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<document id="25FC0CD7A45C73E212CF20A73335E0FD" ID-CLB-Dataset="29502" ID-DOI="10.11646/zootaxa.4484.1.1" ID-GBIF-Dataset="74b777c9-eae0-4770-8c86-dcbb10fb06b3" ID-ISSN="1175-5326" ID-Zenodo-Dep="1437848" ID-ZooBank="D66524CF-9C6D-4DF4-8CA2-B2C9708CF5FD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538125412450" checkinUser="plazi" docAuthor="Grischenko, Andrei V., Gordon, Dennis P. &amp; Melnik, Viacheslav P." docDate="2018" docId="521587E4562E551809EEF8FA884FF85C" docLanguage="en" docName="zootaxa.4484.1.1.pdf" docOrigin="Zootaxa 4484 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Frontohornera frontalis Grischenko &amp; Gordon &amp; Melnik 2018, n. sp." docType="treatment" docVersion="8" lastPageNumber="32" masterDocId="AE2CFF9C563255070979FF888A4AFF87" masterDocTitle="Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining" masterLastPageNumber="91" masterPageNumber="1" pageNumber="29" updateTime="1698557051871" updateUser="ExternalLinkService">
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<mods:title id="77E92BD7A068AEB56DCA19C09CA02871">Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining</mods:title>
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<mods:namePart id="39C8C38D9E52F14264B233C2FCEBBB70">Grischenko, Andrei V.</mods:namePart>
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<mods:namePart id="E899706C1F0E154EE87E1442DA7B0E86">Gordon, Dennis P.</mods:namePart>
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<mods:namePart id="D02A1177D0ECA00A388B5603539B51D3">Melnik, Viacheslav P.</mods:namePart>
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<treatment id="521587E4562E551809EEF8FA884FF85C" ID-DOI="http://doi.org/10.5281/zenodo.5989799" ID-GBIF-Taxon="148403713" ID-Zenodo-Dep="5989799" LSID="urn:lsid:plazi:treatment:521587E4562E551809EEF8FA884FF85C" httpUri="http://treatment.plazi.org/id/521587E4562E551809EEF8FA884FF85C" lastPageId="31" lastPageNumber="32" pageId="28" pageNumber="29">
<subSubSection id="92A66579562E551B09EEF8FA8B25F829" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="DA0336F2562E551B09EEF8FA884EF80A" blockId="28.[151,516,1906,1966]" box="[151,516,1906,1933]" pageId="28" pageNumber="29">
<heading id="814B819E562E551B09EEF8FA884EF80A" bold="true" box="[151,516,1906,1933]" fontSize="11" level="1" pageId="28" pageNumber="29" reason="1">
<emphasis id="E8C8EAE0562E551B09EEF8FA884EF80A" bold="true" box="[151,516,1906,1933]" pageId="28" pageNumber="29">
<taxonomicName id="1DBC4D71562E551B09EEF8FA8BF1F80B" ID-CoL="d57f2ee1-cee9-461d-a0f7-73c72c96d9fc" authority="Grischenko &amp; Gordon &amp; Melnik, 2018" authorityName="Grischenko &amp; Gordon &amp; Melnik" authorityYear="2018" box="[151,443,1906,1932]" class="Stenolaemata" family="Horneridae" genus="Frontohornera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="28" pageNumber="29" phylum="Bryozoa" rank="species" species="frontalis" status="sp. nov.">
<emphasis id="E8C8EAE0562E551B09EEF8FA8BF1F80B" bold="true" box="[151,443,1906,1932]" italics="true" pageId="28" pageNumber="29">Frontohornera frontalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3FB579B562E551B08BBF8FB884EF80A" box="[450,516,1907,1933]" pageId="28" pageNumber="29" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="DA0336F2562E551B09EEF81D8B25F829" blockId="28.[151,516,1906,1966]" box="[151,367,1941,1966]" pageId="28" pageNumber="29">
(
<figureCitation id="42872A77562E551B09E6F81D8B5FF82A" box="[159,277,1941,1966]" captionStart="FIGURE 2" captionStartId="10.[151,250,1774,1796]" captionTargetBox="[151,1436,392,1715]" captionTargetId="figure@10.[151,1436,392,1715]" captionTargetPageId="10" captionText="FIGURE 2. Colonies of some cyclostome bryozoans, in vivo, attached to polymetallic nodules. AE, Pandanipora helix n. gen., n. sp.: A, specimen GLD409, Stn 190; B, specimen GLD412, Stn 262; C, specimen YMG407, Stn 143; D, specimen YMG413, Stn 295; E, specimen GLD 411, Stn 212. F, Tubuliporina sp. indet., specimen YMG1801, Stn 7. G, H, Abyssoecia elevata n. gen., n. sp.: G, specimen GLD409, Stn 196; H, specimen GLD409, Stn 191. I, Discantenna metallica n. sp.: specimen GLD411, Stn 224. J, K, Frontohornera frontalis n. gen., n. sp.: J, specimen YMG407, Stn 124; K, specimen GLD411, Stn 210. L, Alyonushka hystricosa n. gen., n. sp.: specimen GLD409, Stn 199. M, Calyssopora volcano n. gen., n. sp.: specimen YMG1801, Stn 33. N, O, Anyuta anastema n. gen., n. sp.: N, specimen GLD409, Stn 180; O, specimen YMG406, Stn 71. Scale bars: 1 mm." httpUri="https://zenodo.org/record/1437852/files/figure.png" pageId="28" pageNumber="29">Figs 2J, K</figureCitation>
,
<figureCitation id="42872A77562E551B0859F81D8B2DF829" box="[288,359,1941,1966]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStart-2="FIGURE 18" captionStartId-0="29.[151,250,1953,1975]" captionStartId-1="30.[151,250,1724,1746]" captionStartId-2="31.[151,250,1360,1382]" captionTargetBox-0="[151,1436,877,1931]" captionTargetBox-1="[151,1436,193,1703]" captionTargetBox-2="[151,1436,193,1339]" captionTargetId-0="figure@29.[151,1436,877,1931]" captionTargetId-1="figure@30.[151,1436,193,1703]" captionTargetId-2="figure@31.[151,1436,193,1339]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionTargetPageId-2="31" captionText-0="FIGURE 16. Frontohornera frontalis n. gen., n. sp. Holotype specimen, ZIRAS 1/50707, respectively seen in A, frontal and B, abfrontal views. Scale bars: 1 mm." captionText-1="FIGURE 17. Frontohornera frontalis n. gen., n. sp. Holotype, ZIRAS 1/50707, showing details of morphology. A, part of branch distal to gonozooid, showing cancelli and zooidal peristomes; B, part of infertile branch with fewer cancelli; C, D, parts of infertile branch showing peristomial dimorphism; E, part of branch near gonozooid showing numerous cancelli and peristomial dimorphism; F, zooids showing peristomial and apertural dimorphism; G, abfrontal part of branch in vicinity of gonozooid; H, J, K, gonozooid profiles; I, autozooidal peristome showing interior spinules; L, ooeciostome and ooeciopore. Scale bars: A, B, D, E, G, 300 µm; C, H, J, K, 200 µm; F, 150 µm; I, 100 µm; L, 50 µm." captionText-2="FIGURE 18. Frontohornera frontalis n. gen., n. sp. Progressive stages of development of young colonies. AC, specimen GLD411, Stn 215, from different angles; DF, specimen YMG414, Stn 334, from different angles; GI, specimen GLD411, Stn 210, in frontal and abfrontal views and a close-up of the colony base with secondary calcification. Scale bars: AH, 500 µm; I, 250 µm." httpUri-0="https://zenodo.org/record/1437881/files/figure.png" httpUri-1="https://zenodo.org/record/1437883/files/figure.png" httpUri-2="https://zenodo.org/record/1437885/files/figure.png" pageId="28" pageNumber="29">1618</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="92A66579562E551A09EEF8558BA5FE9A" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="DA0336F2562E551B09EEF8558FD3F871" blockId="28.[151,1433,2013,2038]" box="[151,1433,2013,2038]" pageId="28" pageNumber="29">
<emphasis id="E8C8EAE0562E551B09EEF8558BCFF871" bold="true" box="[151,389,2013,2038]" pageId="28" pageNumber="29">Material examined.</emphasis>
<emphasis id="E8C8EAE0562E551B08EBF8558BB2F871" box="[402,504,2013,2038]" italics="true" pageId="28" pageNumber="29">Holotype</emphasis>
: ZIRAS 1/50707, colony detached from nodule, YMG R.V.
<emphasis id="E8C8EAE0562E551B0DB4F8558FD3F871" box="[1229,1433,2013,2038]" italics="true" pageId="28" pageNumber="29">Yuzhmorgeologiya</emphasis>
</paragraph>
<paragraph id="DA0336F2562F551A09EEFF1F8BA5FE9A" blockId="29.[151,1436,151,824]" pageId="29" pageNumber="30">
cruise YMG413, Stn 292,
<date id="AE021032562F551A08B5FF108815FF37" box="[460,607,151,176]" pageId="29" pageNumber="30" value="2014-07-27">27 July 2014</date>
, 13.03353° N, 132.49292° W,
<quantity id="1D449B17562F551A0ACFFF108E44FF28" box="[950,1038,152,176]" metricMagnitude="3" metricUnit="m" metricValue="4.829" pageId="29" pageNumber="30" unit="m" value="4829.0">4829 m</quantity>
.
<emphasis id="E8C8EAE0562F551A0D60FF118ECAFF37" box="[1049,1152,153,176]" italics="true" pageId="29" pageNumber="30">Paratype</emphasis>
: ZIRAS 2/50708, colony attached to nodule particle, YMG R.V.
<emphasis id="E8C8EAE0562F551A0B0BFF3488BAFF52" box="[626,752,188,213]" italics="true" pageId="29" pageNumber="30">Gelendzhik</emphasis>
cruise GLD411, Stn 210,
<date id="AE021032562F551A0D3BFF348EA4FF53" box="[1090,1262,188,213]" pageId="29" pageNumber="30" value="2012-04-20">20 April 2012</date>
, 13.35817° N, 133.27183° W,
<quantity id="1D449B17562F551A0831FF578BE8FF70" box="[328,418,223,248]" metricMagnitude="3" metricUnit="m" metricValue="4.8149999999999995" pageId="29" pageNumber="30" unit="m" value="4815.0">4815 m</quantity>
.
<emphasis id="E8C8EAE0562F551A08D4FF5788C1FF7F" box="[429,651,223,248]" italics="true" pageId="29" pageNumber="30">Additional material</emphasis>
: YMG407, Stn 124; GLD411, Stn 215; YMG414, Stns 329, 334. Total specimens examined six.
</paragraph>
</subSubSection>
<subSubSection id="92A66579562F551A09BEFEAF8E7DFEC7" box="[199,1079,295,320]" pageId="29" pageNumber="30" type="etymology">
<paragraph id="DA0336F2562F551A09BEFEAF8E7DFEC7" blockId="29.[151,1436,151,824]" box="[199,1079,295,320]" pageId="29" pageNumber="30">
<emphasis id="E8C8EAE0562F551A09BEFEAF8B04FEC7" bold="true" box="[199,334,295,320]" pageId="29" pageNumber="30">Etymology.</emphasis>
Latin,
<taxonomicName id="1DBC4D71562F551A08E4FEAF8BB1FEC7" authorityName="Grischenko &amp; Gordon &amp; Melnik" authorityYear="2018" box="[413,507,295,320]" class="Stenolaemata" family="Horneridae" genus="Frontohornera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="29" pageNumber="30" phylum="Bryozoa" rank="species" species="frontalis">
<emphasis id="E8C8EAE0562F551A08E4FEAF8BB1FEC7" box="[413,507,295,320]" italics="true" pageId="29" pageNumber="30">frontalis</emphasis>
</taxonomicName>
, frontal, alluding to the frontally borne gonozooid.
</paragraph>
</subSubSection>
<subSubSection id="92A66579562F551809BEFEC489A9F8CB" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30" type="description">
<paragraph id="DA0336F2562F551A09BEFEC48883FD90" blockId="29.[151,1436,151,824]" pageId="29" pageNumber="30">
<emphasis id="E8C8EAE0562F551A09BEFEC48B13FEE2" bold="true" box="[199,345,332,357]" pageId="29" pageNumber="30">Description.</emphasis>
Colony erect, white, to
<quantity id="1D449B17562F551A0B1FFEC48880FEE3" box="[614,714,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="2.81" pageId="29" pageNumber="30" unit="mm" value="2.81">2.81 mm</quantity>
high,
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broad, dichotomously branching once in a single plane (
<figureCitation id="42872A77562F551A099DFEE78B25FE0F" box="[228,367,367,392]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[151,1436,877,1931]" captionTargetId="figure@29.[151,1436,877,1931]" captionTargetPageId="29" captionText="FIGURE 16. Frontohornera frontalis n. gen., n. sp. Holotype specimen, ZIRAS 1/50707, respectively seen in A, frontal and B, abfrontal views. Scale bars: 1 mm." httpUri="https://zenodo.org/record/1437881/files/figure.png" pageId="29" pageNumber="30">Fig. 16A, B</figureCitation>
), or with short additional bifurcation to produce a third, short branch near colony center (
<figureCitation id="42872A77562F551A0C16FEE78AA4FE2B" captionStart="FIGURE 18" captionStartId="31.[151,250,1360,1382]" captionTargetBox="[151,1436,193,1339]" captionTargetId="figure@31.[151,1436,193,1339]" captionTargetPageId="31" captionText="FIGURE 18. Frontohornera frontalis n. gen., n. sp. Progressive stages of development of young colonies. AC, specimen GLD411, Stn 215, from different angles; DF, specimen YMG414, Stn 334, from different angles; GI, specimen GLD411, Stn 210, in frontal and abfrontal views and a close-up of the colony base with secondary calcification. Scale bars: AH, 500 µm; I, 250 µm." httpUri="https://zenodo.org/record/1437885/files/figure.png" pageId="29" pageNumber="30">Fig. 18G, H</figureCitation>
). Stem diameter 0.32 ×
<quantity id="1D449B17562F551A0B76FE1C8833FE2B" box="[527,633,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="29" pageNumber="30" unit="mm" value="0.37">0.37 mm</quantity>
, widening to colony base (
<quantity id="1D449B17562F551A0AC3FE1C8E2DFE2B" box="[954,1127,404,429]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" metricValueMax="7.6" metricValueMin="6.6" pageId="29" pageNumber="30" unit="mm" value="0.71" valueMax="0.76" valueMin="0.66">0.660.76 mm</quantity>
). With distinct frontal and abfrontal faces (
<figureCitation id="42872A77562F551A0837FE3F8BE8FE57" box="[334,418,439,464]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[151,1436,877,1931]" captionTargetId="figure@29.[151,1436,877,1931]" captionTargetPageId="29" captionText="FIGURE 16. Frontohornera frontalis n. gen., n. sp. Holotype specimen, ZIRAS 1/50707, respectively seen in A, frontal and B, abfrontal views. Scale bars: 1 mm." httpUri="https://zenodo.org/record/1437881/files/figure.png" pageId="29" pageNumber="30">Fig. 16</figureCitation>
), entire surface interior-walled, textured irregularly with minute dimples, granules, ridges and furrows (
<figureCitation id="42872A77562F551A0848FE548BF4FE73" box="[305,446,476,501]" captionStart="FIGURE 17" captionStartId="30.[151,250,1724,1746]" captionTargetBox="[151,1436,193,1703]" captionTargetId="figure@30.[151,1436,193,1703]" captionTargetPageId="30" captionText="FIGURE 17. Frontohornera frontalis n. gen., n. sp. Holotype, ZIRAS 1/50707, showing details of morphology. A, part of branch distal to gonozooid, showing cancelli and zooidal peristomes; B, part of infertile branch with fewer cancelli; C, D, parts of infertile branch showing peristomial dimorphism; E, part of branch near gonozooid showing numerous cancelli and peristomial dimorphism; F, zooids showing peristomial and apertural dimorphism; G, abfrontal part of branch in vicinity of gonozooid; H, J, K, gonozooid profiles; I, autozooidal peristome showing interior spinules; L, ooeciostome and ooeciopore. Scale bars: A, B, D, E, G, 300 µm; C, H, J, K, 200 µm; F, 150 µm; I, 100 µm; L, 50 µm." httpUri="https://zenodo.org/record/1437883/files/figure.png" pageId="29" pageNumber="30">Fig. 17AG</figureCitation>
), with longitudinal linearities only on stem of colony; conspicuous ribbing (nervi) and grooves (sulci) lacking. Pores simple, tiny, sparse.
</paragraph>
<paragraph id="DA0336F2562F551A09BEFDAC88CEFCBF" blockId="29.[151,1436,151,824]" pageId="29" pageNumber="30">
Autozooids in 24 series, their peristomes produced frontally upwards, relatively short along frontolateral and central parts of branch, longest along abfrontolateral margin, with simple circular apertures, no apertural spines (
<figureCitation id="42872A77562F551A09E6FDE48B73FD03" box="[159,313,620,645]" captionStart="FIGURE 17" captionStartId="30.[151,250,1724,1746]" captionTargetBox="[151,1436,193,1703]" captionTargetId="figure@30.[151,1436,193,1703]" captionTargetPageId="30" captionText="FIGURE 17. Frontohornera frontalis n. gen., n. sp. Holotype, ZIRAS 1/50707, showing details of morphology. A, part of branch distal to gonozooid, showing cancelli and zooidal peristomes; B, part of infertile branch with fewer cancelli; C, D, parts of infertile branch showing peristomial dimorphism; E, part of branch near gonozooid showing numerous cancelli and peristomial dimorphism; F, zooids showing peristomial and apertural dimorphism; G, abfrontal part of branch in vicinity of gonozooid; H, J, K, gonozooid profiles; I, autozooidal peristome showing interior spinules; L, ooeciostome and ooeciopore. Scale bars: A, B, D, E, G, 300 µm; C, H, J, K, 200 µm; F, 150 µm; I, 100 µm; L, 50 µm." httpUri="https://zenodo.org/record/1437883/files/figure.png" pageId="29" pageNumber="30">Fig. 17AF, I</figureCitation>
). Peristomes somewhat dimorphic (
<figureCitation id="42872A77562F551A0BB4FDE4897AFD03" box="[717,816,620,645]" captionStart="FIGURE 17" captionStartId="30.[151,250,1724,1746]" captionTargetBox="[151,1436,193,1703]" captionTargetId="figure@30.[151,1436,193,1703]" captionTargetPageId="30" captionText="FIGURE 17. Frontohornera frontalis n. gen., n. sp. Holotype, ZIRAS 1/50707, showing details of morphology. A, part of branch distal to gonozooid, showing cancelli and zooidal peristomes; B, part of infertile branch with fewer cancelli; C, D, parts of infertile branch showing peristomial dimorphism; E, part of branch near gonozooid showing numerous cancelli and peristomial dimorphism; F, zooids showing peristomial and apertural dimorphism; G, abfrontal part of branch in vicinity of gonozooid; H, J, K, gonozooid profiles; I, autozooidal peristome showing interior spinules; L, ooeciostome and ooeciopore. Scale bars: A, B, D, E, G, 300 µm; C, H, J, K, 200 µm; F, 150 µm; I, 100 µm; L, 50 µm." httpUri="https://zenodo.org/record/1437883/files/figure.png" pageId="29" pageNumber="30">Fig. 17F</figureCitation>
), short and uniformly tubular with circular apertures in central face of branches, but strongly elongated, with aperture having shape of teardrop, isosceles triangle or irregular oval along branch margins. Peristome wall interiors with short, thin spinules proximal to aperture (
<figureCitation id="42872A77562F551A0C16FD3C8A8AFD77" captionStart="FIGURE 17" captionStartId="30.[151,250,1724,1746]" captionTargetBox="[151,1436,193,1703]" captionTargetId="figure@30.[151,1436,193,1703]" captionTargetPageId="30" captionText="FIGURE 17. Frontohornera frontalis n. gen., n. sp. Holotype, ZIRAS 1/50707, showing details of morphology. A, part of branch distal to gonozooid, showing cancelli and zooidal peristomes; B, part of infertile branch with fewer cancelli; C, D, parts of infertile branch showing peristomial dimorphism; E, part of branch near gonozooid showing numerous cancelli and peristomial dimorphism; F, zooids showing peristomial and apertural dimorphism; G, abfrontal part of branch in vicinity of gonozooid; H, J, K, gonozooid profiles; I, autozooidal peristome showing interior spinules; L, ooeciostome and ooeciopore. Scale bars: A, B, D, E, G, 300 µm; C, H, J, K, 200 µm; F, 150 µm; I, 100 µm; L, 50 µm." httpUri="https://zenodo.org/record/1437883/files/figure.png" pageId="29" pageNumber="30">
Fig.
<date id="AE021032562F551A09EEFD508A8AFD77" box="[151,192,728,752]" pageId="29" pageNumber="30" value="2017-01">17I</date>
</figureCitation>
). Cancelli most numerous on central part of branch distal to gonozooid (
<figureCitation id="42872A77562F551A0A9EFD5F8E18FD77" box="[999,1106,727,752]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[151,1436,877,1931]" captionTargetId="figure@29.[151,1436,877,1931]" captionTargetPageId="29" captionText="FIGURE 16. Frontohornera frontalis n. gen., n. sp. Holotype specimen, ZIRAS 1/50707, respectively seen in A, frontal and B, abfrontal views. Scale bars: 1 mm." httpUri="https://zenodo.org/record/1437881/files/figure.png" pageId="29" pageNumber="30">Figs 16A</figureCitation>
,
<figureCitation id="42872A77562F551A0D24FD508E8DFD77" box="[1117,1223,728,752]" captionStart="FIGURE 17" captionStartId="30.[151,250,1724,1746]" captionTargetBox="[151,1436,193,1703]" captionTargetId="figure@30.[151,1436,193,1703]" captionTargetPageId="30" captionText="FIGURE 17. Frontohornera frontalis n. gen., n. sp. Holotype, ZIRAS 1/50707, showing details of morphology. A, part of branch distal to gonozooid, showing cancelli and zooidal peristomes; B, part of infertile branch with fewer cancelli; C, D, parts of infertile branch showing peristomial dimorphism; E, part of branch near gonozooid showing numerous cancelli and peristomial dimorphism; F, zooids showing peristomial and apertural dimorphism; G, abfrontal part of branch in vicinity of gonozooid; H, J, K, gonozooid profiles; I, autozooidal peristome showing interior spinules; L, ooeciostome and ooeciopore. Scale bars: A, B, D, E, G, 300 µm; C, H, J, K, 200 µm; F, 150 µm; I, 100 µm; L, 50 µm." httpUri="https://zenodo.org/record/1437883/files/figure.png" pageId="29" pageNumber="30">17A, E, J</figureCitation>
), otherwise sparse, shallow, of varying size. Abfrontal face of colony with outlines of autozooidal tubes (
<figureCitation id="42872A77562F551A0DE2FD748F41FC92" box="[1179,1291,764,789]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[151,1436,877,1931]" captionTargetId="figure@29.[151,1436,877,1931]" captionTargetPageId="29" captionText="FIGURE 16. Frontohornera frontalis n. gen., n. sp. Holotype specimen, ZIRAS 1/50707, respectively seen in A, frontal and B, abfrontal views. Scale bars: 1 mm." httpUri="https://zenodo.org/record/1437881/files/figure.png" pageId="29" pageNumber="30">Figs 16B</figureCitation>
,
<figureCitation id="42872A77562F551A0C65FD748F05FC93" box="[1308,1359,764,788]" captionStart="FIGURE 17" captionStartId="30.[151,250,1724,1746]" captionTargetBox="[151,1436,193,1703]" captionTargetId="figure@30.[151,1436,193,1703]" captionTargetPageId="30" captionText="FIGURE 17. Frontohornera frontalis n. gen., n. sp. Holotype, ZIRAS 1/50707, showing details of morphology. A, part of branch distal to gonozooid, showing cancelli and zooidal peristomes; B, part of infertile branch with fewer cancelli; C, D, parts of infertile branch showing peristomial dimorphism; E, part of branch near gonozooid showing numerous cancelli and peristomial dimorphism; F, zooids showing peristomial and apertural dimorphism; G, abfrontal part of branch in vicinity of gonozooid; H, J, K, gonozooid profiles; I, autozooidal peristome showing interior spinules; L, ooeciostome and ooeciopore. Scale bars: A, B, D, E, G, 300 µm; C, H, J, K, 200 µm; F, 150 µm; I, 100 µm; L, 50 µm." httpUri="https://zenodo.org/record/1437883/files/figure.png" pageId="29" pageNumber="30">17G</figureCitation>
), their longitudinal contours converging proximad.
</paragraph>
<caption id="8EC3667A562F551A09EEF829885EF852" httpUri="https://zenodo.org/record/1437881/files/figure.png" pageId="29" pageNumber="30" startId="29.[151,250,1953,1975]" targetBox="[151,1436,877,1931]" targetPageId="29">
<paragraph id="DA0336F2562F551A09EEF829885EF852" blockId="29.[151,1435,1953,2005]" pageId="29" pageNumber="30">
<emphasis id="E8C8EAE0562F551A09EEF8298B6AF830" bold="true" box="[151,288,1953,1975]" pageId="29" pageNumber="30">FIGURE 16.</emphasis>
<taxonomicName id="1DBC4D71562F551A085EF8298850F830" authority="Grischenko &amp; Gordon &amp; Melnik, 2018" authorityName="Grischenko &amp; Gordon &amp; Melnik" authorityYear="2018" box="[295,538,1953,1975]" class="Stenolaemata" family="Horneridae" genus="Frontohornera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="29" pageNumber="30" phylum="Bryozoa" rank="species" species="frontalis" status="gen. nov.">
<emphasis id="E8C8EAE0562F551A085EF8298850F830" box="[295,538,1953,1975]" italics="true" pageId="29" pageNumber="30">Frontohornera frontalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3FB579B562F551A0B58F82988FBF830" box="[545,689,1953,1975]" pageId="29" pageNumber="30" rank="species">
<emphasis id="E8C8EAE0562F551A0B58F8298820F830" bold="true" box="[545,618,1953,1975]" pageId="29" pageNumber="30">n. gen.</emphasis>
,
<emphasis id="E8C8EAE0562F551A0B0EF82988FBF830" bold="true" box="[631,689,1953,1975]" pageId="29" pageNumber="30">n. sp.</emphasis>
</taxonomicNameLabel>
Holotype specimen, ZIRAS 1/50707, respectively seen in
<emphasis id="E8C8EAE0562F551A0C74F8298F55F830" bold="true" box="[1293,1311,1953,1975]" pageId="29" pageNumber="30">A</emphasis>
, frontal and
<emphasis id="E8C8EAE0562F551A09EEF8378AE2F852" bold="true" box="[151,168,1983,2005]" pageId="29" pageNumber="30">B</emphasis>
, abfrontal views. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="8EC3667A562C551909EEF9348E59F8EC" httpUri="https://zenodo.org/record/1437883/files/figure.png" pageId="30" pageNumber="31" startId="30.[151,250,1724,1746]" targetBox="[151,1436,193,1703]" targetPageId="30">
<paragraph id="DA0336F2562C551909EEF9348E59F8EC" blockId="30.[151,1437,1724,1899]" pageId="30" pageNumber="31">
<emphasis id="E8C8EAE0562C551909EEF9348B68F955" bold="true" box="[151,290,1724,1746]" pageId="30" pageNumber="31">FIGURE 17.</emphasis>
<taxonomicName id="1DBC4D71562C55190853F9348855F955" authority="Grischenko &amp; Gordon &amp; Melnik, 2018" authorityName="Grischenko &amp; Gordon &amp; Melnik" authorityYear="2018" box="[298,543,1724,1746]" class="Stenolaemata" family="Horneridae" genus="Frontohornera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="30" pageNumber="31" phylum="Bryozoa" rank="species" species="frontalis" status="gen. nov.">
<emphasis id="E8C8EAE0562C55190853F9348855F955" box="[298,543,1724,1746]" italics="true" pageId="30" pageNumber="31">Frontohornera frontalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3FB579B562C55190B5EF93488F6F955" box="[551,700,1724,1746]" pageId="30" pageNumber="31" rank="species">
<emphasis id="E8C8EAE0562C55190B5EF934883BF955" bold="true" box="[551,625,1724,1746]" pageId="30" pageNumber="31">n. gen.</emphasis>
,
<emphasis id="E8C8EAE0562C55190BF9F93488F6F955" bold="true" box="[640,700,1724,1746]" pageId="30" pageNumber="31">n. sp.</emphasis>
</taxonomicNameLabel>
Holotype, ZIRAS 1/50707, showing details of morphology.
<emphasis id="E8C8EAE0562C55190C4CF9348F0DF955" bold="true" box="[1333,1351,1724,1746]" pageId="30" pageNumber="31">A</emphasis>
, part of branch distal to gonozooid, showing cancelli and zooidal peristomes;
<emphasis id="E8C8EAE0562C55190A2EF9538922F976" bold="true" box="[855,872,1755,1777]" pageId="30" pageNumber="31">B</emphasis>
, part of infertile branch with fewer cancelli;
<emphasis id="E8C8EAE0562C55190C54F9538F75F976" bold="true" box="[1325,1343,1755,1777]" pageId="30" pageNumber="31">C</emphasis>
,
<emphasis id="E8C8EAE0562C55190C35F9538F14F976" bold="true" box="[1356,1374,1755,1777]" pageId="30" pageNumber="31">D</emphasis>
, parts of infertile branch showing peristomial dimorphism;
<emphasis id="E8C8EAE0562C55190BAAF97188AEF888" bold="true" box="[723,740,1785,1807]" pageId="30" pageNumber="31">E</emphasis>
, part of branch near gonozooid showing numerous cancelli and peristomial dimorphism;
<emphasis id="E8C8EAE0562C551908E2F8908BE2F8A9" bold="true" box="[411,424,1816,1838]" pageId="30" pageNumber="31">F</emphasis>
, zooids showing peristomial and apertural dimorphism;
<emphasis id="E8C8EAE0562C55190A8CF8908E4CF8A9" bold="true" box="[1013,1030,1816,1838]" pageId="30" pageNumber="31">G</emphasis>
, abfrontal part of branch in vicinity of gonozooid;
<emphasis id="E8C8EAE0562C55190868F8BF8B6FF8CA" bold="true" box="[273,293,1847,1869]" pageId="30" pageNumber="31">H</emphasis>
,
<emphasis id="E8C8EAE0562C5519084BF8BF8B75F8CA" bold="true" box="[306,319,1847,1869]" pageId="30" pageNumber="31">J</emphasis>
,
<emphasis id="E8C8EAE0562C55190834F8BF8B2BF8CA" bold="true" box="[333,353,1847,1869]" pageId="30" pageNumber="31">K</emphasis>
, gonozooid profiles;
<emphasis id="E8C8EAE0562C55190B47F8BF8802F8CA" bold="true" box="[574,584,1847,1869]" pageId="30" pageNumber="31">I</emphasis>
, autozooidal peristome showing interior spinules;
<emphasis id="E8C8EAE0562C55190D2BF8BF8E29F8CA" bold="true" box="[1106,1123,1847,1869]" pageId="30" pageNumber="31">L</emphasis>
, ooeciostome and ooeciopore. Scale bars: A, B, D, E, G, 300 µm; C, H, J, K, 200 µm; F, 150 µm; I, 100 µm; L, 50 µm.
</paragraph>
</caption>
<paragraph id="DA0336F2562C551909BEF81B8F11F874" blockId="30.[151,1436,1939,2036]" pageId="30" pageNumber="31">
Gonozooid frontally borne, adjacent to second dichotomy (
<figureCitation id="42872A77562C55190A1DF81B8986F82B" box="[868,972,1939,1964]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[151,1436,877,1931]" captionTargetId="figure@29.[151,1436,877,1931]" captionTargetPageId="29" captionText="FIGURE 16. Frontohornera frontalis n. gen., n. sp. Holotype specimen, ZIRAS 1/50707, respectively seen in A, frontal and B, abfrontal views. Scale bars: 1 mm." httpUri="https://zenodo.org/record/1437881/files/figure.png" pageId="30" pageNumber="31">Fig. 16A</figureCitation>
), subglobular, surface like that of colony but with slightly denser prickled granules (
<figureCitation id="42872A77562C55190BFFF83F897AF848" box="[646,816,1975,2000]" captionStart="FIGURE 17" captionStartId="30.[151,250,1724,1746]" captionTargetBox="[151,1436,193,1703]" captionTargetId="figure@30.[151,1436,193,1703]" captionTargetPageId="30" captionText="FIGURE 17. Frontohornera frontalis n. gen., n. sp. Holotype, ZIRAS 1/50707, showing details of morphology. A, part of branch distal to gonozooid, showing cancelli and zooidal peristomes; B, part of infertile branch with fewer cancelli; C, D, parts of infertile branch showing peristomial dimorphism; E, part of branch near gonozooid showing numerous cancelli and peristomial dimorphism; F, zooids showing peristomial and apertural dimorphism; G, abfrontal part of branch in vicinity of gonozooid; H, J, K, gonozooid profiles; I, autozooidal peristome showing interior spinules; L, ooeciostome and ooeciopore. Scale bars: A, B, D, E, G, 300 µm; C, H, J, K, 200 µm; F, 150 µm; I, 100 µm; L, 50 µm." httpUri="https://zenodo.org/record/1437883/files/figure.png" pageId="30" pageNumber="31">Fig. 17H, J, K</figureCitation>
), frontal peak and distal sides of chamber converging towards short ooeciostome that curves frontalwards (
<figureCitation id="42872A77562C55190B91F85389C9F874" box="[744,899,2011,2036]" captionStart="FIGURE 17" captionStartId="30.[151,250,1724,1746]" captionTargetBox="[151,1436,193,1703]" captionTargetId="figure@30.[151,1436,193,1703]" captionTargetPageId="30" captionText="FIGURE 17. Frontohornera frontalis n. gen., n. sp. Holotype, ZIRAS 1/50707, showing details of morphology. A, part of branch distal to gonozooid, showing cancelli and zooidal peristomes; B, part of infertile branch with fewer cancelli; C, D, parts of infertile branch showing peristomial dimorphism; E, part of branch near gonozooid showing numerous cancelli and peristomial dimorphism; F, zooids showing peristomial and apertural dimorphism; G, abfrontal part of branch in vicinity of gonozooid; H, J, K, gonozooid profiles; I, autozooidal peristome showing interior spinules; L, ooeciostome and ooeciopore. Scale bars: A, B, D, E, G, 300 µm; C, H, J, K, 200 µm; F, 150 µm; I, 100 µm; L, 50 µm." httpUri="https://zenodo.org/record/1437883/files/figure.png" pageId="30" pageNumber="31">Fig.17H, J, K</figureCitation>
), ooeciopore transversely oval (
<figureCitation id="42872A77562C55190D90F8538F04F874" box="[1257,1358,2011,2036]" captionStart="FIGURE 17" captionStartId="30.[151,250,1724,1746]" captionTargetBox="[151,1436,193,1703]" captionTargetId="figure@30.[151,1436,193,1703]" captionTargetPageId="30" captionText="FIGURE 17. Frontohornera frontalis n. gen., n. sp. Holotype, ZIRAS 1/50707, showing details of morphology. A, part of branch distal to gonozooid, showing cancelli and zooidal peristomes; B, part of infertile branch with fewer cancelli; C, D, parts of infertile branch showing peristomial dimorphism; E, part of branch near gonozooid showing numerous cancelli and peristomial dimorphism; F, zooids showing peristomial and apertural dimorphism; G, abfrontal part of branch in vicinity of gonozooid; H, J, K, gonozooid profiles; I, autozooidal peristome showing interior spinules; L, ooeciostome and ooeciopore. Scale bars: A, B, D, E, G, 300 µm; C, H, J, K, 200 µm; F, 150 µm; I, 100 µm; L, 50 µm." httpUri="https://zenodo.org/record/1437883/files/figure.png" pageId="30" pageNumber="31">Fig. 17L</figureCitation>
).
</paragraph>
<caption id="8EC3667A562D551809EEFAD88B7BFA45" httpUri="https://zenodo.org/record/1437885/files/figure.png" pageId="31" pageNumber="32" startId="31.[151,250,1360,1382]" targetBox="[151,1436,193,1339]" targetPageId="31">
<paragraph id="DA0336F2562D551809EEFAD88B7BFA45" blockId="31.[151,1436,1360,1474]" pageId="31" pageNumber="32">
<emphasis id="E8C8EAE0562D551809EEFAD88B68FAE1" bold="true" box="[151,290,1360,1382]" pageId="31" pageNumber="32">FIGURE 18.</emphasis>
<taxonomicName id="1DBC4D71562D55180853FAD88855FAE1" authority="Grischenko &amp; Gordon &amp; Melnik, 2018" authorityName="Grischenko &amp; Gordon &amp; Melnik" authorityYear="2018" box="[298,543,1360,1382]" class="Stenolaemata" family="Horneridae" genus="Frontohornera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="31" pageNumber="32" phylum="Bryozoa" rank="species" species="frontalis" status="gen. nov.">
<emphasis id="E8C8EAE0562D55180853FAD88855FAE1" box="[298,543,1360,1382]" italics="true" pageId="31" pageNumber="32">Frontohornera frontalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3FB579B562D55180B51FAD888F6FAE1" box="[552,700,1360,1382]" pageId="31" pageNumber="32" rank="species">
<emphasis id="E8C8EAE0562D55180B51FAD8883BFAE1" bold="true" box="[552,625,1360,1382]" pageId="31" pageNumber="32">n. gen.</emphasis>
,
<emphasis id="E8C8EAE0562D55180BF9FAD888F6FAE1" bold="true" box="[640,700,1360,1382]" pageId="31" pageNumber="32">n. sp.</emphasis>
</taxonomicNameLabel>
Progressive stages of development of young colonies.
<emphasis id="E8C8EAE0562D55180D85FAD88F67FAE1" bold="true" box="[1276,1325,1360,1382]" pageId="31" pageNumber="32">AC</emphasis>
, specimen GLD411, Stn 215, from different angles;
<emphasis id="E8C8EAE0562D55180B3AFAE78825FA02" bold="true" box="[579,623,1391,1413]" pageId="31" pageNumber="32">DF</emphasis>
, specimen YMG414, Stn 334, from different angles;
<emphasis id="E8C8EAE0562D55180DEDFAE78EF4FA02" bold="true" box="[1172,1214,1391,1413]" pageId="31" pageNumber="32">GI</emphasis>
, specimen GLD411, Stn 210, in frontal and abfrontal views and a close-up of the colony base with secondary calcification. Scale bars: AH, 500 µm; I, 250 µm.
</paragraph>
</caption>
<paragraph id="DA0336F2562D551809BEFA6789BBF9F3" blockId="31.[151,1437,1519,2012]" pageId="31" pageNumber="32">
Ancestrular zooid not seen in isolation; earliest growth stages encountered show lowest peristome of colony (presumed ancestrular peristome), which appears to be wholly erect, though concealed within basal skirt of extrazooidal calcification (
<figureCitation id="42872A77562D551808B5F9BF881AF9C8" box="[460,592,1591,1616]" captionStart="FIGURE 18" captionStartId="31.[151,250,1360,1382]" captionTargetBox="[151,1436,193,1339]" captionTargetId="figure@31.[151,1436,193,1339]" captionTargetPageId="31" captionText="FIGURE 18. Frontohornera frontalis n. gen., n. sp. Progressive stages of development of young colonies. AC, specimen GLD411, Stn 215, from different angles; DF, specimen YMG414, Stn 334, from different angles; GI, specimen GLD411, Stn 210, in frontal and abfrontal views and a close-up of the colony base with secondary calcification. Scale bars: AH, 500 µm; I, 250 µm." httpUri="https://zenodo.org/record/1437885/files/figure.png" pageId="31" pageNumber="32">Fig. 18G, I</figureCitation>
). Peristome of daughter zooid above it in series, but in another colony ancestrular and two daughter peristomes occur as cluster of three (
<figureCitation id="42872A77562D55180A04F9D389A9F9F3" box="[893,995,1627,1652]" captionStart="FIGURE 18" captionStartId="31.[151,250,1360,1382]" captionTargetBox="[151,1436,193,1339]" captionTargetId="figure@31.[151,1436,193,1339]" captionTargetPageId="31" captionText="FIGURE 18. Frontohornera frontalis n. gen., n. sp. Progressive stages of development of young colonies. AC, specimen GLD411, Stn 215, from different angles; DF, specimen YMG414, Stn 334, from different angles; GI, specimen GLD411, Stn 210, in frontal and abfrontal views and a close-up of the colony base with secondary calcification. Scale bars: AH, 500 µm; I, 250 µm." httpUri="https://zenodo.org/record/1437885/files/figure.png" pageId="31" pageNumber="32">Fig. 18B</figureCitation>
).
</paragraph>
<paragraph id="DA0336F2562D551809BEF9F789D8F8A0" blockId="31.[151,1437,1519,2012]" pageId="31" pageNumber="32">
<emphasis id="E8C8EAE0562D551809BEF9F78B83F91F" bold="true" box="[199,457,1663,1688]" pageId="31" pageNumber="32">Measurements (mm).</emphasis>
<typeStatus id="05078850562D551808ADF9F78875F91F" box="[468,575,1663,1688]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
, ZIRAS 1/50707 (
<figureCitation id="42872A77562D55180A61F9F789E2F910" box="[792,936,1663,1688]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[151,1436,877,1931]" captionTargetId="figure@29.[151,1436,877,1931]" captionTargetPageId="29" captionText="FIGURE 16. Frontohornera frontalis n. gen., n. sp. Holotype specimen, ZIRAS 1/50707, respectively seen in A, frontal and B, abfrontal views. Scale bars: 1 mm." httpUri="https://zenodo.org/record/1437881/files/figure.png" pageId="31" pageNumber="32">Fig. 16A, B</figureCitation>
): ZL 0.5690.763 (0.663 ± 0.071). Branch center: PeL 0.0720.197 (0.122 ± 0.042); PeD 0.1410.162 (0.153 ± 0.008); ApD 0.1290.141 (0.137 ± 0.004). Branch margin: PeL 0.1940.526 (0.374 ± 0.102); PeD 0.1100.146 (0.135 ± 0.011); ApL 0.1320.176 (0.156 ± 0.016) (
<emphasis id="E8C8EAE0562D55180988F9648AB5F884" box="[241,255,1772,1795]" italics="true" pageId="31" pageNumber="32">n</emphasis>
= 7); ApW 0.1220.147 (0.132 ± 0.008) (
<emphasis id="E8C8EAE0562D55180B9EF96488BFF884" box="[743,757,1772,1795]" italics="true" pageId="31" pageNumber="32">n</emphasis>
= 7). Gonozooid (
<emphasis id="E8C8EAE0562D55180AB5F9648990F884" box="[972,986,1772,1795]" italics="true" pageId="31" pageNumber="32">n</emphasis>
= 1): GZL 0.744; GZW 0.511; GZH = 0.377; OsL 0.167; OsW 0.132; OsH 0.196; OpL 0.073; OpW 0.071.
</paragraph>
<paragraph id="DA0336F2562D551809EEF8BC89A9F8CB" blockId="31.[151,1437,1519,2012]" box="[151,995,1843,1868]" pageId="31" pageNumber="32">
Non-type specimen GLD411, Stn 210 (
<figureCitation id="42872A77562D55180B25F8BB8897F8CB" box="[604,733,1843,1868]" captionStart="FIGURE 18" captionStartId="31.[151,250,1360,1382]" captionTargetBox="[151,1436,193,1339]" captionTargetId="figure@31.[151,1436,193,1339]" captionTargetPageId="31" captionText="FIGURE 18. Frontohornera frontalis n. gen., n. sp. Progressive stages of development of young colonies. AC, specimen GLD411, Stn 215, from different angles; DF, specimen YMG414, Stn 334, from different angles; GI, specimen GLD411, Stn 210, in frontal and abfrontal views and a close-up of the colony base with secondary calcification. Scale bars: AH, 500 µm; I, 250 µm." httpUri="https://zenodo.org/record/1437885/files/figure.png" pageId="31" pageNumber="32">Fig. 18GI</figureCitation>
): AnPeD 0.144 (
<emphasis id="E8C8EAE0562D55180AE0F8BC89EDF8CC" box="[921,935,1844,1867]" italics="true" pageId="31" pageNumber="32">n</emphasis>
= 1).
</paragraph>
</subSubSection>
<subSubSection id="92A66579562D551809BEF8DF8FC3F813" pageId="31" pageNumber="32" type="discussion">
<paragraph id="DA0336F2562D551809BEF8DF8FC3F813" blockId="31.[151,1437,1519,2012]" pageId="31" pageNumber="32">
<emphasis id="E8C8EAE0562D551809BEF8DF8B71F8F7" bold="true" box="[199,315,1879,1904]" pageId="31" pageNumber="32">Remarks.</emphasis>
<taxonomicName id="1DBC4D71562D5518083AF8DF881BF8F7" authority="Grischenko &amp; Gordon &amp; Melnik, 2018" authorityName="Grischenko &amp; Gordon &amp; Melnik" authorityYear="2018" box="[323,593,1879,1904]" class="Stenolaemata" family="Horneridae" genus="Frontohornera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="31" pageNumber="32" phylum="Bryozoa" rank="species" species="frontalis" status="sp. nov.">
<emphasis id="E8C8EAE0562D5518083AF8DF881BF8F7" box="[323,593,1879,1904]" italics="true" pageId="31" pageNumber="32">Frontohornera frontalis</emphasis>
</taxonomicName>
<emphasis id="E8C8EAE0562D55180B20F8DF88D0F8F7" bold="true" box="[601,666,1879,1904]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="F3FB579B562D55180B20F8DF88D0F8F7" box="[601,666,1879,1904]" pageId="31" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is represented in the collection by six colonies of varying ages and sizes. Only one reproductively mature colony was encountered. All were found attached to polymetallic nodules.
</paragraph>
</subSubSection>
<subSubSection id="92A66579562D551809BEF817884FF85C" pageId="31" pageNumber="32" type="distribution">
<paragraph id="DA0336F2562D551809BEF817884FF85C" blockId="31.[151,1437,1519,2012]" pageId="31" pageNumber="32">
<emphasis id="E8C8EAE0562D551809BEF8178B2AF83F" bold="true" box="[199,352,1951,1976]" pageId="31" pageNumber="32">Distribution.</emphasis>
Recorded from six stations within coordinates 13.03353
<geoCoordinate id="BF885035562D55180A8FF8178EC2F830" box="[1014,1160,1951,1976]" direction="north" orientation="latitude" pageId="31" pageNumber="32" precision="1" value="14.29353">14.29353° N</geoCoordinate>
, 131.87598
<geoCoordinate id="BF885035562D55180C65F8178AF8F85C" direction="west" orientation="longitude" pageId="31" pageNumber="32" precision="1" value="-134.23457">134.23457° W</geoCoordinate>
, at depth range
<quantity id="1D449B17562D5518081BF84B884AF85C" box="[354,512,1987,2012]" metricMagnitude="3" metricUnit="m" metricValue="4.9905" metricValueMax="5.166" metricValueMin="4.8149999999999995" pageId="31" pageNumber="32" unit="m" value="4990.5" valueMax="5166.0" valueMin="4815.0">48155166 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>