185 lines
19 KiB
XML
185 lines
19 KiB
XML
<document id="296A324A2C533F8BAC73349F26E9B1F7" ID-DOI="10.11646/zootaxa.3936.3.10" ID-GBIF-Dataset="61062d56-2d9b-4f53-aca4-6d26a79dfec8" ID-ISSN="1175-5326" ID-Zenodo-Dep="236243" ID-ZooBank="A019200F-65F6-42FF-974C-274BD1213A2C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461016113689" checkinUser="plazi" docAuthor="Schillhammer, Harald" docDate="2015" docId="521C8782620CD63EBE80E67951E99200" docLanguage="en" docName="zt03936p450.pdf" docOrigin="Zootaxa 3936 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gabrius makranczyi Schillhammer, 2015, sp. n." docType="treatment" docVersion="7" lastPageNumber="447" masterDocId="AE25FFFA620DD63FBE17E566500E9501" masterDocTitle="Two new species and additional data of Gabrius Stephens (Coleoptera, Staphylinidae) from the Korean Peninsula" masterLastPageNumber="450" masterPageNumber="446" pageNumber="447" updateTime="1698568845867" updateUser="plazi">
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<mods:title id="43542ECC1282B92B9A2F7532DC2B8CC6">Two new species and additional data of Gabrius Stephens (Coleoptera, Staphylinidae) from the Korean Peninsula</mods:title>
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<mods:namePart id="3D36ED274707F19758CEB17E1E1868D6">Schillhammer, Harald</mods:namePart>
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<mods:title id="FCB9AEBCC263AEED3BDDA1F740FFFF41">Zootaxa</mods:title>
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<mods:date id="BAA7671D9E760461064E6271504FF681">2015</mods:date>
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<mods:number id="1B741E8EB8A7CEEBA36E93DEAA396484">3936</mods:number>
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<mods:identifier id="976E46937BD7BAD33BAF800B99B43BDD" type="DOI">10.11646/zootaxa.3936.3.10</mods:identifier>
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<treatment id="521C8782620CD63EBE80E67951E99200" ID-DOI="http://doi.org/10.5281/zenodo.6122407" ID-GBIF-Taxon="119601693" ID-Zenodo-Dep="6122407" LSID="urn:lsid:plazi:treatment:521C8782620CD63EBE80E67951E99200" httpUri="http://treatment.plazi.org/id/521C8782620CD63EBE80E67951E99200" lastPageNumber="447" pageId="1" pageNumber="447">
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<subSubSection id="92AF651F620CD63EBE80E67951DB963B" box="[151,469,799,826]" pageId="1" pageNumber="447" type="nomenclature">
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<paragraph id="DA0A3694620CD63EBE80E67951DB963B" blockId="1.[151,469,799,826]" box="[151,469,799,826]" pageId="1" pageNumber="447">
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<heading id="814281F8620CD63EBE80E67951DB963B" bold="true" box="[151,469,799,826]" fontSize="11" level="1" pageId="1" pageNumber="447" reason="1">
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<emphasis id="E8C1EA86620CD63EBE80E67951DB963B" bold="true" box="[151,469,799,826]" pageId="1" pageNumber="447">
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<taxonomicName id="1DB54D17620CD63EBE80E67951829638" ID-CoL="6JV4V" box="[151,396,799,825]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="447" phylum="Arthropoda" rank="species" species="makranczyi" status="sp. nov.">
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<emphasis id="E8C1EA86620CD63EBE80E67951829638" bold="true" box="[151,396,799,825]" italics="true" pageId="1" pageNumber="447">Gabrius makranczyi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F3F257FD620CD63EBF84E64651DB963B" box="[403,469,800,826]" pageId="1" pageNumber="447" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="92AF651F620CD63EBE80E603538E96E5" pageId="1" pageNumber="447" type="materials_examined">
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<paragraph id="DA0A3694620CD63EBE80E603538E96E5" blockId="1.[151,1436,869,1793]" pageId="1" pageNumber="447">
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<emphasis id="E8C1EA86620CD63EBE80E60350F1967D" bold="true" box="[151,255,869,892]" pageId="1" pageNumber="447">
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<typeStatus id="050E8836620CD63EBE80E60350F1967D" box="[151,255,869,892]" pageId="1" pageNumber="447" type="holotype">Holotype</typeStatus>
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</emphasis>
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♂: “S-KOREA, Jeollanam-do, Jiri-san, around Nogodan peak, rocky slope at tourist trail with bushes,
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<quantity id="1D4D9B71620CD63EBB5CE6035592967D" box="[1355,1436,869,892]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="447" unit="m" value="1500.0">1500 m</quantity>
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\ soil washing,
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<quantity id="1D4D9B71620CD63EBF57E6EE518A969E" box="[320,388,904,927]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="447" unit="cm" value="20.0">20 cm</quantity>
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deep at rock base under tree (24B),
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<geoCoordinate id="BF815053620CD63EBD09E6EE53AC969E" box="[798,930,904,927]" direction="north" orientation="latitude" pageId="1" pageNumber="447" precision="15" value="35.29333">35°17’36”N</geoCoordinate>
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,
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<geoCoordinate id="BF815053620CD63EBDA7E6EE5431969E" box="[944,1087,904,927]" direction="east" orientation="longitude" pageId="1" pageNumber="447" precision="15" value="127.531944">127°31’55”E</geoCoordinate>
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,
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<date id="AE0B1054620CD63EBA5AE6EE54C7969E" box="[1101,1225,904,927]" pageId="1" pageNumber="447" value="2010-09-15">15.IX.2010</date>
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, Makranczy
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<emphasis id="E8C1EA86620CD63EBB4FE6EE559E969E" box="[1368,1424,903,927]" italics="true" pageId="1" pageNumber="447">et al.</emphasis>
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” (TMB).—
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<typeStatus id="050E8836620CD63EBF16E6CC516196C0" box="[257,367,938,961]" pageId="1" pageNumber="447" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="CCB3FD1D620CD63EBF6BE6CC51A696C0" box="[380,424,938,961]" pageId="1" pageNumber="447" type="generic">1 ♂</specimenCount>
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, with same data as
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<typeStatus id="050E8836620CD63EBC94E6CC52EE96C0" box="[643,736,938,961]" pageId="1" pageNumber="447" type="holotype">holotype</typeStatus>
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(
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<collectionCode id="BCA4AE51620CD63EBCE6E6CC533596C0" box="[753,827,938,961]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="447">NMW</collectionCode>
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);
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<specimenCount id="CCB3FD1D620CD63EBD47E6CC537396C0" box="[848,893,938,961]" pageId="1" pageNumber="447" type="generic">1 ♂</specimenCount>
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: N-KOREA: “
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<collectingCountry id="A2A27604620CD63EBA0AE6CC547096C0" box="[1053,1150,938,961]" name="South Korea" pageId="1" pageNumber="447">KOREA</collectingCountry>
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, no. 692, Kangwon Prov., Mts. Kumgang-san, singled,
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<date id="AE0B1054620CD63EBFDCE6AB524996E5" box="[459,583,973,996]" pageId="1" pageNumber="447" value="1980-09-18">18.IX.1980</date>
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, leg. Tópal & Forró” (TMB).
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</paragraph>
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</subSubSection>
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<subSubSection id="92AF651F620CD63EBED0E69655579398" pageId="1" pageNumber="447" type="diagnosis">
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<paragraph id="DA0A3694620CD63EBED0E69652359128" blockId="1.[151,1436,869,1793]" pageId="1" pageNumber="447">
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<emphasis id="E8C1EA86620CD63EBED0E696513A9106" bold="true" box="[199,308,1008,1031]" pageId="1" pageNumber="447">Diagnosis</emphasis>
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.
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<quantity id="1D4D9B71620CD63EBF53E69651CB9106" box="[324,453,1008,1031]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" metricValueMax="6.3" metricValueMin="5.9" pageId="1" pageNumber="447" unit="mm" value="6.1" valueMax="6.3" valueMin="5.9">5.9–6.3 mm</quantity>
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long (
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<quantity id="1D4D9B71620CD63EBC1AE696526A9106" box="[525,612,1008,1031]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="447" unit="mm" value="3.1">3.1 mm</quantity>
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, abdomen excluded). The diagnosis is based on the
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<typeStatus id="050E8836620CD63EBA8BE69654F29106" box="[1180,1276,1008,1031]" pageId="1" pageNumber="447" type="holotype">holotype</typeStatus>
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, differences in the
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<typeStatus id="050E8836620CD63EBEA9E174512A9128" box="[190,292,1042,1065]" pageId="1" pageNumber="447" type="paratype">paratypes</typeStatus>
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are mentioned separately.
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</paragraph>
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<paragraph id="DA0A3694620CD63EBED0E15353A9916E" blockId="1.[151,1436,869,1793]" pageId="1" pageNumber="447">
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The species hardly differs from
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<taxonomicName id="1DB54D17620CD63EBC0DE1535299914D" box="[538,663,1076,1100]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="447" phylum="Arthropoda" rank="species" species="koreanus">
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<emphasis id="E8C1EA86620CD63EBC0DE1535224914D" box="[538,554,1077,1100]" italics="true" pageId="1" pageNumber="447">G</emphasis>
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.
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<emphasis id="E8C1EA86620CD63EBC20E1525299914D" box="[567,663,1076,1100]" italics="true" pageId="1" pageNumber="447">koreanus</emphasis>
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</taxonomicName>
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both in shape and coloration; head more quadrangular, tempora straight, subparallel, 1.7 times as long as eyes, punctation of elytra slightly denser.
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</paragraph>
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<paragraph id="DA0A3694620CD63EBED0E11C555A901D" blockId="1.[151,1436,869,1793]" pageId="1" pageNumber="447">
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Aedeagus (
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<figureCitation id="428E2A11620CD63EBF56E11C518A9190" box="[321,388,1146,1169]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,268,2008,2031]" captionTargetBox="[158,1429,293,1964]" captionTargetId="figure@2.[158,1429,263,1983]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Aedeagus of 1) Gabrius koreanus; 2) G. makranczyi. Ventral aspect (a), lateral aspect (b), paramere (c, d)." httpUri="https://zenodo.org/record/236244/files/figure.png" pageId="1" pageNumber="447">Fig. 2</figureCitation>
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) with apical portion of median lobe similar to that of
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<taxonomicName id="1DB54D17620CD63EBDC8E11C546E9190" box="[991,1120,1145,1169]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="447" phylum="Arthropoda" rank="species" species="koreanus">
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<emphasis id="E8C1EA86620CD63EBDC8E11C53E19190" box="[991,1007,1146,1169]" italics="true" pageId="1" pageNumber="447">G</emphasis>
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.
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<emphasis id="E8C1EA86620CD63EBDE8E11F546E9190" box="[1023,1120,1145,1169]" italics="true" pageId="1" pageNumber="447">koreanus</emphasis>
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</taxonomicName>
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, but sides bisinuate and less distinctly narrowed towards apex; in lateral view (
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<figureCitation id="428E2A11620CD63EBCDFE1FB530291B5" box="[712,780,1181,1204]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,268,2008,2031]" captionTargetBox="[158,1429,293,1964]" captionTargetId="figure@2.[158,1429,263,1983]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Aedeagus of 1) Gabrius koreanus; 2) G. makranczyi. Ventral aspect (a), lateral aspect (b), paramere (c, d)." httpUri="https://zenodo.org/record/236244/files/figure.png" pageId="1" pageNumber="447">Fig. 2</figureCitation>
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b) with apical portion markedly bent dorsad, especially the sharply pointed apex. Paramere (
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<figureCitation id="428E2A11620CD63EBC16E1A6524791D6" box="[513,585,1216,1239]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,268,2008,2031]" captionTargetBox="[158,1429,293,1964]" captionTargetId="figure@2.[158,1429,263,1983]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Aedeagus of 1) Gabrius koreanus; 2) G. makranczyi. Ventral aspect (a), lateral aspect (b), paramere (c, d)." httpUri="https://zenodo.org/record/236244/files/figure.png" pageId="1" pageNumber="447">Figs 2</figureCitation>
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c, d) hardly widened distally, with a much narrower apical emargination; peg setae similarly arranged but more numerous, basad reaching almost base of apical portion of paramere; apical margin bearing the same thick setae as in
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<taxonomicName id="1DB54D17620CD63EBC15E063528F901D" box="[514,641,1284,1308]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="447" phylum="Arthropoda" rank="species" species="koreanus">
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<emphasis id="E8C1EA86620CD63EBC15E063521C901D" box="[514,530,1285,1308]" italics="true" pageId="1" pageNumber="447">G</emphasis>
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.
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<emphasis id="E8C1EA86620CD63EBC37E062528F901D" box="[544,641,1284,1308]" italics="true" pageId="1" pageNumber="447">koreanus</emphasis>
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</taxonomicName>
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, smaller setae situated almost at base of apical portion of paramere.
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</paragraph>
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<paragraph id="DA0A3694620CD63EBED0E04E5189903E" blockId="1.[151,1436,869,1793]" box="[199,391,1320,1343]" pageId="1" pageNumber="447">Female unknown.</paragraph>
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<paragraph id="DA0A3694620CD63EBED0E02C54D190ED" blockId="1.[151,1436,869,1793]" pageId="1" pageNumber="447">
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<emphasis id="E8C1EA86620CD63EBED0E02C51319060" bold="true" box="[199,319,1354,1377]" pageId="1" pageNumber="447">Variability</emphasis>
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. The slightly teneral
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<typeStatus id="050E8836620CD63EBC3FE02C528D9060" box="[552,643,1354,1377]" pageId="1" pageNumber="447" type="paratype">paratype</typeStatus>
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from
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<collectingCountry id="A2A27604620CD63EBCD0E02C53419060" box="[711,847,1354,1377]" name="South Korea" pageId="1" pageNumber="447">South Korea</collectingCountry>
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looks somewhat different from the syntopic
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<typeStatus id="050E8836620CD63EBB20E02C55979060" box="[1335,1433,1354,1377]" pageId="1" pageNumber="447" type="holotype">holotype</typeStatus>
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: head more distinctly rounded, tempora slightly convex, slightly widened posteriad, sides of pronotum distinctly convex. Paramere (
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<figureCitation id="428E2A11620CD63EBF1AE0F6515E90A6" box="[269,336,1424,1447]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,268,2008,2031]" captionTargetBox="[158,1429,293,1964]" captionTargetId="figure@2.[158,1429,263,1983]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Aedeagus of 1) Gabrius koreanus; 2) G. makranczyi. Ventral aspect (a), lateral aspect (b), paramere (c, d)." httpUri="https://zenodo.org/record/236244/files/figure.png" pageId="1" pageNumber="447">Fig. 2</figureCitation>
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c) with deeper and somewhat narrower apical emargination. Experience has shown that, despite the interspecific similarity,
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<taxonomicName id="1DB54D17620CD63EBF8EE0D751FE90C8" box="[409,496,1457,1481]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="447" phylum="Arthropoda" rank="genus">
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<emphasis id="E8C1EA86620CD63EBF8EE0D751FE90C8" box="[409,496,1457,1481]" italics="true" pageId="1" pageNumber="447">Gabrius</emphasis>
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</taxonomicName>
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species exhibit a sometimes remarkable intraspecific variability (e.g.
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<taxonomicName id="1DB54D17620CD63EBAE4E0D4554990C8" box="[1267,1351,1458,1481]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="450" phylum="Arthropoda" rank="genus">Gabrius</taxonomicName>
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astutus group). Therefore, I have no doubt that the above mentioned
|
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<typeStatus id="050E8836620CD63EBD35E0B3537090ED" box="[802,894,1493,1516]" pageId="1" pageNumber="447" type="paratype">paratype</typeStatus>
|
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is conspecific with the
|
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<typeStatus id="050E8836620CD63EBA6DE0B354D590ED" box="[1146,1243,1493,1516]" pageId="1" pageNumber="447" type="holotype">holotype</typeStatus>
|
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.
|
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</paragraph>
|
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<paragraph id="DA0A3694620CD63EBED0E09E55579398" blockId="1.[151,1436,869,1793]" pageId="1" pageNumber="447">
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<emphasis id="E8C1EA86620CD63EBED0E09E51AA930E" bold="true" box="[199,420,1528,1551]" pageId="1" pageNumber="447">Systematic position</emphasis>
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. This and the previous species share some characters with the
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<taxonomicName id="1DB54D17620CD63EBA5BE0915505930E" box="[1100,1291,1527,1551]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="447" phylum="Arthropoda" rank="species" species="osseticus">
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<emphasis id="E8C1EA86620CD63EBA5BE0915505930E" box="[1100,1291,1527,1551]" italics="true" pageId="1" pageNumber="447">Gabrius osseticus</emphasis>
|
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</taxonomicName>
|
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group and to some extent also with the
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<taxonomicName id="1DB54D17620CD63EBFBAE37F52619330" box="[429,623,1561,1585]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="447" phylum="Arthropoda" rank="species" species="femoralis">
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<emphasis id="E8C1EA86620CD63EBFBAE37F52619330" box="[429,623,1561,1585]" italics="true" pageId="1" pageNumber="447">Gabrius femoralis</emphasis>
|
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</taxonomicName>
|
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group, in particular the shape of the male sternite VIII with extensive semimembranous extension, but also the shape of the paramere. However, the large apical setae of the paramere, which are small or not even developed in those groups, would render the two new species rather strange elements. I will refrain from introducing another species group name until the relationships among
|
||
<taxonomicName id="1DB54D17620CD63EBDA8E3E754189398" box="[959,1046,1665,1689]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E8C1EA86620CD63EBDA8E3E754189398" box="[959,1046,1665,1689]" italics="true" pageId="1" pageNumber="447">Gabrius</emphasis>
|
||
</taxonomicName>
|
||
species are better understood.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92AF651F620CD63EBED0E3C354C693BD" box="[199,1224,1701,1724]" pageId="1" pageNumber="447" type="distribution">
|
||
<paragraph id="DA0A3694620CD63EBED0E3C354C693BD" blockId="1.[151,1436,869,1793]" box="[199,1224,1701,1724]" pageId="1" pageNumber="447">
|
||
<emphasis id="E8C1EA86620CD63EBED0E3C3515C93BD" bold="true" box="[199,338,1701,1724]" pageId="1" pageNumber="447">Distribution</emphasis>
|
||
. The species is at present known from one locality each in North and
|
||
<collectingCountry id="A2A27604620CD63EBA2CE3C354CA93BD" box="[1083,1220,1701,1724]" name="South Korea" pageId="1" pageNumber="447">South Korea</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92AF651F620CD63EBED0E3AE51E99200" pageId="1" pageNumber="447" type="etymology">
|
||
<paragraph id="DA0A3694620CD63EBED0E3AE51E99200" blockId="1.[151,1436,869,1793]" pageId="1" pageNumber="447">
|
||
<emphasis id="E8C1EA86620CD63EBED0E3AE514E93DE" bold="true" box="[199,320,1736,1759]" pageId="1" pageNumber="447">Etymology</emphasis>
|
||
. The species is named after György Makranczy (Budapest), specialist in
|
||
<taxonomicName id="1DB54D17620CD63EBA5CE3AE54CD93DE" box="[1099,1219,1736,1759]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="447" phylum="Arthropoda" rank="subFamily" subFamily="Oxytelinae">Oxytelinae</taxonomicName>
|
||
, who collected most specimens of both new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |