301 lines
51 KiB
XML
301 lines
51 KiB
XML
<document id="865E1BE0471B65E0ECFCA09F171AC06E" ID-CLB-Dataset="29502" ID-DOI="10.11646/zootaxa.4484.1.1" ID-GBIF-Dataset="74b777c9-eae0-4770-8c86-dcbb10fb06b3" ID-ISSN="1175-5326" ID-Zenodo-Dep="1437848" ID-ZooBank="D66524CF-9C6D-4DF4-8CA2-B2C9708CF5FD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538125412450" checkinUser="plazi" docAuthor="Grischenko, Andrei V., Gordon, Dennis P. & Melnik, Viacheslav P." docDate="2018" docId="521587E4562A551B09EEF88F884FFBFB" docLanguage="en" docName="zootaxa.4484.1.1.pdf" docOrigin="Zootaxa 4484 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Abyssoecia elevata Grischenko & Gordon & Melnik 2018, n. sp." docType="treatment" docVersion="8" lastPageNumber="29" masterDocId="AE2CFF9C563255070979FF888A4AFF87" masterDocTitle="Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining" masterLastPageNumber="91" masterPageNumber="1" pageNumber="25" updateTime="1698557051871" updateUser="ExternalLinkService">
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<mods:titleInfo id="D953242F99CB121A0751D31D337F468C">
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<mods:title id="E1FFC25DBE9C26C35496412826C8F9EB">Bryozoa (Cyclostomata and Ctenostomata) from polymetallic nodules in the Russian exploration area, Clarion - Clipperton Fracture Zone, eastern Pacific Ocean-taxon novelty and implications of mining</mods:title>
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<mods:name id="31C3B81DC7CF0872743FF962EE1598E4" type="personal">
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<mods:role id="E738E98F6943FEE30C6069BA7883E262">
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<mods:roleTerm id="2EE9FFED9BCCC66D009F7DBD45603DDF">Author</mods:roleTerm>
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<mods:namePart id="3413C6A9A6F405F28B30AFE2A0ABCA6E">Grischenko, Andrei V.</mods:namePart>
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<mods:roleTerm id="A1F4404F3B1B42F86AAB00E04A0C0D0E">Author</mods:roleTerm>
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<mods:namePart id="B720A898F3F63040E27E57F63E747BCF">Gordon, Dennis P.</mods:namePart>
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<mods:name id="263CE41B3A09AC0FCB3AB6160064D121" type="personal">
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<mods:roleTerm id="1B2F26891C2B13223CBF8D4E73FF7272">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="857E93BFB1127614460E0797DF57FF2B">Melnik, Viacheslav P.</mods:namePart>
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<mods:typeOfResource id="C78D497CA6841B14A279F9D7552AFB0D">text</mods:typeOfResource>
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<mods:titleInfo id="49C4304D297DFF762EE5578BFEC8908E">
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<mods:title id="82470C505C9AB34D27277E5EFF53B8EA">Zootaxa</mods:title>
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<mods:part id="DC21A9CD51752336399376157583507C">
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<mods:date id="FDF05CF2728C7F8EA6138A8BD3135660">2018</mods:date>
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<mods:number id="94C377406A3EF935C4D3158DA75117CD">2018-09-25</mods:number>
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<mods:number id="31B0E548935679315B0009C20323B999">4484</mods:number>
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<mods:detail id="81CE0002F583A3B39906AF3AE0701FD3" type="issue">
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<mods:number id="E0CF5B658A87997340ECB14B72391D05">1</mods:number>
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<mods:classification id="7C4BDB99374F91FF02227AED270E5541">journal article</mods:classification>
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<mods:identifier id="842863E800AA2346A5D06D60749A858B" type="CLB-Dataset">29502</mods:identifier>
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<mods:identifier id="EBBC98C112F6E39F2F623685A78B4B75" type="DOI">10.11646/zootaxa.4484.1.1</mods:identifier>
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<mods:identifier id="217FE9E0053CDA10E309E433680AC2C2" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="BE43E069737004ED4DC3EE62460ECC7E" type="Zenodo-Dep">1437848</mods:identifier>
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<treatment id="521587E4562A551B09EEF88F884FFBFB" ID-DOI="http://doi.org/10.5281/zenodo.5989793" ID-GBIF-Taxon="148403717" ID-Zenodo-Dep="5989793" LSID="urn:lsid:plazi:treatment:521587E4562A551B09EEF88F884FFBFB" httpUri="http://treatment.plazi.org/id/521587E4562A551B09EEF88F884FFBFB" lastPageId="28" lastPageNumber="29" pageId="24" pageNumber="25">
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<subSubSection id="92A66579562A551F09EEF88F8B3FF8C4" pageId="24" pageNumber="25" type="nomenclature">
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<paragraph id="DA0336F2562A551F09EEF88F8B8BF8A5" blockId="24.[151,449,1799,1859]" box="[151,449,1799,1826]" pageId="24" pageNumber="25">
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<heading id="814B819E562A551F09EEF88F8B8BF8A5" bold="true" box="[151,449,1799,1826]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
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<emphasis id="E8C8EAE0562A551F09EEF88F8B8BF8A5" bold="true" box="[151,449,1799,1826]" pageId="24" pageNumber="25">
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<taxonomicName id="1DBC4D71562A551F09EEF88F8B32F8A6" ID-CoL="724c487f-8352-4f25-bcc9-e770c2ac546a" authority="Grischenko & Gordon & Melnik, 2018" authorityName="Grischenko & Gordon & Melnik" authorityYear="2018" box="[151,376,1799,1825]" class="Stenolaemata" family="Oncousoeciidae" genus="Abyssoecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="24" pageNumber="25" phylum="Bryozoa" rank="species" species="elevata" status="sp. nov.">
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<emphasis id="E8C8EAE0562A551F09EEF88F8B32F8A6" bold="true" box="[151,376,1799,1825]" italics="true" pageId="24" pageNumber="25">Abyssoecia elevata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F3FB579B562A551F0806F8808B8BF8A5" box="[383,449,1800,1826]" pageId="24" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="DA0336F2562A551F09EEF8A28B3FF8C4" blockId="24.[151,449,1799,1859]" box="[151,373,1834,1859]" pageId="24" pageNumber="25">
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(
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<figureCitation id="42872A77562A551F09E6F8A28B51F8C5" box="[159,283,1834,1859]" captionStart="FIGURE 2" captionStartId="10.[151,250,1774,1796]" captionTargetBox="[151,1436,392,1715]" captionTargetId="figure@10.[151,1436,392,1715]" captionTargetPageId="10" captionText="FIGURE 2. Colonies of some cyclostome bryozoans, in vivo, attached to polymetallic nodules. A–E, Pandanipora helix n. gen., n. sp.: A, specimen GLD4–09, Stn 190; B, specimen GLD4–12, Stn 262; C, specimen YMG4–07, Stn 143; D, specimen YMG4–13, Stn 295; E, specimen GLD 4–11, Stn 212. F, Tubuliporina sp. indet., specimen YMG18–01, Stn 7. G, H, Abyssoecia elevata n. gen., n. sp.: G, specimen GLD4–09, Stn 196; H, specimen GLD4–09, Stn 191. I, Discantenna metallica n. sp.: specimen GLD4–11, Stn 224. J, K, Frontohornera frontalis n. gen., n. sp.: J, specimen YMG4–07, Stn 124; K, specimen GLD4–11, Stn 210. L, Alyonushka hystricosa n. gen., n. sp.: specimen GLD4–09, Stn 199. M, Calyssopora volcano n. gen., n. sp.: specimen YMG18–01, Stn 33. N, O, Anyuta anastema n. gen., n. sp.: N, specimen GLD4–09, Stn 180; O, specimen YMG4–06, Stn 71. Scale bars: 1 mm." httpUri="https://zenodo.org/record/1437852/files/figure.png" pageId="24" pageNumber="25">Figs 2G, H</figureCitation>
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,
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<figureCitation id="42872A77562A551F085FF8A28B27F8C4" box="[294,365,1834,1859]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStart-2="FIGURE 15" captionStartId-0="25.[151,250,1886,1908]" captionStartId-1="26.[151,250,1852,1874]" captionStartId-2="27.[151,250,1725,1747]" captionTargetBox-0="[151,1436,445,1865]" captionTargetBox-1="[151,1436,193,1830]" captionTargetBox-2="[151,1436,193,1704]" captionTargetId-0="figure@25.[151,1436,445,1865]" captionTargetId-1="figure@26.[151,1436,193,1830]" captionTargetId-2="figure@27.[151,1436,193,1704]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="27" captionText-0="FIGURE 13. Abyssoecia elevata n. gen., n. sp. Holotype and paratype colonies detached from their substrata. A, B, holotype, ZIRAS 1/50703, frontal and profile views; note foramen in colony formed by growth and fusion of lobes around a space; C, D, paratype 1, ZIRAS 2/50704 frontal and abfrontal views; E, F, paratype 2, ZIRAS 3/50705, frontal and profile views. Scale bars: 500 µm." captionText-1="FIGURE 14. Abyssoecia elevata n. gen., n. sp. A–E, paratype 3, ZIRAS 4/50706; F, specimen YMG4–06, Stn 106; G, J, specimen YMG4–13, Stn 302; H, I, N, specimen YMG4–07, Stn 129; K, specimen GLD4–09, Stn 191; L, M, specimen YMG4–04, Stn 44. A, gonozooid at upper left of image with projecting ooeciostome; B, D, distal and distolateral views of gonozooid; C, E, frontal and distal views of ooeciostome with subcircular ooeciopore; F–H, distolateral (F) and distal views of colony margin with developing zooids; I–K, frontal views of short autozooidal peristomes; L–N, ancestrular region of two colonies. Scale bars: A, 500 µm; B, D, F–J, L–N, 200 µm; C, E, 100 µm; K, 50 µm." captionText-2="FIGURE 15. Abyssoecia elevata n. gen., n. sp. Progressive stages of development of young ancestrulate colonies viewed from different angles, showing development of either supportive columns or growth of lateral margins towards substratum as colony grows larger. A–C, specimen GLD4–09, Stn 196; D–F, specimen YMG4–07, Stn 129; G–L, specimen YMG4–04, Stn 44. Scale bars: A, B, D–F, 250 µm; C, 200 µm; G–L, 500 µm." httpUri-0="https://zenodo.org/record/1437875/files/figure.png" httpUri-1="https://zenodo.org/record/1437877/files/figure.png" httpUri-2="https://zenodo.org/record/1437879/files/figure.png" pageId="24" pageNumber="25">13–15</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="92A66579562A551E09EEF8FA8AF2FEE3" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="materials_examined">
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<paragraph id="DA0336F2562A551E09EEF8FA8AF2FEE3" blockId="24.[151,1436,1906,2003]" lastBlockId="25.[151,1436,151,392]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
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<emphasis id="E8C8EAE0562A551F09EEF8FA8BCFF80C" bold="true" box="[151,389,1906,1931]" pageId="24" pageNumber="25">Material examined.</emphasis>
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<emphasis id="E8C8EAE0562A551F08EBF8FA8BB2F80C" box="[402,504,1906,1931]" italics="true" pageId="24" pageNumber="25">Holotype</emphasis>
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: ZIRAS 1/50703, colony detached from nodule, YMG R.V.
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<emphasis id="E8C8EAE0562A551F0DB4F8FA8FD3F80C" box="[1229,1433,1906,1931]" italics="true" pageId="24" pageNumber="25">Yuzhmorgeologiya</emphasis>
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cruise YMG4–13, Stn 302,
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<date id="AE021032562A551F08A2F81F883DF828" box="[475,631,1942,1967]" pageId="24" pageNumber="25" value="2014-07-30">30 July 2014</date>
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, 13.16460° N, 132.52885° W,
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<quantity id="1D449B17562A551F0A98F81F8E77F829" box="[993,1085,1943,1967]" metricMagnitude="3" metricUnit="m" metricValue="4.933999999999999" pageId="24" pageNumber="25" unit="m" value="4934.0">4934 m</quantity>
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.
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<emphasis id="E8C8EAE0562A551F0D32F81F8EF8F829" box="[1099,1202,1943,1966]" italics="true" pageId="24" pageNumber="25">Paratype</emphasis>
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1: ZIRAS 2/50704, colony detached from nodule, YMG R.V.
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<emphasis id="E8C8EAE0562A551F0BEAF832892BF854" box="[659,865,1978,2003]" italics="true" pageId="24" pageNumber="25">Yuzhmorgeologiya</emphasis>
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cruise YMG4–06, Stn 106,
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<date id="AE021032562A551F0DC4F8338FD2F854" box="[1213,1432,1978,2003]" pageId="24" pageNumber="25" value="2006-09-02">2 September 2006</date>
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, 13.26750° N, 134.51797° W,
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<quantity id="1D449B17562B551E0895FF1F880CFF28" box="[492,582,151,176]" metricMagnitude="3" metricUnit="m" metricValue="4.805" pageId="25" pageNumber="26" unit="m" value="4805.0">4805 m</quantity>
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.
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<emphasis id="E8C8EAE0562B551E0B2BFF1188F3FF37" box="[594,697,153,176]" italics="true" pageId="25" pageNumber="26">Paratype</emphasis>
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2: ZIRAS 3/50705, colony attached to particle of nodule, YMG R.V.
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<emphasis id="E8C8EAE0562B551E09ABFF348B1AFF52" box="[210,336,188,213]" italics="true" pageId="25" pageNumber="26">Gelendzhik</emphasis>
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cruise GLD4–09, Stn 191,
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<date id="AE021032562B551E0BEAFF34890CFF53" box="[659,838,188,213]" pageId="25" pageNumber="26" value="2011-01-03">3 January 2011</date>
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, 13.40957° N, 133.47513° W,
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<quantity id="1D449B17562B551E0DD4FF348F42FF53" box="[1197,1288,188,212]" metricMagnitude="3" metricUnit="m" metricValue="4.897" pageId="25" pageNumber="26" unit="m" value="4897.0">4897 m</quantity>
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.
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<emphasis id="E8C8EAE0562B551E0C6CFF358F36FF53" box="[1301,1404,189,212]" italics="true" pageId="25" pageNumber="26">Paratype</emphasis>
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3: ZIRAS 4/50706, colony fragment (with gonozooid) detached from nodule, YMG R.V.
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<emphasis id="E8C8EAE0562B551E0DF8FF578F05FF7F" box="[1153,1359,223,248]" italics="true" pageId="25" pageNumber="26">Yuzhmorgeologiya</emphasis>
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cruise YMG4–13, Stn 282,
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<date id="AE021032562B551E08FDFE8C8853FE9B" box="[388,537,260,285]" pageId="25" pageNumber="26" value="2014-07-12">12 July 2014</date>
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, 13.41580° N, 132.28818° W,
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<quantity id="1D449B17562B551E0A0DFE8C8986FE9B" box="[884,972,260,285]" metricMagnitude="3" metricUnit="m" metricValue="4.946" pageId="25" pageNumber="26" unit="m" value="4946.0">4946 m</quantity>
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.
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<emphasis id="E8C8EAE0562B551E0AAEFE8C8EFEFE9A" box="[983,1204,260,285]" italics="true" pageId="25" pageNumber="26">Additional material</emphasis>
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: YMG4–04, Stn 44; YMG4–06, Stn 99; YMG4–07, Stns 129, 135; GLD4–09, Stn 196; YMG4–13, Stn 313. Total specimens examined 11.
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</paragraph>
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</subSubSection>
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<subSubSection id="92A66579562B551E09BEFEE78E9CFE0F" box="[199,1238,367,392]" pageId="25" pageNumber="26" type="etymology">
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<paragraph id="DA0336F2562B551E09BEFEE78E9CFE0F" blockId="25.[151,1436,151,392]" box="[199,1238,367,392]" pageId="25" pageNumber="26">
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<emphasis id="E8C8EAE0562B551E09BEFEE78B04FE0F" bold="true" box="[199,334,367,392]" pageId="25" pageNumber="26">Etymology.</emphasis>
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Latin,
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<emphasis id="E8C8EAE0562B551E08E4FEE78BB3FE0F" box="[413,505,367,392]" italics="true" pageId="25" pageNumber="26">elevatus</emphasis>
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, raised, alluding to the elevation of colony above the substratum.
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</paragraph>
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</subSubSection>
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<caption id="8EC3667A562B551E09EEF8D68AA6F857" httpUri="https://zenodo.org/record/1437875/files/figure.png" pageId="25" pageNumber="26" startId="25.[151,250,1886,1908]" targetBox="[151,1436,445,1865]" targetPageId="25">
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<paragraph id="DA0336F2562B551E09EEF8D68AA6F857" blockId="25.[151,1436,1886,2000]" pageId="25" pageNumber="26">
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<emphasis id="E8C8EAE0562B551E09EEF8D68B6AF8F3" bold="true" box="[151,288,1886,1908]" pageId="25" pageNumber="26">FIGURE 13.</emphasis>
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<taxonomicName id="1DBC4D71562B551E085FF8D68BA2F8F3" authority="Grischenko & Gordon & Melnik, 2018" authorityName="Grischenko & Gordon & Melnik" authorityYear="2018" box="[294,488,1886,1908]" class="Stenolaemata" family="Oncousoeciidae" genus="Abyssoecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="25" pageNumber="26" phylum="Bryozoa" rank="species" species="elevata" status="gen. nov.">
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<emphasis id="E8C8EAE0562B551E085FF8D68BA2F8F3" box="[294,488,1886,1908]" italics="true" pageId="25" pageNumber="26">Abyssoecia elevata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F3FB579B562B551E0897F8D68836F8F3" box="[494,636,1886,1908]" pageId="25" pageNumber="26" rank="species">
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<emphasis id="E8C8EAE0562B551E0897F8D6887CF8F3" bold="true" box="[494,566,1886,1908]" pageId="25" pageNumber="26">n. gen.</emphasis>
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,
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<emphasis id="E8C8EAE0562B551E0B3BF8D68836F8F3" bold="true" box="[578,636,1886,1908]" pageId="25" pageNumber="26">n. sp.</emphasis>
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</taxonomicNameLabel>
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Holotype and paratype colonies detached from their substrata.
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<emphasis id="E8C8EAE0562B551E0C79F8D68F58F8F3" bold="true" box="[1280,1298,1886,1908]" pageId="25" pageNumber="26">A</emphasis>
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,
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<emphasis id="E8C8EAE0562B551E0C66F8D68F7AF8F3" bold="true" box="[1311,1328,1886,1908]" pageId="25" pageNumber="26">B</emphasis>
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, holotype, ZIRAS 1/50703, frontal and profile views; note foramen in colony formed by growth and fusion of lobes around a space;
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<emphasis id="E8C8EAE0562B551E0C1DF8F58F3CF814" bold="true" box="[1380,1398,1917,1939]" pageId="25" pageNumber="26">C</emphasis>
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,
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<emphasis id="E8C8EAE0562B551E0CFAF8F58FDFF814" bold="true" box="[1411,1429,1917,1939]" pageId="25" pageNumber="26">D</emphasis>
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, paratype 1, ZIRAS 2/50704 frontal and abfrontal views;
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<emphasis id="E8C8EAE0562B551E0BA9F81488ABF835" bold="true" box="[720,737,1948,1970]" pageId="25" pageNumber="26">E</emphasis>
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,
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||
<emphasis id="E8C8EAE0562B551E0B94F81488B0F835" bold="true" box="[749,762,1948,1970]" pageId="25" pageNumber="26">F</emphasis>
|
||
, paratype 2, ZIRAS 3/50705, frontal and profile views. Scale bars: 500 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="8EC3667A5628551D09EEF8B489A3F86C" httpUri="https://zenodo.org/record/1437877/files/figure.png" pageId="26" pageNumber="27" startId="26.[151,250,1852,1874]" targetBox="[151,1436,193,1830]" targetPageId="26">
|
||
<paragraph id="DA0336F25628551D09EEF8B489A3F86C" blockId="26.[151,1436,1852,2027]" pageId="26" pageNumber="27">
|
||
<emphasis id="E8C8EAE05628551D09EEF8B48B68F8D5" bold="true" box="[151,290,1852,1874]" pageId="26" pageNumber="27">FIGURE 14.</emphasis>
|
||
<taxonomicName id="1DBC4D715628551D0855F8B48BBAF8D5" authority="Grischenko & Gordon & Melnik, 2018" authorityName="Grischenko & Gordon & Melnik" authorityYear="2018" box="[300,496,1852,1874]" class="Stenolaemata" family="Oncousoeciidae" genus="Abyssoecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="26" pageNumber="27" phylum="Bryozoa" rank="species" species="elevata" status="gen. nov.">
|
||
<emphasis id="E8C8EAE05628551D0855F8B48BBAF8D5" box="[300,496,1852,1874]" italics="true" pageId="26" pageNumber="27">Abyssoecia elevata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F3FB579B5628551D0880F8B488C5F8D5" box="[505,655,1852,1874]" pageId="26" pageNumber="27" rank="species">
|
||
<emphasis id="E8C8EAE05628551D0880F8B4880EF8D5" bold="true" box="[505,580,1852,1874]" pageId="26" pageNumber="27">n. gen.</emphasis>
|
||
,
|
||
<emphasis id="E8C8EAE05628551D0B2AF8B488C5F8D5" bold="true" box="[595,655,1852,1874]" pageId="26" pageNumber="27">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
A–E, paratype 3, ZIRAS 4/50706; F, specimen YMG4–06, Stn 106; G, J, specimen YMG4–13, Stn 302; H, I, N, specimen YMG4–07, Stn 129; K, specimen GLD4–09, Stn 191; L, M, specimen YMG4–04, Stn 44.
|
||
<emphasis id="E8C8EAE05628551D081FF8F18B32F808" bold="true" box="[358,376,1913,1935]" pageId="26" pageNumber="27">A</emphasis>
|
||
, gonozooid at upper left of image with projecting ooeciostome;
|
||
<emphasis id="E8C8EAE05628551D0D65F8F18E67F808" bold="true" box="[1052,1069,1913,1935]" pageId="26" pageNumber="27">B</emphasis>
|
||
,
|
||
<emphasis id="E8C8EAE05628551D0D45F8F18E04F808" bold="true" box="[1084,1102,1913,1935]" pageId="26" pageNumber="27">D</emphasis>
|
||
, distal and distolateral views of gonozooid;
|
||
<emphasis id="E8C8EAE05628551D0876F8108B6BF829" bold="true" box="[271,289,1944,1966]" pageId="26" pageNumber="27">C</emphasis>
|
||
,
|
||
<emphasis id="E8C8EAE05628551D0854F8108B74F829" bold="true" box="[301,318,1944,1966]" pageId="26" pageNumber="27">E</emphasis>
|
||
, frontal and distal views of ooeciostome with subcircular ooeciopore;
|
||
<emphasis id="E8C8EAE05628551D0D7FF8108E7CF829" bold="true" box="[1030,1078,1944,1966]" pageId="26" pageNumber="27">F–H</emphasis>
|
||
, distolateral (F) and distal views of colony margin with developing zooids;
|
||
<emphasis id="E8C8EAE05628551D0B4FF83E882BF84B" bold="true" box="[566,609,1974,1996]" pageId="26" pageNumber="27">I–K</emphasis>
|
||
, frontal views of short autozooidal peristomes;
|
||
<emphasis id="E8C8EAE05628551D0D2EF83E8ECCF84B" bold="true" box="[1111,1158,1974,1996]" pageId="26" pageNumber="27">L–N</emphasis>
|
||
, ancestrular region of two colonies. Scale bars: A, 500 µm; B, D, F–J, L–N, 200 µm; C, E, 100 µm; K, 50 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="8EC3667A5629551C09EEF93588ACF8A8" httpUri="https://zenodo.org/record/1437879/files/figure.png" pageId="27" pageNumber="28" startId="27.[151,250,1725,1747]" targetBox="[151,1436,193,1704]" targetPageId="27">
|
||
<paragraph id="DA0336F25629551C09EEF93588ACF8A8" blockId="27.[151,1437,1725,1839]" pageId="27" pageNumber="28">
|
||
<emphasis id="E8C8EAE05629551C09EEF9358B55F954" bold="true" box="[151,287,1725,1747]" pageId="27" pageNumber="28">FIGURE 15.</emphasis>
|
||
<taxonomicName id="1DBC4D715629551C085CF9358BACF954" authority="Grischenko & Gordon & Melnik, 2018" authorityName="Grischenko & Gordon & Melnik" authorityYear="2018" box="[293,486,1725,1747]" class="Stenolaemata" family="Oncousoeciidae" genus="Abyssoecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclostomatida" pageId="27" pageNumber="28" phylum="Bryozoa" rank="species" species="elevata" status="gen. nov.">
|
||
<emphasis id="E8C8EAE05629551C085CF9358BACF954" box="[293,486,1725,1747]" italics="true" pageId="27" pageNumber="28">Abyssoecia elevata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F3FB579B5629551C0895F9358832F954" box="[492,632,1725,1747]" pageId="27" pageNumber="28" rank="species">
|
||
<emphasis id="E8C8EAE05629551C0895F9358879F954" bold="true" box="[492,563,1725,1747]" pageId="27" pageNumber="28">n. gen.</emphasis>
|
||
,
|
||
<emphasis id="E8C8EAE05629551C0B46F9358832F954" bold="true" box="[575,632,1725,1747]" pageId="27" pageNumber="28">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Progressive stages of development of young ancestrulate colonies viewed from different angles, showing development of either supportive columns or growth of lateral margins towards substratum as colony grows larger.
|
||
<emphasis id="E8C8EAE05629551C085EF9738B12F896" bold="true" box="[295,344,1787,1809]" pageId="27" pageNumber="28">A–C</emphasis>
|
||
, specimen GLD4–09, Stn 196;
|
||
<emphasis id="E8C8EAE05629551C0BD1F9738899F896" bold="true" box="[680,723,1787,1809]" pageId="27" pageNumber="28">D–F</emphasis>
|
||
, specimen YMG4–07, Stn 129;
|
||
<emphasis id="E8C8EAE05629551C0D52F9738E16F896" bold="true" box="[1067,1116,1787,1809]" pageId="27" pageNumber="28">G–L</emphasis>
|
||
, specimen YMG4–04, Stn 44. Scale bars: A, B, D–F, 250 µm; C, 200 µm; G–L, 500 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="92A665795629551B09BEF8D48B91FC4F" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="description">
|
||
<paragraph id="DA0336F25629551B09BEF8D48E68FE9A" blockId="27.[151,1436,1884,2017]" lastBlockId="28.[151,1437,151,1149]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
|
||
<emphasis id="E8C8EAE05629551C09BEF8D48B13F8F2" bold="true" box="[199,345,1884,1909]" pageId="27" pageNumber="28">Description.</emphasis>
|
||
Colony suberect to erect, flabellate (
|
||
<figureCitation id="42872A775629551C0A69F8D4892DF8F2" box="[784,871,1884,1909]" captionStart="FIGURE 13" captionStartId="25.[151,250,1886,1908]" captionTargetBox="[151,1436,445,1865]" captionTargetId="figure@25.[151,1436,445,1865]" captionTargetPageId="25" captionText="FIGURE 13. Abyssoecia elevata n. gen., n. sp. Holotype and paratype colonies detached from their substrata. A, B, holotype, ZIRAS 1/50703, frontal and profile views; note foramen in colony formed by growth and fusion of lobes around a space; C, D, paratype 1, ZIRAS 2/50704 frontal and abfrontal views; E, F, paratype 2, ZIRAS 3/50705, frontal and profile views. Scale bars: 500 µm." httpUri="https://zenodo.org/record/1437875/files/figure.png" pageId="27" pageNumber="28">Fig. 13</figureCitation>
|
||
), pale lemon yellow (
|
||
<figureCitation id="42872A775629551C0D15F8D48EA7F8F2" box="[1132,1261,1884,1909]" captionStart="FIGURE 2" captionStartId="10.[151,250,1774,1796]" captionTargetBox="[151,1436,392,1715]" captionTargetId="figure@10.[151,1436,392,1715]" captionTargetPageId="10" captionText="FIGURE 2. Colonies of some cyclostome bryozoans, in vivo, attached to polymetallic nodules. A–E, Pandanipora helix n. gen., n. sp.: A, specimen GLD4–09, Stn 190; B, specimen GLD4–12, Stn 262; C, specimen YMG4–07, Stn 143; D, specimen YMG4–13, Stn 295; E, specimen GLD 4–11, Stn 212. F, Tubuliporina sp. indet., specimen YMG18–01, Stn 7. G, H, Abyssoecia elevata n. gen., n. sp.: G, specimen GLD4–09, Stn 196; H, specimen GLD4–09, Stn 191. I, Discantenna metallica n. sp.: specimen GLD4–11, Stn 224. J, K, Frontohornera frontalis n. gen., n. sp.: J, specimen YMG4–07, Stn 124; K, specimen GLD4–11, Stn 210. L, Alyonushka hystricosa n. gen., n. sp.: specimen GLD4–09, Stn 199. M, Calyssopora volcano n. gen., n. sp.: specimen YMG18–01, Stn 33. N, O, Anyuta anastema n. gen., n. sp.: N, specimen GLD4–09, Stn 180; O, specimen YMG4–06, Stn 71. Scale bars: 1 mm." httpUri="https://zenodo.org/record/1437852/files/figure.png" pageId="27" pageNumber="28">Fig. 2G, H</figureCitation>
|
||
), supported by founding zooids and additional pillars derived from autozooids (
|
||
<figureCitation id="42872A775629551C0A11F80889A4F81F" box="[872,1006,1920,1945]" captionStart="FIGURE 13" captionStartId="25.[151,250,1886,1908]" captionTargetBox="[151,1436,445,1865]" captionTargetId="figure@25.[151,1436,445,1865]" captionTargetPageId="25" captionText="FIGURE 13. Abyssoecia elevata n. gen., n. sp. Holotype and paratype colonies detached from their substrata. A, B, holotype, ZIRAS 1/50703, frontal and profile views; note foramen in colony formed by growth and fusion of lobes around a space; C, D, paratype 1, ZIRAS 2/50704 frontal and abfrontal views; E, F, paratype 2, ZIRAS 3/50705, frontal and profile views. Scale bars: 500 µm." httpUri="https://zenodo.org/record/1437875/files/figure.png" pageId="27" pageNumber="28">Figs 13B, F</figureCitation>
|
||
,
|
||
<figureCitation id="42872A775629551C0A80F8088E0EF81E" box="[1017,1092,1920,1945]" captionStart="FIGURE 15" captionStartId="27.[151,250,1725,1747]" captionTargetBox="[151,1436,193,1704]" captionTargetId="figure@27.[151,1436,193,1704]" captionTargetPageId="27" captionText="FIGURE 15. Abyssoecia elevata n. gen., n. sp. Progressive stages of development of young ancestrulate colonies viewed from different angles, showing development of either supportive columns or growth of lateral margins towards substratum as colony grows larger. A–C, specimen GLD4–09, Stn 196; D–F, specimen YMG4–07, Stn 129; G–L, specimen YMG4–04, Stn 44. Scale bars: A, B, D–F, 250 µm; C, 200 µm; G–L, 500 µm." httpUri="https://zenodo.org/record/1437879/files/figure.png" pageId="27" pageNumber="28">15C–I</figureCitation>
|
||
); up to
|
||
<quantity id="1D449B175629551C0DE1F8088EB6F81F" box="[1176,1276,1920,1945]" metricMagnitude="-3" metricUnit="m" metricValue="5.77" pageId="27" pageNumber="28" unit="mm" value="5.77">5.77 mm</quantity>
|
||
long and
|
||
<quantity id="1D449B175629551C0C12F8098A89F83B" metricMagnitude="-3" metricUnit="m" metricValue="9.09" pageId="27" pageNumber="28" unit="mm" value="9.09">9.09 mm</quantity>
|
||
wide. Arcuate lateral margins of large colonies tending to curve proximad, occasionally meeting around a space or spaces, creating open calyciform shape (
|
||
<figureCitation id="42872A775629551C0BB5F8408962F866" box="[716,808,1992,2017]" captionStart="FIGURE 2" captionStartId="10.[151,250,1774,1796]" captionTargetBox="[151,1436,392,1715]" captionTargetId="figure@10.[151,1436,392,1715]" captionTargetPageId="10" captionText="FIGURE 2. Colonies of some cyclostome bryozoans, in vivo, attached to polymetallic nodules. A–E, Pandanipora helix n. gen., n. sp.: A, specimen GLD4–09, Stn 190; B, specimen GLD4–12, Stn 262; C, specimen YMG4–07, Stn 143; D, specimen YMG4–13, Stn 295; E, specimen GLD 4–11, Stn 212. F, Tubuliporina sp. indet., specimen YMG18–01, Stn 7. G, H, Abyssoecia elevata n. gen., n. sp.: G, specimen GLD4–09, Stn 196; H, specimen GLD4–09, Stn 191. I, Discantenna metallica n. sp.: specimen GLD4–11, Stn 224. J, K, Frontohornera frontalis n. gen., n. sp.: J, specimen YMG4–07, Stn 124; K, specimen GLD4–11, Stn 210. L, Alyonushka hystricosa n. gen., n. sp.: specimen GLD4–09, Stn 199. M, Calyssopora volcano n. gen., n. sp.: specimen YMG18–01, Stn 33. N, O, Anyuta anastema n. gen., n. sp.: N, specimen GLD4–09, Stn 180; O, specimen YMG4–06, Stn 71. Scale bars: 1 mm." httpUri="https://zenodo.org/record/1437852/files/figure.png" pageId="27" pageNumber="28">Figs 2G</figureCitation>
|
||
,
|
||
<figureCitation id="42872A775629551C0A4FF8418920F866" box="[822,874,1993,2017]" captionStart="FIGURE 13" captionStartId="25.[151,250,1886,1908]" captionTargetBox="[151,1436,445,1865]" captionTargetId="figure@25.[151,1436,445,1865]" captionTargetPageId="25" captionText="FIGURE 13. Abyssoecia elevata n. gen., n. sp. Holotype and paratype colonies detached from their substrata. A, B, holotype, ZIRAS 1/50703, frontal and profile views; note foramen in colony formed by growth and fusion of lobes around a space; C, D, paratype 1, ZIRAS 2/50704 frontal and abfrontal views; E, F, paratype 2, ZIRAS 3/50705, frontal and profile views. Scale bars: 500 µm." httpUri="https://zenodo.org/record/1437875/files/figure.png" pageId="27" pageNumber="28">13A</figureCitation>
|
||
). Supportive pillars of about 4–8 connate zooids, each budded in such a way as to be bent, with proximal portion subvertical, descending towards substratum (e.g.
|
||
<figureCitation id="42872A77562E551B09EEFF348AB0FF52" box="[151,250,188,213]" captionStart="FIGURE 15" captionStartId="27.[151,250,1725,1747]" captionTargetBox="[151,1436,193,1704]" captionTargetId="figure@27.[151,1436,193,1704]" captionTargetPageId="27" captionText="FIGURE 15. Abyssoecia elevata n. gen., n. sp. Progressive stages of development of young ancestrulate colonies viewed from different angles, showing development of either supportive columns or growth of lateral margins towards substratum as colony grows larger. A–C, specimen GLD4–09, Stn 196; D–F, specimen YMG4–07, Stn 129; G–L, specimen YMG4–04, Stn 44. Scale bars: A, B, D–F, 250 µm; C, 200 µm; G–L, 500 µm." httpUri="https://zenodo.org/record/1437879/files/figure.png" pageId="28" pageNumber="29">Fig. 15F</figureCitation>
|
||
), and frontodistal part forming peristomial tubes (e.g.
|
||
<figureCitation id="42872A77562E551B0A1EFF34898EFF52" box="[871,964,188,213]" captionStart="FIGURE 15" captionStartId="27.[151,250,1725,1747]" captionTargetBox="[151,1436,193,1704]" captionTargetId="figure@27.[151,1436,193,1704]" captionTargetPageId="27" captionText="FIGURE 15. Abyssoecia elevata n. gen., n. sp. Progressive stages of development of young ancestrulate colonies viewed from different angles, showing development of either supportive columns or growth of lateral margins towards substratum as colony grows larger. A–C, specimen GLD4–09, Stn 196; D–F, specimen YMG4–07, Stn 129; G–L, specimen YMG4–04, Stn 44. Scale bars: A, B, D–F, 250 µm; C, 200 µm; G–L, 500 µm." httpUri="https://zenodo.org/record/1437879/files/figure.png" pageId="28" pageNumber="29">
|
||
Fig.
|
||
<date id="AE021032562E551B0AE2FF34898EFF52" box="[923,964,188,213]" pageId="28" pageNumber="29" value="2015-01">15I</date>
|
||
</figureCitation>
|
||
). No kenozooids. Surface calcification of colony forming sinuous concentric striae (
|
||
<figureCitation id="42872A77562E551B0BD3FF57896BFF7F" box="[682,801,223,248]" captionStart="FIGURE 13" captionStartId="25.[151,250,1886,1908]" captionTargetBox="[151,1436,445,1865]" captionTargetId="figure@25.[151,1436,445,1865]" captionTargetPageId="25" captionText="FIGURE 13. Abyssoecia elevata n. gen., n. sp. Holotype and paratype colonies detached from their substrata. A, B, holotype, ZIRAS 1/50703, frontal and profile views; note foramen in colony formed by growth and fusion of lobes around a space; C, D, paratype 1, ZIRAS 2/50704 frontal and abfrontal views; E, F, paratype 2, ZIRAS 3/50705, frontal and profile views. Scale bars: 500 µm." httpUri="https://zenodo.org/record/1437875/files/figure.png" pageId="28" pageNumber="29">Figs 13D</figureCitation>
|
||
,
|
||
<figureCitation id="42872A77562E551B0A41FF6889E4FF7F" box="[824,942,224,248]" captionStart="FIGURE 14" captionStartId="26.[151,250,1852,1874]" captionTargetBox="[151,1436,193,1830]" captionTargetId="figure@26.[151,1436,193,1830]" captionTargetPageId="26" captionText="FIGURE 14. Abyssoecia elevata n. gen., n. sp. A–E, paratype 3, ZIRAS 4/50706; F, specimen YMG4–06, Stn 106; G, J, specimen YMG4–13, Stn 302; H, I, N, specimen YMG4–07, Stn 129; K, specimen GLD4–09, Stn 191; L, M, specimen YMG4–04, Stn 44. A, gonozooid at upper left of image with projecting ooeciostome; B, D, distal and distolateral views of gonozooid; C, E, frontal and distal views of ooeciostome with subcircular ooeciopore; F–H, distolateral (F) and distal views of colony margin with developing zooids; I–K, frontal views of short autozooidal peristomes; L–N, ancestrular region of two colonies. Scale bars: A, 500 µm; B, D, F–J, L–N, 200 µm; C, E, 100 µm; K, 50 µm." httpUri="https://zenodo.org/record/1437877/files/figure.png" pageId="28" pageNumber="29">14A, E–J</figureCitation>
|
||
) and having fibrous (planar-spherulitic) microstructure (
|
||
<figureCitation id="42872A77562E551B0830FE8C8BF8FE9B" box="[329,434,260,285]" captionStart="FIGURE 14" captionStartId="26.[151,250,1852,1874]" captionTargetBox="[151,1436,193,1830]" captionTargetId="figure@26.[151,1436,193,1830]" captionTargetPageId="26" captionText="FIGURE 14. Abyssoecia elevata n. gen., n. sp. A–E, paratype 3, ZIRAS 4/50706; F, specimen YMG4–06, Stn 106; G, J, specimen YMG4–13, Stn 302; H, I, N, specimen YMG4–07, Stn 129; K, specimen GLD4–09, Stn 191; L, M, specimen YMG4–04, Stn 44. A, gonozooid at upper left of image with projecting ooeciostome; B, D, distal and distolateral views of gonozooid; C, E, frontal and distal views of ooeciostome with subcircular ooeciopore; F–H, distolateral (F) and distal views of colony margin with developing zooids; I–K, frontal views of short autozooidal peristomes; L–N, ancestrular region of two colonies. Scale bars: A, 500 µm; B, D, F–J, L–N, 200 µm; C, E, 100 µm; K, 50 µm." httpUri="https://zenodo.org/record/1437877/files/figure.png" pageId="28" pageNumber="29">Fig. 14K</figureCitation>
|
||
); pseudopores small, simple, sparse, evenly distributed.
|
||
</paragraph>
|
||
<paragraph id="DA0336F2562E551B09BEFEAF8B3DFE57" blockId="28.[151,1437,151,1149]" pageId="28" pageNumber="29">
|
||
Frontal surface with evenly distributed autozooidal peristomes, all obliquely inclined in direction of colony growth, generally longer towards colony margin, fully formed apertures mostly circular (
|
||
<figureCitation id="42872A77562E551B0DB0FEC48F6AFEE2" box="[1225,1312,332,357]" captionStart="FIGURE 14" captionStartId="26.[151,250,1852,1874]" captionTargetBox="[151,1436,193,1830]" captionTargetId="figure@26.[151,1436,193,1830]" captionTargetPageId="26" captionText="FIGURE 14. Abyssoecia elevata n. gen., n. sp. A–E, paratype 3, ZIRAS 4/50706; F, specimen YMG4–06, Stn 106; G, J, specimen YMG4–13, Stn 302; H, I, N, specimen YMG4–07, Stn 129; K, specimen GLD4–09, Stn 191; L, M, specimen YMG4–04, Stn 44. A, gonozooid at upper left of image with projecting ooeciostome; B, D, distal and distolateral views of gonozooid; C, E, frontal and distal views of ooeciostome with subcircular ooeciopore; F–H, distolateral (F) and distal views of colony margin with developing zooids; I–K, frontal views of short autozooidal peristomes; L–N, ancestrular region of two colonies. Scale bars: A, 500 µm; B, D, F–J, L–N, 200 µm; C, E, 100 µm; K, 50 µm." httpUri="https://zenodo.org/record/1437877/files/figure.png" pageId="28" pageNumber="29">Fig.14J</figureCitation>
|
||
), calcified terminal diaphragms rare. New zooids budded from part of distobasal wall of preceding zooid (
|
||
<figureCitation id="42872A77562E551B0DB9FEE78F0CFE0F" box="[1216,1350,367,392]" captionStart="FIGURE 14" captionStartId="26.[151,250,1852,1874]" captionTargetBox="[151,1436,193,1830]" captionTargetId="figure@26.[151,1436,193,1830]" captionTargetPageId="26" captionText="FIGURE 14. Abyssoecia elevata n. gen., n. sp. A–E, paratype 3, ZIRAS 4/50706; F, specimen YMG4–06, Stn 106; G, J, specimen YMG4–13, Stn 302; H, I, N, specimen YMG4–07, Stn 129; K, specimen GLD4–09, Stn 191; L, M, specimen YMG4–04, Stn 44. A, gonozooid at upper left of image with projecting ooeciostome; B, D, distal and distolateral views of gonozooid; C, E, frontal and distal views of ooeciostome with subcircular ooeciopore; F–H, distolateral (F) and distal views of colony margin with developing zooids; I–K, frontal views of short autozooidal peristomes; L–N, ancestrular region of two colonies. Scale bars: A, 500 µm; B, D, F–J, L–N, 200 µm; C, E, 100 µm; K, 50 µm." httpUri="https://zenodo.org/record/1437877/files/figure.png" pageId="28" pageNumber="29">Fig. 14G, H</figureCitation>
|
||
), hence zooids partly overlapping. Interior wall surface of imbricated foliated fabric of wedge-shaped crystallites; no spinules (
|
||
<figureCitation id="42872A77562E551B087BFE3F8B20FE57" box="[258,362,439,464]" captionStart="FIGURE 14" captionStartId="26.[151,250,1852,1874]" captionTargetBox="[151,1436,193,1830]" captionTargetId="figure@26.[151,1436,193,1830]" captionTargetPageId="26" captionText="FIGURE 14. Abyssoecia elevata n. gen., n. sp. A–E, paratype 3, ZIRAS 4/50706; F, specimen YMG4–06, Stn 106; G, J, specimen YMG4–13, Stn 302; H, I, N, specimen YMG4–07, Stn 129; K, specimen GLD4–09, Stn 191; L, M, specimen YMG4–04, Stn 44. A, gonozooid at upper left of image with projecting ooeciostome; B, D, distal and distolateral views of gonozooid; C, E, frontal and distal views of ooeciostome with subcircular ooeciopore; F–H, distolateral (F) and distal views of colony margin with developing zooids; I–K, frontal views of short autozooidal peristomes; L–N, ancestrular region of two colonies. Scale bars: A, 500 µm; B, D, F–J, L–N, 200 µm; C, E, 100 µm; K, 50 µm." httpUri="https://zenodo.org/record/1437877/files/figure.png" pageId="28" pageNumber="29">Fig. 14K</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="DA0336F2562E551B09BEFE548BFCFDBB" blockId="28.[151,1437,151,1149]" pageId="28" pageNumber="29">
|
||
Gonozooid simple, transversely reniform (
|
||
<figureCitation id="42872A77562E551B0BCEFE54896EFE73" box="[695,804,476,501]" captionStart="FIGURE 14" captionStartId="26.[151,250,1852,1874]" captionTargetBox="[151,1436,193,1830]" captionTargetId="figure@26.[151,1436,193,1830]" captionTargetPageId="26" captionText="FIGURE 14. Abyssoecia elevata n. gen., n. sp. A–E, paratype 3, ZIRAS 4/50706; F, specimen YMG4–06, Stn 106; G, J, specimen YMG4–13, Stn 302; H, I, N, specimen YMG4–07, Stn 129; K, specimen GLD4–09, Stn 191; L, M, specimen YMG4–04, Stn 44. A, gonozooid at upper left of image with projecting ooeciostome; B, D, distal and distolateral views of gonozooid; C, E, frontal and distal views of ooeciostome with subcircular ooeciopore; F–H, distolateral (F) and distal views of colony margin with developing zooids; I–K, frontal views of short autozooidal peristomes; L–N, ancestrular region of two colonies. Scale bars: A, 500 µm; B, D, F–J, L–N, 200 µm; C, E, 100 µm; K, 50 µm." httpUri="https://zenodo.org/record/1437877/files/figure.png" pageId="28" pageNumber="29">Fig. 14A</figureCitation>
|
||
), wedged in among autozooidal peristomes at colony margin, moderately densely pseudoporous, narrow ooeciostome directed outwards, with more or less circular ooeciopore (
|
||
<figureCitation id="42872A77562E551B085AFDAC8BE3FDBB" box="[291,425,548,573]" captionStart="FIGURE 14" captionStartId="26.[151,250,1852,1874]" captionTargetBox="[151,1436,193,1830]" captionTargetId="figure@26.[151,1436,193,1830]" captionTargetPageId="26" captionText="FIGURE 14. Abyssoecia elevata n. gen., n. sp. A–E, paratype 3, ZIRAS 4/50706; F, specimen YMG4–06, Stn 106; G, J, specimen YMG4–13, Stn 302; H, I, N, specimen YMG4–07, Stn 129; K, specimen GLD4–09, Stn 191; L, M, specimen YMG4–04, Stn 44. A, gonozooid at upper left of image with projecting ooeciostome; B, D, distal and distolateral views of gonozooid; C, E, frontal and distal views of ooeciostome with subcircular ooeciopore; F–H, distolateral (F) and distal views of colony margin with developing zooids; I–K, frontal views of short autozooidal peristomes; L–N, ancestrular region of two colonies. Scale bars: A, 500 µm; B, D, F–J, L–N, 200 µm; C, E, 100 µm; K, 50 µm." httpUri="https://zenodo.org/record/1437877/files/figure.png" pageId="28" pageNumber="29">Fig. 14B–E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="DA0336F2562E551B09BEFDCF8EF6FD4A" blockId="28.[151,1437,151,1149]" pageId="28" pageNumber="29">
|
||
Ancestrula (
|
||
<figureCitation id="42872A77562E551B082BFDCF8B95FDE7" box="[338,479,583,608]" captionStart="FIGURE 14" captionStartId="26.[151,250,1852,1874]" captionTargetBox="[151,1436,193,1830]" captionTargetId="figure@26.[151,1436,193,1830]" captionTargetPageId="26" captionText="FIGURE 14. Abyssoecia elevata n. gen., n. sp. A–E, paratype 3, ZIRAS 4/50706; F, specimen YMG4–06, Stn 106; G, J, specimen YMG4–13, Stn 302; H, I, N, specimen YMG4–07, Stn 129; K, specimen GLD4–09, Stn 191; L, M, specimen YMG4–04, Stn 44. A, gonozooid at upper left of image with projecting ooeciostome; B, D, distal and distolateral views of gonozooid; C, E, frontal and distal views of ooeciostome with subcircular ooeciopore; F–H, distolateral (F) and distal views of colony margin with developing zooids; I–K, frontal views of short autozooidal peristomes; L–N, ancestrular region of two colonies. Scale bars: A, 500 µm; B, D, F–J, L–N, 200 µm; C, E, 100 µm; K, 50 µm." httpUri="https://zenodo.org/record/1437877/files/figure.png" pageId="28" pageNumber="29">Figs 14L–N</figureCitation>
|
||
,
|
||
<figureCitation id="42872A77562E551B0894FDCF8816FDE7" box="[493,604,583,608]" captionStart="FIGURE 15" captionStartId="27.[151,250,1725,1747]" captionTargetBox="[151,1436,193,1704]" captionTargetId="figure@27.[151,1436,193,1704]" captionTargetPageId="27" captionText="FIGURE 15. Abyssoecia elevata n. gen., n. sp. Progressive stages of development of young ancestrulate colonies viewed from different angles, showing development of either supportive columns or growth of lateral margins towards substratum as colony grows larger. A–C, specimen GLD4–09, Stn 196; D–F, specimen YMG4–07, Stn 129; G–L, specimen YMG4–04, Stn 44. Scale bars: A, B, D–F, 250 µm; C, 200 µm; G–L, 500 µm." httpUri="https://zenodo.org/record/1437879/files/figure.png" pageId="28" pageNumber="29">15D–I, K</figureCitation>
|
||
), suberect, inclined at 50–60° angle to substratum. Protoecium typically flared at base, narrowing at base of peristome but no distinction in surface calcification of needle-like crystallites; peristome tending to be angled to one side (
|
||
<figureCitation id="42872A77562E551B0BF5FD0788BFFD2F" box="[652,757,655,680]" captionStart="FIGURE 14" captionStartId="26.[151,250,1852,1874]" captionTargetBox="[151,1436,193,1830]" captionTargetId="figure@26.[151,1436,193,1830]" captionTargetPageId="26" captionText="FIGURE 14. Abyssoecia elevata n. gen., n. sp. A–E, paratype 3, ZIRAS 4/50706; F, specimen YMG4–06, Stn 106; G, J, specimen YMG4–13, Stn 302; H, I, N, specimen YMG4–07, Stn 129; K, specimen GLD4–09, Stn 191; L, M, specimen YMG4–04, Stn 44. A, gonozooid at upper left of image with projecting ooeciostome; B, D, distal and distolateral views of gonozooid; C, E, frontal and distal views of ooeciostome with subcircular ooeciopore; F–H, distolateral (F) and distal views of colony margin with developing zooids; I–K, frontal views of short autozooidal peristomes; L–N, ancestrular region of two colonies. Scale bars: A, 500 µm; B, D, F–J, L–N, 200 µm; C, E, 100 µm; K, 50 µm." httpUri="https://zenodo.org/record/1437877/files/figure.png" pageId="28" pageNumber="29">Figs 14L</figureCitation>
|
||
,
|
||
<figureCitation id="42872A77562E551B0A7AFD078907FD2F" box="[771,845,655,680]" captionStart="FIGURE 15" captionStartId="27.[151,250,1725,1747]" captionTargetBox="[151,1436,193,1704]" captionTargetId="figure@27.[151,1436,193,1704]" captionTargetPageId="27" captionText="FIGURE 15. Abyssoecia elevata n. gen., n. sp. Progressive stages of development of young ancestrulate colonies viewed from different angles, showing development of either supportive columns or growth of lateral margins towards substratum as colony grows larger. A–C, specimen GLD4–09, Stn 196; D–F, specimen YMG4–07, Stn 129; G–L, specimen YMG4–04, Stn 44. Scale bars: A, B, D–F, 250 µm; C, 200 µm; G–L, 500 µm." httpUri="https://zenodo.org/record/1437879/files/figure.png" pageId="28" pageNumber="29">15H, I</figureCitation>
|
||
). Later daughter zooids tending to form descending outgrowths to substratum, helping to support ancestrula as colony grows; no basal kenozooids.
|
||
</paragraph>
|
||
<paragraph id="DA0336F2562E551B09BEFD5F8939FCBF" blockId="28.[151,1437,151,1149]" pageId="28" pageNumber="29">
|
||
<emphasis id="E8C8EAE0562E551B09BEFD5F8B8CFD77" bold="true" box="[199,454,727,752]" pageId="28" pageNumber="29">Measurements (mm).</emphasis>
|
||
<typeStatus id="05078850562E551B08B4FD5F8872FD77" box="[461,568,727,752]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
|
||
, ZIRAS 1/50703 (
|
||
<figureCitation id="42872A77562E551B0A70FD5F89D9FD77" box="[777,915,727,752]" captionStart="FIGURE 13" captionStartId="25.[151,250,1886,1908]" captionTargetBox="[151,1436,445,1865]" captionTargetId="figure@25.[151,1436,445,1865]" captionTargetPageId="25" captionText="FIGURE 13. Abyssoecia elevata n. gen., n. sp. Holotype and paratype colonies detached from their substrata. A, B, holotype, ZIRAS 1/50703, frontal and profile views; note foramen in colony formed by growth and fusion of lobes around a space; C, D, paratype 1, ZIRAS 2/50704 frontal and abfrontal views; E, F, paratype 2, ZIRAS 3/50705, frontal and profile views. Scale bars: 500 µm." httpUri="https://zenodo.org/record/1437875/files/figure.png" pageId="28" pageNumber="29">Fig. 13A–B</figureCitation>
|
||
): Colony length 3.28, width 4.88, height 2.79; ZL 0.597–0.802 (0.711 ± 0.074); PeL 0.062–0.191 (0.108 ± 0.037); PeD 0.148–0.185 (0.171 ± 0.012); ApL 0.136–0.165 (0.149 ± 0.008); ApW 0.124–0.143 (0.133 ± 0.006).
|
||
</paragraph>
|
||
<paragraph id="DA0336F2562E551B09BEFCCD8BA5FC07" blockId="28.[151,1437,151,1149]" pageId="28" pageNumber="29">
|
||
Paratype 3, ZIRAS 4/50706 (
|
||
<figureCitation id="42872A77562E551B0B65FCCC88E2FCDB" box="[540,680,836,861]" captionStart="FIGURE 14" captionStartId="26.[151,250,1852,1874]" captionTargetBox="[151,1436,193,1830]" captionTargetId="figure@26.[151,1436,193,1830]" captionTargetPageId="26" captionText="FIGURE 14. Abyssoecia elevata n. gen., n. sp. A–E, paratype 3, ZIRAS 4/50706; F, specimen YMG4–06, Stn 106; G, J, specimen YMG4–13, Stn 302; H, I, N, specimen YMG4–07, Stn 129; K, specimen GLD4–09, Stn 191; L, M, specimen YMG4–04, Stn 44. A, gonozooid at upper left of image with projecting ooeciostome; B, D, distal and distolateral views of gonozooid; C, E, frontal and distal views of ooeciostome with subcircular ooeciopore; F–H, distolateral (F) and distal views of colony margin with developing zooids; I–K, frontal views of short autozooidal peristomes; L–N, ancestrular region of two colonies. Scale bars: A, 500 µm; B, D, F–J, L–N, 200 µm; C, E, 100 µm; K, 50 µm." httpUri="https://zenodo.org/record/1437877/files/figure.png" pageId="28" pageNumber="29">Fig. 14A–E</figureCitation>
|
||
): Gonozooid (
|
||
<emphasis id="E8C8EAE0562E551B0A37FCCD8916FCDB" box="[846,860,837,860]" italics="true" pageId="28" pageNumber="29">n</emphasis>
|
||
= 1): GZL 0.703; GZW 0.880; GZH 0.383; OsL 0.155; OsD 0.133; OpD 0.111.
|
||
</paragraph>
|
||
<paragraph id="DA0336F2562E551B09BEFC058B91FC4F" blockId="28.[151,1437,151,1149]" pageId="28" pageNumber="29">
|
||
Non-type specimen YMG4–04, Stn 44 (
|
||
<figureCitation id="42872A77562E551B0BF3FC048953FC23" box="[650,793,908,933]" captionStart="FIGURE 14" captionStartId="26.[151,250,1852,1874]" captionTargetBox="[151,1436,193,1830]" captionTargetId="figure@26.[151,1436,193,1830]" captionTargetPageId="26" captionText="FIGURE 14. Abyssoecia elevata n. gen., n. sp. A–E, paratype 3, ZIRAS 4/50706; F, specimen YMG4–06, Stn 106; G, J, specimen YMG4–13, Stn 302; H, I, N, specimen YMG4–07, Stn 129; K, specimen GLD4–09, Stn 191; L, M, specimen YMG4–04, Stn 44. A, gonozooid at upper left of image with projecting ooeciostome; B, D, distal and distolateral views of gonozooid; C, E, frontal and distal views of ooeciostome with subcircular ooeciopore; F–H, distolateral (F) and distal views of colony margin with developing zooids; I–K, frontal views of short autozooidal peristomes; L–N, ancestrular region of two colonies. Scale bars: A, 500 µm; B, D, F–J, L–N, 200 µm; C, E, 100 µm; K, 50 µm." httpUri="https://zenodo.org/record/1437877/files/figure.png" pageId="28" pageNumber="29">Figs 14L–M</figureCitation>
|
||
,
|
||
<figureCitation id="42872A77562E551B0A5FFC048937FC22" box="[806,893,908,933]" captionStart="FIGURE 15" captionStartId="27.[151,250,1725,1747]" captionTargetBox="[151,1436,193,1704]" captionTargetId="figure@27.[151,1436,193,1704]" captionTargetPageId="27" captionText="FIGURE 15. Abyssoecia elevata n. gen., n. sp. Progressive stages of development of young ancestrulate colonies viewed from different angles, showing development of either supportive columns or growth of lateral margins towards substratum as colony grows larger. A–C, specimen GLD4–09, Stn 196; D–F, specimen YMG4–07, Stn 129; G–L, specimen YMG4–04, Stn 44. Scale bars: A, B, D–F, 250 µm; C, 200 µm; G–L, 500 µm." httpUri="https://zenodo.org/record/1437879/files/figure.png" pageId="28" pageNumber="29">15G–K</figureCitation>
|
||
): Ancestrula (
|
||
<emphasis id="E8C8EAE0562E551B0D62FC058E63FC23" box="[1051,1065,909,932]" italics="true" pageId="28" pageNumber="29">n</emphasis>
|
||
= 1): AnH 0.903; AnPeD 0.221; AnPrL 0.247; AnPrW 0.234.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92A66579562E551B09BEFC5C8B3DFBB2" pageId="28" pageNumber="29" type="discussion">
|
||
<paragraph id="DA0336F2562E551B09BEFC5C8B3DFBB2" blockId="28.[151,1437,151,1149]" pageId="28" pageNumber="29">
|
||
<emphasis id="E8C8EAE0562E551B09BEFC5C8B71FC6A" bold="true" box="[199,315,980,1005]" pageId="28" pageNumber="29">Remarks.</emphasis>
|
||
Eleven colonies, representing a range of sizes, were isolated from the nodule samples; only one was fertile. The elevated, flabellate form of the colony is consistent, with the pillar-like supports providing stability as the colony expands.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92A66579562E551B09BEFBB7884FFBFB" pageId="28" pageNumber="29" type="distribution">
|
||
<paragraph id="DA0336F2562E551B09BEFBB7884FFBFB" blockId="28.[151,1437,151,1149]" pageId="28" pageNumber="29">
|
||
<emphasis id="E8C8EAE0562E551B09BEFBB78B2AFBDF" bold="true" box="[199,352,1087,1112]" pageId="28" pageNumber="29">Distribution.</emphasis>
|
||
Recorded from 10 stations within coordinates 13.09421–
|
||
<geoCoordinate id="BF885035562E551B0A8DFBC88EC2FBD0" box="[1012,1160,1088,1112]" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="1" value="13.49873">13.49873° N</geoCoordinate>
|
||
, 130.80695–
|
||
<geoCoordinate id="BF885035562E551B0C62FBB78AF8FBFB" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="1" value="-134.7224">134.72239° W</geoCoordinate>
|
||
, at depth range
|
||
<quantity id="1D449B17562E551B081BFBEC884AFBFB" box="[354,512,1124,1149]" metricMagnitude="3" metricUnit="m" metricValue="4.9185" metricValueMax="5.05" metricValueMin="4.787" pageId="28" pageNumber="29" unit="m" value="4918.5" valueMax="5050.0" valueMin="4787.0">4787–5050 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |