552,265,291]" pageId="4" pageNumber="37">J. Romero-Rincon</collectorName>
and
<collectorName id="766C5311CB5CFFA0E477FE95E898FED1" box="[618,733,265,291]" pageId="4" pageNumber="37">R. De La</collectorName>
Cruz leg.;
<collectedFrom id="6885753ACB5CFFA0E571FE95EE86FED1" box="[876,1219,265,291]" pageId="4" pageNumber="37">inside a decaying fallen log</collectedFrom>
;
<collectingMethod id="02D84ED0CB5CFFA0E2CEFE95EB70FEB4" pageId="4" pageNumber="37">daytime hand collection</collectingMethod>
;
<specimenCode id="8B3F9EBCCB5CFFA0E75AFEB0E875FEB4" box="[327,560,300,326]" pageId="4" pageNumber="37">MHN-UPN-MD-96</specimenCode>
</materialsCitation>
<materialsCitation id="6BF13C9ACB5CFFA0E44AFEB0EEA0FE9B" collectedFrom="under a decaying fallen log" collectingDate="2022-04-29" collectingDateMax="2022-09-29" collectingDateMin="2022-09-26" collectingMethod="night time hand collection" collectionCode="R" collectorName="J. Romero-Rincon" country="Colombia" elevation="2253" latitude="4.617133" location="Parque Natural Chicaque" longLatPrecision="1" longitude="-74.313835" municipality="San Antonio del Tequendama" pageId="4" pageNumber="37" specimenCode="MHN-UPN-MD-99" specimenCount="1" specimenCount-female="1" stateProvince="Cundinamarca">
<specimenCount id="CD9FFD4ECB5CFFA0E44AFEB0E8CFFEB5" box="[599,650,300,327]" count="1" pageId="4" pageNumber="37" type="female">1 ♀</specimenCount>
; same locality as for preceding;
<date id="AF271007CB5CFFA0E22CFEB0EE9FFEB4" box="[1073,1242,300,326]" pageId="4" pageNumber="37" value="2022-04-29">
<collectingDate id="BF63E9EFCB5CFFA0E22CFEB0EE9FFEB4" box="[1073,1242,300,326]" pageId="4" pageNumber="37" value="2022-04-29">29 Apr. 2022</collectingDate>
</date>
;
<collectorName id="766C5311CB5CFFA0E2F0FEB1EB57FE9B" pageId="4" pageNumber="37">J. Romero-Rincon</collectorName>
leg.;
<collectedFrom id="6885753ACB5CFFA0E745FED3E8E6FE9B" box="[344,675,335,362]" pageId="4" pageNumber="37">under a decaying fallen log</collectedFrom>
;
<collectingMethod id="02D84ED0CB5CFFA0E4ACFED3E9AAFE9B" box="[689,1007,335,361]" pageId="4" pageNumber="37">night time hand collection</collectingMethod>
;
<specimenCode id="8B3F9EBCCB5CFFA0E5E1FED3EEA0FE9B" box="[1020,1253,335,361]" pageId="4" pageNumber="37">MHN-UPN-MD-99</specimenCode>
</materialsCitation>
<materialsCitation id="6BF13C9ACB5CFFA0E31FFED3EECEFE42" collectedFrom="inside a decaying fallen log" collectingDate="2022-06-25" collectingDateMax="2022-06-26" collectingDateMin="2022-06-25" collectingMethod="night time hand collection" collectionCode="R" collectorName="J. Romero-Rincon &amp; R. De La Cruz" country="Colombia" elevation="2253" latitude="4.617133" location="Parque Natural Chicaque" longLatPrecision="1" longitude="-74.313835" municipality="San Antonio del Tequendama" pageId="4" pageNumber="37" specimenCode="MHN-UPN-MD-98" specimenCount="6" specimenCount-female="1" specimenCount-immature="4" specimenCount-male="1" stateProvince="Cundinamarca">
<specimenCount id="CD9FFD4ECB5CFFA0E31FFED3EF77FE98" box="[1282,1330,335,362]" count="1" pageId="4" pageNumber="37" type="male">1 ♂</specimenCount>
,
<specimenCount id="CD9FFD4ECB5CFFA0E35CFED3EF2AFE98" box="[1345,1391,335,362]" count="1" pageId="4" pageNumber="37" type="female">1 ♀</specimenCount>
,
<specimenCount id="CD9FFD4ECB5CFFA0E6A0FEEFEB10FE7E" box="[189,341,370,397]" count="4" pageId="4" pageNumber="37" type="immature">4 immatures</specimenCount>
; same locality as for preceding;
<date id="AF271007CB5CFFA0E4C7FEEEE9E6FE7F" box="[730,931,370,397]" pageId="4" pageNumber="37" value="2022-06-25" valueMax="2022-06-26" valueMin="2022-06-25">
<collectingDate id="BF63E9EFCB5CFFA0E4C7FEEEE9E6FE7F" box="[730,931,370,397]" pageId="4" pageNumber="37" value="2022-06-25" valueMax="2022-06-26" valueMin="2022-06-25">2526 Jun. 2022</collectingDate>
</date>
;
<collectorName id="766C5311CB5CFFA0E5ACFEEFEECFFE7E" box="[945,1162,370,397]" pageId="4" pageNumber="37">J. Romero-Rincon</collectorName>
and
<collectorName id="766C5311CB5CFFA0E2DBFEEFEF33FE7F" box="[1222,1398,371,397]" pageId="4" pageNumber="37">R. De La Cruz</collectorName>
leg.;
<collectedFrom id="6885753ACB5CFFA0E6E6FE0AE80FFE42" box="[251,586,406,432]" pageId="4" pageNumber="37">inside a decaying fallen log</collectedFrom>
;
<collectingMethod id="02D84ED0CB5CFFA0E445FE0AE9D0FE42" box="[600,917,406,432]" pageId="4" pageNumber="37">night time hand collection</collectingMethod>
;
<specimenCode id="8B3F9EBCCB5CFFA0E5BEFE0AEECEFE42" box="[931,1163,406,432]" pageId="4" pageNumber="37">MHN-UPN-MD-98</specimenCode>
</materialsCitation>
<materialsCitation id="6BF13C9ACB5CFFA0E2B4FE0AE98DFE04" collectedFrom="under a decaying fallen log" collectingDate="2022-06-25" collectingDateMax="2022-06-26" collectingDateMin="2022-06-25" collectingMethod="daytime hand collection" collectionCode="R" collectorName="J. Romero-Rincon &amp; R. De La Cruz" country="Colombia" elevation="2253" latitude="4.617133" location="Parque Natural Chicaque" longLatPrecision="1" longitude="-74.313835" municipality="San Antonio del Tequendama" pageId="4" pageNumber="37" specimenCode="MHNUC-MD-412" specimenCount="2" specimenCount-immature="1" specimenCount-male="1" stateProvince="Cundinamarca">
<specimenCount id="CD9FFD4ECB5CFFA0E2B4FE0AEE9CFE42" box="[1193,1241,406,432]" count="1" pageId="4" pageNumber="37" type="male">1 ♂</specimenCount>
,
<specimenCount id="CD9FFD4ECB5CFFA0E2F5FE0AEF31FE42" box="[1256,1396,406,432]" count="1" pageId="4" pageNumber="37" type="immature">1 immature</specimenCount>
; same locality as for preceding;
<date id="AF271007CB5CFFA0E42AFE25E944FE21" box="[567,769,441,468]" pageId="4" pageNumber="37" value="2022-06-25" valueMax="2022-06-26" valueMin="2022-06-25">
<collectingDate id="BF63E9EFCB5CFFA0E42AFE25E944FE21" box="[567,769,441,468]" pageId="4" pageNumber="37" value="2022-06-25" valueMax="2022-06-26" valueMin="2022-06-25">2526 Jun. 2022</collectingDate>
</date>
;
<collectorName id="766C5311CB5CFFA0E512FE25E9ADFE21" box="[783,1000,441,467]" pageId="4" pageNumber="37">J. Romero-Rincon</collectorName>
and
<collectorName id="766C5311CB5CFFA0E23BFE25EE92FE21" box="[1062,1239,441,467]" pageId="4" pageNumber="37">R. De La Cruz</collectorName>
leg.;
<collectedFrom id="6885753ACB5CFFA0E300FE25EBEFFE04" pageId="4" pageNumber="37">under a decaying fallen log</collectedFrom>
;
<collectingMethod id="02D84ED0CB5CFFA0E7A4FE40E898FE04" box="[441,733,476,502]" pageId="4" pageNumber="37">daytime hand collection</collectingMethod>
;
<specimenCode id="8B3F9EBCCB5CFFA0E4F6FE40E98DFE04" box="[747,968,476,502]" pageId="4" pageNumber="37">MHNUC-MD-412</specimenCode>
</materialsCitation>
<materialsCitation id="6BF13C9ACB5CFFA0E5FAFE40E879FDCE" collectedFrom="under a decaying fallen log" collectingDate="2023-06-19" collectingDateMax="2022-06-26" collectingDateMin="2022-06-25" collectingMethod="night time hand collection" collectionCode="R" collectorName="J. Romero-Rincon" country="Colombia" elevation="2253" latitude="4.617133" location="Parque Natural Chicaque" longLatPrecision="1" longitude="-74.313835" municipality="San Antonio del Tequendama" pageId="4" pageNumber="37" specimenCode="MHN-UPN-MD-163" specimenCount="3" specimenCount-female="2" specimenCount-immature="1" stateProvince="Cundinamarca">
<specimenCount id="CD9FFD4ECB5CFFA0E5FAFE40EE68FE05" box="[999,1069,476,503]" count="2" pageId="4" pageNumber="37" type="female">2 ♀♀</specimenCount>
,
<specimenCount id="CD9FFD4ECB5CFFA0E223FE40EE8EFE04" box="[1086,1227,476,502]" count="1" pageId="4" pageNumber="37" type="immature">1 immature</specimenCount>
; same locality as for preceding;
<date id="AF271007CB5CFFA0E78CFE63E875FDEB" box="[401,560,511,538]" pageId="4" pageNumber="37" value="2023-06-19">
<collectingDate id="BF63E9EFCB5CFFA0E78CFE63E875FDEB" box="[401,560,511,538]" pageId="4" pageNumber="37" value="2023-06-19">19 Jun. 2023</collectingDate>
</date>
;
<collectorName id="766C5311CB5CFFA0E423FD9CE95DFDEB" box="[574,792,511,538]" pageId="4" pageNumber="37">J. Romero-Rincon</collectorName>
leg.;
<collectedFrom id="6885753ACB5CFFA0E542FE63EEEEFDEB" box="[863,1195,511,538]" pageId="4" pageNumber="37">under a decaying fallen log</collectedFrom>
;
<collectingMethod id="02D84ED0CB5CFFA0E2A7FE63EB72FDCE" pageId="4" pageNumber="37">night time hand collection</collectingMethod>
;
<specimenCode id="8B3F9EBCCB5CFFA0E759FDBEE879FDCE" box="[324,572,546,572]" pageId="4" pageNumber="37">MHN-UPN-MD-163</specimenCode>
</materialsCitation>
<materialsCitation id="6BF13C9ACB5CFFA0E445FDBFEB67FD71" collectedFrom="under a decaying fallen log" collectingDate="2023-08-16" collectingDateMax="2022-06-26" collectingDateMin="2022-06-25" collectingMethod="daytime hand collection" collectionCode="R" collectorName="J. Romero-Rincon &amp; H. Reip" country="Colombia" elevation="2253" latitude="4.617133" location="Parque Natural Chicaque" longLatPrecision="1" longitude="-74.313835" municipality="San Antonio del Tequendama" pageId="4" pageNumber="37" specimenCode="MHN-UPN- MD-169" specimenCount="4" specimenCount-immature="3" specimenCount-male="1" stateProvince="Cundinamarca">
<specimenCount id="CD9FFD4ECB5CFFA0E445FDBFE8CDFDCF" box="[600,648,547,573]" count="1" pageId="4" pageNumber="37" type="male">1 ♂</specimenCount>
,
<specimenCount id="CD9FFD4ECB5CFFA0E48BFDBFE96BFDCE" box="[662,814,546,573]" count="3" pageId="4" pageNumber="37" type="immature">3 immatures</specimenCount>
; same locality as for preceding;
<date id="AF271007CB5CFFA0E2AEFDBEEF1DFDCF" box="[1203,1368,546,573]" pageId="4" pageNumber="37" value="2023-08-16">
<collectingDate id="BF63E9EFCB5CFFA0E2AEFDBEEF1DFDCF" box="[1203,1368,546,573]" pageId="4" pageNumber="37" value="2023-08-16">16 Aug. 2023</collectingDate>
</date>
;
<collectorName id="766C5311CB5CFFA0E37BFDBFEB3FFD92" pageId="4" pageNumber="37">J. Romero-Rincon</collectorName>
and
<collectorName id="766C5311CB5CFFA0E7AAFDDAE851FD92" box="[439,532,582,608]" pageId="4" pageNumber="37">H. Reip</collectorName>
leg.;
<collectedFrom id="6885753ACB5CFFA0E446FDDAE9E3FD92" box="[603,934,582,608]" pageId="4" pageNumber="37">under a decaying fallen log</collectedFrom>
;
<collectingMethod id="02D84ED0CB5CFFA0E5A9FDDAEE92FD92" box="[948,1239,582,608]" pageId="4" pageNumber="37">daytime hand collection</collectingMethod>
;
<specimenCode id="8B3F9EBCCB5CFFA0E2F9FDDAEB67FD71" pageId="4" pageNumber="37">MHN-UPNMD-169</specimenCode>
</materialsCitation>
<materialsCitation id="6BF13C9ACB5CFFA0E72BFDF5E91DFD3B" collectedFrom="under bark of fallen log" collectingDate="2022-04-28" collectingDateMax="2022-04-29" collectingDateMin="2022-04-28" collectingMethod="daytime hand collection" collectorName="J. Romero-Rincon" country="Colombia" elevation="2155" latitude="4.6142335" location="Parque Natural Chicaque" longLatPrecision="1" longitude="-74.314316" municipality="San Antonio del Tequendama" pageId="4" pageNumber="37" specimenCode="MHN-UPN-MD-97" specimenCount="1" specimenCount-female="1" stateProvince="Cundinamarca">
<specimenCount id="CD9FFD4ECB5CFFA0E72BFDF5EB25FD71" box="[310,352,617,643]" count="1" pageId="4" pageNumber="37" type="female">1 ♀</specimenCount>
;
<collectingRegion id="195DF825CB5CFFA0E771FDF5E85FFD71" box="[364,538,617,643]" country="Colombia" name="Cundinamarca" pageId="4" pageNumber="37">Cundinamarca</collectingRegion>
,
<collectingMunicipality id="3B42ACBDCB5CFFA0E439FDF5E93CFD71" box="[548,889,617,643]" pageId="4" pageNumber="37">San Antonio del Tequendama</collectingMunicipality>
,
<location id="DE46601CCB5CFFA0E59FFDF5EEE2FD71" LSID="urn:lsid:plazi:treatment:533087D1CB5AFFA3E473F9A2EE24FCA9:DE46601CCB5CFFA0E59FFDF5EEE2FD71" box="[898,1191,617,643]" country="Colombia" latitude="4.6142335" longLatPrecision="1" longitude="-74.314316" municipality="San Antonio del Tequendama" name="Parque Natural Chicaque" pageId="4" pageNumber="37" stateProvince="Cundinamarca">Parque Natural Chicaque</location>
,
<locationDeviation id="CA942FE5CB5CFFA0E2ADFDF5EF37FD71" box="[1200,1394,617,643]" pageId="4" pageNumber="37">secondary forest</locationDeviation>
; [
<geoCoordinate id="BEAD5000CB5CFFA0E6D9FD17EB20FD55" box="[196,357,651,679]" degrees="04" direction="north" minutes="36.854" orientation="latitude" pageId="4" pageNumber="37" precision="1" value="4.6142335">04º36.854 N</geoCoordinate>
,
<geoCoordinate id="BEAD5000CB5CFFA0E76EFD17E858FD55" box="[371,541,651,679]" degrees="74" direction="west" minutes="18.859" orientation="longitude" pageId="4" pageNumber="37" precision="1" value="-74.314316">74°18.859 W</geoCoordinate>
];
<elevation id="50B4D1F4CB5CFFA0E42FFD10E88BFD54" box="[562,718,652,679]" metricMagnitude="3" metricUnit="m" metricValue="2.155" pageId="4" pageNumber="37" unit="m" value="2155.0">
<quantity id="1C619B22CB5CFFA0E42FFD10E8C8FD55" box="[562,653,652,679]" metricMagnitude="3" metricUnit="m" metricValue="2.155" pageId="4" pageNumber="37" unit="m" value="2155.0">2155 m</quantity>
a.s.l.
</elevation>
;
<date id="AF271007CB5CFFA0E4C3FD10E9ECFD54" box="[734,937,652,678]" pageId="4" pageNumber="37" value="2022-04-28" valueMax="2022-04-29" valueMin="2022-04-28">
<collectingDate id="BF63E9EFCB5CFFA0E4C3FD10E9ECFD54" box="[734,937,652,678]" pageId="4" pageNumber="37" value="2022-04-28" valueMax="2022-04-29" valueMin="2022-04-28">2829 Apr. 2022</collectingDate>
</date>
;
<collectorName id="766C5311CB5CFFA0E5AAFD11EED5FD54" box="[951,1168,652,679]" pageId="4" pageNumber="37">J. Romero-Rincon</collectorName>
leg.;
<collectedFrom id="6885753ACB5CFFA0E2C8FD10EB77FD3B" pageId="4" pageNumber="37">under bark of fallen log</collectedFrom>
;
<collectingMethod id="02D84ED0CB5CFFA0E75DFD33E827FD3B" box="[320,610,687,713]" pageId="4" pageNumber="37">daytime hand collection</collectingMethod>
;
<specimenCode id="8B3F9EBCCB5CFFA0E472FD33E91DFD3B" box="[623,856,687,713]" pageId="4" pageNumber="37">MHN-UPN-MD-97</specimenCode>
</materialsCitation>
<materialsCitation id="6BF13C9ACB5CFFA0E568FD33EB52FCE3" collectedFrom="under a decaying fallen log" collectingDate="2022-07-30" collectingDateMax="2022-07-31" collectingDateMin="2022-07-30" collectingMethod="night time hand collection" collectorName="J. Romero-Rincon" country="Colombia" elevation="2155" latitude="4.6142335" location="Parque Natural Chicaque" longLatPrecision="1" longitude="-74.314316" municipality="San Antonio del Tequendama" pageId="4" pageNumber="37" specimenCode="MHN-UPN- MD-95" specimenCount="1" specimenCount-male="1" stateProvince="Cundinamarca">
<specimenCount id="CD9FFD4ECB5CFFA0E568FD33E9E3FD38" box="[885,934,687,714]" count="1" pageId="4" pageNumber="37" type="male">1 ♂</specimenCount>
; same locality as for preceding;
<date id="AF271007CB5CFFA0E330FD33EB69FD1F" pageId="4" pageNumber="37" value="2022-07-30" valueMax="2022-07-31" valueMin="2022-07-30">
<collectingDate id="BF63E9EFCB5CFFA0E330FD33EB69FD1F" pageId="4" pageNumber="37" value="2022-07-30" valueMax="2022-07-31" valueMin="2022-07-30">3031 Jul. 2022</collectingDate>
</date>
;
<collectorName id="766C5311CB5CFFA0E725FD4FE84AFD1E" box="[312,527,722,749]" pageId="4" pageNumber="37">J. Romero-Rincon</collectorName>
leg.;
<collectedFrom id="6885753ACB5CFFA0E44FFD4EE9D0FD1E" box="[594,917,722,749]" pageId="4" pageNumber="37">under a decaying fallen log</collectedFrom>
;
<collectingMethod id="02D84ED0CB5CFFA0E5BCFD4EEE9CFD1E" box="[929,1241,722,748]" pageId="4" pageNumber="37">night time hand collection</collectingMethod>
;
<specimenCode id="8B3F9EBCCB5CFFA0E2F9FD4FEB52FCE3" pageId="4" pageNumber="37">MHN-UPNMD-95</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="8FE6664FCB5BFFA7E6A0F835E902F815" ID-DOI="http://doi.org/10.5281/zenodo.14622253" ID-Zenodo-Dep="14622253" httpUri="https://zenodo.org/record/14622253/files/figure.png" pageId="3" pageNumber="36" startId="3.[189,232,1961,1987]" targetBox="[227,1202,978,1932]" targetPageId="3">
<paragraph id="DB2636C7CB5BFFA7E6A0F835E902F815" blockId="3.[189,1399,1961,2023]" pageId="3" pageNumber="36">
<emphasis id="E9EDEAD5CB5BFFA7E6A0F835EB4EF831" bold="true" box="[189,267,1961,1987]" pageId="3" pageNumber="36">Fig. 1.</emphasis>
Habitus of
<taxonomicName id="1C994D44CB5BFFA7E785F835E8C3F831" authorityName="Romero-Rincon &amp; Alvear &amp; Martínez-Torres &amp; Robles-Piñeros" authorityYear="2025" box="[408,646,1961,1987]" class="Diplopoda" family="Comodesmidae" genus="Inodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="mesibovi" status="sp. nov.">
<emphasis id="E9EDEAD5CB5BFFA7E785F835E8C3F831" box="[408,646,1961,1987]" italics="true" pageId="3" pageNumber="36">Inodesmus mesibovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2DE57AECB5BFFA7E490F836E8AEF836" box="[653,747,1962,1988]" pageId="3" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
, male paratype (above; MHNUC-MD-276) and female paratype (below; MHNUC-MD-276). Scale bar: 1 mm.
</paragraph>
</caption>
<subSubSection id="9383654CCB5CFFA3E6A0FCB3E800FE7F" lastPageId="7" lastPageNumber="40" pageId="4" pageNumber="37" type="description">
<paragraph id="DB2636C7CB5CFFA0E6A0FCB3EB0BFCBB" blockId="4.[189,334,815,841]" box="[189,334,815,841]" pageId="4" pageNumber="37">
<heading id="806E81ABCB5CFFA0E6A0FCB3EB0BFCBB" bold="true" box="[189,334,815,841]" fontSize="11" level="3" pageId="4" pageNumber="37" reason="3">
<emphasis id="E9EDEAD5CB5CFFA0E6A0FCB3EB0BFCBB" bold="true" box="[189,334,815,841]" pageId="4" pageNumber="37">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DB2636C7CB5CFFA0E6A0FCC2EEAEFC6E" blockId="4.[189,1399,862,924]" pageId="4" pageNumber="37">
MEASUREMENTS. Male with head + 19 rings, female with head + 20 rings (
<figureCitation id="43A22A42CB5CFFA0E205FCC2EE19FC8A" box="[1048,1116,862,888]" captionStart="Fig" captionStartId="3.[189,232,1961,1987]" captionTargetBox="[227,1202,978,1932]" captionTargetId="figure-222@3.[189,1399,966,1947]" captionTargetPageId="3" captionText="Fig. 1. Habitus of Inodesmus mesibovi sp. nov., male paratype (above; MHNUC-MD-276) and female paratype (below; MHNUC-MD-276). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14622253" httpUri="https://zenodo.org/record/14622253/files/figure.png" pageId="4" pageNumber="37">Fig. 1</figureCitation>
). Average measurement of adult
<typeStatus id="04228865CB5CFFA0E703FC1EEB14FC6E" box="[286,337,898,924]" pageId="4" pageNumber="37">type</typeStatus>
specimens: male/female ca 10/
<quantity id="1C619B22CB5CFFA0E4DCFC1DE96CFC6E" box="[705,809,897,924]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="4" pageNumber="37" unit="mm" value="11.5">11.5 mm</quantity>
long; maximum width ca 0.9/
<quantity id="1C619B22CB5CFFA0E291FC1DEEA2FC6E" box="[1164,1255,897,924]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="37" unit="mm" value="1.2">1.2 mm</quantity>
.
</paragraph>
<paragraph id="DB2636C7CB5CFFA0E6A0FC27EED9FC0B" blockId="4.[189,1398,955,1017]" pageId="4" pageNumber="37">COLORATION. Metazonites, prozonites, and collum deep reddish brown 41 to deep brown 56; head, legs (prefemur and coxa pale greenish yellow 104), and antennae light reddish brown 42.</paragraph>
<paragraph id="DB2636C7CB5CFFA0E6A0FB85EBF7FB6E" blockId="4.[185,1399,1048,1180]" pageId="4" pageNumber="37">
HEAD. Wider than high; narrower than collum; facing downwards (
<figureCitation id="43A22A42CB5CFFA0E5DEFB84EE07FBC0" box="[963,1090,1048,1074]" captionStart="Fig" captionStartId="5.[189,232,1785,1811]" captionTargetBox="[188,1397,532,1759]" captionTargetId="figure-243@5.[172,1414,518,1775]" captionTargetPageId="5" captionText="Fig. 3. Inodesmus mesibovi sp. nov., male holotype (A, CD, HI; ICN-MD-2913) and male paratype (B, EG; MHN-UPN-MD-168). A. Ventrolateral view of head. B. Dorsolateral view of the anterior part of the body. C. Lateral view of the anterior part of the body. D. Midbody sternite. E. Sculpture of the prozonite. F. Ozopore. G. Lateral view of the posterior part of the body. H. Ventral view of telson. I. Spinnerets. Abbreviations: a = anterior part of prozonite; b = posterior part of prozonite; L = limbus; lo = lobe; o = ozopore; r = ridge; s = spherical knobs; t = tooth-like lobe. Scale bars: A, H = 0.2 mm; B, G = 0.5 mm; C = 1 mm; D = 0.1 mm; EF = 0.02 mm; I = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14622259" httpUri="https://zenodo.org/record/14622259/files/figure.png" pageId="4" pageNumber="37">Fig. 3AC</figureCitation>
); setose clypeus (
<figureCitation id="43A22A42CB5CFFA0E311FB84EF2CFBC0" box="[1292,1385,1048,1074]" captionStart="Fig" captionStartId="5.[189,232,1785,1811]" captionTargetBox="[188,1397,532,1759]" captionTargetId="figure-243@5.[172,1414,518,1775]" captionTargetPageId="5" captionText="Fig. 3. Inodesmus mesibovi sp. nov., male holotype (A, CD, HI; ICN-MD-2913) and male paratype (B, EG; MHN-UPN-MD-168). A. Ventrolateral view of head. B. Dorsolateral view of the anterior part of the body. C. Lateral view of the anterior part of the body. D. Midbody sternite. E. Sculpture of the prozonite. F. Ozopore. G. Lateral view of the posterior part of the body. H. Ventral view of telson. I. Spinnerets. Abbreviations: a = anterior part of prozonite; b = posterior part of prozonite; L = limbus; lo = lobe; o = ozopore; r = ridge; s = spherical knobs; t = tooth-like lobe. Scale bars: A, H = 0.2 mm; B, G = 0.5 mm; C = 1 mm; D = 0.1 mm; EF = 0.02 mm; I = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14622259" httpUri="https://zenodo.org/record/14622259/files/figure.png" pageId="4" pageNumber="37">Fig. 3A</figureCitation>
). Vertex microvillose and microgranulate. Antennae (
<figureCitation id="43A22A42CB5CFFA0E50AFBA7E9D0FBA4" box="[791,917,1083,1110]" captionStart="Fig" captionStartId="5.[189,232,1785,1811]" captionTargetBox="[188,1397,532,1759]" captionTargetId="figure-243@5.[172,1414,518,1775]" captionTargetPageId="5" captionText="Fig. 3. Inodesmus mesibovi sp. nov., male holotype (A, CD, HI; ICN-MD-2913) and male paratype (B, EG; MHN-UPN-MD-168). A. Ventrolateral view of head. B. Dorsolateral view of the anterior part of the body. C. Lateral view of the anterior part of the body. D. Midbody sternite. E. Sculpture of the prozonite. F. Ozopore. G. Lateral view of the posterior part of the body. H. Ventral view of telson. I. Spinnerets. Abbreviations: a = anterior part of prozonite; b = posterior part of prozonite; L = limbus; lo = lobe; o = ozopore; r = ridge; s = spherical knobs; t = tooth-like lobe. Scale bars: A, H = 0.2 mm; B, G = 0.5 mm; C = 1 mm; D = 0.1 mm; EF = 0.02 mm; I = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14622259" httpUri="https://zenodo.org/record/14622259/files/figure.png" pageId="4" pageNumber="37">Fig. 3AC</figureCitation>
) short, stout, clavate, densely setose, and held close to head. Antennomere relative lengths as follows: 6&gt; 1&gt; 5&gt; 4= 2&gt; 3&gt; 7. Antennae with 4 sensory apical cones.
</paragraph>
<paragraph id="DB2636C7CB5CFFA2E6A0FB20E950FC6E" blockId="4.[189,1400,1212,1274]" lastBlockId="6.[189,1400,265,925]" lastPageId="6" lastPageNumber="39" pageId="4" pageNumber="37">
TRUNK. Anterior and posterior margins of collum broadly convex; corners rounded and hidden under paranota of second ring, collum with an irregular pattern of two groups of rounded tubercles. At first glance, pattern continues on remaining body rings, with a supposed transverse medial separation on each tergite (
<figureCitation id="43A22A42CB5DFFA1E744FEB0EB9CFEB4" box="[345,473,300,326]" captionStart="Fig" captionStartId="5.[189,232,1785,1811]" captionTargetBox="[188,1397,532,1759]" captionTargetId="figure-243@5.[172,1414,518,1775]" captionTargetPageId="5" captionText="Fig. 3. Inodesmus mesibovi sp. nov., male holotype (A, CD, HI; ICN-MD-2913) and male paratype (B, EG; MHN-UPN-MD-168). A. Ventrolateral view of head. B. Dorsolateral view of the anterior part of the body. C. Lateral view of the anterior part of the body. D. Midbody sternite. E. Sculpture of the prozonite. F. Ozopore. G. Lateral view of the posterior part of the body. H. Ventral view of telson. I. Spinnerets. Abbreviations: a = anterior part of prozonite; b = posterior part of prozonite; L = limbus; lo = lobe; o = ozopore; r = ridge; s = spherical knobs; t = tooth-like lobe. Scale bars: A, H = 0.2 mm; B, G = 0.5 mm; C = 1 mm; D = 0.1 mm; EF = 0.02 mm; I = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14622259" httpUri="https://zenodo.org/record/14622259/files/figure.png" pageId="5" pageNumber="38">Fig. 3BC</figureCitation>
). Ring 2 tergite largest, extending basally, laterally, and posteriorly; extension of paranota of ring 2 incrassate without well-defined tubercles (
<figureCitation id="43A22A42CB5DFFA1E5A9FED3EE70FE9B" box="[948,1077,335,361]" captionStart="Fig" captionStartId="5.[189,232,1785,1811]" captionTargetBox="[188,1397,532,1759]" captionTargetId="figure-243@5.[172,1414,518,1775]" captionTargetPageId="5" captionText="Fig. 3. Inodesmus mesibovi sp. nov., male holotype (A, CD, HI; ICN-MD-2913) and male paratype (B, EG; MHN-UPN-MD-168). A. Ventrolateral view of head. B. Dorsolateral view of the anterior part of the body. C. Lateral view of the anterior part of the body. D. Midbody sternite. E. Sculpture of the prozonite. F. Ozopore. G. Lateral view of the posterior part of the body. H. Ventral view of telson. I. Spinnerets. Abbreviations: a = anterior part of prozonite; b = posterior part of prozonite; L = limbus; lo = lobe; o = ozopore; r = ridge; s = spherical knobs; t = tooth-like lobe. Scale bars: A, H = 0.2 mm; B, G = 0.5 mm; C = 1 mm; D = 0.1 mm; EF = 0.02 mm; I = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14622259" httpUri="https://zenodo.org/record/14622259/files/figure.png" pageId="5" pageNumber="38">Fig. 3BC</figureCitation>
). Posterior rings (
<figureCitation id="43A22A42CB5DFFA1E317FED3EF2CFE9B" box="[1290,1385,335,361]" captionStart="Fig" captionStartId="5.[189,232,1785,1811]" captionTargetBox="[188,1397,532,1759]" captionTargetId="figure-243@5.[172,1414,518,1775]" captionTargetPageId="5" captionText="Fig. 3. Inodesmus mesibovi sp. nov., male holotype (A, CD, HI; ICN-MD-2913) and male paratype (B, EG; MHN-UPN-MD-168). A. Ventrolateral view of head. B. Dorsolateral view of the anterior part of the body. C. Lateral view of the anterior part of the body. D. Midbody sternite. E. Sculpture of the prozonite. F. Ozopore. G. Lateral view of the posterior part of the body. H. Ventral view of telson. I. Spinnerets. Abbreviations: a = anterior part of prozonite; b = posterior part of prozonite; L = limbus; lo = lobe; o = ozopore; r = ridge; s = spherical knobs; t = tooth-like lobe. Scale bars: A, H = 0.2 mm; B, G = 0.5 mm; C = 1 mm; D = 0.1 mm; EF = 0.02 mm; I = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14622259" httpUri="https://zenodo.org/record/14622259/files/figure.png" pageId="5" pageNumber="38">Fig. 3G</figureCitation>
), up to ring 18, with a lateral row of three small tubercles, neither enlarged nor forming pseudo-paranota. Ozopores (
<figureCitation id="43A22A42CB5DFFA1E75FFE0AEBDBFE42" box="[322,414,406,432]" captionStart="Fig" captionStartId="5.[189,232,1785,1811]" captionTargetBox="[188,1397,532,1759]" captionTargetId="figure-243@5.[172,1414,518,1775]" captionTargetPageId="5" captionText="Fig. 3. Inodesmus mesibovi sp. nov., male holotype (A, CD, HI; ICN-MD-2913) and male paratype (B, EG; MHN-UPN-MD-168). A. Ventrolateral view of head. B. Dorsolateral view of the anterior part of the body. C. Lateral view of the anterior part of the body. D. Midbody sternite. E. Sculpture of the prozonite. F. Ozopore. G. Lateral view of the posterior part of the body. H. Ventral view of telson. I. Spinnerets. Abbreviations: a = anterior part of prozonite; b = posterior part of prozonite; L = limbus; lo = lobe; o = ozopore; r = ridge; s = spherical knobs; t = tooth-like lobe. Scale bars: A, H = 0.2 mm; B, G = 0.5 mm; C = 1 mm; D = 0.1 mm; EF = 0.02 mm; I = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14622259" httpUri="https://zenodo.org/record/14622259/files/figure.png" pageId="5" pageNumber="38">Fig. 3F</figureCitation>
) very small, internal closing not evident, located just above or in middle of last lateral row of tubercles (
<figureCitation id="43A22A42CB5DFFA1E7C1FE25E87CFE21" box="[476,569,441,467]" captionStart="Fig" captionStartId="5.[189,232,1785,1811]" captionTargetBox="[188,1397,532,1759]" captionTargetId="figure-243@5.[172,1414,518,1775]" captionTargetPageId="5" captionText="Fig. 3. Inodesmus mesibovi sp. nov., male holotype (A, CD, HI; ICN-MD-2913) and male paratype (B, EG; MHN-UPN-MD-168). A. Ventrolateral view of head. B. Dorsolateral view of the anterior part of the body. C. Lateral view of the anterior part of the body. D. Midbody sternite. E. Sculpture of the prozonite. F. Ozopore. G. Lateral view of the posterior part of the body. H. Ventral view of telson. I. Spinnerets. Abbreviations: a = anterior part of prozonite; b = posterior part of prozonite; L = limbus; lo = lobe; o = ozopore; r = ridge; s = spherical knobs; t = tooth-like lobe. Scale bars: A, H = 0.2 mm; B, G = 0.5 mm; C = 1 mm; D = 0.1 mm; EF = 0.02 mm; I = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14622259" httpUri="https://zenodo.org/record/14622259/files/figure.png" pageId="5" pageNumber="38">Fig. 3G</figureCitation>
). Ozopores internally and externally bordered by microtubercules, with neither evident elevation nor porostele. Pore formula normal (5, 7, 9, 10, 12, 13, 1519). Diplosternites (
<figureCitation id="43A22A42CB5EFFA2E6DBFEB0EB67FEB4" box="[198,290,300,326]" captionStart="Fig" captionStartId="5.[189,232,1785,1811]" captionTargetBox="[188,1397,532,1759]" captionTargetId="figure-243@5.[172,1414,518,1775]" captionTargetPageId="5" captionText="Fig. 3. Inodesmus mesibovi sp. nov., male holotype (A, CD, HI; ICN-MD-2913) and male paratype (B, EG; MHN-UPN-MD-168). A. Ventrolateral view of head. B. Dorsolateral view of the anterior part of the body. C. Lateral view of the anterior part of the body. D. Midbody sternite. E. Sculpture of the prozonite. F. Ozopore. G. Lateral view of the posterior part of the body. H. Ventral view of telson. I. Spinnerets. Abbreviations: a = anterior part of prozonite; b = posterior part of prozonite; L = limbus; lo = lobe; o = ozopore; r = ridge; s = spherical knobs; t = tooth-like lobe. Scale bars: A, H = 0.2 mm; B, G = 0.5 mm; C = 1 mm; D = 0.1 mm; EF = 0.02 mm; I = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14622259" httpUri="https://zenodo.org/record/14622259/files/figure.png" pageId="6" pageNumber="39">Fig. 3D</figureCitation>
) with transverse impression deeper than longitudinal impression, anterior sternites with eight acicular setae, posterior sternites on ring 5 with bumps bearing six acicular setae each. Legs (
<figureCitation id="43A22A42CB5EFFA2E312FED3EB5EFE7F" captionStart="Fig" captionStartId="5.[189,232,1785,1811]" captionTargetBox="[188,1397,532,1759]" captionTargetId="figure-243@5.[172,1414,518,1775]" captionTargetPageId="5" captionText="Fig. 3. Inodesmus mesibovi sp. nov., male holotype (A, CD, HI; ICN-MD-2913) and male paratype (B, EG; MHN-UPN-MD-168). A. Ventrolateral view of head. B. Dorsolateral view of the anterior part of the body. C. Lateral view of the anterior part of the body. D. Midbody sternite. E. Sculpture of the prozonite. F. Ozopore. G. Lateral view of the posterior part of the body. H. Ventral view of telson. I. Spinnerets. Abbreviations: a = anterior part of prozonite; b = posterior part of prozonite; L = limbus; lo = lobe; o = ozopore; r = ridge; s = spherical knobs; t = tooth-like lobe. Scale bars: A, H = 0.2 mm; B, G = 0.5 mm; C = 1 mm; D = 0.1 mm; EF = 0.02 mm; I = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14622259" httpUri="https://zenodo.org/record/14622259/files/figure.png" pageId="6" pageNumber="39">Fig. 3B C, GH</figureCitation>
) short and stout; relative lengths of podomeres: tarsus&gt;(prefemur≥ femur)&gt;tibia&gt;postfemur; claw about as long as postfemur. Spiracles not evident. Telson facing downwards. Paraprocts parallel to substrate and almost flat (
<figureCitation id="43A22A42CB5EFFA2E40BFE25E8DBFE21" box="[534,670,441,467]" captionStart="Fig" captionStartId="5.[189,232,1785,1811]" captionTargetBox="[188,1397,532,1759]" captionTargetId="figure-243@5.[172,1414,518,1775]" captionTargetPageId="5" captionText="Fig. 3. Inodesmus mesibovi sp. nov., male holotype (A, CD, HI; ICN-MD-2913) and male paratype (B, EG; MHN-UPN-MD-168). A. Ventrolateral view of head. B. Dorsolateral view of the anterior part of the body. C. Lateral view of the anterior part of the body. D. Midbody sternite. E. Sculpture of the prozonite. F. Ozopore. G. Lateral view of the posterior part of the body. H. Ventral view of telson. I. Spinnerets. Abbreviations: a = anterior part of prozonite; b = posterior part of prozonite; L = limbus; lo = lobe; o = ozopore; r = ridge; s = spherical knobs; t = tooth-like lobe. Scale bars: A, H = 0.2 mm; B, G = 0.5 mm; C = 1 mm; D = 0.1 mm; EF = 0.02 mm; I = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14622259" httpUri="https://zenodo.org/record/14622259/files/figure.png" pageId="6" pageNumber="39">Fig. 3GH</figureCitation>
). Epiproct (
<figureCitation id="43A22A42CB5EFFA2E531FE25E9E1FE21" box="[812,932,441,467]" captionStart="Fig" captionStartId="5.[189,232,1785,1811]" captionTargetBox="[188,1397,532,1759]" captionTargetId="figure-243@5.[172,1414,518,1775]" captionTargetPageId="5" captionText="Fig. 3. Inodesmus mesibovi sp. nov., male holotype (A, CD, HI; ICN-MD-2913) and male paratype (B, EG; MHN-UPN-MD-168). A. Ventrolateral view of head. B. Dorsolateral view of the anterior part of the body. C. Lateral view of the anterior part of the body. D. Midbody sternite. E. Sculpture of the prozonite. F. Ozopore. G. Lateral view of the posterior part of the body. H. Ventral view of telson. I. Spinnerets. Abbreviations: a = anterior part of prozonite; b = posterior part of prozonite; L = limbus; lo = lobe; o = ozopore; r = ridge; s = spherical knobs; t = tooth-like lobe. Scale bars: A, H = 0.2 mm; B, G = 0.5 mm; C = 1 mm; D = 0.1 mm; EF = 0.02 mm; I = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14622259" httpUri="https://zenodo.org/record/14622259/files/figure.png" pageId="6" pageNumber="39">Fig. 3HI</figureCitation>
) distally projected, but short, flattened dorsoventrally, with four inconspicuous setae (spinnerets), each spinneret with a single low sheath, each seta inside a circular, deep, walled depression (
<figureCitation id="43A22A42CB5EFFA2E53AFE63E93FFDEB" box="[807,890,511,537]" captionStart="Fig" captionStartId="5.[189,232,1785,1811]" captionTargetBox="[188,1397,532,1759]" captionTargetId="figure-243@5.[172,1414,518,1775]" captionTargetPageId="5" captionText="Fig. 3. Inodesmus mesibovi sp. nov., male holotype (A, CD, HI; ICN-MD-2913) and male paratype (B, EG; MHN-UPN-MD-168). A. Ventrolateral view of head. B. Dorsolateral view of the anterior part of the body. C. Lateral view of the anterior part of the body. D. Midbody sternite. E. Sculpture of the prozonite. F. Ozopore. G. Lateral view of the posterior part of the body. H. Ventral view of telson. I. Spinnerets. Abbreviations: a = anterior part of prozonite; b = posterior part of prozonite; L = limbus; lo = lobe; o = ozopore; r = ridge; s = spherical knobs; t = tooth-like lobe. Scale bars: A, H = 0.2 mm; B, G = 0.5 mm; C = 1 mm; D = 0.1 mm; EF = 0.02 mm; I = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14622259" httpUri="https://zenodo.org/record/14622259/files/figure.png" pageId="6" pageNumber="39">Fig. 3I</figureCitation>
). Hypoproct (
<figureCitation id="43A22A42CB5EFFA2E203FE63EE3FFDEB" box="[1054,1146,511,537]" captionStart="Fig" captionStartId="5.[189,232,1785,1811]" captionTargetBox="[188,1397,532,1759]" captionTargetId="figure-243@5.[172,1414,518,1775]" captionTargetPageId="5" captionText="Fig. 3. Inodesmus mesibovi sp. nov., male holotype (A, CD, HI; ICN-MD-2913) and male paratype (B, EG; MHN-UPN-MD-168). A. Ventrolateral view of head. B. Dorsolateral view of the anterior part of the body. C. Lateral view of the anterior part of the body. D. Midbody sternite. E. Sculpture of the prozonite. F. Ozopore. G. Lateral view of the posterior part of the body. H. Ventral view of telson. I. Spinnerets. Abbreviations: a = anterior part of prozonite; b = posterior part of prozonite; L = limbus; lo = lobe; o = ozopore; r = ridge; s = spherical knobs; t = tooth-like lobe. Scale bars: A, H = 0.2 mm; B, G = 0.5 mm; C = 1 mm; D = 0.1 mm; EF = 0.02 mm; I = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14622259" httpUri="https://zenodo.org/record/14622259/files/figure.png" pageId="6" pageNumber="39">Fig. 3H</figureCitation>
) subtrapeziform with a slightly convex anterior margin. Tegument with microsculpture, especially along posterior edge of metazonite, anterior edge of prozonite, head, collum, lateral part of metazonite of second ring, edges of ozopores, and posterior area of telson (
<figureCitation id="43A22A42CB5EFFA2E48DFDF5E89DFD71" box="[656,728,617,643]" captionStart="Fig" captionStartId="5.[189,232,1785,1811]" captionTargetBox="[188,1397,532,1759]" captionTargetId="figure-243@5.[172,1414,518,1775]" captionTargetPageId="5" captionText="Fig. 3. Inodesmus mesibovi sp. nov., male holotype (A, CD, HI; ICN-MD-2913) and male paratype (B, EG; MHN-UPN-MD-168). A. Ventrolateral view of head. B. Dorsolateral view of the anterior part of the body. C. Lateral view of the anterior part of the body. D. Midbody sternite. E. Sculpture of the prozonite. F. Ozopore. G. Lateral view of the posterior part of the body. H. Ventral view of telson. I. Spinnerets. Abbreviations: a = anterior part of prozonite; b = posterior part of prozonite; L = limbus; lo = lobe; o = ozopore; r = ridge; s = spherical knobs; t = tooth-like lobe. Scale bars: A, H = 0.2 mm; B, G = 0.5 mm; C = 1 mm; D = 0.1 mm; EF = 0.02 mm; I = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14622259" httpUri="https://zenodo.org/record/14622259/files/figure.png" pageId="6" pageNumber="39">Fig. 3</figureCitation>
). Microsculpture mostly as a cellular mesh with narrow irregular folds. Integument further elevated into tubercles of different sizes and shapes on head, collum, tergites, metatergites, and telson (
<figureCitation id="43A22A42CB5EFFA2E45DFD33E943FD3B" box="[576,774,687,713]" captionStart="Fig" captionStartId="5.[189,232,1785,1811]" captionTargetBox="[188,1397,532,1759]" captionTargetId="figure-243@5.[172,1414,518,1775]" captionTargetPageId="5" captionText="Fig. 3. Inodesmus mesibovi sp. nov., male holotype (A, CD, HI; ICN-MD-2913) and male paratype (B, EG; MHN-UPN-MD-168). A. Ventrolateral view of head. B. Dorsolateral view of the anterior part of the body. C. Lateral view of the anterior part of the body. D. Midbody sternite. E. Sculpture of the prozonite. F. Ozopore. G. Lateral view of the posterior part of the body. H. Ventral view of telson. I. Spinnerets. Abbreviations: a = anterior part of prozonite; b = posterior part of prozonite; L = limbus; lo = lobe; o = ozopore; r = ridge; s = spherical knobs; t = tooth-like lobe. Scale bars: A, H = 0.2 mm; B, G = 0.5 mm; C = 1 mm; D = 0.1 mm; EF = 0.02 mm; I = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14622259" httpUri="https://zenodo.org/record/14622259/files/figure.png" pageId="6" pageNumber="39">Fig. 3AC, GH</figureCitation>
); some tubercles with a single small and acicular seta. Cell boundaries in posterior part of metazonite not extending basally past limbus (
<figureCitation id="43A22A42CB5EFFA2E2BFFD4EEF45FD1F" box="[1186,1280,722,749]" captionStart="Fig" captionStartId="5.[189,232,1785,1811]" captionTargetBox="[188,1397,532,1759]" captionTargetId="figure-243@5.[172,1414,518,1775]" captionTargetPageId="5" captionText="Fig. 3. Inodesmus mesibovi sp. nov., male holotype (A, CD, HI; ICN-MD-2913) and male paratype (B, EG; MHN-UPN-MD-168). A. Ventrolateral view of head. B. Dorsolateral view of the anterior part of the body. C. Lateral view of the anterior part of the body. D. Midbody sternite. E. Sculpture of the prozonite. F. Ozopore. G. Lateral view of the posterior part of the body. H. Ventral view of telson. I. Spinnerets. Abbreviations: a = anterior part of prozonite; b = posterior part of prozonite; L = limbus; lo = lobe; o = ozopore; r = ridge; s = spherical knobs; t = tooth-like lobe. Scale bars: A, H = 0.2 mm; B, G = 0.5 mm; C = 1 mm; D = 0.1 mm; EF = 0.02 mm; I = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14622259" httpUri="https://zenodo.org/record/14622259/files/figure.png" pageId="6" pageNumber="39">Fig. 3E</figureCitation>
). Primary limbus element with a regular set of rounded lobes and tooth-like lobes on the secondary element (
<figureCitation id="43A22A42CB5EFFA2E6DBFC85EB62FCC1" box="[198,295,793,819]" captionStart="Fig" captionStartId="5.[189,232,1785,1811]" captionTargetBox="[188,1397,532,1759]" captionTargetId="figure-243@5.[172,1414,518,1775]" captionTargetPageId="5" captionText="Fig. 3. Inodesmus mesibovi sp. nov., male holotype (A, CD, HI; ICN-MD-2913) and male paratype (B, EG; MHN-UPN-MD-168). A. Ventrolateral view of head. B. Dorsolateral view of the anterior part of the body. C. Lateral view of the anterior part of the body. D. Midbody sternite. E. Sculpture of the prozonite. F. Ozopore. G. Lateral view of the posterior part of the body. H. Ventral view of telson. I. Spinnerets. Abbreviations: a = anterior part of prozonite; b = posterior part of prozonite; L = limbus; lo = lobe; o = ozopore; r = ridge; s = spherical knobs; t = tooth-like lobe. Scale bars: A, H = 0.2 mm; B, G = 0.5 mm; C = 1 mm; D = 0.1 mm; EF = 0.02 mm; I = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14622259" httpUri="https://zenodo.org/record/14622259/files/figure.png" pageId="6" pageNumber="39">Fig. 3E</figureCitation>
). Prozonites sharply demarcated from metazonites (
<figureCitation id="43A22A42CB5EFFA2E5B2FC85EE18FCC1" box="[943,1117,793,819]" captionStart="Fig" captionStartId="5.[189,232,1785,1811]" captionTargetBox="[188,1397,532,1759]" captionTargetId="figure-243@5.[172,1414,518,1775]" captionTargetPageId="5" captionText="Fig. 3. Inodesmus mesibovi sp. nov., male holotype (A, CD, HI; ICN-MD-2913) and male paratype (B, EG; MHN-UPN-MD-168). A. Ventrolateral view of head. B. Dorsolateral view of the anterior part of the body. C. Lateral view of the anterior part of the body. D. Midbody sternite. E. Sculpture of the prozonite. F. Ozopore. G. Lateral view of the posterior part of the body. H. Ventral view of telson. I. Spinnerets. Abbreviations: a = anterior part of prozonite; b = posterior part of prozonite; L = limbus; lo = lobe; o = ozopore; r = ridge; s = spherical knobs; t = tooth-like lobe. Scale bars: A, H = 0.2 mm; B, G = 0.5 mm; C = 1 mm; D = 0.1 mm; EF = 0.02 mm; I = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14622259" httpUri="https://zenodo.org/record/14622259/files/figure.png" pageId="6" pageNumber="39">Fig. 3BC, G</figureCitation>
). Anterior part of prozonite (
<emphasis id="E9EDEAD5CB5EFFA2E704FCA0EB6DFCA4" bold="true" box="[281,296,828,854]" pageId="6" pageNumber="39">a</emphasis>
) with small irregular subcardiform units arranged in transverse “rows” placed on a smooth background. A transverse ridge (
<emphasis id="E9EDEAD5CB5EFFA2E42DFCC3E878FC8B" bold="true" box="[560,573,863,889]" pageId="6" pageNumber="39">r</emphasis>
) is marked by a row of cell-like units. Posterior part of the prozonite (
<emphasis id="E9EDEAD5CB5EFFA2E340FCC3EF28FC8B" bold="true" box="[1373,1389,863,889]" pageId="6" pageNumber="39">b</emphasis>
) with a smooth surface and lacking spherical knobs.
</paragraph>
<caption id="8FE6664FCB5CFFA0E6A0F835E96EF815" ID-DOI="http://doi.org/10.5281/zenodo.14622257" ID-Zenodo-Dep="14622257" httpUri="https://zenodo.org/record/14622257/files/figure.png" pageId="4" pageNumber="37" startId="4.[189,232,1961,1987]" targetBox="[189,1366,1332,1937]" targetPageId="4">
<paragraph id="DB2636C7CB5CFFA0E6A0F835E96EF815" blockId="4.[189,1399,1961,2023]" pageId="4" pageNumber="37">
<emphasis id="E9EDEAD5CB5CFFA0E6A0F835EB4EF831" bold="true" box="[189,267,1961,1987]" pageId="4" pageNumber="37">Fig. 2.</emphasis>
<taxonomicName id="1C994D44CB5CFFA0E70FF835EBBAF831" authorityName="Romero-Rincon &amp; Alvear &amp; Martínez-Torres &amp; Robles-Piñeros" authorityYear="2025" box="[274,511,1961,1987]" class="Diplopoda" family="Comodesmidae" genus="Inodesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="mesibovi" status="sp. nov.">
<emphasis id="E9EDEAD5CB5CFFA0E70FF835EBBAF831" box="[274,511,1961,1987]" italics="true" pageId="4" pageNumber="37">Inodesmus mesibovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2DE57AECB5CFFA0E41AF836E821F836" box="[519,612,1962,1988]" pageId="4" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (ICN-MD-2913).
<emphasis id="E9EDEAD5CB5CFFA0E5EFF835EE42F831" bold="true" box="[1010,1031,1961,1987]" pageId="4" pageNumber="37">A</emphasis>
. Ventral view of the gonopods.
<emphasis id="E9EDEAD5CB5CFFA0E6A0F851EA95F815" bold="true" box="[189,208,1997,2023]" pageId="4" pageNumber="37">B</emphasis>
. Left gonopod in ventral view. Scale bars: 0.1 mm.
</paragraph>
</caption>
<caption id="8FE6664FCB5DFFA1E6A0F965EE44F815" ID-DOI="http://doi.org/10.5281/zenodo.14622259" ID-Zenodo-Dep="14622259" httpUri="https://zenodo.org/record/14622259/files/figure.png" pageId="5" pageNumber="38" startId="5.[189,232,1785,1811]" targetBox="[188,1397,532,1759]" targetPageId="5">
<paragraph id="DB2636C7CB5DFFA1E6A0F965EE44F815" blockId="5.[189,1399,1785,2023]" pageId="5" pageNumber="38">
<emphasis id="E9EDEAD5CB5DFFA1E6A0F965EB4EF8E1" bold="true" box="[189,267,1785,1811]" pageId="5" pageNumber="38">Fig. 3.</emphasis>
<taxonomicName id="1C994D44CB5DFFA1E709F965E847F8E1" authorityName="Romero-Rincon &amp; Alvear &amp; Martínez-Torres &amp; Robles-Piñeros" authorityYear="2025" box="[276,514,1785,1811]" class="Diplopoda" family="Comodesmidae" genus="Inodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="38" phylum="Arthropoda" rank="species" species="mesibovi" status="sp. nov.">
<emphasis id="E9EDEAD5CB5DFFA1E709F965E847F8E1" box="[276,514,1785,1811]" italics="true" pageId="5" pageNumber="38">Inodesmus mesibovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2DE57AECB5DFFA1E416F966E82CF8E6" box="[523,617,1786,1812]" pageId="5" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (A, CD, HI; ICN-MD-2913) and male paratype (B, EG; MHN-UPN-MD-168).
<emphasis id="E9EDEAD5CB5DFFA1E454F880E81BF8C4" bold="true" box="[585,606,1820,1846]" pageId="5" pageNumber="38">A</emphasis>
. Ventrolateral view of head.
<emphasis id="E9EDEAD5CB5DFFA1E5DEF880E992F8C4" bold="true" box="[963,983,1820,1846]" pageId="5" pageNumber="38">B</emphasis>
. Dorsolateral view of the anterior part of the body.
<emphasis id="E9EDEAD5CB5DFFA1E794F8A3EBD8F8AB" bold="true" box="[393,413,1855,1881]" pageId="5" pageNumber="38">C</emphasis>
. Lateral view of the anterior part of the body.
<emphasis id="E9EDEAD5CB5DFFA1E5D8F8DCE99CF8A8" bold="true" box="[965,985,1856,1882]" pageId="5" pageNumber="38">D</emphasis>
. Midbody sternite.
<emphasis id="E9EDEAD5CB5DFFA1E2DEF8DCEE93F8A8" bold="true" box="[1219,1238,1856,1882]" pageId="5" pageNumber="38">E</emphasis>
. Sculpture of the prozonite.
<emphasis id="E9EDEAD5CB5DFFA1E775F8FFEB3CF88F" bold="true" box="[360,377,1891,1917]" pageId="5" pageNumber="38">F</emphasis>
. Ozopore.
<emphasis id="E9EDEAD5CB5DFFA1E7E0F8FEE856F88E" bold="true" box="[509,531,1890,1916]" pageId="5" pageNumber="38">G</emphasis>
. Lateral view of the posterior part of the body.
<emphasis id="E9EDEAD5CB5DFFA1E25BF8FFEE19F88F" bold="true" box="[1094,1116,1891,1917]" pageId="5" pageNumber="38">H</emphasis>
. Ventral view of telson.
<emphasis id="E9EDEAD5CB5DFFA1E6A0F81AEA8DF852" bold="true" box="[189,200,1926,1952]" pageId="5" pageNumber="38">I</emphasis>
. Spinnerets. Abbreviations:
<emphasis id="E9EDEAD5CB5DFFA1E409F81AE866F852" bold="true" box="[532,547,1926,1952]" pageId="5" pageNumber="38">a</emphasis>
= anterior part of prozonite;
<emphasis id="E9EDEAD5CB5DFFA1E567F81AE9E6F852" bold="true" box="[890,931,1926,1952]" pageId="5" pageNumber="38">b =</emphasis>
posterior part of prozonite;
<emphasis id="E9EDEAD5CB5DFFA1E2F0F81AEF5DF852" bold="true" box="[1261,1304,1926,1952]" pageId="5" pageNumber="38">L =</emphasis>
limbus;
<emphasis id="E9EDEAD5CB5DFFA1E6A0F835EAA8F831" bold="true" box="[189,237,1961,1987]" pageId="5" pageNumber="38">lo =</emphasis>
lobe;
<emphasis id="E9EDEAD5CB5DFFA1E725F835EB02F831" bold="true" box="[312,327,1961,1987]" pageId="5" pageNumber="38">o</emphasis>
= ozopore;
<emphasis id="E9EDEAD5CB5DFFA1E7C4F835EBA3F831" bold="true" box="[473,486,1961,1987]" pageId="5" pageNumber="38">r</emphasis>
= ridge;
<emphasis id="E9EDEAD5CB5DFFA1E449F835E81AF831" bold="true" box="[596,607,1961,1987]" pageId="5" pageNumber="38">s</emphasis>
= spherical knobs;
<emphasis id="E9EDEAD5CB5DFFA1E554F835E928F831" bold="true" box="[841,877,1961,1987]" pageId="5" pageNumber="38">t =</emphasis>
tooth-like lobe. Scale bars: A, H = 0.2 mm; B, G = 0.5 mm; C = 1 mm; D = 0.1 mm; EF = 0.02 mm; I = 0.05 mm.
</paragraph>
</caption>
<caption id="8FE6664FCB5EFFA2E6A0F819EB17F814" ID-DOI="http://doi.org/10.5281/zenodo.14622263" ID-Zenodo-Dep="14622263" httpUri="https://zenodo.org/record/14622263/files/figure.png" pageId="6" pageNumber="39" startId="6.[189,232,1925,1951]" targetBox="[231,1366,973,1898]" targetPageId="6">
<paragraph id="DB2636C7CB5EFFA2E6A0F819EB17F814" blockId="6.[186,1399,1925,2023]" pageId="6" pageNumber="39">
<emphasis id="E9EDEAD5CB5EFFA2E6A0F819EB4CF86D" bold="true" box="[189,265,1925,1951]" pageId="6" pageNumber="39">Fig. 4.</emphasis>
Teratologies in
<taxonomicName id="1C994D44CB5EFFA2E7A2F819E8EEF86D" authorityName="Romero-Rincon &amp; Alvear &amp; Martínez-Torres &amp; Robles-Piñeros" authorityYear="2025" box="[447,683,1925,1951]" class="Diplopoda" family="Comodesmidae" genus="Inodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" species="mesibovi" status="sp. nov.">
<emphasis id="E9EDEAD5CB5EFFA2E7A2F819E8EEF86D" box="[447,683,1925,1951]" italics="true" pageId="6" pageNumber="39">Inodesmus mesibovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2DE57AECB5EFFA2E4AEF81AE94AF852" box="[691,783,1926,1952]" pageId="6" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
, female paratype (MHN-UPN-MD-177).
<emphasis id="E9EDEAD5CB5EFFA2E2EFF819EF42F86D" bold="true" box="[1266,1287,1925,1951]" pageId="6" pageNumber="39">A</emphasis>
. Anterior part of body.
<emphasis id="E9EDEAD5CB5EFFA2E749F835EB2DF831" bold="true" box="[340,360,1961,1987]" pageId="6" pageNumber="39">B</emphasis>
. Middle part of the body. The numbers 6 and 9 correspond to the body ring number. Scale bar: 0.5 mm.
</paragraph>
</caption>
<paragraph id="DB2636C7CB5FFFA3E6A0FE95E800FE7F" blockId="7.[189,1399,265,397]" pageId="7" pageNumber="40">
GONOPOD. Oval aperture, rim widely raised laterally. Telopodite (
<figureCitation id="43A22A42CB5FFFA3E5A3FE95EE41FED1" box="[958,1028,265,291]" captionStart="Fig" captionStartId="4.[189,232,1961,1987]" captionTargetBox="[189,1366,1332,1937]" captionTargetId="figure-413@4.[189,1399,1332,1937]" captionTargetPageId="4" captionText="Fig. 2. Inodesmus mesibovi sp. nov., male holotype (ICN-MD-2913). A. Ventral view of the gonopods. B. Left gonopod in ventral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14622257" httpUri="https://zenodo.org/record/14622257/files/figure.png" pageId="7" pageNumber="40">Fig. 2</figureCitation>
) long, reaching base of leg pair 5 (
<figureCitation id="43A22A42CB5FFFA3E6C6FEB0EB72FEB4" box="[219,311,300,326]" captionStart="Fig" captionStartId="5.[189,232,1785,1811]" captionTargetBox="[188,1397,532,1759]" captionTargetId="figure-243@5.[172,1414,518,1775]" captionTargetPageId="5" captionText="Fig. 3. Inodesmus mesibovi sp. nov., male holotype (A, CD, HI; ICN-MD-2913) and male paratype (B, EG; MHN-UPN-MD-168). A. Ventrolateral view of head. B. Dorsolateral view of the anterior part of the body. C. Lateral view of the anterior part of the body. D. Midbody sternite. E. Sculpture of the prozonite. F. Ozopore. G. Lateral view of the posterior part of the body. H. Ventral view of telson. I. Spinnerets. Abbreviations: a = anterior part of prozonite; b = posterior part of prozonite; L = limbus; lo = lobe; o = ozopore; r = ridge; s = spherical knobs; t = tooth-like lobe. Scale bars: A, H = 0.2 mm; B, G = 0.5 mm; C = 1 mm; D = 0.1 mm; EF = 0.02 mm; I = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14622259" httpUri="https://zenodo.org/record/14622259/files/figure.png" pageId="7" pageNumber="40">Fig. 3C</figureCitation>
). Basal portion of telopodite with a blunt, basally directed projection arising posteromedially to junction with
<emphasis id="E9EDEAD5CB5FFFA3E79FFED3EBD8FE9B" bold="true" box="[386,413,335,361]" pageId="7" pageNumber="40">cx</emphasis>
, several short setae in basal portion of telopodite, and three large setae in a row on lateral edge of apical tab (
<figureCitation id="43A22A42CB5FFFA3E7F3FEEEE873FE7F" box="[494,566,370,397]" captionStart="Fig" captionStartId="4.[189,232,1961,1987]" captionTargetBox="[189,1366,1332,1937]" captionTargetId="figure-413@4.[189,1399,1332,1937]" captionTargetPageId="4" captionText="Fig. 2. Inodesmus mesibovi sp. nov., male holotype (ICN-MD-2913). A. Ventral view of the gonopods. B. Left gonopod in ventral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14622257" httpUri="https://zenodo.org/record/14622257/files/figure.png" pageId="7" pageNumber="40">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9383654CCB5FFFA3E6A0FE31EE8EFD19" pageId="7" pageNumber="40" type="discussion">
<paragraph id="DB2636C7CB5FFFA3E6A0FE31EB6AFE35" blockId="7.[189,303,429,455]" box="[189,303,429,455]" pageId="7" pageNumber="40">
<heading id="806E81ABCB5FFFA3E6A0FE31EB6AFE35" bold="true" box="[189,303,429,455]" fontSize="11" level="3" pageId="7" pageNumber="40" reason="3">
<emphasis id="E9EDEAD5CB5FFFA3E6A0FE31EB6AFE35" bold="true" box="[189,303,429,455]" pageId="7" pageNumber="40">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="DB2636C7CB5FFFA3E6A0FE47EE8EFD19" blockId="7.[188,1400,475,748]" pageId="7" pageNumber="40">
A notable case of “helicomerism” was observed in a female of
<taxonomicName id="1C994D44CB5FFFA3E5C6FE40EE24FE07" authorityName="Romero-Rincon &amp; Alvear &amp; Martínez-Torres &amp; Robles-Piñeros" authorityYear="2025" box="[987,1121,475,501]" class="Diplopoda" family="Comodesmidae" genus="Inodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="40" phylum="Arthropoda" rank="species" species="mesibovi" status="sp. nov.">
<emphasis id="E9EDEAD5CB5FFFA3E5C6FE40EE24FE07" box="[987,1121,475,501]" italics="true" pageId="7" pageNumber="40">I. mesibovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2DE57AECB5FFFA3E273FE40EE8BFE04" box="[1134,1230,476,502]" pageId="7" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
in which the segmental anomaly breaks the serial arrangement of the body rings (
<bibRefCitation id="BF084B36CB5FFFA3E5FFFE62EF51FDEA" author="Demange J. - M. &amp; Pereira L. A." box="[994,1300,510,536]" pageId="7" pageNumber="40" pagination="261 - 267" refId="ref5886" refString="Demange J. - M. &amp; Pereira L. A. 1980. Deux anomalies segmentaires chez deux especes de geophilomorphes du Perou (Myriapoda: Chilopoda). Senckenberginiana Biologica 60: 261 - 267." type="journal article" year="1980">Demange &amp; Pereira 1980</bibRefCitation>
;
<bibRefCitation id="BF084B36CB5FFFA3E33DFE62EB38FDC9" author="Minelli A. &amp; Pasqual C." pageId="7" pageNumber="40" pagination="135 - 141" refId="ref6746" refString="Minelli A. &amp; Pasqual C. 1986. On some abnormal specimens of centipedes (Chilopoda). Lavori della Societa veneziana di Scienze naturali 11: 135 - 141." type="journal article" year="1986">Minelli &amp; Pasqual 1986</bibRefCitation>
;
<bibRefCitation id="BF084B36CB5FFFA3E794FDBDE8D8FDC9" author="Lesniewska M. &amp; Bonato L. &amp; Minelli A. &amp; Fusco G." box="[393,669,545,572]" pageId="7" pageNumber="40" pagination="417 - 426" refId="ref6535" refString="Lesniewska M., Bonato L., Minelli A. &amp; Fusco G. 2009. Trunk anomalies in the centipede Stigmatogaster subterranea provide insight into late-embryonic segmentation. Arthropod Structure &amp; Development 38: 417 - 426. https: // doi. org / 10.1016 / j. asd. 2009.05.001" type="journal article" year="2009">
Leśniewska
<emphasis id="E9EDEAD5CB5FFFA3E406FDBEE80AFDC9" box="[539,591,545,571]" italics="true" pageId="7" pageNumber="40">et al</emphasis>
. 2009
</bibRefCitation>
). In this specimen (MHN-UPN-MD-177), the tergite of ring 8 is entirely divided into two sections, a smaller isolated portion on the left and one on the right side of the metazonite that is completely fused with ring 7, giving a diagonal shape to the posterior part of the segment (
<figureCitation id="43A22A42CB5FFFA3E72FFD17EBD4FD57" box="[306,401,651,677]" captionStart="Fig" captionStartId="6.[189,232,1925,1951]" captionTargetBox="[231,1366,973,1898]" captionTargetId="figure-343@6.[189,1399,958,1912]" captionTargetPageId="6" captionText="Fig. 4. Teratologies inInodesmus mesibovi sp. nov., female paratype (MHN-UPN-MD-177). A. Anterior part of body. B. Middle part of the body. The numbers 6 and 9 correspond to the body ring number. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14622263" httpUri="https://zenodo.org/record/14622263/files/figure.png" pageId="7" pageNumber="40">Fig. 4B</figureCitation>
). Additionally, another teratological feature is observed in the collum, in which an abnormal longitudinal transverse partition is evident (
<figureCitation id="43A22A42CB5FFFA3E533FD32E9C9FD3A" box="[814,908,686,712]" captionStart="Fig" captionStartId="6.[189,232,1925,1951]" captionTargetBox="[231,1366,973,1898]" captionTargetId="figure-343@6.[189,1399,958,1912]" captionTargetPageId="6" captionText="Fig. 4. Teratologies inInodesmus mesibovi sp. nov., female paratype (MHN-UPN-MD-177). A. Anterior part of body. B. Middle part of the body. The numbers 6 and 9 correspond to the body ring number. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14622263" httpUri="https://zenodo.org/record/14622263/files/figure.png" pageId="7" pageNumber="40">Fig. 4A</figureCitation>
). However, it cannot be dismissed that the damage may have been influenced by external factors beyond the scope of development.
</paragraph>
</subSubSection>
<subSubSection id="9383654CCB5FFFA3E6A0FC8FEE24FCA9" box="[189,1121,787,860]" pageId="7" pageNumber="40" type="distribution">
<paragraph id="DB2636C7CB5FFFA3E6A0FC8FEB13FCDF" blockId="7.[189,342,787,813]" box="[189,342,787,813]" pageId="7" pageNumber="40">
<heading id="806E81ABCB5FFFA3E6A0FC8FEB13FCDF" bold="true" box="[189,342,787,813]" fontSize="11" level="3" pageId="7" pageNumber="40" reason="3">
<emphasis id="E9EDEAD5CB5FFFA3E6A0FC8FEB13FCDF" bold="true" box="[189,342,787,813]" pageId="7" pageNumber="40">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DB2636C7CB5FFFA3E6A0FCDDEE24FCA9" blockId="7.[189,1121,833,860]" box="[189,1121,833,860]" pageId="7" pageNumber="40">
Only known from Parque Natural Chicaque,
<collectingRegion id="195DF825CB5FFFA3E4D1FCDDE93FFCA9" box="[716,890,833,859]" country="Colombia" name="Cundinamarca" pageId="7" pageNumber="40">Cundinamarca</collectingRegion>
,
<collectingCountry id="A38E7657CB5FFFA3E59BFCDDE9BFFCA9" box="[902,1018,833,859]" name="Colombia" pageId="7" pageNumber="40">Colombia</collectingCountry>
(
<figureCitation id="43A22A42CB5FFFA3E217FCDDEE15FCA9" box="[1034,1104,833,859]" captionStart="Fig" captionStartId="11.[189,232,1151,1177]" captionTargetBox="[182,1410,282,1138]" captionTargetId="figure-243@11.[174,1414,265,1142]" captionTargetPageId="11" captionText="Fig. 8. Distribution map of the known species of Inodesmus Cook, 1896 in Colombia." figureDoi="http://doi.org/10.5281/zenodo.14622271" httpUri="https://zenodo.org/record/14622271/files/figure.png" pageId="7" pageNumber="40">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>